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<document id="05FECF7F7C0B22C395D9377E189897E4" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFFD22530CC07997FCD35015" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus subapicalis Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub 2021, stat. nov." docType="treatment" docVersion="3" lastPageNumber="585" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="584" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B3402D97807DC3AEFADD825E286D85AD">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="C5DDB676BBFDDBEAF4C4B45A06FB5636">Suksai, Benjamart</mods:namePart>
<mods:nameIdentifier id="23D1B853121CFFD0F07C29F111AE56BC" type="ORCID">0000-0002-2472-6100</mods:nameIdentifier>
<mods:nameIdentifier id="DC501373CBCCDD1D2FAA91789E1CF4B5" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B18C7582060B34C8775934FDEEF16D7A">Gustafson, Grey T.</mods:namePart>
<mods:affiliation id="6A7E2F5269E5520BC502CBB68F95CD78">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="A1AA02EE4269E06085895455511C077F">Sites, Robert W.</mods:namePart>
<mods:nameIdentifier id="B84189AD8975C64E45B270672EEAFC32" type="ORCID">0000-0002-3895-813X</mods:nameIdentifier>
<mods:affiliation id="484BB92859F23D442DDDA1E2E80C2258">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:role id="A29B65151717C91EC3053BA1F3828440">
<mods:roleTerm id="608901D464A72B829DD8000C7A43EF6C">Author</mods:roleTerm>
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<mods:namePart id="305306117AED4A6EE3DE65E545CD3E7F">Sangpradub, Narumon</mods:namePart>
<mods:nameIdentifier id="8F7951214749C93907C15B5F21F33E7B" type="ORCID">0000-0002-2472-6100</mods:nameIdentifier>
<mods:affiliation id="C6A3804C0584236E732DC17EC96FF9D2">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
<mods:nameIdentifier id="994DB5C118462E0B5203D34179F4CDB0" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:title id="E6BA47CF66E9409162AE82A8ED3AF5C4">Zootaxa</mods:title>
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<mods:part id="7ED19461A7D2F5F3438F524A7D8B4725">
<mods:date id="6DCB36504EF3249BC1F480E49C1F040B">2021</mods:date>
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<mods:number id="65F1CA93C1E8BF9CB1F4E386C0D1AE68">2021-06-25</mods:number>
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<mods:number id="77FF722714DE10529FF2D83D02A1052D">4991</mods:number>
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<mods:number id="110EB5259580B4A93E37BE2DC2B5C880">3</mods:number>
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<mods:identifier id="92357E6C36E8DA7B07EC125E490A147B" type="DOI">10.11646/zootaxa.4991.3.8</mods:identifier>
<mods:identifier id="67E8BFCECB47358038C7F530C0804663" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C30F655CFFFD225C0CC07997FD4F50B3" box="[151,656,947,974]" pageId="23" pageNumber="584" type="nomenclature">
<paragraph id="8BAA36D7FFFD225C0CC07997FD4F50B3" blockId="23.[151,656,947,1009]" box="[151,656,947,974]" pageId="23" pageNumber="584">
<heading id="D0E281BBFFFD225C0CC07997FD4F50B3" bold="true" box="[151,656,947,974]" fontSize="11" level="1" pageId="23" pageNumber="584" reason="1">
<emphasis id="B961EAC5FFFD225C0CC07997FD4F50B3" bold="true" box="[151,656,947,974]" pageId="23" pageNumber="584">
<taxonomicName id="4C154D54FFFD225C0CC07997FDC650B3" authority="(Ochs, 1930)" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[151,537,947,974]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="subapicalis" status="stat. nov.">
<emphasis id="B961EAC5FFFD225C0CC07997FEAA50B3" bold="true" box="[151,373,947,974]" italics="true" pageId="23" pageNumber="584">Patrus subapicalis</emphasis>
(
<bibRefCitation id="EF844B26FFFD225C0DD27997FDCE50B3" author="Ochs" box="[389,529,948,974]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25257BEFFFD225C0E767997FD4F50B3" box="[545,656,948,974]" pageId="23" pageNumber="584">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFD225C0CC079F4FDD0508C" box="[151,527,983,1009]" pageId="23" pageNumber="584" type="description">
<paragraph id="8BAA36D7FFFD225C0CC079F4FDD0508C" blockId="23.[151,656,947,1009]" box="[151,527,983,1009]" pageId="23" pageNumber="584">
(
<figureCitation id="132E2A52FFFD225C0CC879F4FEDC508C" box="[159,259,983,1009]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="23" pageNumber="584">Figs. 2H</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0D5879F4FEEC508C" box="[271,307,983,1009]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="23" pageNumber="584">3H</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0D6879F4FEBB508C" box="[319,356,983,1009]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="23" pageNumber="584">4H</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0D3879F4FE52508C" box="[367,397,983,1009]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="23" pageNumber="584">12</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0DCE79F4FE13508C" box="[409,460,983,1009]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="23" pageNumber="584">13A</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0D8F79F4FDD8508C" box="[472,519,983,1009]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="23" pageNumber="584">14B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFD225C0CC07E3DFB8E5725" pageId="23" pageNumber="584" type="reference_group">
