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<document id="3997155EBE5C9B7552C85E7F80D8B3C1" ID-DOI="10.5281/zenodo.280552" ID-GBIF-Dataset="0e833bb1-2998-430a-9447-865578f47ff8" ID-ISSN="1175-5326" ID-Zenodo-Dep="280552" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460461116901" checkinUser="plazi" docAuthor="Dooley, James K. &amp; Iwatsuki, Yukio" docDate="2012" docId="03BC87C5FFD3896C47CF67EDFCEDF8F4" docLanguage="en" docName="zt03249p038.pdf" docOrigin="Zootaxa 3249" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Branchiostegus saitoi Dooley &amp; Iwatsuki, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FF85FFBDFFD2896F47586079FFFCFFA5" masterDocTitle="A new species of deepwater tilefish (Percoidea: Branchiostegidae) from the Philippines, with a brief discussion of the status of tilefish systematics" masterLastPageNumber="38" masterPageNumber="31" pageNumber="32" updateTime="1698303942547" updateUser="plazi">
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<mods:title id="98B4AA30C8DB6F8EA2496D7B17EB99AC">A new species of deepwater tilefish (Percoidea: Branchiostegidae) from the Philippines, with a brief discussion of the status of tilefish systematics</mods:title>
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<mods:namePart id="C26C25565E3503D14636559AA4237B38">Dooley, James K.</mods:namePart>
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<mods:namePart id="B6BB9088D671D0E2045C6F047E52DBEA">Iwatsuki, Yukio</mods:namePart>
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<mods:date id="DA6CC33A6BFA029129261336F65C0730">2012</mods:date>
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<treatment id="03BC87C5FFD3896C47CF67EDFCEDF8F4" ID-DOI="http://doi.org/10.5281/zenodo.5678692" ID-GBIF-Taxon="119512428" ID-Zenodo-Dep="5678692" LSID="urn:lsid:plazi:treatment:03BC87C5FFD3896C47CF67EDFCEDF8F4" httpUri="http://treatment.plazi.org/id/03BC87C5FFD3896C47CF67EDFCEDF8F4" lastPageId="3" lastPageNumber="34" pageId="1" pageNumber="32">
<subSubSection id="C30F6558FFD3896E47CF67EDFDFDF808" box="[151,513,1939,1966]" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="8BAA36D3FFD3896E47CF67EDFDFDF808" blockId="1.[151,513,1939,1966]" box="[151,513,1939,1966]" pageId="1" pageNumber="32">
<heading id="D0E281BFFFD3896E47CF67EDFDFDF808" bold="true" box="[151,513,1939,1966]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
<emphasis id="B961EAC1FFD3896E47CF67EDFDFDF808" bold="true" box="[151,513,1939,1966]" pageId="1" pageNumber="32">
<taxonomicName id="4C154D50FFD3896E47CF67EDFE60F80B" ID-CoL="N6R6" box="[151,412,1940,1966]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="saitoi" status="sp. nov.">
<emphasis id="B961EAC1FFD3896E47CF67EDFE60F80B" bold="true" box="[151,412,1940,1966]" italics="true" pageId="1" pageNumber="32">Branchiostegus saitoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257BAFFD3896E46FC67EAFDFDF808" box="[420,513,1939,1965]" pageId="1" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30F6558FFD3896D47CF67A2FB38FC12" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32" type="materials_examined">
<paragraph id="8BAA36D3FFD3896D47CF67A2FC71FF72" blockId="1.[151,1435,2010,2035]" lastBlockId="2.[151,1436,151,215]" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32">
<emphasis id="B961EAC1FFD3896E47CF67A2FEF7F856" bold="true" box="[151,267,2011,2035]" pageId="1" pageNumber="32">
<typeStatus id="54AE8871FFD3896E47CF67A2FEFAF856" box="[151,262,2011,2035]" pageId="1" pageNumber="32" type="holotype">Holotype</typeStatus>
.
