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<document id="6C1BED92134E1C355E3EBAE1AB586844" ID-DOI="10.5281/zenodo.194087" ID-GBIF-Dataset="b0c9cf9c-4bbd-47d6-a260-75a3d2f16b72" ID-ISSN="1175-5326" ID-Zenodo-Dep="194087" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189116326" checkinUser="plazi" docAuthor="Liu, Qifei &amp; Yang, Ding" docDate="2010" docId="6327462BFFCCFFBEFF1BF533FA7ABA15" docLanguage="en" docName="zt02401p060.pdf" docOrigin="Zootaxa 2401" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Brithura" docType="key" docVersion="9" lastPageNumber="53" masterDocId="9F1E3E53FFCCFFBFFF8CF24EFFE6BC5A" masterDocTitle="Four new species of the genus Brithura Edwards from China, with an updated key to world species (Diptera, Tipulidae)" masterLastPageNumber="60" masterPageNumber="52" pageNumber="52" updateTime="1698236378629" updateUser="plazi">
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<mods:title id="BC2DF301A1BE507C072A1782A80C093D">Four new species of the genus Brithura Edwards from China, with an updated key to world species (Diptera, Tipulidae)</mods:title>
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<mods:namePart id="8F8069178CD4649AAA8456A042320B42">Liu, Qifei</mods:namePart>
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<mods:date id="35E938B2314203C803CF64C7EED21563">2010</mods:date>
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<treatment id="6327462BFFCCFFBEFF1BF533FA7ABA15" ID-DOI="http://doi.org/10.5281/zenodo.6210815" ID-GBIF-Taxon="124956725" ID-Zenodo-Dep="6210815" LSID="urn:lsid:plazi:treatment:6327462BFFCCFFBEFF1BF533FA7ABA15" httpUri="http://treatment.plazi.org/id/6327462BFFCCFFBEFF1BF533FA7ABA15" lastPageId="1" lastPageNumber="53" pageId="0" pageNumber="52">
<subSubSection id="A394A4B6FFCCFFBFFF1BF533FDC0BBCD" box="[151,550,1917,1943]" pageId="0" pageNumber="52" type="nomenclature">
<paragraph id="EB31F73DFFCCFFBFFF1BF533FDC0BBCD" blockId="0.[151,550,1917,1943]" box="[151,550,1917,1943]" pageId="0" pageNumber="52">
<heading id="B0794051FFCCFFBFFF1BF533FDC0BBCD" bold="true" box="[151,550,1917,1943]" fontSize="11" level="1" pageId="0" pageNumber="52" reason="1">
<emphasis id="D9FA2B2FFFCCFFBFFF1BF533FDC0BBCD" bold="true" box="[151,550,1917,1943]" pageId="0" pageNumber="52">
Key to species of genus
<taxonomicName id="2C8E8CBEFFCCFFBFFE30F533FDC0BBCD" ID-CoL="87FCJ" box="[444,550,1917,1943]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="D9FA2B2FFFCCFFBFFE30F533FDC0BBCD" bold="true" box="[444,550,1917,1943]" italics="true" pageId="0" pageNumber="52">Brithura</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A394A4B6FFCCFFBEFF1BF58BFA7ABA15" lastPageId="1" lastPageNumber="53" pageId="0" pageNumber="52" type="key">
<keyStep id="507AEF98FFCCFFBEFF1BF58BFA7DBCF5" lastPageId="1" lastPageNumber="53" pageId="0" pageNumber="52">
<paragraph id="EB31F73DFFCCFFBFFF1BF58BFA7DBB86" blockId="0.[151,1435,1989,2012]" box="[151,1435,1989,2012]" pageId="0" pageNumber="52">
<keyLead id="507F5408FFCCFFBFFF1BF58BFA7DBB86" box="[151,1435,1989,2012]" pageId="0" pageNumber="52">1. Abdominal tergites with big blackish brown spots at lateral margins .......................................................................... 2</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF2D6FA7DBCF5" blockId="1.[151,1436,152,1615]" box="[151,1435,152,175]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF2D6FA7DBCF5" box="[151,1435,152,175]" pageId="1" pageNumber="53">- Abdominal tergites without big spots at lateral margins ............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF2F6FA7ABD55" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF2F6FA7ABCB5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF2F6FA7ABCB5" pageId="1" pageNumber="53">
2. Wing dark grayish brown with some light golden areas; male hypopygium with outer gonostylus thick, laterally with three extensions at tip, medially with pointed spine at bottom (
<figureCitation id="73B5EBB8FFCDFFBEFCEDF296FC49BCB5" box="[865,943,216,239]" captionStart="FIGURES 9 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="1" pageNumber="53">Fig. 12</figureCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFC34F296FAA6BCB5" box="[952,1344,216,239]" italics="true" pageId="1" pageNumber="53">
..........................................
