147 lines
18 KiB
XML
147 lines
18 KiB
XML
<document id="3AC996790706A83C32CDE97D112D0D76" ID-DOI="10.5281/zenodo.194087" ID-GBIF-Dataset="b0c9cf9c-4bbd-47d6-a260-75a3d2f16b72" ID-ISSN="1175-5326" ID-Zenodo-Dep="194087" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189116326" checkinUser="plazi" docAuthor="Liu, Qifei & Yang, Ding" docDate="2010" docId="6327462BFFC8FFBBFF1BF3D3FC14BB80" docLanguage="en" docName="zt02401p060.pdf" docOrigin="Zootaxa 2401" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Brithura aureola Liu & Yang, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="9F1E3E53FFCCFFBFFF8CF24EFFE6BC5A" masterDocTitle="Four new species of the genus Brithura Edwards from China, with an updated key to world species (Diptera, Tipulidae)" masterLastPageNumber="60" masterPageNumber="52" pageNumber="56" updateTime="1698236378629" updateUser="plazi">
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<mods:mods id="10D174372A0E238B51E5270CBA92636A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="617EA78D3CE789BBD0419547AA7C6F28">
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<mods:title id="722AA6E99C7061E9595746E380CEE22A">Four new species of the genus Brithura Edwards from China, with an updated key to world species (Diptera, Tipulidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="21C68BA4ABC75C549DBDEE0822E03290" type="personal">
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<mods:roleTerm id="24D51728378FB420790213A5ADE4E748">Author</mods:roleTerm>
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<mods:namePart id="CA45D41134C33E711F645DC13E3EFA6B">Liu, Qifei</mods:namePart>
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<mods:name id="1B91DEF038A9874347F0A4E2F01B8D47" type="personal">
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<mods:roleTerm id="F93ED06D76A42C2F3B70A78ABCCC1A45">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="854B7458FAC1EC3078C03F61040D10AC">Yang, Ding</mods:namePart>
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<mods:typeOfResource id="7C05651656CEAECDE979FC0DD2D06042">text</mods:typeOfResource>
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<mods:title id="DFFDB72CD67645B5C7D2AF78022A4625">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DBE6954192EDAF2CEBD97031BFB283D2">
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<mods:date id="E79D399D8563631B415145D53182F50E">2010</mods:date>
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<mods:detail id="3E0DE04FECF762DF56863C0A2AC3D192" type="volume">
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<mods:number id="D255A18F0B7EED67FFF45063D32E07B3">2401</mods:number>
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<mods:start id="A4B24695E1793A1D231AF76421B5D4AC">52</mods:start>
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<mods:end id="DBDA678C341D324D5D3EDFC7EE63D793">60</mods:end>
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<mods:classification id="539B69252BAA5BE6D12C5D56AFAB6058">journal article</mods:classification>
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<mods:identifier id="DCAEAB4C8C5A526B8D249D013EA721E3" type="DOI">10.5281/zenodo.194087</mods:identifier>
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<mods:identifier id="659C3CABA43580E6F48578DAC73D7531" type="GBIF-Dataset">b0c9cf9c-4bbd-47d6-a260-75a3d2f16b72</mods:identifier>
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<mods:identifier id="603A905B3B520241CBBB1C384BBB7D56" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B380C049A6ADC76D38E7658F0E75B4C2" type="Zenodo-Dep">194087</mods:identifier>
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<treatment id="6327462BFFC8FFBBFF1BF3D3FC14BB80" ID-DOI="http://doi.org/10.5281/zenodo.6210821" ID-GBIF-Taxon="124956730" ID-Zenodo-Dep="6210821" LSID="urn:lsid:plazi:treatment:6327462BFFC8FFBBFF1BF3D3FC14BB80" httpUri="http://treatment.plazi.org/id/6327462BFFC8FFBBFF1BF3D3FC14BB80" lastPageNumber="56" pageId="4" pageNumber="56">
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<subSubSection id="A394A4B6FFC8FFBBFF1BF3D3FEA2BD86" pageId="4" pageNumber="56" type="nomenclature">
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<paragraph id="EB31F73DFFC8FFBBFF1BF3D3FE2FBDED" blockId="4.[151,457,413,476]" box="[151,457,413,439]" pageId="4" pageNumber="56">