<paragraph id="8BAA36D7FFFD225C0CC07E3DFC1D574B" blockId="23.[151,1105,1054,1113]" box="[151,962,1054,1078]" pageId="23" pageNumber="584">
<treatmentCitationGroup id="AB0511F9FFFD225C0CC07E3DFC1D574B" box="[151,962,1054,1078]" pageId="23" pageNumber="584">
<treatmentCitation id="0AB410C6FFFD225C0CC07E3DFD3B574B" author="Ochs" box="[151,740,1054,1078]" page="20" pageId="23" pageNumber="584" year="1930">
<taxonomicName id="4C154D54FFFD225C0CC07E3DFD3B574B" authority="Ochs, 1930: 20" authorityName="Ochs" authorityPageNumber="20" authorityYear="1930" box="[151,740,1054,1078]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="subSpecies" species="apicalis" subGenus="Patrus" subSpecies="subapicalis">
<emphasis id="B961EAC5FFFD225C0CC07E3DFEC6574B" box="[151,281,1054,1078]" italics="true" pageId="23" pageNumber="584">Orectochilus</emphasis>
(
<emphasis id="B961EAC5FFFD225C0D7F7E3CFEB3574B" box="[296,364,1055,1078]" italics="true" pageId="23" pageNumber="584">Patrus</emphasis>
)
<emphasis id="B961EAC5FFFD225C0D2D7E3DFD9B574B" box="[378,580,1054,1078]" italics="true" pageId="23" pageNumber="584">apicalis subapicalis</emphasis>
<bibRefCitation id="EF844B26FFFD225C0E1D7E3DFD3B574B" author="Ochs" box="[586,740,1054,1078]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36D7FFFD225C0CC07E62FB8E5725" blockId="23.[151,1105,1054,1113]" box="[151,1105,1088,1113]" pageId="23" pageNumber="584">
<treatmentCitationGroup id="AB0511F9FFFD225C0CC07E62FB8E5725" box="[151,1105,1088,1113]" pageId="23" pageNumber="584">
<taxonomicName id="4C154D54FFFD225C0CC07E62FDF75724" authority="Ochs, 1930" authorityName="Ochs" authorityYear="1930" box="[151,552,1088,1113]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="subSpecies" species="apicalis" subSpecies="subapicalis">
<emphasis id="B961EAC5FFFD225C0CC07E62FE745725" box="[151,427,1088,1112]" italics="true" pageId="23" pageNumber="584">Patrus apicalis subapicalis</emphasis>
<bibRefCitation id="EF844B26FFFD225C0DE67E63FDF75724" author="Ochs" box="[433,552,1088,1113]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFFD225C0E657E63FCC25724" author="Hajek, J. &amp; Fery, H." box="[562,797,1088,1113]" page="28" pageId="23" pageNumber="584" year="2017">
<bibRefCitation id="EF844B26FFFD225C0E657E63FCC25724" author="Hajek, J. &amp; Fery, H." box="[562,797,1088,1113]" pageId="23" pageNumber="584" pagination="1810" refId="ref21748" refString="Hajek, J. &amp; Fery, H. (2017) Family Gyrinidae Latreille, 1810. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Leiden, Brill, pp. 22 - 29." type="book chapter" year="2017">Hájek &amp; Fery 2017: 28</bibRefCitation>
</treatmentCitation>
(new combination, catalogue)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFD225C0CC07EA7FAB9544C" pageId="23" pageNumber="584" type="materials_examined">
<paragraph id="8BAA36D7FFFD225C0CC07EA7FE2657E2" blockId="23.[151,1437,1156,2015]" box="[151,505,1156,1183]" pageId="23" pageNumber="584">
<materialsCitation id="3B7D3C8AFFFD225C0CC07EA7FE2657E2" box="[151,505,1156,1183]" country="Laos" location="Nam Nat'" pageId="23" pageNumber="584" specimenCount="1" typeStatus="holotype">
<emphasis id="B961EAC5FFFD225C0CC07EA7FE9A57E3" bold="true" box="[151,325,1156,1182]" pageId="23" pageNumber="584">
<typeStatus id="54AE8875FFFD225C0CC07EA7FF0D57E3" box="[151,210,1156,1182]" pageId="23" pageNumber="584">Type</typeStatus>
locality. ‘
</emphasis>
<location id="8ECA600CFFFD225C0D117EA6FE6857E3" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C0D117EA6FE6857E3" box="[326,439,1156,1183]" country="Laos" name="Nam Nat'" pageId="23" pageNumber="584">Nam Nat’</location>
,
<collectingCountry id="F3027647FFFD225C0D947EA6FE2657E2" box="[451,505,1157,1183]" name="Laos" pageId="23" pageNumber="584">Laos</collectingCountry>
</materialsCitation>
</paragraph>
<paragraph id="8BAA36D7FFFD225C0C907E84FF2E579E" blockId="23.[151,1437,1156,2015]" pageId="23" pageNumber="584">
<emphasis id="B961EAC5FFFD225C0C907E84FE2757BD" bold="true" box="[199,504,1190,1217]" pageId="23" pageNumber="584">Type material examined.</emphasis>
<typeStatus id="54AE8875FFFD225C0E547E84FD5157BC" box="[515,654,1191,1217]" pageId="23" pageNumber="584" type="paratype">PARATYPE</typeStatus>
female SMF C 9548, additional label information not provided to authors.