<taxonomicName id="2C8E8CBEFFCDFFBEFB58F296FAA6BCB5" ID-CoL="NBNW" box="[1236,1344,216,239]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">B. aureola</taxonomicName>
</emphasis>
<emphasis id="D9FA2B2FFFCDFFBEFACBF299FA7ABCB5" bold="true" box="[1351,1436,215,239]" pageId="1" pageNumber="53">
<taxonomicNameLabel id="C2C99654FFCDFFBEFACBF299FA7ABCB5" box="[1351,1436,215,239]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF2B6FA7ABD55" blockId="1.[151,1436,152,1615]" box="[151,1436,247,271]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF2B6FA7ABD55" box="[151,1436,247,271]" pageId="1" pageNumber="53">
- Wing yellowish; male hypopygium with outer spine of outer gonostylus conspicuous (
<figureCitation id="73B5EBB8FFCDFFBEFBECF2B6FB79BD55" box="[1120,1183,248,271]" captionStart="FIGURES 5 8" captionStartId="3.[151,275,889,913]" captionTargetBox="[425,1165,206,853]" captionTargetId="figure@3.[407,1180,194,868]" captionTargetPageId="3" captionText="FIGURES 5 8. Brithura shii sp. nov. Male. 5. Hypopygium, lateral view; 6. ninth tergite, dorsal view; 7. inner gonostylus, lateral view; 8. outer gonostylus, lateral view. Scale bar of 5, 6 = 0.5 mm, 7, 8 = 0.25 mm." httpUri="https://zenodo.org/record/194089/files/figure.png" pageId="1" pageNumber="53">Fig. 8</figureCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFB24F2B6FAA6BD55" box="[1192,1344,248,271]" italics="true" pageId="1" pageNumber="53">
............
<taxonomicName id="2C8E8CBEFFCDFFBEFB70F2B6FAA6BD55" ID-CoL="NBPD" box="[1276,1344,248,271]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="shii" status="sp. nov.">B. shii</taxonomicName>
</emphasis>
<emphasis id="D9FA2B2FFFCDFFBEFACBF2B9FA7ABD55" bold="true" box="[1351,1436,247,271]" pageId="1" pageNumber="53">
<taxonomicNameLabel id="C2C99654FFCDFFBEFACBF2B9FA7ABD55" box="[1351,1436,247,271]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF356FA7DBD15" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF356FA7DBD75" blockId="1.[151,1436,152,1615]" box="[151,1435,280,303]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF356FA7DBD75" box="[151,1435,280,303]" pageId="1" pageNumber="53">3. All femora with conspicuous subterminal ring............................................................................................................. 4</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF376FA7DBD15" blockId="1.[151,1436,152,1615]" box="[151,1435,312,335]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF376FA7DBD15" box="[151,1435,312,335]" pageId="1" pageNumber="53">- Posterior femur without subterminal ring................................................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF316FA7DBDD5" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF316FA7DBD35" blockId="1.[151,1436,152,1615]" box="[151,1435,344,367]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF316FA7DBD35" box="[151,1435,344,367]" pageId="1" pageNumber="53">4. Femora with tips yellow, subterminal ring brown or black .......................................................................................... 5</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF336FA7DBDD5" blockId="1.[151,1436,152,1615]" box="[151,1435,376,399]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF336FA7DBDD5" box="[151,1435,376,399]" pageId="1" pageNumber="53">- Femora with tips brown or black, subterminal ring yellow.......................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF3D6FA7ABE55" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF3D6FA7ABD95" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF3D6FA7ABD95" pageId="1" pageNumber="53">
5. Femora brownish yellow, apical ring yellow, preceded by a narrow black ring; outer gonostylus with two dissimilar spines at middle outer side (