</emphasis>
Deposited at University of Miyazaki, Division of Fisheries Science, Miyazaki,
<collectingCountry id="F3027643FFD3896E43C067A2FB27F857" box="[1176,1243,2011,2034]" name="Japan" pageId="1" pageNumber="32">Japan</collectingCountry>
; [
<collectionCode id="ED04AE16FFD3896E43B667A3FAC1F857" box="[1262,1341,2010,2034]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="1" pageNumber="32">MUFS</collectionCode>
36081], male,
<quantity id="4CED9B36FFD0896D478660E3FEC1FF17" box="[222,317,154,178]" metricMagnitude="-1" metricUnit="m" metricValue="3.29" pageId="2" pageNumber="33" unit="mm" value="329.0">329 mm</quantity>
SL; hook and line, Batangas, Laiya,
<collectingCountry id="F3027643FFD0896D45B560E3FC92FF17" box="[749,878,154,178]" name="Philippines" pageId="2" pageNumber="33">Philippines</collectingCountry>
, 13
<geoCoordinate id="EE215014FFD0896D44CF60EEFC03FF17" box="[919,1023,151,178]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="92" value="0.64000005">0 38.4 N</geoCoordinate>
,
<geoCoordinate id="EE215014FFD0896D435560E3FB6EFF17" box="[1037,1170,151,178]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="1" value="121.00711">121 0 25.6 E</geoCoordinate>
;
<date id="FFAB1013FFD0896D43F860E3FAB8FF17" box="[1184,1348,154,178]" pageId="2" pageNumber="33" value="2011-04-14">14 April 2011</date>
;
<quantity id="4CED9B36FFD0896D420A60E3FA60FF17" box="[1362,1436,154,178]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="33" unit="m" value="210.0">210 m</quantity>
depth, rocky-sand bottom; photographed fresh, tissue sample taken.
</paragraph>
<caption id="DF6A665BFFD0896D47CF628BFD4AFC82" httpUri="https://zenodo.org/record/280553/files/figure.png" pageId="2" pageNumber="33" targetBox="[234,1358,285,728]" targetPageId="2">
<paragraph id="8BAA36D3FFD0896D47CF628BFD4AFC82" blockId="2.[151,1436,754,807]" pageId="2" pageNumber="33">
<emphasis id="B961EAC1FFD0896D47CF628BFEEFFCAD" bold="true" box="[151,275,754,777]" pageId="2" pageNumber="33">FIGURE 1.</emphasis>
Fresh colouration of 329 mm SL (holotype, MUFS 36081) specimen of
<taxonomicName id="4C154D50FFD0896D44AB628AFBBBFCAD" box="[1011,1095,755,776]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="saitoi">
<emphasis id="B961EAC1FFD0896D44AB628AFBBBFCAD" box="[1011,1095,755,776]" italics="true" pageId="2" pageNumber="33">B. saitoi</emphasis>
</taxonomicName>
caught by hook and line, 14 May 2011, Batangas, Laiya, Philippines; photo by J. Saito.
</paragraph>
</caption>
<paragraph id="8BAA36D3FFD0896D479F632DFB38FC12" blockId="2.[151,1436,852,2032]" pageId="2" pageNumber="33">
<emphasis id="B961EAC1FFD0896D479F632DFEC7FCC9" bold="true" box="[199,315,852,876]" pageId="2" pageNumber="33">
<typeStatus id="54AE8871FFD0896D479F632DFECAFCC9" box="[199,310,852,876]" pageId="2" pageNumber="33" type="paratype">Paratype</typeStatus>
.
