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<document id="2A0280545CFA8B29F0B6E627714962E6" ID-DOI="10.11646/zootaxa.3619.1.5" ID-GBIF-Dataset="2b59518c-4b07-4628-bde3-40b67bb154ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="220118" ID-ZooBank="CA83F7A5-7849-43D4-BE3A-778937872F10" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460577212236" checkinUser="plazi" docAuthor="Derka, Tomáš &amp; Figueroa, J. Manuel Tierno De" docDate="2013" docId="03BC87B1FF96C45FFF556F62FC1A6AB2" docLanguage="en" docName="zt03619p078.pdf.imf" docOrigin="Zootaxa 3619 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enderleina khazeni Derka &amp; Figueroa, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="76" masterDocId="FF85FFC9FF97C45EFFC26D52FF876D46" masterDocTitle="Enderleina khazeni n. sp. (Plecoptera, Perlidae), a new stonefly from Venezuelan Guayana" masterLastPageNumber="78" masterPageNumber="75" pageNumber="76" updateTime="1698323481542" updateUser="plazi">
<mods:mods id="40108CBB50EFAA685AE27A4A3184E0CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9CFE9140E6A117EB3CE5FB9B5333F16">
<mods:title id="F88D816C452E0EFAF3E24A521BC3236D">Enderleina khazeni n. sp. (Plecoptera, Perlidae), a new stonefly from Venezuelan Guayana</mods:title>
</mods:titleInfo>
<mods:name id="4046AE617AB3A6C3A9710ABD5F89262F" type="personal">
<mods:role id="9923C38377319055E3713DD23D717161">
<mods:roleTerm id="FFEA2898C8DAF5E53D9379E2904AD601">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B8D72D661EFACDC9B776E148508F431">Derka, Tomáš</mods:namePart>
</mods:name>
<mods:name id="9720C35E81A28A495DF3FC46AC6B6102" type="personal">
<mods:role id="D0DE48926AE9681E82A9322A35F68AD7">
<mods:roleTerm id="A80E2014E7AB6A702B622A878F207BAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="34EAFF7839F116067FF24BDC39247ECE">Figueroa, J. Manuel Tierno De</mods:namePart>
</mods:name>
<mods:typeOfResource id="3ED5FA0FE80FD3F1BA910DCEE6E31D8C">text</mods:typeOfResource>
<mods:relatedItem id="768B724AC3DB8FF073F06B17DBB3DD48" type="host">
<mods:titleInfo id="5317C590A87439E5F17B69E61321A6C8">
<mods:title id="C1B28994AD2C149E4D86B0E4F0E6C583">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="01DC4285F790E6ABCE60F35117A8FB8E">
<mods:date id="185451A2E9B15413A6821AE28C463D49">2013</mods:date>
<mods:detail id="3CBF7C092519E217A5577A657ABF45EB" type="volume">
<mods:number id="8722D1CCBE14685415570042C8FA5C8D">3619</mods:number>
</mods:detail>
<mods:detail id="EB79EDAC063AFD01216E546A22FA4F64" type="issue">
<mods:number id="431D7DFA8862BC09A0EA094A534FE131">1</mods:number>
</mods:detail>
<mods:extent id="BCB9C77F2B6B233F16678BC289BDA32E" unit="page">
<mods:start id="8CFB5B74F480200403764A59A9797BEE">75</mods:start>
<mods:end id="235489E08CC854F96FFEA00AD772EA03">78</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="775F31EF819F90AEA97F6B734881675C">journal article</mods:classification>
<mods:identifier id="3CCA720D31D7E868CBCCDF4ABE7830A4" type="DOI">10.11646/zootaxa.3619.1.5</mods:identifier>
<mods:identifier id="ECE461E47D956C5699DD054D3A701A29" type="GBIF-Dataset">2b59518c-4b07-4628-bde3-40b67bb154ba</mods:identifier>
<mods:identifier id="9E18707C99C4F4F68ABDA12429C509FC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="93013928DB3B1B62B3D99CDDBAA72567" type="Zenodo-Dep">220118</mods:identifier>
<mods:identifier id="43E6A1F5E90F08C86713ED051433BDDF" type="ZooBank">CA83F7A5-7849-43D4-BE3A-778937872F10</mods:identifier>
</mods:mods>