<bibRefCitation id="8F1F8ACCFFCDFFBEFE7DF3F6FC9FBD95" author="Edwards" box="[497,889,440,463]" pageId="1" pageNumber="60" refString="Edwards, F. W. (1916) New and little-known Tipulidae, chiefly from Formosa. Annals and Magazine of Natural History, (8) 18, 245 - 269." type="journal article" year="1916" yearSuffix="p">Edwards 1916, p264, Pl. XII. fig. 12</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFC0FF3F6FB68BD95" box="[899,1166,440,463]" italics="true" pageId="1" pageNumber="53">........................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFB1EF3F6FA7ABD95" ID-CoL="NBNZ" authority="Edwards, 1916" authorityName="Edwards" authorityYear="1916" box="[1170,1436,440,463]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="D9FA2B2FFFCDFFBEFB1EF3F6FB13BD95" box="[1170,1269,440,463]" italics="true" pageId="1" pageNumber="53">B. crassa</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB70F3F6FA7ABD95" author="Edwards" box="[1276,1436,440,463]" pageId="1" pageNumber="60" refString="Edwards, F. W. (1916) New and little-known Tipulidae, chiefly from Formosa. Annals and Magazine of Natural History, (8) 18, 245 - 269." type="journal article" year="1916">Edwards, 1916</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF396FA7ABE55" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF396FA7ABE55" pageId="1" pageNumber="53">
- Femora light brown with a narrow virtually terminal yellow ring, preceded by a subequal brown ring; outer gonosty- lus not as above (
<bibRefCitation id="8F1F8ACCFFCDFFBEFEF3F3B6FD09BE55" author="Alexander" box="[383,751,504,527]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1964) New or little-known Tipulidae from eastern Asia (Diptera). LIII. Philippine Journal of Science, 93, 77 - 130." type="journal article" year="1964" yearSuffix="p">Alexander 1964, p87, Pl. I. fig. 29</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFD7BF3B6FBD1BE55" box="[759,1079,504,527]" italics="true" pageId="1" pageNumber="53">................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBB6F3B6FA7ABE55" ID-CoL="68R4J" authority="(Alexander, 1964)" authorityName="Alexander" authorityYear="1964" baseAuthorityName="Alexander" baseAuthorityYear="1964" box="[1082,1436,504,527]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="nielseniana">
<emphasis id="D9FA2B2FFFCDFFBEFBB6F3B6FB35BE55" box="[1082,1235,504,527]" italics="true" pageId="1" pageNumber="53">B. nielseniana</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB6DF3B6FA73BE55" author="Alexander" box="[1249,1429,504,527]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1964) New or little-known Tipulidae from eastern Asia (Diptera). LIII. Philippine Journal of Science, 93, 77 - 130." type="journal article" year="1964">Alexander, 1964</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF056FA7DBE15" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF056FA7DBE75" blockId="1.[151,1436,152,1615]" box="[151,1435,536,559]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF056FA7DBE75" box="[151,1435,536,559]" pageId="1" pageNumber="53">6. Abdominal tergites dark brown ................................................................................................................................... 7</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF076FA7DBE15" blockId="1.[151,1436,152,1615]" box="[151,1435,568,591]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF076FA7DBE15" box="[151,1435,568,591]" pageId="1" pageNumber="53">- Abdominal tergites not as above .................................................................................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF016FA7DBE95" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF016FA7ABED5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF016FA7ABED5" pageId="1" pageNumber="53">
7. Scutellum obscure yellow; pleura uniform in color except by pale dorsal surface of pleurotergal tubercle; outer spine of outer gonostylus large and conspicuous (
<bibRefCitation id="8F1F8ACCFFCDFFBEFD19F036FBE8BED5" author="Alexander" box="[661,1038,632,655]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1935) New or little-known Tipulidae from eastern Asia (Diptera). XXV. Philippine Journal of Science, 57, 81 - 148." type="journal article" year="1935" yearSuffix="p">Alexander 1935, p90, Pl. II. fig. 25</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFBAFF036FBD8BED5" box="[1059,1086,632,655]" italics="true" pageId="1" pageNumber="53">....</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBCFF036FA7ABED5" ID-CoL="NBP3" authority="(Alexander, 1935)" authorityName="Alexander" authorityYear="1935" baseAuthorityName="Alexander" baseAuthorityYear="1935" box="[1091,1436,632,655]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="fracticosta">