</paragraph>
<paragraph id="8BAA36D7FFFD225C0C907ECFFAB9544C" blockId="23.[151,1437,1156,2015]" pageId="23" pageNumber="584">
<emphasis id="B961EAC5FFFD225C0C907ECFFDDD567B" bold="true" box="[199,514,1260,1286]" pageId="23" pageNumber="584">Other material examined.</emphasis>
<materialsCitation id="3B7D3C8AFFFD225C0E5B7ECFFDA45636" collectingDate="2004-05-18" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1296" latitude="18.516666" location="Ban Khun Klang" longLatPrecision="1275" longitude="98.51667" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<emphasis id="B961EAC5FFFD225C0E5B7ECFFC1A567B" bold="true" box="[524,965,1260,1286]" pageId="23" pageNumber="584">
<collectingCountry id="F3027647FFFD225C0E5B7ECFFD78567B" box="[524,679,1260,1286]" name="Thailand" pageId="23" pageNumber="584">THAILAND</collectingCountry>
:
<collectingRegion id="49D1F835FFFD225C0EE17ECFFC1F567B" box="[694,960,1260,1286]" country="Thailand" name="Chiang Mai" pageId="23" pageNumber="584">Chiang Mai Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFFD225C0F987ECFFB2F567B" box="[975,1264,1260,1286]" pageId="23" pageNumber="584">Doi Inthanon Natl. Park.</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFD225C08AA7ECEFAA2567B" box="[1277,1405,1260,1287]" pageId="23" pageNumber="584">Mae Klang</collectingMunicipality>
at
<location id="8ECA600CFFFD225C0CC07F2CFE815654" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C0CC07F2CFE815654" box="[151,350,1295,1321]" country="Thailand" county="Doi Inthanon Natl. Park." latitude="18.516666" longLatPrecision="1275" longitude="98.51667" municipality="Mae Klang" name="Ban Khun Klang" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Ban Khun Klang</location>
,
<geoCoordinate id="EE215010FFFD225C0D3B7F2CFE125654" box="[364,461,1295,1321]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="23" pageNumber="584" precision="925" value="18.516666">18°31’N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C0D807F2CFDE85654" box="[471,567,1295,1321]" degrees="98" direction="east" minutes="31" orientation="longitude" pageId="23" pageNumber="584" precision="925" value="98.51667">98°31’E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0E117F2CFD015654" box="[582,734,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.2959999999999998" pageId="23" pageNumber="584" unit="m" value="1296.0">
<quantity id="4CED9B32FFFD225C0E117F2CFD415654" box="[582,670,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.2959999999999998" pageId="23" pageNumber="584" unit="m" value="1296.0">1296 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0EB97F2CFCBC5654" box="[750,867,1295,1321]" pageId="23" pageNumber="584" value="2004-05-18">
<collectingDate id="EFEFE9FFFFFD225C0EB97F2CFCBC5654" box="[750,867,1295,1321]" pageId="23" pageNumber="584" value="2004-05-18">18.V.2004</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFD225C0FF47F2CFB8A5654" box="[931,1109,1295,1321]" pageId="23" pageNumber="584">A. Vitheepradit</collectorName>
,
<collectorName id="26E05301FFFD225C08347F2CFB3F5654" box="[1123,1248,1295,1321]" pageId="23" pageNumber="584">T. Prommi</collectorName>
,
<collectorName id="26E05301FFFD225C08B87F2CFA815654" box="[1263,1374,1295,1321]" pageId="23" pageNumber="584">P. Laudee</collectorName>
&amp;
<collectorName id="26E05301FFFD225C09DE7F2CFE815636" pageId="23" pageNumber="584">P. Thamasenanupap</collectorName>
, L-694,
<specimenCount id="9D13FD5EFFFD225C0DEB7F12FDC15636" box="[444,542,1329,1355]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0E797F12FDAB5636" box="[558,628,1329,1355]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0ED07F11FC2C5613" collectingDate="2002-04-03" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
same locality except for,
<location id="8ECA600CFFFD225C0FF57F12FB1A5636" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C0FF57F12FB1A5636" box="[930,1221,1329,1356]" country="Thailand" latitude="18.516666" longLatPrecision="1275" longitude="98.45" name="Huai Sai Luang Waterfall" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Huai Sai Luang Waterfall</location>
,
<geoCoordinate id="EE215010FFFD225C08987F12FAEF5636" box="[1231,1328,1329,1355]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="23" pageNumber="584" precision="925" value="18.516666">18°31’N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C096F7F12FA475636" box="[1336,1432,1329,1355]" degrees="98" direction="east" minutes="27" orientation="longitude" pageId="23" pageNumber="584" precision="925" value="98.45">98°27’E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0CC07F77FEF55613" box="[151,298,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="23" pageNumber="584" unit="m" value="1060.0">
<quantity id="4CED9B32FFFD225C0CC07F77FF325612" box="[151,237,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="23" pageNumber="584" unit="m" value="1060.0">1060 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0D607F77FE785613" box="[311,423,1364,1390]" pageId="23" pageNumber="584" value="2002-04-03">
<collectingDate id="EFEFE9FFFFFD225C0D607F77FE785613" box="[311,423,1364,1390]" pageId="23" pageNumber="584" value="2002-04-03">3.IV.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-321,
<specimenCount id="9D13FD5EFFFD225C0F637F77FC4A5613" box="[820,917,1364,1390]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0FF27F77FC345613" box="[933,1003,1364,1390]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0FA87F76FD5956EC" collectingDate="2002-12-07" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C094D7F77FA475613" box="[1306,1432,1364,1390]" pageId="23" pageNumber="584" value="2002-12-07">