</emphasis>
Deposited at the Division of Fishers, National Museum of Sciences, Tokyo,
<collectingCountry id="F3027643FFD0896D4382632CFAE2FCC9" box="[1242,1310,853,876]" name="Japan" pageId="2" pageNumber="33">Japan</collectingCountry>
; [NSMT-P 106562], male,
<quantity id="4CED9B36FFD0896D460E6303FE36FC37" box="[342,458,890,914]" metricMagnitude="-1" metricUnit="m" metricValue="3.156" pageId="2" pageNumber="33" unit="mm" value="315.6">315.6 mm</quantity>
SL; hook and line, Batangas, Laiya,
<collectingCountry id="F3027643FFD0896D44246303FC01FC37" box="[892,1021,890,914]" name="Philippines" pageId="2" pageNumber="33">Philippines</collectingCountry>
, 13
<geoCoordinate id="EE215014FFD0896D437F630EFB73FC37" box="[1063,1167,887,914]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="92" value="0.64000005">0 38.4 N</geoCoordinate>
, 121
<geoCoordinate id="EE215014FFD0896D439F630EFAD0FC37" box="[1223,1324,887,914]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="92" value="0.425">0 25.5 E</geoCoordinate>
;
<date id="FFAB1013FFD0896D42636303FF2DFC12" pageId="2" pageNumber="33" value="2011-04-14">14 April 2011</date>
;
<quantity id="4CED9B36FFD0896D478463E6FEDFFC12" box="[220,291,927,951]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="33" unit="m" value="220.0">220 m</quantity>
depth, rocky-sand bottom; no fresh photograph taken; muscle tissue sample taken.
</paragraph>
</subSubSection>
<subSubSection id="C30F6558FFD0896D479F63BBFCB9F89F" pageId="2" pageNumber="33" type="diagnosis">
<paragraph id="8BAA36D3FFD0896D479F63BBFF18FB5F" blockId="2.[151,1436,852,2032]" pageId="2" pageNumber="33">
<emphasis id="B961EAC1FFD0896D479F63BBFEBDFC7E" bold="true" box="[199,321,962,987]" pageId="2" pageNumber="33">Diagnosis.</emphasis>
This species can easily be distinguished from its congeners by the following combination of characters: a great body depth (2829% SL; other species usually 27% SL) longer head length (3031% SL; other species usually 28%), head profile oblique, about 130 degree angle (other species usually 95130 degrees); head depth (2627% SL; other species usually 26% SL); eyes high on head (interorbital width 3134% HL); small orbital diameter 2425% HL, only
<taxonomicName id="4C154D50FFD0896D4697642DFD7EFBCE" box="[463,642,1107,1131]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="sawakinensis">
<emphasis id="B961EAC1FFD0896D4697642DFD7EFBCE" box="[463,642,1107,1131]" italics="true" pageId="2" pageNumber="33">B. sawakinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D50FFD0896D45E0642DFCEFFBCE" box="[696,787,1107,1131]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="albus">
<emphasis id="B961EAC1FFD0896D45E0642DFCEFFBCE" box="[696,787,1107,1131]" italics="true" pageId="2" pageNumber="33">B. albus</emphasis>
</taxonomicName>
with small orbital diameters, 2324% HL) long predorsal length (35% SL; other species usually 3032% SL, rarely 3334% SL), deep body (28-29% SL; other species usually 2427%, rarely, only in
<taxonomicName id="4C154D50FFD0896D468C64E5FDA3FB16" box="[468,607,1180,1203]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="B961EAC1FFD0896D468C64E5FDA3FB16" box="[468,607,1180,1203]" italics="true" pageId="2" pageNumber="33">B. japonicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D50FFD0896D45CC64E5FCB6FB16" box="[660,842,1179,1203]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="semifasciatus">
<emphasis id="B961EAC1FFD0896D45CC64E5FCB6FB16" box="[660,842,1179,1203]" italics="true" pageId="2" pageNumber="33">B. semifasciatus</emphasis>
</taxonomicName>
, 2829% SL); wide body (1415% SL; other species usually 1113% SL); number of first arch gill rakers low 18 + 1 rudimentary, most other species 1824, modally 1922.