<emphasis id="D9FA2B2FFFCDFFBEFBCFF036FB35BED5" box="[1091,1235,632,655]" italics="true" pageId="1" pageNumber="53">B. fracticosta</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB6DF036FA72BED5" author="Alexander" box="[1249,1428,632,655]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1935) New or little-known Tipulidae from eastern Asia (Diptera). XXV. Philippine Journal of Science, 57, 81 - 148." type="journal article" year="1935">Alexander, 1935</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF0D6FA7DBE95" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF0D6FA7DBE95" pageId="1" pageNumber="53">- Scutellum dusky brown; pleura with variegated with silvery, additional to pale pleurotergal tubercle; outer gonosty- lus not as above ............................................................................................................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF096FA7ABF15" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF096FA7ABF55" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF096FA7ABF55" pageId="1" pageNumber="53">
8. Antennal flagellum brownish yellow; femora yellowish brown, with base yellow; outer gonostylus with a short slen- der rod at outer margin (
<bibRefCitation id="8F1F8ACCFFCDFFBEFE46F0B6FCAEBF55" author="Alexander" box="[458,840,760,783]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1929) New or little-known Tipulidae from eastern Asia (Diptera). IV. Philippine Journal of Science, 40, 317 - 348." type="journal article" year="1929" yearSuffix="p">Alexander 1929, p317, Pl II. fig. 19</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFCD7F0B6FB63BF55" box="[859,1157,760,783]" italics="true" pageId="1" pageNumber="53">............................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFB09F0B6FA7ABF55" ID-CoL="NBPC" authority="Alexander, 1929" authorityName="Alexander" authorityYear="1929" box="[1157,1436,760,783]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="sancta">
<emphasis id="D9FA2B2FFFCDFFBEFB09F0B6FB03BF55" box="[1157,1253,760,783]" italics="true" pageId="1" pageNumber="53">B. sancta</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB67F0B6FA7ABF55" author="Alexander" box="[1259,1436,760,783]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1929) New or little-known Tipulidae from eastern Asia (Diptera). IV. Philippine Journal of Science, 40, 317 - 348." type="journal article" year="1929">Alexander, 1929</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF156FA7ABF15" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF156FA7ABF15" pageId="1" pageNumber="53">
- Antennal flagellum uniformly dark brown; femora yellow; outer spine of outer gonostylus small and thin (Alexan- der 1935, Pl. II. fig. 26)
<emphasis id="D9FA2B2FFFCDFFBEFE47F176FBDBBF15" box="[459,1085,824,847]" italics="true" pageId="1" pageNumber="53">..............................................................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBC9F176FA7ABF15" ID-CoL="NBP4" authority="Alexander, 1925" authorityName="Alexander" authorityYear="1925" box="[1093,1436,824,847]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="fractistigma">
<emphasis id="D9FA2B2FFFCDFFBEFBC9F176FB02BF15" box="[1093,1252,824,847]" italics="true" pageId="1" pageNumber="53">B. fractistigma</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB67F176FA7ABF15" author="Alexander" box="[1259,1436,824,847]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1925) New or little-known Tipulidae (Diptera). XXVII. Palaearctic species. Annals and Magazine of Natural History, (9) 15, 385 - 408." type="journal article" year="1925">Alexander, 1925</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF116FA7ABF95" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF116FA7ABFD5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF116FA7ABFD5" pageId="1" pageNumber="53">
9. Abdominal tergites chestnut brown?strips of prescutum disintegrated into two lines of subrectangular brown spots; outer gonostylus laterally with three processes (