<treatment id="03BC87B1FF96C45FFF556F62FC1A6AB2" ID-DOI="http://doi.org/10.5281/zenodo.6145946" ID-GBIF-Taxon="120682238" ID-Zenodo-Dep="6145946" LSID="urn:lsid:plazi:treatment:03BC87B1FF96C45FFF556F62FC1A6AB2" httpUri="http://treatment.plazi.org/id/03BC87B1FF96C45FFF556F62FC1A6AB2" lastPageNumber="76" pageId="1" pageNumber="76">
<subSubSection id="C30F652CFF96C45FFF556F62FE8A6F2F" pageId="1" pageNumber="76" type="nomenclature">
<paragraph id="8BAA36A7FF96C45FFF556F62FD7A6F01" blockId="1.[151,765,559,617]" box="[151,765,559,584]" pageId="1" pageNumber="76">
<heading id="D0E281CBFF96C45FFF556F62FD7A6F01" bold="true" box="[151,765,559,584]" fontSize="10" level="2" pageId="1" pageNumber="76" reason="2">
<emphasis id="B961EAB5FF96C45FFF556F62FD7A6F01" bold="true" box="[151,765,559,584]" pageId="1" pageNumber="76">
<taxonomicName id="4C154D24FF96C45FFF556F62FEE96F0E" ID-CoL="39QVT" box="[151,366,560,584]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni" status="sp. nov.">
<emphasis id="B961EAB5FF96C45FFF556F62FEE96F0E" bold="true" box="[151,366,560,584]" italics="true" pageId="1" pageNumber="76">Enderleina khazeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257CEFF96C45FFEB76F7DFE366F01" box="[373,433,559,583]" pageId="1" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
Derka &amp; Tierno de Figueroa
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF556F00FE8A6F2F" blockId="1.[151,765,559,617]" box="[151,269,594,617]" pageId="1" pageNumber="76">
<figureCitation id="132E2A22FF96C45FFF556F00FF5C6F2F" box="[151,219,594,617]" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Figs. 1</figureCitation>
a–d.
</paragraph>
</subSubSection>
<subSubSection id="C30F652CFF96C45FFF556FC6FC9D6E3A" pageId="1" pageNumber="76" type="materials_examined">
<paragraph id="8BAA36A7FF96C45FFF556FC6FE036E52" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF556FC6FEC76FEA" bold="true" box="[151,320,660,684]" pageId="1" pageNumber="76">
<typeStatus id="54AE8805FF96C45FFF556FC6FF486FEA" box="[151,207,660,684]" pageId="1" pageNumber="76">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AE8805FF96C45FFE886FC7FE366FEA" box="[330,433,661,684]" pageId="1" pageNumber="76" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3027637FF96C45FFE7C6FC7FDA96FEA" box="[446,558,661,684]" name="Venezuela" pageId="1" pageNumber="76">Venezuela</collectingCountry>
, Bolívar Province, Gran Sabana, Río Tarota, below the road to
<collectingCountry id="F3027637FF96C45FFB356FC7FAFE6FEA" box="[1271,1401,661,684]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="76">Santa Elena</collectingCountry>
de Uairén,
<quantity id="4CED9B42FF96C45FFF336FEAFECA6F89" box="[241,333,696,719]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="1" pageNumber="76" unit="m" value="1320.0">1,320 m</quantity>
a.s.l.,
<geoCoordinate id="EE215060FF96C45FFE596FEAFDB16F89" box="[411,566,696,719]" direction="north" orientation="latitude" pageId="1" pageNumber="76" precision="15" value="5.8158336">5º 48’ 57’’ N</geoCoordinate>
,
<geoCoordinate id="EE215060FF96C45FFD876FEAFD646F89" box="[581,739,696,719]" direction="west" orientation="longitude" pageId="1" pageNumber="76" precision="15" value="-61.41889">61º 25’ 8’’ W</geoCoordinate>
,
<date id="FFAB1067FF96C45FFD306FEAFCF56F89" box="[754,882,696,719]" pageId="1" pageNumber="76" value="2011-11-19">19-XI-2011</date>
,
<specimenCount id="9D13FD2EFF96C45FFC426FEAFC576F89" box="[896,976,696,719]" pageId="1" pageNumber="76" type="male">1 male</specimenCount>
, Derka leg.