</paragraph>
<paragraph id="8BAA36D3FFD0896D479F657EFCB9F89F" blockId="2.[151,1436,852,2032]" pageId="2" pageNumber="33">
Fresh colouration includes: Upper body rosy red with flecks of pearly-silver scales and pearly pinkish-silver lower body and anterior lower throat regions; dorsal to pectoral to about the lateral line with a large golden yellow area with a silver central patch; lower head region bluish-white from about a horizontal line from lower cheek above the preopercular angle to the pectoral fin base; lateral head metallic golden from anterior orbit to about midorbit dorsally (
<figureCitation id="132E2A56FFD0896D466665EEFE7CFA0A" box="[318,384,1431,1455]" captionStart="FIGURE 1" captionStartId="2.[151,250,754,777]" captionTargetBox="[234,1358,285,728]" captionTargetId="figure@2.[228,1365,268,729]" captionTargetPageId="2" captionText="FIGURE 1. Fresh colouration of 329 mm SL (holotype, MUFS 36081) specimen of B. saitoi caught by hook and line, 14 May 2011, Batangas, Laiya, Philippines; photo by J. Saito." httpUri="https://zenodo.org/record/280553/files/figure.png" pageId="2" pageNumber="33">Fig. 1</figureCitation>
);
<taxonomicName id="4C154D50FFD0896D46C265E1FE04FA0A" box="[410,504,1432,1455]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="saitoi">
<emphasis id="B961EAC1FFD0896D46C265E1FE04FA0A" box="[410,504,1432,1455]" italics="true" pageId="2" pageNumber="33">B. saitoi</emphasis>
</taxonomicName>
also has distinct head, tail and body fresh colouration: a prominent dark predorsal ridge overlying yellow-orange pigment reaching to a vertical above the anterior orbital rim (
<figureCitation id="132E2A56FFD0896D439B65C3FACEFA77" box="[1219,1330,1466,1490]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,754,777]" captionStartId-1="3.[151,250,1563,1586]" captionStartId-2="4.[151,250,1142,1165]" captionTargetBox-0="[234,1358,285,728]" captionTargetBox-1="[151,1436,846,1542]" captionTargetBox-2="[168,1424,191,1120]" captionTargetId-0="figure@2.[228,1365,268,729]" captionTargetId-1="figure@3.[151,1436,846,1542]" captionTargetId-2="figure@4.[139,1424,161,1121]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Fresh colouration of 329 mm SL (holotype, MUFS 36081) specimen of B. saitoi caught by hook and line, 14 May 2011, Batangas, Laiya, Philippines; photo by J. Saito." captionText-1="FIGURE 2. Freshly preserved holotype 329 mm SL; photo by the second author." captionText-2="FIGURE 3. Head of the holotype of B. saitoi (329 mm SL); photo by J. Saito." httpUri-0="https://zenodo.org/record/280553/files/figure.png" httpUri-1="https://zenodo.org/record/280554/files/figure.png" httpUri-2="https://zenodo.org/record/280555/files/figure.png" pageId="2" pageNumber="33">Figs. 13</figureCitation>
); (a dark more prominent predorsal ridge found only in
<taxonomicName id="4C154D50FFD0896D45F46599FCA1FA52" box="[684,861,1503,1527]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="semifasciatus">
<emphasis id="B961EAC1FFD0896D45F46599FCA1FA52" box="[684,861,1503,1527]" italics="true" pageId="2" pageNumber="33">B semifasciatus</emphasis>
</taxonomicName>
from West Africa, but it lacks underlying yelloworange colour); upper lip orange-yellow (over maxillary) and white (over premaxillary and mandible) iris goldengreen; snout reddish; lateral head from above about mid orbit to top of head red; skin over dorsal tips of branchiostegals characteristically bright yellow (
<figureCitation id="132E2A56FFD0896D45C86633FD2AF9C7" box="[656,726,1610,1634]" captionStart="FIGURE 3" captionStartId="4.[151,250,1142,1165]" captionTargetBox="[168,1424,191,1120]" captionTargetId="figure@4.[139,1424,161,1121]" captionTargetPageId="4" captionText="FIGURE 3. Head of the holotype of B. saitoi (329 mm SL); photo by J. Saito." httpUri="https://zenodo.org/record/280555/files/figure.png" pageId="2" pageNumber="33">Fig. 3</figureCitation>