<collectingDate id="EFEFE9FFFFFD225C094D7F77FA475613" box="[1306,1432,1364,1390]" pageId="23" pageNumber="584" value="2002-12-07">7.XII.2002</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C0D027F54FE7056EC" box="[341,431,1399,1425]" pageId="23" pageNumber="584" type="male">3 males</specimenCount>
,
<specimenCount id="9D13FD5EFFFD225C0DEC7F55FDF756EC" box="[443,552,1398,1425]" pageId="23" pageNumber="584" type="female">5 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0E6F7F54FDA156EC" box="[568,638,1399,1425]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0EC57F54FE8E56CE" collectingDate="2002-05-08" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="2" specimenCount-male="2" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0FF87F54FBC956EC" box="[943,1046,1399,1425]" pageId="23" pageNumber="584" value="2002-05-08">
<collectingDate id="EFEFE9FFFFFD225C0FF87F54FBC956EC" box="[943,1046,1399,1425]" pageId="23" pageNumber="584" value="2002-05-08">8.V.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-396,
<specimenCount id="9D13FD5EFFFD225C0CC07FBAFF2E56CE" box="[151,241,1433,1459]" pageId="23" pageNumber="584" type="male">2 males</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0D537FBAFE9656CE" box="[260,329,1433,1459]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFD225C0D377FBAFD7656CE" box="[352,681,1433,1459]" collectingDate="2002-05-08" collectionCode="THNHM, UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" isTailEnumeration="true" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Mai">
<specimenCount id="9D13FD5EFFFD225C0D377FBAFE0F56CE" box="[352,464,1433,1459]" pageId="23" pageNumber="584" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0DB57FBAFD9256CE" box="[482,589,1433,1459]" pageId="23" pageNumber="584">THNHM</collectionCode>
,
<collectionCode id="ED04AE12FFFD225C0E0B7FBAFD7E56CE" box="[604,673,1433,1459]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0EEF7FB9FE1E56AA" collectingDate="2003-01-13" collectionCode="UMC, THNHM" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" isTailEnumeration="true" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0F887FBAFB9256CE" box="[991,1101,1433,1459]" pageId="23" pageNumber="584" value="2003-01-13">
<collectingDate id="EFEFE9FFFFFD225C0F887FBAFB9256CE" box="[991,1101,1433,1459]" pageId="23" pageNumber="584" value="2003-01-13">13.I.2003</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C09747FBAFAA056CE" box="[1315,1407,1433,1459]" pageId="23" pageNumber="584" type="male">2 males</specimenCount>
,
<specimenCount id="9D13FD5EFFFD225C09D97FBAFF3156AB" pageId="23" pageNumber="584" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0CA97F9FFE9D56AB" box="[254,322,1468,1494]" pageId="23" pageNumber="584">UMC</collectionCode>
,
<collectionCode id="ED04AE12FFFD225C0D1A7F9EFE6656AA" box="[333,441,1469,1495]" pageId="23" pageNumber="584">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0D9B7F9EFB2456AB" box="[460,1275,1468,1495]" collectingDate="2003-03-14" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi &amp; P. Laudee &amp; P. Thamasenanupap" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1060" isTailEnumeration="true" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0EBF7F9FFCB756AB" box="[744,872,1468,1494]" pageId="23" pageNumber="584" value="2003-03-14">
<collectingDate id="EFEFE9FFFFFD225C0EBF7F9FFCB756AB" box="[744,872,1468,1494]" pageId="23" pageNumber="584" value="2003-03-14">14.III.2003</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C08667F9FFB4156AB" box="[1073,1182,1468,1494]" pageId="23" pageNumber="584" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C08FA7F9FFB2C56AB" box="[1197,1267,1468,1494]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C09507F9EFDF75566" collectingDate="2003-04-03" collectionCode="UMC" collectorName="A. Vitheepradit &amp; R. W. Sites &amp; K. Seetaphan" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1100" isTailEnumeration="true" latitude="18.516666" location="Mae Pan Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
same locality except for,
<location id="8ECA600CFFFD225C0D4F7FFCFE2F5684" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C0D4F7FFCFE2F5684" box="[280,496,1503,1529]" country="Thailand" latitude="18.516666" longLatPrecision="1275" longitude="98.45" name="Mae Pan Waterfall" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Mae Pan Waterfall</location>
,
<geoCoordinate id="EE215010FFFD225C0DAA7FFCFD825684" box="[509,605,1503,1529]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="23" pageNumber="584" precision="925" value="18.516666">18°31’N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C0E317FFCFD195684" box="[614,710,1503,1529]" degrees="98" direction="east" minutes="27" orientation="longitude" pageId="23" pageNumber="584" precision="925" value="98.45">98°27’E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0E837FFCFCB75684" box="[724,872,1503,1529]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="584" unit="m" value="1100.0">
<quantity id="4CED9B32FFFD225C0E837FFCFCF65684" box="[724,809,1503,1529]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="584" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0F217FFCFC385684" box="[886,999,1503,1529]" pageId="23" pageNumber="584" value="2003-04-03">
<collectingDate id="EFEFE9FFFFFD225C0F217FFCFC385684" box="[886,999,1503,1529]" pageId="23" pageNumber="584" value="2003-04-03">3.IV.2003</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFD225C08737FFCFB0A5684" box="[1060,1237,1503,1529]" pageId="23" pageNumber="584">A. Vitheepradit</collectorName>
,
<collectorName id="26E05301FFFD225C08B57FFCFA865684" box="[1250,1369,1503,1529]" pageId="23" pageNumber="584">R.W. Sites</collectorName>
&amp;
<collectorName id="26E05301FFFD225C09D67FFCFED25566" pageId="23" pageNumber="584">K. Seetaphan</collectorName>
, L-442,
<specimenCount id="9D13FD5EFFFD225C0D3D7C22FE145566" box="[362,459,1537,1563]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0D8D7C22FDFF5566" box="[474,544,1537,1563]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0E647C21FA475542" collectingDate="2002-03-21" collectionCode="UMC, THNHM" collectorName="R. W. Sites &amp; A. Vitheepradit &amp; K. Kirawanich" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1460" isTailEnumeration="true" latitude="18.533333" location="Siriphum Waterfall" longLatPrecision="1275" longitude="98.51667" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="3" specimenCount-male="3" stateProvince="Chiang Mai">
same locality except for,
<location id="8ECA600CFFFD225C0F1B7C22FBFD5566" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C0F1B7C22FBFD5566" box="[844,1058,1537,1563]" country="Thailand" latitude="18.533333" longLatPrecision="1275" longitude="98.51667" name="Siriphum Waterfall" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Siriphum Waterfall</location>
(lowest level),
<geoCoordinate id="EE215010FFFD225C08877C22FAEF5566" box="[1232,1328,1537,1563]" degrees="18" direction="north" minutes="32" orientation="latitude" pageId="23" pageNumber="584" precision="925" value="18.533333">18°32’N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C096F7C22FA485566" box="[1336,1431,1537,1563]" degrees="98" direction="east" minutes="31" orientation="longitude" pageId="23" pageNumber="584" precision="925" value="98.51667">98°31’E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0CC07C07FEF15543" box="[151,302,1572,1599]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="23" pageNumber="584" unit="m" value="1460.0">
<quantity id="4CED9B32FFFD225C0CC07C07FF305542" box="[151,239,1572,1599]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="23" pageNumber="584" unit="m" value="1460.0">1460 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0D6A7C07FE625543" box="[317,445,1572,1598]" pageId="23" pageNumber="584" value="2002-03-21">
<collectingDate id="EFEFE9FFFFFD225C0D6A7C07FE625543" box="[317,445,1572,1598]" pageId="23" pageNumber="584" value="2002-03-21">21.III.2002</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFD225C0DA97C06FDA65543" box="[510,633,1572,1599]" pageId="23" pageNumber="584">R.W. Sites</collectorName>
,
<collectorName id="26E05301FFFD225C0ED27C07FCEC5543" box="[645,819,1572,1598]" pageId="23" pageNumber="584">A. Vitheepradit</collectorName>
&amp;
<collectorName id="26E05301FFFD225C0F0B7C06FBD85543" box="[860,1031,1572,1599]" pageId="23" pageNumber="584">K. Kirawanich</collectorName>
, L-315,
<specimenCount id="9D13FD5EFFFD225C083E7C07FB1D5543" box="[1129,1218,1572,1598]" pageId="23" pageNumber="584" type="male">3 males</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C08847C07FAC85543" box="[1235,1303,1572,1598]" pageId="23" pageNumber="584">UMC</collectionCode>
,
<collectionCode id="ED04AE12FFFD225C09737C06FA4F5542" box="[1316,1424,1573,1599]" pageId="23" pageNumber="584">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0CC07C64FBBF551C" box="[151,1120,1607,1633]" collectingDate="2002-05-09" collectionCode="UMC" collectorName="R. W. Sites &amp; A. Vitheepradit &amp; K. Kirawanich" country="Thailand" county="Doi Inthanon Natl. Park." elevation="1460" isTailEnumeration="true" latitude="18.533333" location="Siriphum Waterfall" longLatPrecision="1275" longitude="98.51667" municipality="Mae Klang" pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0DE67C64FDC7551C" box="[433,536,1607,1633]" pageId="23" pageNumber="584" value="2002-05-09">
<collectingDate id="EFEFE9FFFFFD225C0DE67C64FDC7551C" box="[433,536,1607,1633]" pageId="23" pageNumber="584" value="2002-05-09">9.V.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-401,
<specimenCount id="9D13FD5EFFFD225C0FF67C64FBDC551C" box="[929,1027,1607,1633]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C08457C64FB87551C" box="[1042,1112,1607,1633]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C083C7C64FAF055FE" collectingDate="2002-05-05" collectionCode="KKU, UMC" country="Thailand" elevation="1100" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="2" specimenCount-male="2" stateProvince="Chiang Mai">
<collectingMunicipality id="6BCEACADFFFD225C083C7C64FAB2551C" box="[1131,1389,1607,1633]" pageId="23" pageNumber="584">Doi Suthep Natl. Park.</collectingMunicipality>
,
<location id="8ECA600CFFFD225C092F7C64FEEF55FE" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C092F7C64FEEF55FE" country="Thailand" latitude="18.8" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." name="Sai Yoi Waterfall" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Sai Yoi Waterfall</location>
,
<geoCoordinate id="EE215010FFFD225C0D6C7C4AFE4355FE" box="[315,412,1641,1667]" degrees="18" direction="north" minutes="48" orientation="latitude" pageId="23" pageNumber="584" precision="925" value="18.8">18°48’N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C0DF47C4AFDDB55F9" box="[419,516,1641,1668]" degrees="98" direction="east" minutes="55" orientation="longitude" pageId="23" pageNumber="584" precision="925" value="98.916664">98°55’E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0E477C4AFD7E55FE" box="[528,673,1641,1668]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="584" unit="m" value="1100.0">
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a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0EF97C4AFCCA55F9" box="[686,789,1641,1668]" pageId="23" pageNumber="584" value="2002-05-05">
<collectingDate id="EFEFE9FFFFFD225C0EF97C4AFCCA55F9" box="[686,789,1641,1668]" pageId="23" pageNumber="584" value="2002-05-05">5.V.2002</collectingDate>
</date>
, leg. CMU team, L-402,
<specimenCount id="9D13FD5EFFFD225C08677C4AFB5855FE" box="[1072,1159,1641,1667]" pageId="23" pageNumber="584" type="male">2 males</specimenCount>
(
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,
<collectionCode id="ED04AE12FFFD225C08B47C4AFAF855FE" box="[1251,1319,1641,1667]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFD225C096C7C4AFF3555DB" collectingDate="2002-05-05" collectionCode="KKU" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<specimenCount id="9D13FD5EFFFD225C096C7C4AFA4355FE" box="[1339,1436,1641,1667]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0CC87CAFFF3D55DB" box="[159,226,1676,1702]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="23" pageNumber="584">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0CAF7CAEFB7A55DB" box="[248,1189,1676,1703]" collectingDate="2002-09-24" collectionCode="UMC" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0E4C7CAFFD4255DB" box="[539,669,1676,1702]" pageId="23" pageNumber="584" value="2002-09-24">
<collectingDate id="EFEFE9FFFFFD225C0E4C7CAFFD4255DB" box="[539,669,1676,1702]" pageId="23" pageNumber="584" value="2002-09-24">24.IX.2002</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C0F397CAFFC1655DB" box="[878,969,1676,1702]" pageId="23" pageNumber="584" type="male">4 males</specimenCount>
,
<specimenCount id="9D13FD5EFFFD225C0F807CAFFB9A55DB" box="[983,1093,1676,1702]" pageId="23" pageNumber="584" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C08007CAFFB4255DB" box="[1111,1181,1676,1702]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C08E47CAEFEB555B4" collectingDate="2002-09-24" collectionCode="KKU" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
same locality except for,
<specimenCount id="9D13FD5EFFFD225C0C927C8CFECF55B4" box="[197,272,1711,1737]" pageId="23" pageNumber="584" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0D487C8CFEBD55B4" box="[287,354,1711,1737]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="23" pageNumber="584">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0D227C8CFBA055B4" box="[373,1151,1711,1737]" collectingDate="2002-10-08" collectionCode="UMC" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0ED87C8CFD2555B4" box="[655,762,1711,1737]" pageId="23" pageNumber="584" value="2002-10-08">
<collectingDate id="EFEFE9FFFFFD225C0ED87C8CFD2555B4" box="[655,762,1711,1737]" pageId="23" pageNumber="584" value="2002-10-08">8.X.2002</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C0F977C8CFBFD55B4" box="[960,1058,1711,1737]" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C08667C8CFBA855B4" box="[1073,1143,1711,1737]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C08DD7C8CFCD35596" collectingDate="2002-11-16" collectionCode="UMC" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C0CC07CF2FEC55596" box="[151,282,1745,1771]" pageId="23" pageNumber="584" value="2002-11-16">
<collectingDate id="EFEFE9FFFFFD225C0CC07CF2FEC55596" box="[151,282,1745,1771]" pageId="23" pageNumber="584" value="2002-11-16">16.XI.2002</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C0DB27CF2FDEB5596" box="[485,564,1745,1771]" pageId="23" pageNumber="584" type="male">1 male</specimenCount>
,
<specimenCount id="9D13FD5EFFFD225C0E167CF2FD715596" box="[577,686,1745,1771]" pageId="23" pageNumber="584" type="female">4 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0EE97CF2FCDB5596" box="[702,772,1745,1771]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0F4E7CF1FE9E5473" collectingDate="2002-12-10" collectionCode="UMC" country="Thailand" elevation="1100" isTailEnumeration="true" latitude="18.8" location="Sai Yoi Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="23" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
same locality except for,
<date id="FFAB1017FFFD225C08607CF2FB1D5596" box="[1079,1218,1745,1771]" pageId="23" pageNumber="584" value="2002-12-10">
<collectingDate id="EFEFE9FFFFFD225C08607CF2FB1D5596" box="[1079,1218,1745,1771]" pageId="23" pageNumber="584" value="2002-12-10">10.XII.2002</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFD225C09D97CF2FF3C5473" pageId="23" pageNumber="584" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C0CA47CD7FEE65473" box="[243,313,1780,1806]" pageId="23" pageNumber="584">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFD225C0D197CD6FABD544C" collectingDate="2013-04-07" collectionCode="KKU" collectorName="K. Rattanachan" country="Thailand" county="Kongma-Maesa Biosphere Reserve" elevation="1213" isTailEnumeration="true" latitude="18.812939" location="Mueang" longLatPrecision="1" longitude="98.90622" municipality="Suthep Subdistrict" pageId="23" pageNumber="584" specimenCount="3" specimenCount-female="3" stateProvince="Chiang Mai">
same locality except for,
<collectingCounty id="62CB4E5BFFFD225C0E3C7CD6FC225472" box="[619,1021,1780,1807]" pageId="23" pageNumber="584">Kongma-Maesa Biosphere Reserve</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFD225C085E7CD7FB045473" box="[1033,1243,1780,1806]" pageId="23" pageNumber="584">Suthep Subdistrict</collectingMunicipality>
,
<location id="8ECA600CFFFD225C08B07CD6FA9C5472" LSID="urn:lsid:plazi:treatment:03BC87C1FFFD22530CC07997FCD35015:8ECA600CFFFD225C08B07CD6FA9C5472" box="[1255,1347,1781,1807]" country="Thailand" county="Kongma-Maesa Biosphere Reserve" latitude="18.812939" longLatPrecision="1" longitude="98.90622" municipality="Suthep Subdistrict" name="Mueang" pageId="23" pageNumber="584" stateProvince="Chiang Mai">Mueang</location>
<collectingRegion id="49D1F835FFFD225C091C7CD7FEFD544C" country="Thailand" name="Chiang Mai" pageId="23" pageNumber="584">Chiang Mai District</collectingRegion>
,
<geoCoordinate id="EE215010FFFD225C0D7A7D35FE06544C" box="[301,473,1814,1841]" degrees="18" direction="north" minutes="48" orientation="latitude" pageId="23" pageNumber="584" precision="1" seconds="46.58" value="18.812939">18°48’46.58”N</geoCoordinate>
<geoCoordinate id="EE215010FFFD225C0DB77D35FD53544C" box="[480,652,1814,1841]" degrees="98" direction="east" minutes="54" orientation="longitude" pageId="23" pageNumber="584" precision="1" seconds="22.39" value="98.90622">98°54’22.39”E</geoCoordinate>
,
<elevation id="0038D1E4FFFD225C0ECF7D34FCF6544C" box="[664,809,1815,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.2129999999999999" pageId="23" pageNumber="584" unit="m" value="1213.0">
<quantity id="4CED9B32FFFD225C0ECF7D34FD32544C" box="[664,749,1815,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.2129999999999999" pageId="23" pageNumber="584" unit="m" value="1213.0">1213 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFD225C0F627D34FC79544C" box="[821,934,1815,1841]" pageId="23" pageNumber="584" value="2013-04-07">
<collectingDate id="EFEFE9FFFFFD225C0F627D34FC79544C" box="[821,934,1815,1841]" pageId="23" pageNumber="584" value="2013-04-07">7.IV.2013</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFD225C0FB57D34FB4E544C" box="[994,1169,1815,1841]" pageId="23" pageNumber="584">K. Rattanachan</collectorName>
,
<specimenCount id="9D13FD5EFFFD225C08CB7D34FAD7544C" box="[1180,1288,1815,1841]" pageId="23" pageNumber="584" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFFD225C09407D34FA85544C" box="[1303,1370,1815,1841]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="23" pageNumber="584">KKU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFD22530C907D1AFE5E512C" lastPageId="24" lastPageNumber="585" pageId="23" pageNumber="584" type="diagnosis">
<paragraph id="8BAA36D7FFFD22530C907D1AFDA152BB" blockId="23.[151,1437,1156,2015]" lastBlockId="24.[151,1437,151,872]" lastPageId="24" lastPageNumber="585" pageId="23" pageNumber="584">
<emphasis id="B961EAC5FFFD225C0C907D1AFE9D542E" bold="true" box="[199,322,1849,1875]" pageId="23" pageNumber="584">Diagnosis.</emphasis>
Size: Male:
<quantity id="4CED9B32FFFD225C0D877D1AFDAB5429" box="[464,628,1849,1876]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.27" metricValueMin="1.18" pageId="23" pageNumber="584" unit="mm" value="12.25" valueMax="12.7" valueMin="11.8">11.8–12.7 mm</quantity>
, female:
<quantity id="4CED9B32FFFD225C0E8C7D1AFCAC5429" box="[731,883,1849,1876]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.25" metricValueMin="0.97" pageId="23" pageNumber="584" unit="mm" value="11.1" valueMax="12.5" valueMin="9.7">9.7–12.5 mm</quantity>
; body form in lateral view weakly dorsoventrally convex; elytral glabrous region of male (
<figureCitation id="132E2A52FFFD225C0E367D7FFD1A540B" box="[609,709,1884,1910]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="23" pageNumber="584">Figs. 2H</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0E867D7FFCDC540B" box="[721,771,1884,1910]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="23" pageNumber="584">12A</figureCitation>
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to considerably wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; female elytral glabrous region (
<figureCitation id="132E2A52FFFD225C0F247D82FC0854C6" box="[883,983,1953,1979]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="23" pageNumber="584">Figs. 3H</figureCitation>
,
<figureCitation id="132E2A52FFFD225C0FB57D82FBCE54C6" box="[994,1041,1953,1979]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="23" pageNumber="584">12E</figureCitation>
) dissimilar to male, broad for nearentire length occupying 10/11 elytral width, lateral margins straight for 3/4 length creating parallel-sided appearance, abruptly acuminate medially in apical 1/4, posterolateral corners produced into points, posterior margin evenly convex toward suture; in lateral view width of pubescence margin at humeral region 2x as broad as that extending posterior; elytral apex of both sexes similar, obliquely truncate without apicolateral sinuation, epipleural angle distinct, weakly rounded; male protibia (
<figureCitation id="132E2A52FFF222530E007ADCFD725264" box="[599,685,255,281]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="24" pageNumber="585">Fig. 4H</figureCitation>
) wedge-shaped, without basal constriction, protibial apex obliquely truncate without prominent distolateral angle; male protarsus (
<figureCitation id="132E2A52FFF222530F1C7B02FC7D5246" box="[843,930,289,315]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="24" pageNumber="585">Fig. 4H</figureCitation>
) ca. 3/4 length of protibia, protarsomeres 1–4 mostly similar in width; mesoventrite often without medial pit; aedeagus (
<figureCitation id="132E2A52FFF222530FB37B67FBB35223" box="[996,1132,324,350]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="24" pageNumber="585">Fig. 12B–D</figureCitation>
) with median lobe weakly constricted medially, parallel-sided in apical 1/3 before acumination, acumination short, 1/5 length of median lobe, apex triangular; female reproductive tract (
<figureCitation id="132E2A52FFF222530ED77BAAFD3752DE" box="[640,744,393,419]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="24" pageNumber="585">Fig. 12G</figureCitation>
) without additional sclerotized structure between gonocoxae (
<figureCitation id="132E2A52FFF222530CF77B8FFEDC52BB" box="[160,259,428,454]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="24" pageNumber="585">Fig. 12F</figureCitation>
), fertilization duct as in
<figureCitation id="132E2A52FFF222530E447B8FFDA552BB" box="[531,634,428,454]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="24" pageNumber="585">Fig. 12H</figureCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFF222530C907BEDFE5E512C" blockId="24.[151,1437,151,872]" pageId="24" pageNumber="585">
<emphasis id="B961EAC5FFF222530C907BEDFE1B5295" bold="true" box="[199,452,462,488]" pageId="24" pageNumber="585">Differential diagnosis</emphasis>
.