<figureCitation id="73B5EBB8FFCDFFBEFD4FF136FCF7BFD5" box="[707,785,888,911]" captionStart="FIGURES 17 20" captionStartId="7.[151,271,1127,1151]" captionTargetBox="[419,1180,282,1087]" captionTargetId="figure@7.[393,1193,269,1106]" captionTargetPageId="7" captionText="FIGURES 17 20. Brithura stigmosa sp. nov. Male. 17. Hypopygium, lateral view; 18. ninth tergite, dorsal view; 19. inner gonostylus, lateral view; 20. outer gonostylus, lateral view. Scale bar of 17, 18 = 0.5 mm, 19, 20 = 0.25 mm." httpUri="https://zenodo.org/record/194092/files/figure.png" pageId="1" pageNumber="53">Fig. 20</figureCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFC94F136FAA6BFD5" box="[792,1344,888,911]" italics="true" pageId="1" pageNumber="53">
................................................................
<taxonomicName id="2C8E8CBEFFCDFFBEFB48F136FAA6BFD5" ID-CoL="692SJ" box="[1220,1344,888,911]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="stigmosa" status="sp. nov.">B. stigmosa</taxonomicName>
</emphasis>
<emphasis id="D9FA2B2FFFCDFFBEFACBF139FA7ABFD5" bold="true" box="[1351,1436,887,911]" pageId="1" pageNumber="53">
<taxonomicNameLabel id="C2C99654FFCDFFBEFACBF139FA7ABFD5" box="[1351,1436,887,911]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF1D6FA7ABF95" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF1D6FA7ABF95" pageId="1" pageNumber="53">
- Abdominal tergites bright orange; strips of prescutum continuous; outer gonostylus outer gonostylus at near apex with a cylindrical finger-like lobe on outer margin (
<bibRefCitation id="8F1F8ACCFFCDFFBEFD67F1F6FBCBBF95" author="Alexander" box="[747,1069,952,975]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1971) New or little-known species of Asiatic Tipulidae (Diptera). IV. Transactions of the American Entomological Society, 97, 361 - 412." type="journal article" year="1971" yearSuffix="p">Alexander 1971, p378, fig. 15</bibRefCitation>
).
<taxonomicName id="2C8E8CBEFFCDFFBEFBC7F1F6FA7ABF95" ID-CoL="NBP9" authority="Alexander, 1971" authorityName="Alexander" authorityYear="1971" box="[1099,1436,952,975]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="keiliniana">
<emphasis id="D9FA2B2FFFCDFFBEFBC7F1F6FB35BF95" box="[1099,1235,952,975]" italics="true" pageId="1" pageNumber="53">B. keiliniana</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB6DF1F6FA72BF95" author="Alexander" box="[1249,1428,952,975]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1971) New or little-known species of Asiatic Tipulidae (Diptera). IV. Transactions of the American Entomological Society, 97, 361 - 412." type="journal article" year="1971">Alexander, 1971</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF196FA7DB855" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF196FA7ABFB5" blockId="1.[151,1436,152,1615]" box="[151,1436,984,1007]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF196FA7ABFB5" box="[151,1436,984,1007]" pageId="1" pageNumber="53">10. Fore and middle femora with conspicuous subterminal ring...................................................................................... 11</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF1B6FA7DB855" blockId="1.[151,1436,152,1615]" box="[151,1435,1016,1039]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF1B6FA7DB855" box="[151,1435,1016,1039]" pageId="1" pageNumber="53">- Femora without subterminal ring................................................................................................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF656FA7DB835" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF656FA7AB815" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF656FA7AB815" pageId="1" pageNumber="53">
11. Ground color of abdomen light brownish orange; outer gonostylus apically bent forward greatly with a short subtri- angular posterior process (
<bibRefCitation id="8F1F8ACCFFCDFFBEFE6DF676FCFEB815" author="Liu" box="[481,792,1080,1103]" pageId="1" pageNumber="60" refString="Liu, Q.; Yang, D. (2009) Two new species of the genus Brithura Edwards from China, with a key to world species (Diptera, Tipulidae). Zootaxa 1991, 51 - 56." type="journal article" year="2009" yearSuffix="p">Liu &amp; Yang 2009, p54, fig. 5</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFCAAF676FB32B815" box="[806,1236,1080,1103]" italics="true" pageId="1" pageNumber="53">
........................................