); dorsal fin membrane translucent pink, yellow distally along entire margin of the fin; and anal fin membranes translucent to pinkish white; pelvics white with overlying yellow; dorsal upper pectoral rays dusky, ventral half yellow; caudal characteristically coloured among all congeners (although somewhat similar to
<taxonomicName id="4C154D50FFD0896D46AD66C1FDA9F96A" box="[501,597,1719,1743]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="wardi">
<emphasis id="B961EAC1FFD0896D46AD66C1FDA9F96A" box="[501,597,1719,1743]" italics="true" pageId="2" pageNumber="33">B. wardi</emphasis>
</taxonomicName>
); a central translucent horizontal band, most of the area dorsal to the band bright yellow, upper margin black with outer margin white; below the clear central caudal band also bright yellow for about one third of the ventral caudal area, followed by a little more than third of the caudal black then a little less than a third of the ventral caudal white (
<figureCitation id="132E2A56FFD0896D45D0675BFD29F89F" box="[648,725,1826,1850]" captionStart="FIGURE 1" captionStartId="2.[151,250,754,777]" captionTargetBox="[234,1358,285,728]" captionTargetId="figure@2.[228,1365,268,729]" captionTargetPageId="2" captionText="FIGURE 1. Fresh colouration of 329 mm SL (holotype, MUFS 36081) specimen of B. saitoi caught by hook and line, 14 May 2011, Batangas, Laiya, Philippines; photo by J. Saito." httpUri="https://zenodo.org/record/280553/files/figure.png" pageId="2" pageNumber="33">Figs. 1</figureCitation>
,
<figureCitation id="132E2A56FFD0896D45BB675BFD0DF89F" box="[739,753,1826,1850]" captionStart="FIGURE 2" captionStartId="3.[151,250,1563,1586]" captionTargetBox="[151,1436,846,1542]" captionTargetId="figure@3.[151,1436,846,1542]" captionTargetPageId="3" captionText="FIGURE 2. Freshly preserved holotype 329 mm SL; photo by the second author." httpUri="https://zenodo.org/record/280554/files/figure.png" pageId="2" pageNumber="33">2</figureCitation>
and
<figureCitation id="132E2A56FFD0896D447F675BFCCBF89F" box="[807,823,1826,1850]" captionStart="FIGURE 5" captionStartId="5.[151,250,1755,1778]" captionTargetBox="[151,1436,151,1733]" captionTargetId="figure@5.[151,1436,127,1733]" captionTargetPageId="5" captionText="FIGURE 5. Freshly preserved colouration of the tail of B. saitoi, 329 mm SL (holotype) photo by the second author." httpUri="https://zenodo.org/record/280557/files/figure.png" pageId="2" pageNumber="33">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F6558FFD0896C479F673FFC0BFD50" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33" type="description">
<paragraph id="8BAA36D3FFD0896C479F673FFC0BFD50" blockId="2.[151,1436,852,2032]" lastBlockId="3.[151,1436,151,758]" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33">
<emphasis id="B961EAC1FFD0896D479F673FFEAEF8FA" bold="true" box="[199,338,1862,1887]" pageId="2" pageNumber="33">Description</emphasis>
(parentheses denotes
<typeStatus id="54AE8871FFD0896D451E6731FD55F8C5" box="[582,681,1864,1888]" pageId="2" pageNumber="33" type="paratype">paratype</typeStatus>
, if different from
<typeStatus id="54AE8871FFD0896D4436673EFC2EF8FA" box="[878,978,1863,1887]" pageId="2" pageNumber="33" type="holotype">holotype</typeStatus>
) Dorsal fin rays VII,15, first dorsal spine 1.4 into second spine, dorsal fin origin over pectoral fin base, dorsal long and continuous, greatest height of dorsal fin 13% SL (11%), base length 58% SL (56% SL), antepenultimate ray elongate 18% SL; anal fin rays II,12, anal fin origin under and between dorsal fin rays 4 and 5, dorsal long and continuous, greatest height 12.6% SL (10% SL), base length 28% SL, penultimate ray elongate, 10% SL; pectoral fin rays 18, length 26% SL; pelvic fin I,5, length 14% SL (15%); caudal margin double scalloped; (from radiographs) principal caudal fin rays 17; procurrent caudal rays 10 (upper) + 9 (lower); 1 uroneural and parhypural, 3 epurals, and 2+3 autogenous hypurals; preopercular angle about 90 0 with fine serrae (about 52) on upper limb, a slight indention near preopercle angle (like