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<heading id="B0794051FFC8FFBBFF1BF3D3FE2FBDED" bold="true" box="[151,457,413,439]" fontSize="11" level="1" pageId="4" pageNumber="56" reason="1">
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<emphasis id="D9FA2B2FFFC8FFBBFF1BF3D3FE2FBDED" bold="true" box="[151,457,413,439]" pageId="4" pageNumber="56">
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<taxonomicName id="2C8E8CBEFFC8FFBBFF1BF3D3FE83BDED" ID-CoL="NBNW" box="[151,357,413,439]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
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<emphasis id="D9FA2B2FFFC8FFBBFF1BF3D3FE83BDED" bold="true" box="[151,357,413,439]" italics="true" pageId="4" pageNumber="56">Brithura aureola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C2C99654FFC8FFBBFEE0F3D3FE2FBDED" box="[364,457,413,439]" pageId="4" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF1BF38CFEA2BD86" blockId="4.[151,457,413,476]" box="[151,324,450,476]" pageId="4" pageNumber="56">
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(
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<figureCitation id="73B5EBB8FFC8FFBBFF13F38CFF15BD86" box="[159,243,450,476]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1967,1991]" captionTargetBox="[187,1396,197,1966]" captionTargetId="figure@2.[187,1412,164,1973]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Wing (male). 1. Brithura shii sp. nov.; 2. Brithura aureola sp. nov.; 3. Brithura flaviflagellum sp. nov.; 4. Brithura stigmosa sp. nov.;" httpUri="https://zenodo.org/record/194088/files/figure.png" pageId="4" pageNumber="56">Figs. 2</figureCitation>
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,
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<figureCitation id="73B5EBB8FFC8FFBBFF73F38CFEDDBD86" box="[255,315,450,476]" captionStart="FIGURES 9 – 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 – 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="4" pageNumber="56">9–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF1BF043FD6BBEE6" pageId="4" pageNumber="56" type="diagnosis">
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<paragraph id="EB31F73DFFC8FFBBFF1BF043FD6BBEE6" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF1BF043FEF1BE7D" bold="true" box="[151,279,525,551]" pageId="4" pageNumber="56">Diagnosis.</emphasis>
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Antennal flagellum yellowish brown, tip of vertical tubercle brownish yellow. Wing dark grayish brown with some light golden areas, costal border opposite stigma bulged and broken in male. Legs yellow with tips black. Tergites of abdomen velvet-yellow, black spots at lateral margins; sternites yellowish brown with lateral margins black. Outer lobe of outer gonostylus with three extensions at tip, the middle one sharp; inner lobe bent forward with a small point.
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</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF48F089FE42BA8E" pageId="4" pageNumber="56" type="description">
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<paragraph id="EB31F73DFFC8FFBBFF48F089FC14BEB8" blockId="4.[151,1437,525,2010]" box="[196,1010,711,738]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF48F089FEBABEBB" bold="true" box="[196,348,711,737]" pageId="4" pageNumber="56">Description.</emphasis>
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Male. Body length 32.0–34.5 mm, wing 22.0–24.0 mm.
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</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF48F0A3FB65BF2D" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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Head. Rostrum dark brownish black, with stout blackish brown nasus. Vertex and occiput blackish. Vertical tubercle dark brown with tip brownish yellow. Orbit brownish. Hairs on head light brown. Antenna
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<quantity id="2C765AD8FFC8FFBBFA01F15CFF23BF08" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="56" unit="mm" value="6.0">6 mm</quantity>
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long; scape blackish brown; pedicel brownish; flagellum brownish yellow. Probocis dark reddish brown; first segment of palpus black, others blackish brown, joint yellow; setae short, brown.