<taxonomicName id="2C8E8CBEFFCDFFBEFBBAF676FB32B815" ID-CoL="NBP8" authority="Liu &amp; Yang, 2009" authorityName="Liu &amp; Yang" authorityYear="2009" box="[1078,1236,1080,1103]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="jinpingensis">B. jinpingensis</taxonomicName>
</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB57F676FA7AB815" author="Liu" box="[1243,1436,1080,1103]" pageId="1" pageNumber="60" refString="Liu, Q.; Yang, D. (2009) Two new species of the genus Brithura Edwards from China, with a key to world species (Diptera, Tipulidae). Zootaxa 1991, 51 - 56." type="journal article" year="2009">Liu &amp; Yang, 2009</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF616FA7DB835" blockId="1.[151,1436,152,1615]" box="[151,1435,1112,1135]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF616FA7DB835" box="[151,1435,1112,1135]" pageId="1" pageNumber="53">- Ground color of abdomen dark brown or brownish black.......................................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF636FA7AB8B5" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF636FA7AB8F5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF636FA7AB8F5" pageId="1" pageNumber="53">
12. Femora yellow; outer gonostylus anteriorly with a long slender rod (
<bibRefCitation id="8F1F8ACCFFCDFFBEFC2CF636FB05B8D5" author="Alexander" box="[928,1251,1144,1167]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1971) New or little-known species of Asiatic Tipulidae (Diptera). IV. Transactions of the American Entomological Society, 97, 361 - 412." type="journal article" year="1971" yearSuffix="p">Alexander 1971, p378, fig. 16</bibRefCitation>
) ........................
<emphasis id="D9FA2B2FFFCDFFBEFF42F6D6FBADB8F5" box="[206,1099,1176,1199]" italics="true" pageId="1" pageNumber="53">......................................................................................................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBC1F6D6FA7AB8F5" ID-CoL="NBNX" authority="(Alexander, 1971)" authorityName="Alexander" authorityYear="1971" baseAuthorityName="Alexander" baseAuthorityYear="1971" box="[1101,1436,1176,1199]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="brulleana">
<emphasis id="D9FA2B2FFFCDFFBEFBC1F6D6FB35B8F5" box="[1101,1235,1176,1199]" italics="true" pageId="1" pageNumber="53">B. brulleana</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB6DF6D6FA72B8F5" author="Alexander" box="[1249,1428,1176,1199]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1971) New or little-known species of Asiatic Tipulidae (Diptera). IV. Transactions of the American Entomological Society, 97, 361 - 412." type="journal article" year="1971">Alexander, 1971</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF6F6FA7AB8B5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF6F6FA7AB8B5" pageId="1" pageNumber="53">
- Femora reddish-brown; outer gonostylus slightly bent forward, basal outer margin with an erect slender rod (
<bibRefCitation id="8F1F8ACCFFCDFFBEFF42F696FDB3B8B5" author="Edwards" box="[206,597,1240,1263]" pageId="1" pageNumber="60" refString="Edwards, F. W. (1916) New and little-known Tipulidae, chiefly from Formosa. Annals and Magazine of Natural History, (8) 18, 245 - 269." type="journal article" year="1916" yearSuffix="p">Edwards 1916, p263, Pl. XII. fig. 10</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFDE8F696FBBEB8B5" box="[612,1112,1240,1263]" italics="true" pageId="1" pageNumber="53">...........................................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBD6F696FA7AB8B5" ID-CoL="NBP7" authority="(Brunetti, 1913)" authorityName="Brunetti" authorityYear="1913" baseAuthorityName="Brunetti" baseAuthorityYear="1913" box="[1114,1436,1240,1263]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="imperfecta">
<emphasis id="D9FA2B2FFFCDFFBEFBD6F696FB0FB8B5" box="[1114,1257,1240,1263]" italics="true" pageId="1" pageNumber="53">B. imperfecta</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB7BF696FA72B8B5" author="Brunetti" box="[1271,1428,1240,1263]" pageId="1" pageNumber="60" refString="Brunetti, E. (1913) New and interesting Diptera from the eastern Himalayas. Records of the Indian Museum, 9, 255 - 277." type="journal article" year="1913">Brunetti, 1913</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF6B6FA7DB935" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF6B6FA7AB975" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF6B6FA7AB975" pageId="1" pageNumber="53">
13. Tip of vertical tubercle polished, light yellow; abdominal tergites uniformly orange, outer lateral angles not pale ....
<emphasis id="D9FA2B2FFFCDFFBEFF42F756FBD0B975" box="[206,1078,1304,1327]" italics="true" pageId="1" pageNumber="53">...................................................................................................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBB7F756FA7AB975" ID-CoL="NBNV" authority="(Alexander, 1935)" authorityName="Alexander" authorityYear="1935" baseAuthorityName="Alexander" baseAuthorityYear="1935" box="[1083,1436,1304,1327]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="argyrospila">
<emphasis id="D9FA2B2FFFCDFFBEFBB7F756FB34B975" box="[1083,1234,1304,1327]" italics="true" pageId="1" pageNumber="53">B. argyrospila</emphasis>
(
<bibRefCitation id="8F1F8ACCFFCDFFBEFB6DF756FA72B975" author="Alexander" box="[1249,1428,1304,1327]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1935) New or little-known Tipulidae from eastern Asia (Diptera). XXV. Philippine Journal of Science, 57, 81 - 148." type="journal article" year="1935">Alexander, 1935</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF776FA7DB935" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF776FA7DB935" pageId="1" pageNumber="53">- Vertical tubercle uniformly darkened; color of abdominal tergites not as above, outer lateral angles of basal abdomi- nal tergites pale ........................................................................................................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF736FA7DB995" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF736FA7AB9F5" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF736FA7AB9F5" pageId="1" pageNumber="53">
14. Abdominal tergites cinnamon brown; outer gonostylus with two processes terminating in long slender rod ..............
<emphasis id="D9FA2B2FFFCDFFBEFF42F7D6FBB9B9F5" box="[206,1119,1432,1455]" italics="true" pageId="1" pageNumber="53">.........................................................................................................................................</emphasis>
<taxonomicName id="2C8E8CBEFFCDFFBEFBD3F7D6FA7AB9F5" ID-CoL="NBPB" authority="Alexander, 1927" authorityName="Alexander" authorityYear="1927" box="[1119,1436,1432,1455]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="nymphica">
<emphasis id="D9FA2B2FFFCDFFBEFBD3F7D6FB03B9F5" box="[1119,1253,1432,1455]" italics="true" pageId="1" pageNumber="53">B. nymphica</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB67F7D6FA7AB9F5" author="Alexander" box="[1259,1436,1432,1455]" pageId="1" pageNumber="60" refString="Alexander, C. P. (1927) Undescribed crane-flies from the Holarctic region in the United States National Museum. Proceedings of the United States National Museum, 72 (2), 1 - 17." type="journal article" year="1927">Alexander, 1927</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF7F6FA7DB995" blockId="1.[151,1436,152,1615]" box="[151,1435,1464,1487]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF7F6FA7DB995" box="[151,1435,1464,1487]" pageId="1" pageNumber="53">- Abdominal tergites orange; outer gonostylus not as above ........................................................................................ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="507AEF98FFCDFFBEFF1BF796FA7ABA15" pageId="1" pageNumber="53">
<paragraph id="EB31F73DFFCDFFBEFF1BF796FA7ABA55" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF796FA7ABA55" pageId="1" pageNumber="53">
15. Mediotergite brown with posterior margin yellow; outer gonostylus bent forward slightly at tip, laterally with a con- spicuous spine (
<figureCitation id="73B5EBB8FFCDFFBEFEFEF7B6FE59BA55" box="[370,447,1528,1551]" captionStart="FIGURES 13 16" captionStartId="6.[151,270,1239,1263]" captionTargetBox="[447,1140,431,1206]" captionTargetId="figure@6.[430,1157,418,1218]" captionTargetPageId="6" captionText="FIGURES 13 16. Brithura flaviflagellum sp. nov. Male. 13. Hypopygium, lateral view; 14. ninth tergite, dorsal view; 15. inner gonostylus, lateral view; 16. outer gonostylus, lateral view. Scale bar of 13, 14 = 0.5 mm, 15, 16 = 0.25 mm." httpUri="https://zenodo.org/record/194091/files/figure.png" pageId="1" pageNumber="53">Fig. 16</figureCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFE47F7B6FAA6BA55" box="[459,1344,1528,1551]" italics="true" pageId="1" pageNumber="53">
..........................................................................................................
<taxonomicName id="2C8E8CBEFFCDFFBEFB1EF7B6FAA6BA55" ID-CoL="NBP2" box="[1170,1344,1528,1551]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="flaviflagellum" status="sp. nov.">B. flaviflagellum</taxonomicName>
</emphasis>
<emphasis id="D9FA2B2FFFCDFFBEFACBF7B9FA7ABA55" bold="true" box="[1351,1436,1527,1551]" pageId="1" pageNumber="53">
<taxonomicNameLabel id="C2C99654FFCDFFBEFACBF7B9FA7ABA55" box="[1351,1436,1527,1551]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EB31F73DFFCDFFBEFF1BF456FA7ABA15" blockId="1.[151,1436,152,1615]" pageId="1" pageNumber="53">
<keyLead id="507F5408FFCDFFBEFF1BF456FA7ABA15" pageId="1" pageNumber="53">
- Mediotergite yellow with posterior margin yellowish brown; outer gonostylus with two lobe-like processes (
<bibRefCitation id="8F1F8ACCFFCDFFBEFAD5F456FE53BA15" author="Liu" pageId="1" pageNumber="60" refString="Liu, Q.; Yang, D. (2009) Two new species of the genus Brithura Edwards from China, with a key to world species (Diptera, Tipulidae). Zootaxa 1991, 51 - 56." type="journal article" year="2009" yearSuffix="p">Liu &amp; Yang 2009, p55, fig. 9</bibRefCitation>
)
<emphasis id="D9FA2B2FFFCDFFBEFE32F476FB32BA15" box="[446,1236,1592,1615]" italics="true" pageId="1" pageNumber="53">
.............................................................................................
<taxonomicName id="2C8E8CBEFFCDFFBEFBBCF476FB32BA15" ID-CoL="NBP6" authority="Liu &amp; Yang, 2009" authorityName="Liu &amp; Yang" authorityYear="2009" box="[1072,1236,1592,1615]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="guangxiensis">B. guangxiensis</taxonomicName>
</emphasis>
<bibRefCitation id="8F1F8ACCFFCDFFBEFB57F476FA7ABA15" author="Liu" box="[1243,1436,1592,1615]" pageId="1" pageNumber="60" refString="Liu, Q.; Yang, D. (2009) Two new species of the genus Brithura Edwards from China, with a key to world species (Diptera, Tipulidae). Zootaxa 1991, 51 - 56." type="journal article" year="2009">Liu &amp; Yang, 2009</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>