<taxonomicName id="4C154D50FFD1896C42DC609DFF28FEBA" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="albus">
<emphasis id="B961EAC1FFD1896C42DC609DFF28FEBA" italics="true" pageId="3" pageNumber="34">B. albus</emphasis>
</taxonomicName>
), no serrae on lower limb of preopercle; scales ctenoid (in pockets) over most of body, cycloid in head region; scale rows to orbital rim 11 (12) and with free margins, (not imbedded as in
<taxonomicName id="4C154D50FFD1896C43596155FB72FEE6" box="[1025,1166,300,323]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="B961EAC1FFD1896C43596155FB72FEE6" box="[1025,1166,300,323]" italics="true" pageId="3" pageNumber="34">B. japonicus</emphasis>
</taxonomicName>
); 7 opercular free scale rows, no enlarged spine at opercle angle (only a single blunt spine); 7 (8); scale rows above the lateral line; scale rows below the lateral line 23 (24); interoperculum not scaled; 49 (50) pored lateral line scales to hypural crease plus 1 (2) on tail; first arch gill rakers (upper limb + lower limb) 7 +11 + 1 rudimentary; total gill rakers on all four arches 49 (51) (incl. rudimentary gill rakers); vertebrae 24 (10+14); predorsal fin support formula 0-0-2-; preopercular length 78% HL; (snout to upper margin of operculum); head profile oblique about 50o angle (
<figureCitation id="132E2A56FFD1896C424D6198FAA8FE5C" box="[1301,1364,481,505]" captionStart="FIGURE 2" captionStartId="3.[151,250,1563,1586]" captionTargetBox="[151,1436,846,1542]" captionTargetId="figure@3.[151,1436,846,1542]" captionTargetPageId="3" captionText="FIGURE 2. Freshly preserved holotype 329 mm SL; photo by the second author." httpUri="https://zenodo.org/record/280554/files/figure.png" pageId="3" pageNumber="34">Fig.2</figureCitation>
); well developed supraoccipital crest; jaws terminal and slightly inferior, upper jaw protrusile; maxillary extends posteriorly to below mid-pupil (
<figureCitation id="132E2A56FFD1896C46E96250FDE0FDE4" box="[433,540,553,577]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,754,777]" captionStartId-1="3.[151,250,1563,1586]" captionStartId-2="4.[151,250,1142,1165]" captionTargetBox-0="[234,1358,285,728]" captionTargetBox-1="[151,1436,846,1542]" captionTargetBox-2="[168,1424,191,1120]" captionTargetId-0="figure@2.[228,1365,268,729]" captionTargetId-1="figure@3.[151,1436,846,1542]" captionTargetId-2="figure@4.[139,1424,161,1121]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Fresh colouration of 329 mm SL (holotype, MUFS 36081) specimen of B. saitoi caught by hook and line, 14 May 2011, Batangas, Laiya, Philippines; photo by J. Saito." captionText-1="FIGURE 2. Freshly preserved holotype 329 mm SL; photo by the second author." captionText-2="FIGURE 3. Head of the holotype of B. saitoi (329 mm SL); photo by J. Saito." httpUri-0="https://zenodo.org/record/280553/files/figure.png" httpUri-1="https://zenodo.org/record/280554/files/figure.png" httpUri-2="https://zenodo.org/record/280555/files/figure.png" pageId="3" pageNumber="34">Figs. 1 3</figureCitation>
); upper jaw with fine canine teeth in 2 3 rows on each side widening anteriorly to a patch on either side of jaw symphysis; with an enlarged tooth at the rear of jaw; lower jaw similarly with rows of fine canines terminating in a patch on either side of the symphysis; no teeth on palatine, vomer, tongue; pharyngeal canine teeth well developed 5 mandibular pores unilaterally (as with most congeners); first haemal spine over second anal-fin spine with parhypophyses fused medially into an arch for the rear projection of the swimbladder, first haemal spine forming a broad blade ventrally (varies among congeners).
</paragraph>
</subSubSection>
<caption id="DF6A665BFFD1896C47CF6662FC2CF994" box="[151,976,1563,1586]" httpUri="https://zenodo.org/record/280554/files/figure.png" pageId="3" pageNumber="34" targetBox="[151,1436,846,1542]" targetPageId="3">
<paragraph id="8BAA36D3FFD1896C47CF6662FC2CF994" blockId="3.[151,976,1563,1586]" box="[151,976,1563,1586]" pageId="3" pageNumber="34">
<emphasis id="B961EAC1FFD1896C47CF6662FEEFF994" bold="true" box="[151,275,1563,1586]" pageId="3" pageNumber="34">FIGURE 2.</emphasis>
Freshly preserved holotype 329 mm SL; photo by the second author.
</paragraph>
</caption>
<subSubSection id="C30F6558FFD1896C479F6618FD85F964" pageId="3" pageNumber="34" type="distribution">
<paragraph id="8BAA36D3FFD1896C479F6618FD85F964" blockId="3.[151,1436,1630,1873]" pageId="3" pageNumber="34">
<emphasis id="B961EAC1FFD1896C479F6618FEA5F9DF" bold="true" box="[199,345,1633,1658]" pageId="3" pageNumber="34">Distribution</emphasis>
. Known only from Bayangas, Laiya,
<collectingCountry id="F3027643FFD1896C44416618FC6BF9DC" box="[793,919,1633,1657]" name="Philippines" pageId="3" pageNumber="34">Philippines</collectingCountry>
(13
<geoCoordinate id="EE215014FFD1896C44906627FBC4F9DC" box="[968,1080,1630,1657]" direction="north" orientation="latitude" pageId="3" pageNumber="34" precision="92" value="0.64166665">0 38.5 N</geoCoordinate>
, 121
<geoCoordinate id="EE215014FFD1896C43296627FB2AF9DC" box="[1137,1238,1630,1657]" direction="east" orientation="longitude" pageId="3" pageNumber="34" precision="92" value="0.42666668">0 25.6 E</geoCoordinate>
) at 210220 M depths, it is possibly endemic; but as it is very rare, and probably inhabits burrows as do other tilefishes,
<taxonomicName id="4C154D50FFD1896C426666FFFA60F938" box="[1342,1436,1670,1693]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="saitoi">
<emphasis id="B961EAC1FFD1896C426666FFFA60F938" box="[1342,1436,1670,1693]" italics="true" pageId="3" pageNumber="34">B. saitoi</emphasis>
</taxonomicName>
may ultimately be found at other localities.
</paragraph>
</subSubSection>
<subSubSection id="C30F6558FFD1896C479F66B4FCEDF8F4" pageId="3" pageNumber="34" type="etymology">
<paragraph id="8BAA36D3FFD1896C479F66B4FCEDF8F4" blockId="3.[151,1436,1630,1873]" pageId="3" pageNumber="34">
<emphasis id="B961EAC1FFD1896C479F66B4FEB2F940" bold="true" box="[199,334,1741,1765]" pageId="3" pageNumber="34">Etymology.</emphasis>
A new species of
<taxonomicName id="4C154D50FFD1896C454366B4FD35F940" box="[539,713,1741,1765]" class="Actinopterygii" family="Malacanthidae" genus="Branchiostegus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B961EAC1FFD1896C454366B4FD35F940" box="[539,713,1741,1765]" italics="true" pageId="3" pageNumber="34">Branchiostegus</emphasis>
</taxonomicName>
was first caught off Batangas Laiya,
<collectingCountry id="F3027643FFD1896C433366B4FB15F940" box="[1131,1257,1741,1765]" name="Philippines" pageId="3" pageNumber="34">Philippines</collectingCountry>
in 2009, photographed and eaten. Two years passed before two more specimens were caught by J. Saito, an amateur angler from
<collectingCountry id="F3027643FFD1896C47CF676FFF2BF888" box="[151,215,1814,1837]" name="Japan" pageId="3" pageNumber="34">Japan</collectingCountry>
in April, 2011. Without J. Saitos considerable efforts and interest, this species would have remained unknown. The new species is herein named in his honor.
</paragraph>
</subSubSection>
</treatment>
</document>