<typeStatus id="54AE8805FF96C45FFB9F6FEAFB386F89" box="[1117,1215,696,719]" pageId="1" pageNumber="76" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F3027637FF96C45FFB0F6FEAFAB96F89" box="[1229,1342,696,719]" name="Venezuela" pageId="1" pageNumber="76">Venezuela</collectingCountry>
, Bolívar Province, Gran Sabana, Río Torón, above Salto Toroncito,
<quantity id="4CED9B42FF96C45FFCD36F88FCEE6FB7" box="[785,873,730,753]" metricMagnitude="3" metricUnit="m" metricValue="1.384" pageId="1" pageNumber="76" unit="m" value="1384.0">1,384 m</quantity>
a. s. l.,
<geoCoordinate id="EE215060FF96C45FFC7C6F88FBCA6FB7" box="[958,1101,730,753]" direction="north" orientation="latitude" pageId="1" pageNumber="76" precision="15" value="5.7269444">5º 43’ 37’’ N</geoCoordinate>
,
<geoCoordinate id="EE215060FF96C45FFB9B6F88FB7E6FB7" box="[1113,1273,730,753]" direction="west" orientation="longitude" pageId="1" pageNumber="76" precision="15" value="-61.609722">61º 36’ 35’’ W</geoCoordinate>
,
<date id="FFAB1067FF96C45FFAC66F88FA046FB7" box="[1284,1411,730,753]" pageId="1" pageNumber="76" value="2011-11-20">20-XI-2011</date>
,
<specimenCount id="9D13FD2EFF96C45FFA4D6F88FF496E52" pageId="1" pageNumber="76" type="male">1 male</specimenCount>
, Semelbauer leg.
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF056E72FB0A6E1F" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
The
<typeStatus id="54AE8805FF96C45FFF3C6E72FEDC6E71" box="[254,347,800,823]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
and
<typeStatus id="54AE8805FF96C45FFE5C6E72FE7D6E71" box="[414,506,800,823]" pageId="1" pageNumber="76" type="paratype">paratype</typeStatus>
are deposited in the Collection of the
<emphasis id="B961EAB5FF96C45FFC0F6E72FA9F6E71" box="[973,1304,800,823]" italics="true" pageId="1" pageNumber="76">Musée Cantonal de Zoologie</emphasis>
(Lausanne,
<collectingCountry id="F3027637FF96C45FFF556E10FE9F6E1F" box="[151,280,834,857]" name="Switzerland" pageId="1" pageNumber="76">Switzerland</collectingCountry>
) with the registration numbers, 69029 for the
<typeStatus id="54AE8805FF96C45FFCC06E10FCD86E1F" box="[770,863,834,857]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
and 69030 for the
<typeStatus id="54AE8805FF96C45FFBE86E10FB0E6E1F" box="[1066,1161,834,857]" pageId="1" pageNumber="76" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF056E36FC9D6E3A" blockId="1.[151,1437,660,2036]" box="[199,794,868,892]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF056E36FED96E3A" bold="true" box="[199,350,868,892]" pageId="1" pageNumber="76">
<typeStatus id="54AE8805FF96C45FFF056E36FF786E3A" box="[199,255,868,892]" pageId="1" pageNumber="76">Type</typeStatus>
locality.
</emphasis>
Río Tarota, Bolívar Province,
<collectingCountry id="F3027637FF96C45FFD676E37FC916E3A" box="[677,790,869,892]" name="Venezuela" pageId="1" pageNumber="76">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F652CFF96C45FFF056ED5FDDA6E87" pageId="1" pageNumber="76" type="etymology">
<paragraph id="8BAA36A7FF96C45FFF056ED5FDDA6E87" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF056ED5FEC06ED9" bold="true" box="[199,327,903,927]" pageId="1" pageNumber="76">Etymology.</emphasis>
The species is named after Frank Khazen, a great person, Gran Sabana expert and friend of TD, who unexpectedly passed away in
<date id="FFAB1067FF96C45FFE166EF8FDDE6E87" box="[468,601,938,961]" pageId="1" pageNumber="76" value="2012-03">March 2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F652CFF96C45FFF056E9EFC1A6AB2" pageId="1" pageNumber="76" type="description">
<paragraph id="8BAA36A7FF96C45FFF056E9EFC3869BF" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF056E9EFED56EA2" bold="true" box="[199,338,972,996]" pageId="1" pageNumber="76">Description.</emphasis>
Male
<typeStatus id="54AE8805FF96C45FFE556E9FFE736EA2" box="[407,500,973,996]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
(
<figureCitation id="132E2A22FF96C45FFDC66E9FFDC66EA2" box="[516,577,973,996]" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
a–d): Total length:
<quantity id="4CED9B42FF96C45FFCD06E9FFCE76EA2" box="[786,864,973,996]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="76" unit="mm" value="12.0">12 mm</quantity>
, Forewing:
<quantity id="4CED9B42FF96C45FFC206E9FFBC26EA2" box="[994,1093,973,996]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="76" unit="mm" value="12.6">12.6 mm</quantity>
. General colour brownish black except the prothorax (both dorsally and ventrally) that is orange in the living specimen and yellow once preserved (
<figureCitation id="132E2A22FF96C45FFAB26EA2FF25696F" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
a). Antennae black, lighter proximally. Three ocelli, the anterior one smaller. Mouth parts light brown. Wings infuscated with brown. Legs, abdomen and cerci black. Cerci with long and abundant setae. Abdomen hairy (
<figureCitation id="132E2A22FF96C45FFB066967FA87690A" box="[1220,1280,1077,1100]" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
b–c). Sternum IX wider than long (1.25 wide/long) with long dark bristles and a pale, rounded and flattened hammer (
<quantity id="4CED9B42FF96C45FFAF5690AFA1B6929" box="[1335,1436,1112,1135]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="76" unit="mm" value="0.15">0.15 mm</quantity>
diameter) (
<figureCitation id="132E2A22FF96C45FFECC6928FECA69D7" box="[270,333,1146,1169]" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
c). Hammer set on a dark band with two elongated, adjacent, pale areas. Paraproct laterally pointed. Aedeagus with a median, triangular shaped sclerite and a pair of long, slender basolateral spines (
<figureCitation id="132E2A22FF96C45FFB0269CFFB7969F2" box="[1216,1278,1181,1204]" captionStart="FIGURE 1" captionStartId="2.[151,255,1920,1944]" captionTargetBox="[219,1344,207,1919]" captionTargetId="figure@2.[215,1370,194,1919]" captionTargetPageId="2" captionText="FIGURE 1. Enderleina khazeni n. sp. Male; a) habitus of a preserved individual; b) terminalia, dorsal; c) terminalia, ventral (slightly oblique), showing the IX sternum; d) penial armature and basal part of the aedeagus (tubular membranous part not shown)." httpUri="https://zenodo.org/record/220119/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
d). Each spine bearing ventrally an enlarged, subapical fan-shaped structure which terminates in a blade-like structure extending to midlength of the median sclerite. Aedeagus with numerous spine-like setae.
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF056857FE836827" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
The male
<typeStatus id="54AE8805FF96C45FFEFF6857FE1E685A" box="[317,409,1285,1308]" pageId="1" pageNumber="76" type="paratype">paratype</typeStatus>
differs from the
<typeStatus id="54AE8805FF96C45FFDA66857FD46685A" box="[612,705,1285,1308]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
slightly in size (total length:
<quantity id="4CED9B42FF96C45FFBDF6857FBF0685A" box="[1053,1143,1285,1308]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="1" pageNumber="76" unit="mm" value="9.3">9.3 mm</quantity>
, forewing:
<quantity id="4CED9B42FF96C45FFB3E6857FAE6685A" box="[1276,1377,1285,1308]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="1" pageNumber="76" unit="mm" value="12.4">12.4 mm</quantity>
) and coloration (mouth parts and ventral part of the head pale brown-orange, and body slightly paler ventrally than the
<typeStatus id="54AE8805FF96C45FFF556818FF7E6827" box="[151,249,1354,1377]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF05683EFD746A01" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF05683EFEBE68C2" bold="true" box="[199,313,1388,1412]" pageId="1" pageNumber="76">Affinities:</emphasis>
The male of
<taxonomicName id="4C154D24FF96C45FFE14683FFDCD68C2" box="[470,586,1389,1412]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFE14683FFDCD68C2" box="[470,586,1389,1412]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
can be clearly distinguished from the males of
<emphasis id="B961EAB5FF96C45FFBA4683FFACE68C2" box="[1126,1353,1389,1412]" italics="true" pageId="1" pageNumber="76">
<taxonomicName id="4C154D24FF96C45FFBA4683FFB6268C2" box="[1126,1253,1389,1412]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">E. preclara</taxonomicName>
,
<taxonomicName id="4C154D24FF96C45FFB36683FFACE68C2" box="[1268,1353,1389,1412]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="yano">E. yano</taxonomicName>
</emphasis>
and
<taxonomicName id="4C154D24FF96C45FFA47683FFF7968E1" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFA47683FFF7968E1" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
mainly by the shape of the penial sclerites and of the paraprocts (
<bibRefCitation id="EF844B56FF96C45FFC7D68C2FBC268E1" author="Jewett" box="[959,1093,1424,1447]" pageId="1" pageNumber="76" refString="Jewett, S. G. Jr. (1960) Notes and descriptions concerning Brazilian stoneflies (Plecoptera). Arquivos do Museu Nacional, 50, 167 - 183." type="journal article" year="1960">Jewett 1960</bibRefCitation>
;
<bibRefCitation id="EF844B56FF96C45FFB9268C2FB4068E1" author="Stark" box="[1104,1223,1424,1447]" pageId="1" pageNumber="76" refString="Stark, B. P. (1989) The genus Enderleina (Plecoptera: Perlidae). Aquatic Insects, 11 (3), 153 - 160. http: // dx. doi. org / 10.1080 / 01650428909361364" type="journal article" year="1989">Stark 1989</bibRefCitation>
; Ribeiro-Ferreira &amp;
<bibRefCitation id="EF844B56FF96C45FFF5568E0FE99688F" author="Rafael" box="[151,286,1458,1481]" pageId="1" pageNumber="76" refString="Ribeiro, J. M. F. &amp; Rafael, J. A. (2005) A key to adult Amazonian stonefly genera with new geographical records of Enderleina Jewett for Brazil and first description of the E. froehlichi Ribeiro-Ferreira female (Insecta: Plecoptera). Zootaxa, 1096, 61 - 67." type="journal article" year="2005">Rafael 2005</bibRefCitation>
). The spines are narrowed in the apical third of the penial sclerites in
<taxonomicName id="4C154D24FF96C45FFBD368E1FB0D688C" box="[1041,1162,1459,1482]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFBD368E1FB0D688C" box="[1041,1162,1459,1482]" italics="true" pageId="1" pageNumber="76">E. preclara</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C154D24FF96C45FFAD668E1FAE0688C" box="[1300,1383,1459,1482]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="yano">
<emphasis id="B961EAB5FF96C45FFAD668E1FAE0688C" box="[1300,1383,1459,1482]" italics="true" pageId="1" pageNumber="76">E. yano</emphasis>
</taxonomicName>
they are more gradually tapered, in
<taxonomicName id="4C154D24FF96C45FFE276887FDEC68AA" box="[485,619,1493,1516]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFE276887FDEC68AA" box="[485,619,1493,1516]" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
they are considerably stronger, and in
<taxonomicName id="4C154D24FF96C45FFBD36887FB0668AA" box="[1041,1153,1493,1516]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFBD36887FB0668AA" box="[1041,1153,1493,1516]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
they have a characteristic shape ventrally extending as an enlarged fan-shaped structure, dorsally terminating in a blade-shaped structure. Moreover, dorsally the spines in
<taxonomicName id="4C154D24FF96C45FFDC26B49FDF66B74" box="[512,625,1563,1586]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFDC26B49FDF66B74" box="[512,625,1563,1586]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
reach near the apex of the median sclerite, whereas in
<taxonomicName id="4C154D24FF96C45FFB106B49FACA6B74" box="[1234,1357,1563,1586]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFB106B49FACA6B74" box="[1234,1357,1563,1586]" italics="true" pageId="1" pageNumber="76">E. preclara</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D24FF96C45FFA476B49FF796B12" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFA476B49FF796B12" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
these structures are considerably shorter. The paraprocts are laterally pointed in
<taxonomicName id="4C154D24FF96C45FFB556B6FFA8B6B12" box="[1175,1292,1597,1620]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFB556B6FFA8B6B12" box="[1175,1292,1597,1620]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
, not slightly notched at the tip as in
<taxonomicName id="4C154D24FF96C45FFE526B32FE666B31" box="[400,481,1632,1655]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="yano">
<emphasis id="B961EAB5FF96C45FFE526B32FE666B31" box="[400,481,1632,1655]" italics="true" pageId="1" pageNumber="76">E. yano</emphasis>
</taxonomicName>
, they lack a subapical denticle as in
<taxonomicName id="4C154D24FF96C45FFCA86B32FC686B31" box="[874,1007,1632,1655]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFCA86B32FC686B31" box="[874,1007,1632,1655]" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
, and they are not bulbous at the apex as in
<taxonomicName id="4C154D24FF96C45FFF716BD1FEAB6BDC" box="[179,300,1667,1690]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFF716BD1FEAB6BDC" box="[179,300,1667,1690]" italics="true" pageId="1" pageNumber="76">E. preclara</emphasis>
</taxonomicName>
.
<taxonomicName id="4C154D24FF96C45FFEF86BD1FD8E6BDC" box="[314,521,1667,1690]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFEF86BD1FD8E6BDC" box="[314,521,1667,1690]" italics="true" pageId="1" pageNumber="76">Enderleina khazeni</emphasis>
</taxonomicName>
can be also distinguished from
<taxonomicName id="4C154D24FF96C45FFCA36BD1FC346BDC" box="[865,947,1667,1690]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="yano">
<emphasis id="B961EAB5FF96C45FFCA36BD1FC346BDC" box="[865,947,1667,1690]" italics="true" pageId="1" pageNumber="76">E. yano</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D24FF96C45FFC2A6BD1FBEA6BDC" box="[1000,1133,1667,1690]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFC2A6BD1FBEA6BDC" box="[1000,1133,1667,1690]" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
by their coloration:
<taxonomicName id="4C154D24FF96C45FFA886BD1FA1B6BDC" box="[1354,1436,1667,1690]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="yano">
<emphasis id="B961EAB5FF96C45FFA886BD1FA1B6BDC" box="[1354,1436,1667,1690]" italics="true" pageId="1" pageNumber="76">E. yano</emphasis>
</taxonomicName>
has a narrow orange V-mark connecting the antennal bases and anterior ocellus, and the basal half of the femora are yellow (according to
<bibRefCitation id="EF844B56FF96C45FFEBB6B9AFE776B99" author="Stark" box="[377,496,1736,1759]" pageId="1" pageNumber="76" refString="Stark, B. P. (1989) The genus Enderleina (Plecoptera: Perlidae). Aquatic Insects, 11 (3), 153 - 160. http: // dx. doi. org / 10.1080 / 01650428909361364" type="journal article" year="1989">Stark 1989</bibRefCitation>
), and
<taxonomicName id="4C154D24FF96C45FFDEC6B9AFD356B99" box="[558,690,1736,1759]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="froehlichi">
<emphasis id="B961EAB5FF96C45FFDEC6B9AFD356B99" box="[558,690,1736,1759]" italics="true" pageId="1" pageNumber="76">E. froehlichi</emphasis>
</taxonomicName>
has a pale brown body.
<taxonomicName id="4C154D24FF96C45FFC706B9AFB0E6B99" box="[946,1161,1736,1759]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFC706B9AFB0E6B99" box="[946,1161,1736,1759]" italics="true" pageId="1" pageNumber="76">Enderleina preclara</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C154D24FF96C45FFAD06B9AFA066B99" box="[1298,1409,1736,1759]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFAD06B9AFA066B99" box="[1298,1409,1736,1759]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
in colour, but the latter has two large pale areas laterally on sternum IX, not mentioned or illustrated by
<bibRefCitation id="EF844B56FF96C45FFB0A6BB8FADD6A47" author="Jewett" box="[1224,1370,1770,1793]" pageId="1" pageNumber="76" refString="Jewett, S. G. Jr. (1960) Notes and descriptions concerning Brazilian stoneflies (Plecoptera). Arquivos do Museu Nacional, 50, 167 - 183." type="journal article" year="1960">Jewett (1960)</bibRefCitation>
in the description of
<taxonomicName id="4C154D24FF96C45FFEF46A5FFE366A62" box="[310,433,1805,1828]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFEF46A5FFE366A62" box="[310,433,1805,1828]" italics="true" pageId="1" pageNumber="76">E. preclara</emphasis>
</taxonomicName>
. Despite the previous list of differences,
<taxonomicName id="4C154D24FF96C45FFCB86A5FFC6B6A62" box="[890,1004,1805,1828]" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="khazeni">
<emphasis id="B961EAB5FF96C45FFCB86A5FFC6B6A62" box="[890,1004,1805,1828]" italics="true" pageId="1" pageNumber="76">E. khazeni</emphasis>
</taxonomicName>
seems to be more closely related to
<taxonomicName id="4C154D24FF96C45FFA476A5FFF756A01" class="Insecta" family="Perlidae" genus="Enderleina" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="preclara">
<emphasis id="B961EAB5FF96C45FFA476A5FFF756A01" italics="true" pageId="1" pageNumber="76">E. preclara</emphasis>
</taxonomicName>
, and it is the species found in closest proximity.
</paragraph>
<paragraph id="8BAA36A7FF96C45FFF056A00FC1A6AB2" blockId="1.[151,1437,660,2036]" pageId="1" pageNumber="76">
<emphasis id="B961EAB5FF96C45FFF056A00FE006A2C" bold="true" box="[199,391,1874,1898]" pageId="1" pageNumber="76">Ecological notes.</emphasis>
The
<typeStatus id="54AE8805FF96C45FFE036A00FD996A2F" box="[449,542,1874,1897]" pageId="1" pageNumber="76" type="holotype">holotype</typeStatus>
was captured by light trapping. The
<typeStatus id="54AE8805FF96C45FFC766A00FC656A2F" box="[948,994,1874,1897]" pageId="1" pageNumber="76">type</typeStatus>
locality is ca.
<quantity id="4CED9B42FF96C45FFB466A00FB3A6A2F" box="[1156,1213,1874,1897]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="76" unit="m" value="10.0">10 m</quantity>
wide and up to
<quantity id="4CED9B42FF96C45FFAB36A00FA1B6A2F" box="[1393,1436,1874,1897]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="76" unit="m" value="1.0">1 m</quantity>
deep with riffles and small cascades. The substrate consists primarily of boulders (megalithal) and stones (macro- and mesolithal) and some aquatic macrophytes. Water temperature was 21 °C,
<collectingCountry id="F3027637FF96C45FFC716ACAFC536AE9" box="[947,980,1944,1967]" name="Philippines" pageId="1" pageNumber="76">pH</collectingCountry>
= 5.89. The
<typeStatus id="54AE8805FF96C45FFB9E6ACAFB3F6AE9" box="[1116,1208,1944,1967]" pageId="1" pageNumber="76" type="paratype">paratype</typeStatus>
was captured netting on the bank of a medium sized river with stony substrate (mega- and mesolithal), water temperature 20.1°C,
<collectingCountry id="F3027637FF96C45FFAF76AE8FAD26A97" box="[1333,1365,1978,2001]" name="Philippines" pageId="1" pageNumber="76">pH</collectingCountry>
= 5.6. Both rivers are surrounded by savannas and remnants of riparian forests.
</paragraph>
</subSubSection>
</treatment>
</document>
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