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</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF48F1CCFDDDB905" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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Thorax. Ground color chestnut brown. Pronotum dark reddish brown; prescutum yellowish brown with three brown longitudinal stripes, lateral ones each with a line of yellow hairs; lateral margins of prescutum yellowish brown; scutellum yellowish brown; scutum black with long hairs; mediotergite blackish; pleura chiefly chestnut brown, pleurotergal tubercle small, pointed, with silvery pubescence on dorsal surface. Hairs on thorax long, brown. Coxae and trochanters brown; femora yellowish brown with tips black; tibiae yellowish brown; tarsi blackish brown; tibial spurs 1–1–2. Setae on coxae and trochanters yellow, on other segments of legs brown. Wing (
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<figureCitation id="73B5EBB8FFC8FFBBFD97F62CFD82B826" box="[539,612,1122,1148]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1967,1991]" captionTargetBox="[187,1396,197,1966]" captionTargetId="figure@2.[187,1412,164,1973]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Wing (male). 1. Brithura shii sp. nov.; 2. Brithura aureola sp. nov.; 3. Brithura flaviflagellum sp. nov.; 4. Brithura stigmosa sp. nov.;" httpUri="https://zenodo.org/record/194088/files/figure.png" pageId="4" pageNumber="56">Fig. 2</figureCitation>
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) dark grayish brown with costal region opposite stigma dilated and broken. Cells c and sc yellow; stigma brown. A black spot at origin of Rs; area from stigma to cell dm along cord yellowish brown, surrounded by light golden suffusion; cell br yellowish, with brown spots at base and center; cell bm brown with yellow strip in middle and yellow spot in outer end; cell dm yellow with brown spot at bottom; cell r4, cell m2, cell m3 with light yellow spot at margin of wing; cell cua2 and cell a1 with two yellow spots at middle. Veins yellow, margined with yellow. Halter with stem yellowish brown, knob more blackish but yellowish brown at tip.
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</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF48F724FD9FB995" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">Abdomen. Ground color velvet-like brownish yellow. Tergite 1 blackish brown, tergites 2–8 velvet-like brownish yellow with big blackish brown spots at lateral margins. Venter yellowish brown with brownish lateral margin. Setae on abdomen brown.</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF48F794FBA7BAF5" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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Hypopygium (
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<figureCitation id="73B5EBB8FFC8FFBBFEF8F794FE1CB9AE" box="[372,506,1498,1524]" captionStart="FIGURES 9 – 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 – 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="4" pageNumber="56">Figs. 9–12</figureCitation>
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). Tergite 9 (
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<figureCitation id="73B5EBB8FFC8FFBBFD02F794FD01B9AE" box="[654,743,1498,1524]" captionStart="FIGURES 9 – 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 – 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="4" pageNumber="56">Fig. 10</figureCitation>
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) as wide as long, caudal margin with a fairly shallow Vshaped incision that is fringed with the yellow setae, medially a pair of blades adhered to each other; sternite 9 ventrally with a light brown knob and a small process at each side; outer gonostylus (
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<figureCitation id="73B5EBB8FFC8FFBBFB06F46BFB06BA65" box="[1162,1248,1573,1599]" captionStart="FIGURES 9 – 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 – 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="4" pageNumber="56">Fig. 12</figureCitation>
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) thick, laterally with three extensions at tip, middle one pointed, medially with a relative long extension bent forward with a small dorsal pointed spine; inner gonostylus (
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<figureCitation id="73B5EBB8FFC8FFBBFD24F43EFD1DBAD0" box="[680,763,1648,1674]" captionStart="FIGURES 9 – 12" captionStartId="5.[151,270,1010,1034]" captionTargetBox="[413,1168,207,963]" captionTargetId="figure@5.[393,1193,197,989]" captionTargetPageId="5" captionText="FIGURES 9 – 12. Brithura aureola sp. nov. Male. 9. Hypopygium, lateral view; 10. ninth tergite, dorsal view; 11. inner gonostylus, lateral view; 12. outer gonostylus, lateral view. Scale bar of 9, 10 = 0.5 mm, 11, 12 = 0.25 mm." httpUri="https://zenodo.org/record/194090/files/figure.png" pageId="4" pageNumber="56">Fig. 11</figureCitation>
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) bent at middle, tip relative flat with two small processes, a long sulcus on outer surface, setae on inner gonostylus long, yellowish brown.
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</paragraph>
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<paragraph id="EB31F73DFFC8FFBBFF48F4F4FE42BA8E" blockId="4.[151,1437,525,2010]" box="[196,420,1722,1748]" pageId="4" pageNumber="56">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF4AF4AEFD8ABB45" pageId="4" pageNumber="56" type="materials_examined">
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<paragraph id="EB31F73DFFC8FFBBFF4AF4AEFD8ABB45" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF4AF4AEFE99BAA0" bold="true" box="[198,383,1760,1786]" pageId="4" pageNumber="56">
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<typeStatus id="3435499FFFC8FFBBFF4AF4AEFEE5BAA0" box="[198,259,1760,1786]" pageId="4" pageNumber="56">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="3435499FFFC8FFBBFE06F4AEFE11BAA0" box="[394,503,1760,1786]" pageId="4" pageNumber="56" type="holotype">Holotype</typeStatus>
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male, Sichuan: Meigu, Wahou (
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<quantity id="2C765AD8FFC8FFBBFC0CF4AEFC07BAA0" box="[896,993,1760,1786]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="56" unit="m" value="1900.0">1900 m</quantity>
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), 2008.