<taxonomicName id="4C154D54FFF222530D877BECFD7F5294" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[464,672,463,489]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFF222530D877BECFD7F5294" box="[464,672,463,489]" italics="true" pageId="24" pageNumber="585">Patrus subapicalis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C154D54FFF222530F327BECFC085294" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[869,983,463,489]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFF222530F327BECFC085294" box="[869,983,463,489]" italics="true" pageId="24" pageNumber="585">P. apicalis</emphasis>
</taxonomicName>
, with which it was previously considered conspecific. They can be distinguished from each other by the differential diagnosis provided under
<taxonomicName id="4C154D54FFF2225309497BD1FA495176" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1310,1430,497,523]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFF2225309497BD1FA495176" box="[1310,1430,497,523]" italics="true" pageId="24" pageNumber="585">P. apicalis</emphasis>
</taxonomicName>
. Females of this species are also similar to
<taxonomicName id="4C154D54FFF222530E3D7836FD395153" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[618,742,533,558]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
<emphasis id="B961EAC5FFF222530E3D7836FD395153" box="[618,742,533,558]" italics="true" pageId="24" pageNumber="585">P. nanensis</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFF222530EBA7837FC9A5153" bold="true" box="[749,837,532,558]" pageId="24" pageNumber="585">
<taxonomicNameLabel id="A25257BEFFF222530EBA7837FC9A5153" box="[749,837,532,558]" pageId="24" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished by the differential diagnosis for the latter species.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFF222530C90787AFF0751A6" pageId="24" pageNumber="585" type="distribution">
<paragraph id="8BAA36D7FFF222530C90787AFF0751A6" blockId="24.[151,1437,151,872]" pageId="24" pageNumber="585">
<emphasis id="B961EAC5FFF222530C90787AFEBE510E" bold="true" box="[199,353,601,627]" pageId="24" pageNumber="585">Distribution.</emphasis>
<taxonomicName id="4C154D54FFF222530D3D7879FDE0510E" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[362,575,601,627]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFF222530D3D7879FDE0510E" box="[362,575,601,627]" italics="true" pageId="24" pageNumber="585">Patrus subapicalis</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="F3027647FFF222530FFC7879FC3E5109" box="[939,993,602,628]" name="Laos" pageId="24" pageNumber="585">Laos</collectingCountry>
and was previously known only from there. This is the first record of
<taxonomicName id="4C154D54FFF222530E55785EFD4151EB" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[514,670,636,662]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFF222530E55785EFD4151EB" box="[514,670,636,662]" italics="true" pageId="24" pageNumber="585">P. subapicalis</emphasis>
</taxonomicName>
from
<collectingCountry id="F3027647FFF222530EB3785FFC9451EB" box="[740,843,636,662]" name="Thailand" pageId="24" pageNumber="585">Thailand</collectingCountry>
. Within
<collectingCountry id="F3027647FFF222530FFC785FFBCE51EB" box="[939,1041,636,662]" name="Thailand" pageId="24" pageNumber="585">Thailand</collectingCountry>
, it is known only from the Thanon Thong Chai mountain range (
<figureCitation id="132E2A52FFF222530DBC78BCFD8C51C4" box="[491,595,671,697]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="24" pageNumber="585">Fig. 14B</figureCitation>
), where it has been collected sympatrically with
<taxonomicName id="4C154D54FFF2225308DE78BCFB2751C4" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[1161,1272,671,697]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFF2225308DE78BCFB2751C4" box="[1161,1272,671,697]" italics="true" pageId="24" pageNumber="585">P. garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFF22253095678BCFA8251C4" bold="true" box="[1281,1373,671,697]" pageId="24" pageNumber="585">
<taxonomicNameLabel id="A25257BEFFF22253095678BCFA8251C4" box="[1281,1373,671,697]" pageId="24" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132E2A52FFF22253093878BCFF1451A6" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="24" pageNumber="585">Fig. 13A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFF222530C9078C7FCD35015" pageId="24" pageNumber="585" type="discussion">
<paragraph id="8BAA36D7FFF222530C9078C7FCD35015" blockId="24.[151,1437,151,872]" pageId="24" pageNumber="585">
<emphasis id="B961EAC5FFF222530C9078C7FE935183" bold="true" box="[199,332,740,766]" pageId="24" pageNumber="585">Discussion.</emphasis>
<taxonomicName id="4C154D54FFF222530D0D78C6FDEC5183" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[346,563,740,766]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFF222530D0D78C6FDEC5183" box="[346,563,740,766]" italics="true" pageId="24" pageNumber="585">Patrus subapicalis</emphasis>
</taxonomicName>
clearly exhibits species-level morphological differences from
<taxonomicName id="4C154D54FFF22253097678C6FA425183" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1313,1437,740,766]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFF22253097678C6FA425183" box="[1313,1437,740,766]" italics="true" pageId="24" pageNumber="585">P. apicalis</emphasis>
</taxonomicName>
particularly in terms of the median lobe of the aedeagus. The results of the phylogenetic analyses also clearly support these two species as being distinct from one another (
<figureCitation id="132E2A52FFF222530F36790AFC7A503E" box="[865,933,809,835]" captionStart="FIGURE 1" captionStartId="4.[151,260,1414,1440]" captionTargetBox="[165,1277,439,1361]" captionTargetId="figure-96@4.[159,1428,415,1390]" captionTargetPageId="4" captionText="FIGURE 1. Result of the single-locus Generalized Mixed Yule Coalescent (GMYC) species delimitation analysis using COI. Black branches denote interspecific-level divergences while red branches indicate intraspecific coalescence events. Values above nodes show posterior probability support for the ultrametric COI tree generated through BEAST used in the GMYC species delimitation analysis (see supplementary Fig. S2). Farthest outgroup not shown." figureDoi="http://doi.org/10.5281/zenodo.5042658" httpUri="https://zenodo.org/record/5042658/files/figure.png" pageId="24" pageNumber="585">Fig. 1</figureCitation>
and supplementary
<figureCitation id="132E2A52FFF2225308D8790AFAD3503E" box="[1167,1292,809,835]" captionStart="FIGURE 1" captionStartId="4.[151,260,1414,1440]" captionTargetBox="[165,1277,439,1361]" captionTargetId="figure-96@4.[159,1428,415,1390]" captionTargetPageId="4" captionText="FIGURE 1. Result of the single-locus Generalized Mixed Yule Coalescent (GMYC) species delimitation analysis using COI. Black branches denote interspecific-level divergences while red branches indicate intraspecific coalescence events. Values above nodes show posterior probability support for the ultrametric COI tree generated through BEAST used in the GMYC species delimitation analysis (see supplementary Fig. S2). Farthest outgroup not shown." figureDoi="http://doi.org/10.5281/zenodo.5042658" httpUri="https://zenodo.org/record/5042658/files/figure.png" pageId="24" pageNumber="585">Figs. S1–S</figureCitation>
<figureCitation id="132E2A52FFF22253095B790AFAC3503E" box="[1292,1308,809,835]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="24" pageNumber="585">2</figureCitation>
). For these reasons, we here elevate
<taxonomicName id="4C154D54FFF222530DF9796DFD95501A" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[430,586,845,871]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="585" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFF222530DF9796DFD95501A" box="[430,586,845,871]" italics="true" pageId="24" pageNumber="585">P. subapicalis</emphasis>
</taxonomicName>
to species status.
</paragraph>
</subSubSection>
</treatment>
</document>
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