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<date id="9F30D1FDFFC8FFBBFBC9F4AEFB46BAA0" box="[1093,1184,1760,1786]" pageId="4" pageNumber="56" value="1916-07">VII. 16</date>
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, light, Jinjing Wang.
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<typeStatus id="3435499FFFC8FFBBFF1BF54BFF18BB45" box="[151,254,1797,1823]" pageId="4" pageNumber="56" type="paratype">Paratype</typeStatus>
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<specimenCount id="FD883CB4FFC8FFBBFE88F54BFEB0BB45" box="[260,342,1797,1823]" pageId="4" pageNumber="56" type="male">1 male</specimenCount>
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, same data as
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<typeStatus id="3435499FFFC8FFBBFE72F54BFD8EBB45" box="[510,616,1797,1823]" pageId="4" pageNumber="56" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF48F564FDD4BB1E" box="[196,562,1834,1860]" pageId="4" pageNumber="56" type="distribution">
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<paragraph id="EB31F73DFFC8FFBBFF48F564FDD4BB1E" blockId="4.[151,1437,525,2010]" box="[196,562,1834,1860]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF48F564FE82BB1E" bold="true" box="[196,356,1834,1860]" pageId="4" pageNumber="56">Distribution.</emphasis>
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<collectingCountry id="9399B7ADFFC8FFBBFEE0F564FE54BB1E" box="[364,434,1834,1860]" name="China" pageId="4" pageNumber="56">China</collectingCountry>
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(Sichuan).
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</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF48F51EFC0FBBEE" pageId="4" pageNumber="56" type="discussion">
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<paragraph id="EB31F73DFFC8FFBBFF48F51EFC0FBBEE" blockId="4.[151,1437,525,2010]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF48F51EFEDBBB30" bold="true" box="[196,317,1872,1898]" pageId="4" pageNumber="56">Remarks.</emphasis>
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This new species is similar to
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<taxonomicName id="2C8E8CBEFFC8FFBBFD24F51EFD15BB33" box="[680,755,1872,1897]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="shii">
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<emphasis id="D9FA2B2FFFC8FFBBFD24F51EFD15BB33" box="[680,755,1872,1897]" italics="true" pageId="4" pageNumber="56">B. shii</emphasis>
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</taxonomicName>
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in having the similar color pattern of the abdomen. It can be easily separated from
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<taxonomicName id="2C8E8CBEFFC8FFBBFE4CF538FDEABBD5" box="[448,524,1910,1935]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="shii">
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<emphasis id="D9FA2B2FFFC8FFBBFE4CF538FDEABBD5" box="[448,524,1910,1935]" italics="true" pageId="4" pageNumber="56">B. shii</emphasis>
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</taxonomicName>
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by the color pattern of the wing and structure of the hypopygium. In
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<emphasis id="D9FA2B2FFFC8FFBBFAC4F538FA7ABBD5" box="[1352,1436,1910,1935]" italics="true" pageId="4" pageNumber="56">
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<taxonomicName id="2C8E8CBEFFC8FFBBFAC4F538FA7EBBD5" box="[1352,1432,1910,1935]" class="Insecta" family="Tipulidae" genus="Brithura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="shii">B. shii</taxonomicName>
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,
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</emphasis>
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the wing is yellowish, and the outer gonostylus has only one spine at tip.
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</paragraph>
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</subSubSection>
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<subSubSection id="A394A4B6FFC8FFBBFF48F58EFC14BB80" box="[196,1010,1984,2010]" pageId="4" pageNumber="56" type="etymology">
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<paragraph id="EB31F73DFFC8FFBBFF48F58EFC14BB80" blockId="4.[151,1437,525,2010]" box="[196,1010,1984,2010]" pageId="4" pageNumber="56">
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<emphasis id="D9FA2B2FFFC8FFBBFF48F58EFEB7BB80" bold="true" box="[196,337,1984,2010]" pageId="4" pageNumber="56">Etymology.</emphasis>
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The species name is based on the golden patterned wing.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |