290 lines
38 KiB
XML
290 lines
38 KiB
XML
<document id="32FDCB5B2469FB7EBCF22E90A6671D3A" ID-CLB-Dataset="298741" ID-DOI="10.11646/zootaxa.5471.4.2" ID-GBIF-Dataset="2370cfa6-ebc7-4962-ae93-ade18759c5cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190291" ID-ZooBank="A625B488-6A82-44F4-99AD-A66CAB9FC996" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718893826382" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="E56187A31F17FFE26C9EFB57FCB14EC5" docLanguage="en" docName="zootaxa.5471.4.2.pdf" docOrigin="Zootaxa 5471 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loeblites anaiensis Jałoszyński 2024, sp. n." docType="treatment" docVersion="6" lastPageNumber="425" masterDocId="1958FFDB1F16FFE16C09FF98FF914C6C" masterDocTitle="Three new species, one subspecies, and new records of the Oriental genus Loeblites Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="432" masterPageNumber="422" pageNumber="423" updateTime="1719007836080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6DE9A6CDF21458840B32C4A10C2993F6">Three new species, one subspecies, and new records of the Oriental genus Loeblites Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="E5E9FA53062DEA1AD6D53591B520A660">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="4573666235D18EABF3B400C02B05EE2D">Zootaxa</mods:title>
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<mods:date id="BC3AAF9E02F95DC9AC904EDF92D6C5AB">2024</mods:date>
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<mods:number id="88A8A8BE8C419056D9681A766F821978">2024-06-20</mods:number>
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<treatment id="E56187A31F17FFE26C9EFB57FCB14EC5" ID-DOI="http://doi.org/10.5281/zenodo.12190844" ID-GBIF-Taxon="233317985" ID-Zenodo-Dep="12190844" LSID="urn:lsid:plazi:treatment:E56187A31F17FFE26C9EFB57FCB14EC5" httpUri="http://treatment.plazi.org/id/E56187A31F17FFE26C9EFB57FCB14EC5" lastPageId="3" lastPageNumber="425" pageId="1" pageNumber="423">
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<subSubSection id="25D2653E1F17FFE06C9EFB57FE564885" box="[151,455,1231,1258]" pageId="1" pageNumber="423" type="nomenclature">
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<paragraph id="6D7736B51F17FFE06C9EFB57FE564885" blockId="1.[151,455,1231,1294]" box="[151,455,1231,1258]" pageId="1" pageNumber="423">
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<heading id="363F81D91F17FFE06C9EFB57FE564885" bold="true" box="[151,455,1231,1258]" fontSize="11" level="1" pageId="1" pageNumber="423" reason="1">
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<taxonomicName id="AAC84D361F17FFE06C9EFB57FE564885" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[151,455,1231,1258]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="423" phylum="Arthropoda" rank="species" species="anaiensis" status="sp. nov.">
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<emphasis id="5FBCEAA71F17FFE06C9EFB57FE564885" bold="true" box="[151,455,1231,1258]" pageId="1" pageNumber="423">
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<emphasis id="5FBCEAA71F17FFE06C9EFB57FEEF4885" bold="true" box="[151,382,1231,1258]" italics="true" pageId="1" pageNumber="423">Loeblites anaiensis</emphasis>
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<taxonomicNameLabel id="448F57DC1F17FFE06D8CFB57FE564885" box="[389,455,1231,1257]" pageId="1" pageNumber="423" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="25D2653E1F17FFE06C9EFB6BFE344961" box="[151,421,1267,1294]" pageId="1" pageNumber="423" type="description">
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<paragraph id="6D7736B51F17FFE06C9EFB6BFE344961" blockId="1.[151,455,1231,1294]" box="[151,421,1267,1294]" pageId="1" pageNumber="423">
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(
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<figureCitation id="F5F32A301F17FFE06C96FB6BFE954962" box="[159,260,1267,1294]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="1" pageNumber="423">Figs 1, 5</figureCitation>
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,
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<figureCitation id="F5F32A301F17FFE06D19FB6BFEDB4961" box="[272,330,1267,1293]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="1" pageNumber="423">9–13</figureCitation>
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,
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<figureCitation id="F5F32A301F17FFE06D5FFB6BFEE54961" box="[342,372,1267,1293]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="1" pageNumber="423">29</figureCitation>
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,
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<figureCitation id="F5F32A301F17FFE06D89FB6BFE0C4961" box="[384,413,1267,1293]" captionStart="FIGURE 32" captionStartId="7.[151,250,1999,2023]" captionTargetBox="[177,1411,849,1976]" captionTargetId="figure-234@7.[177,1411,849,1976]" captionTargetPageId="7" captionText="FIGURE 32. Distribution of nominal species of Loeblites." figureDoi="http://doi.org/10.5281/zenodo.12190305" httpUri="https://zenodo.org/record/12190305/files/figure.png" pageId="1" pageNumber="423">32</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="25D2653E1F17FFE06C9EFAA3FEE84989" pageId="1" pageNumber="423" type="materials_examined">
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<paragraph id="6D7736B51F17FFE06C9EFAA3FEE84989" blockId="1.[151,1437,1339,2014]" pageId="1" pageNumber="423">
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<emphasis id="5FBCEAA71F17FFE06C9EFAA3FEF54939" bold="true" box="[151,356,1339,1365]" pageId="1" pageNumber="423">Material studied.</emphasis>
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||
<materialsCitation id="DDA03CE81F17FFE06D63FAA3FE0B49AD" ID-GBIF-Occurrence="4898138308" collectingDate="2006-06-01" collectionCode="MHNG" collectorName="P. Schwendinger" country="Indonesia" county="West Sumatra Province" determinerName="P. Jaloszynski" elevation="500" latitude="-0.47722223" location="Anai Vallei" longLatPrecision="21" longitude="100.35389" municipality="Anai Vallei" pageId="1" pageNumber="423" specimenCount="1" specimenCount-male="1" stateProvince="SUMATRA" typeStatus="holotype">
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<emphasis id="5FBCEAA71F17FFE06D63FAA3FC2E4939" bold="true" box="[362,959,1339,1365]" pageId="1" pageNumber="423">
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<typeStatus id="B27388171F17FFE06D63FAA3FE4A4939" box="[362,475,1339,1365]" pageId="1" pageNumber="423" type="holotype">Holotype</typeStatus>
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: INDONESIA (West Sumatra Province):
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||
</emphasis>
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<specimenCount id="7BCEFD3C1F17FFE06FCCFAA4FC4D493A" box="[965,988,1340,1366]" count="1" pageId="1" pageNumber="423" type="male">♂</specimenCount>
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, two labels: “
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||
<emphasis id="5FBCEAA71F17FFE0687DFAA3FB6D4939" bold="true" box="[1140,1276,1339,1365]" pageId="1" pageNumber="423">Sum-06/03:</emphasis>
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<emphasis id="5FBCEAA71F17FFE0690AFAA3FA1F4939" bold="true" box="[1283,1422,1339,1365]" pageId="1" pageNumber="423">
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<collectingRegion id="AF0CF8571F17FFE0690AFAA3FA1F4939" box="[1283,1422,1339,1365]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="423">SUMATRA</collectingRegion>
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</emphasis>
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||
/
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||
<collectingCounty id="84164E391F17FFE06C9EFAF8FE0C4915" box="[151,413,1375,1402]" pageId="1" pageNumber="423">West Sumatra Province</collectingCounty>
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||
/
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||
<collectingMunicipality id="8D13ACCF1F17FFE06DB9FAC7FDA14915" box="[432,560,1375,1401]" pageId="1" pageNumber="423">Anai Vallei</collectingMunicipality>
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,
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<locationDeviation id="7CC52F971F17FFE06E32FAC7FCFB4915" box="[571,874,1375,1401]" pageId="1" pageNumber="423">
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<quantity id="AA309B501F17FFE06E32FAC7FDE24915" box="[571,627,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="423" unit="km" value="6.0">6 km</quantity>
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||
S of / Padangpanjang
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</locationDeviation>
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,
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||
<geoCoordinate id="08FC50721F17FFE06F7CFAC7FC764915" box="[885,999,1375,1401]" degrees="0" direction="south" minutes="28" orientation="latitude" pageId="1" pageNumber="423" precision="15" seconds="38" value="-0.47722223">0°28'38''S</geoCoordinate>
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, /
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<geoCoordinate id="08FC50721F17FFE06809FAC7FB014915" box="[1024,1168,1375,1401]" degrees="100" direction="east" minutes="21" orientation="longitude" pageId="1" pageNumber="423" precision="15" seconds="14" value="100.35389">100°21'14''E</geoCoordinate>
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,
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<quantity id="AA309B501F17FFE06892FAC7FB734916" box="[1179,1250,1375,1402]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="423" unit="m" value="500.0">
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<elevation id="E6E5D1861F17FFE06892FAC7FB734916" box="[1179,1250,1375,1402]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="423" unit="m" value="500.0">500 m</elevation>
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</quantity>
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/ primary forest,
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||
<date id="197610751F17FFE06C9EFA1BFE9849F1" box="[151,265,1411,1437]" pageId="1" pageNumber="423" value="2006-06-01">
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<collectingDate id="0932E99D1F17FFE06C9EFA1BFE9849F1" box="[151,265,1411,1437]" pageId="1" pageNumber="423" value="2006-06-01">1.VI.2006</collectingDate>
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</date>
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/ leg.
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||
<collectorName id="C03D53631F17FFE06D45FA1CFD9849F1" box="[332,521,1411,1438]" pageId="1" pageNumber="423">P. Schwendinger</collectorName>
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” [white], “
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||
<taxonomicName id="AAC84D361F17FFE06E8AFA1BFC8049F1" authorityName="Franz" authorityYear="1986" box="[643,785,1411,1437]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="423" phylum="Arthropoda" rank="genus">
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<emphasis id="5FBCEAA71F17FFE06E8AFA1BFC8049F1" box="[643,785,1411,1437]" italics="true" pageId="1" pageNumber="423">LOEBLITES</emphasis>
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</taxonomicName>
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||
/
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<taxonomicName id="AAC84D361F17FFE06F2CFA1CFC1E49F1" authorityName="Jałoszyński" authorityYear="2024" box="[805,911,1412,1437]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="423" phylum="Arthropoda" rank="species" species="anaiensis">
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<emphasis id="5FBCEAA71F17FFE06F2CFA1CFC1E49F1" box="[805,911,1412,1437]" italics="true" pageId="1" pageNumber="423">anaiensis</emphasis>
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</taxonomicName>
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m. / det. P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (
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<collectionCode id="0BD9AE701F17FFE06D3FFA3FFE0349AD" box="[310,402,1447,1473]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="423" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
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||
.
|
||
<materialsCitation id="DDA03CE81F17FFE06DAAFA3FFEE44989" ID-GBIF-Occurrence="4898138309" collectingDate="2006-06-01" collectionCode="MHNG" collectorName="P. Schwendinger" country="Indonesia" county="West Sumatra Province" determinerName="P. Jaloszynski" elevation="500" latitude="-0.47722223" location="Padangpanjang" longLatPrecision="21" longitude="100.35389" municipality="Anai Vallei" pageId="1" pageNumber="423" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="paratype">
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||
<emphasis id="5FBCEAA71F17FFE06DAAFA3FFD8A49AD" bold="true" box="[419,539,1447,1473]" pageId="1" pageNumber="423">Paratypes</emphasis>
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||
(4 exx.):
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||
<specimenCount id="7BCEFD3C1F17FFE06E8AFA3FFD3C49AE" box="[643,685,1447,1474]" count="1" pageId="1" pageNumber="423" type="male">1 ♂</specimenCount>
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||
,
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||
<specimenCount id="7BCEFD3C1F17FFE06EBEFA3FFD6749AE" box="[695,758,1447,1474]" count="3" pageId="1" pageNumber="423" type="female">3 ♀♀</specimenCount>
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||
, same data as for holotype, each with yellow “PARATYPUS” label (
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<collectionCode id="0BD9AE701F17FFE06CD5FA53FEA74989" box="[220,310,1483,1509]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="423" type="Museum">MHNG</collectionCode>
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, cPJ)
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</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25D2653E1F17FFE06CCEFA77FD254A19" pageId="1" pageNumber="423" type="diagnosis">
|
||
<paragraph id="6D7736B51F17FFE06CCEFA77FD254A19" blockId="1.[151,1437,1339,2014]" pageId="1" pageNumber="423">
|
||
<emphasis id="5FBCEAA71F17FFE06CCEFA77FED34A65" bold="true" box="[199,322,1519,1545]" pageId="1" pageNumber="423">Diagnosis.</emphasis>
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Pronotum with fine, inconspicuous punctures; aedeagus (
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||
<figureCitation id="F5F32A301F17FFE06FDAFA77FB894A65" box="[979,1048,1519,1545]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="1" pageNumber="423">Fig. 9</figureCitation>
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) slender, with conspicuously long parameres, endophallus (
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<figureCitation id="F5F32A301F17FFE06DBBF98BFD954A41" box="[434,516,1555,1581]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="1" pageNumber="423">Fig. 13</figureCitation>
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) symmetrical, with strongly oblique lateral sclerites and with sclerotized median structure distal to lateral sclerites about as long as broad, with concave anterior margin; spermatheca in lateral view (
|
||
<figureCitation id="F5F32A301F17FFE06C96F9C3FF604A19" box="[159,241,1627,1653]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="1" pageNumber="423">Fig. 29</figureCitation>
|
||
) subtrapezoidal, about as long as broad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25D2653E1F17FFE26CCEF9E7FDBF4DC1" lastPageId="3" lastPageNumber="425" pageId="1" pageNumber="423" type="description">
|
||
<paragraph id="6D7736B51F17FFE06CCEF9E7FDB94AD2" blockId="1.[151,1437,1339,2014]" pageId="1" pageNumber="423">
|
||
<emphasis id="5FBCEAA71F17FFE06CCEF9E7FEC84AF5" bold="true" box="[199,345,1663,1689]" pageId="1" pageNumber="423">Description.</emphasis>
|
||
Body of male (
|
||
<figureCitation id="F5F32A301F17FFE06E07F9E7FDC04AF5" box="[526,593,1663,1689]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="1" pageNumber="423">Fig. 1</figureCitation>
|
||
) slender and strongly convex, moderately dark brown, covered with light brown vestiture. BL
|
||
<quantity id="AA309B501F17FFE06D77F93BFDB54AD2" box="[382,548,1699,1726]" metricMagnitude="-3" metricUnit="m" metricValue="2.49" metricValueMax="2.5" metricValueMin="2.48" pageId="1" pageNumber="423" unit="mm" value="2.49" valueMax="2.5" valueMin="2.48">2.48–2.50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6D7736B51F17FFE06CCEF95FFE8A4BFA" blockId="1.[151,1437,1339,2014]" pageId="1" pageNumber="423">
|
||
Head in dorsal view transverse, broadest at eyes, HL
|
||
<quantity id="AA309B501F17FFE06F4BF95FFC7A4A8E" box="[834,1003,1735,1762]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="1" pageNumber="423" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
|
||
, HW
|
||
<quantity id="AA309B501F17FFE0683CF95FFB4E4A8E" box="[1077,1247,1735,1762]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="1" pageNumber="423" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
; vertex strongly transverse and evenly convex, anteriorly confluent with frons; frons flattened between prominent supraantennal tubercles; tempora in dorsal view distinctly longer than eyes; eyes small, oval, with slightly concave posteroventral margin, finely faceted. Vertex and frons covered with fine, inconspicuous punctures and sparse, moderately long, suberect setae. Antennae long and slender, slightly shorter than BL, AnL
|
||
<quantity id="AA309B501F17FFE06FCAF8CFFBF64B1E" box="[963,1127,1879,1906]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" metricValueMax="2.13" metricValueMin="2.03" pageId="1" pageNumber="423" unit="mm" value="2.08" valueMax="2.13" valueMin="2.03">2.03–2.13 mm</quantity>
|
||
, all antennomeres elongate, as in
|
||
<figureCitation id="F5F32A301F17FFE06CDBF8E3FE874BFA" box="[210,278,1915,1942]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="1" pageNumber="423">Fig. 5</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6D7736B51F17FFE36CCEF838FE664C95" blockId="1.[151,1437,1339,2014]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="424" pageId="1" pageNumber="423">
|
||
Pronotum elongate, about as broad as head, broadest slightly in front of anterior third; PL
|
||
<quantity id="AA309B501F17FFE068CFF807FABC4BD6" box="[1222,1325,1951,1978]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="423" unit="mm" value="0.7">0.70 mm</quantity>
|
||
, PW
|
||
<quantity id="AA309B501F17FFE06962F807FF574BB2" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="423" unit="mm" value="0.55">0.55 mm</quantity>
|
||
. Anterior margin broadly rounded, sides strongly rounded in anterior third, posteriorly broadly and weakly constricted; posterior margin nearly straight. Four dorsal antebasal pits large and deep, two lateral pits at each side distinct. Pronotum with fine, inconspicuous punctures, those behind antebasal pits not differing from those on disc; setae long, sparse and suberect.
|
||
</paragraph>
|
||
<caption id="39B7663D1F14FFE36C9EF9DEFBAE4AEE" ID-DOI="http://doi.org/10.5281/zenodo.12190293" ID-Zenodo-Dep="12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="2" pageNumber="424" startId="2.[151,264,1606,1630]" targetBox="[156,1431,276,1573]" targetPageId="2" targetType="figure">
|
||
<paragraph id="6D7736B51F14FFE36C9EF9DEFBAE4AEE" blockId="2.[151,1436,1606,1666]" pageId="2" pageNumber="424">
|
||
<emphasis id="5FBCEAA71F14FFE36C9EF9DEFEAB4A32" bold="true" box="[151,314,1606,1630]" pageId="2" pageNumber="424">FIGURES 1–8.</emphasis>
|
||
Males of
|
||
<taxonomicName id="AAC84D361F14FFE36D96F9DEFDF34A32" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[415,610,1606,1630]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="424" phylum="Arthropoda" rank="species" species="anaiensis" status="sp. nov.">
|
||
<emphasis id="5FBCEAA71F14FFE36D96F9DEFDF34A32" box="[415,610,1606,1630]" italics="true" pageId="2" pageNumber="424">Loeblites anaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="448F57DC1F14FFE36E6EF9DEFD0C4A31" box="[615,669,1606,1629]" pageId="2" pageNumber="424" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="5FBCEAA71F14FFE36EA2F9DEFD404A32" bold="true" box="[683,721,1606,1630]" pageId="2" pageNumber="424">1, 5</emphasis>
|
||
);
|
||
<taxonomicName id="AAC84D361F14FFE36EEFF9DFFC034A32" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[742,914,1606,1630]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="424" phylum="Arthropoda" rank="species" species="umphangensis" status="sp. n.">
|
||
<emphasis id="5FBCEAA71F14FFE36EEFF9DFFC034A32" box="[742,914,1606,1630]" italics="true" pageId="2" pageNumber="424">L. umphangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="448F57DC1F14FFE36F9EF9DEFC5C4A31" box="[919,973,1606,1629]" pageId="2" pageNumber="424" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="5FBCEAA71F14FFE36FD2F9DEFB904A32" bold="true" box="[987,1025,1606,1630]" pageId="2" pageNumber="424">2, 6</emphasis>
|
||
);
|
||
<taxonomicName id="AAC84D361F14FFE3681FF9DFFAAE4A32" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[1046,1343,1606,1630]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="424" phylum="Arthropoda" rank="subSpecies" species="pseudominor" status="sp. n." subSpecies="pseudominor">
|
||
<emphasis id="5FBCEAA71F14FFE3681FF9DFFAAE4A32" box="[1046,1343,1606,1630]" italics="true" pageId="2" pageNumber="424">L. pseudominor pseudominor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="448F57DC1F14FFE3694CF9DEFAEA4A31" box="[1349,1403,1606,1629]" pageId="2" pageNumber="424" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="5FBCEAA71F14FFE36980F9DEFF354AEE" bold="true" pageId="2" pageNumber="424">3, 7</emphasis>
|
||
); and
|
||
<taxonomicName id="AAC84D361F14FFE36CECF9F3FD934AEE" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[229,514,1642,1666]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="424" phylum="Arthropoda" rank="subSpecies" species="pseudominor" status="subsp. n." subSpecies="sumatrensis">
|
||
<emphasis id="5FBCEAA71F14FFE36CECF9F3FD934AEE" box="[229,514,1642,1666]" italics="true" pageId="2" pageNumber="424">L. pseudominor sumatrensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="448F57DC1F14FFE36E01F9F2FDF34AED" box="[520,610,1642,1666]" pageId="2" pageNumber="424" rank="subSpecies">subsp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="5FBCEAA71F14FFE36E78F9F2FD064AEE" bold="true" box="[625,663,1642,1666]" pageId="2" pageNumber="424">4, 8</emphasis>
|
||
). Dorsal habitus (
|
||
<emphasis id="5FBCEAA71F14FFE36F44F9F2FCE24AEE" bold="true" box="[845,883,1642,1666]" pageId="2" pageNumber="424">1–4</emphasis>
|
||
) and antenna (
|
||
<emphasis id="5FBCEAA71F14FFE36802F9F2FBA04AEE" bold="true" box="[1035,1073,1642,1666]" pageId="2" pageNumber="424">5–8</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6D7736B51F14FFE36CCEF94CFDB04B37" blockId="2.[151,1436,1748,2027]" pageId="2" pageNumber="424">
|
||
Elytra more convex than pronotum, oval, broadest distinctly in front of middle; EL
|
||
<quantity id="AA309B501F14FFE36859F94CFB624A83" box="[1104,1267,1748,1775]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="2" pageNumber="424" unit="mm" value="1.375" valueMax="1.4" valueMin="1.35">1.35–1.40 mm</quantity>
|
||
, EW
|
||
<quantity id="AA309B501F14FFE36925F94CFF544B7F" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="2" pageNumber="424" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.85–0.88 mm</quantity>
|
||
, EI 1.54–1.65. Humeral calli weakly elevated, elongate; apex of each elytron subtriangular, sharply-angled but with blunt tip. Punctures more distinct than those on head and pronotum, but superficial, diffuse and inconspicuous; setae similar to those on pronotum.
|
||
</paragraph>
|
||
<paragraph id="6D7736B51F14FFE36CCEF8FDFB8E4BCE" blockId="2.[151,1436,1748,2027]" pageId="2" pageNumber="424">Legs long and slender; femora abruptly clavate, dorsal longitudinal grooves on femoral clava complete and with distinct glandular opening at highest site of each femur; all tibiae nearly straight.</paragraph>
|
||
<paragraph id="6D7736B51F14FFE26CCEF834FAE44D71" blockId="2.[151,1436,1748,2027]" lastBlockId="3.[151,1437,151,681]" lastPageId="3" lastPageNumber="425" pageId="2" pageNumber="424">
|
||
Aedeagus (
|
||
<figureCitation id="F5F32A301F14FFE36D4DF834FE244BAA" box="[324,437,1964,1990]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="2" pageNumber="424">Figs 9–13</figureCitation>
|
||
) strongly elongate and slender; AeL
|
||
<quantity id="AA309B501F14FFE36F59F834FC644BAB" box="[848,1013,1964,1991]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="2" pageNumber="424" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
|
||
; median lobe in ventral view broadest near base and narrowing to subtriangular apex, with its distal margin evenly rounded; endophallus (
|
||
<figureCitation id="F5F32A301F14FFE3690EF848FAC84B86" box="[1287,1369,2000,2026]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="2" pageNumber="424">Fig. 13</figureCitation>
|
||
) with pair of strongly elongate and oblique lateral sclerites (
|
||
<figureCitation id="F5F32A301F15FFE26EFBFF0FFCD74CDD" box="[754,838,151,177]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="3" pageNumber="425">Fig. 13</figureCitation>
|
||
;
|
||
<emphasis id="5FBCEAA71F15FFE26F58FF0FFCE14CDD" box="[849,880,151,177]" italics="true" pageId="3" pageNumber="425">lsc</emphasis>
|
||
) distally connected with large median sclerotized structure, the latter about as long as broad and with distal margin concave, proximally lateral sclerites connected with sclerotized median proximal complex, with its proximal margin rounded and strongly convex; parameres conspicuously long, longer than median lobe and their apices far exceeding apex of median lobe, lacking setae.
|
||
</paragraph>
|
||
<paragraph id="6D7736B51F15FFE26CCEFEBFFDBF4DC1" blockId="3.[151,1437,151,681]" pageId="3" pageNumber="425">
|
||
Female. Externally similar to male, with antennae indistinctly shorter in relation to body; BL
|
||
<quantity id="AA309B501F15FFE268F9FEBEFA094D2D" box="[1264,1432,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.49" metricValueMax="2.5" metricValueMin="2.48" pageId="3" pageNumber="425" unit="mm" value="2.49" valueMax="2.5" valueMin="2.48">2.48–2.50 mm</quantity>
|
||
; HL
|
||
<quantity id="AA309B501F15FFE26CCAFED2FEF84D09" box="[195,361,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="3" pageNumber="425" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
|
||
, HW
|
||
<quantity id="AA309B501F15FFE26DA5FED2FDC04D09" box="[428,593,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="3" pageNumber="425" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
, AnL 1.95–2.00 mm; PL
|
||
<quantity id="AA309B501F15FFE26F78FED3FC494D09" box="[881,984,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="425" unit="mm" value="0.7">0.70 mm</quantity>
|
||
, PW
|
||
<quantity id="AA309B501F15FFE2681FFED2FB2C4D09" box="[1046,1213,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="3" pageNumber="425" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
; EL
|
||
<quantity id="AA309B501F15FFE268FBFED2FA094D09" box="[1266,1432,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.38" metricValueMin="1.35" pageId="3" pageNumber="425" unit="mm" value="1.365" valueMax="1.38" valueMin="1.35">1.35–1.38 mm</quantity>
|
||
, EW
|
||
<quantity id="AA309B501F15FFE26CC4FEF6FEE74DE5" box="[205,374,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="3" pageNumber="425" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.85–0.88 mm</quantity>
|
||
, EI 1.57–1.59. Spermatheca (
|
||
<figureCitation id="F5F32A301F15FFE26ED8FEF7FCB74DE5" box="[721,806,367,393]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="3" pageNumber="425">Fig. 29</figureCitation>
|
||
) in lateral view
|
||
<quantity id="AA309B501F15FFE26FE5FEF6FBF34DE5" box="[1004,1122,366,393]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="425" unit="mm" value="0.075">0.075 mm</quantity>
|
||
wide, subtrapezoidal, with conspicuously broad distal ‘pocket’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25D2653E1F15FFE26CCEFE2EFD7D4DBD" box="[199,748,438,465]" pageId="3" pageNumber="425" type="distribution">
|
||
<paragraph id="6D7736B51F15FFE26CCEFE2EFD7D4DBD" blockId="3.[151,1437,151,681]" box="[199,748,438,465]" pageId="3" pageNumber="425">
|
||
<emphasis id="5FBCEAA71F15FFE26CCEFE2EFEF04DBC" bold="true" box="[199,353,438,464]" pageId="3" pageNumber="425">Distribution.</emphasis>
|
||
Central western
|
||
<collectingRegion id="AF0CF8571F15FFE26E16FE2FFDED4DBD" box="[543,636,439,465]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="425">Sumatra</collectingRegion>
|
||
(
|
||
<figureCitation id="F5F32A301F15FFE26E85FE2FFD714DBD" box="[652,736,439,465]" captionStart="FIGURE 32" captionStartId="7.[151,250,1999,2023]" captionTargetBox="[177,1411,849,1976]" captionTargetId="figure-234@7.[177,1411,849,1976]" captionTargetPageId="7" captionText="FIGURE 32. Distribution of nominal species of Loeblites." figureDoi="http://doi.org/10.5281/zenodo.12190305" httpUri="https://zenodo.org/record/12190305/files/figure.png" pageId="3" pageNumber="425">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25D2653E1F15FFE26CCEFE43FD714D99" box="[199,736,475,501]" pageId="3" pageNumber="425" type="etymology">
|
||
<paragraph id="6D7736B51F15FFE26CCEFE43FD714D99" blockId="3.[151,1437,151,681]" box="[199,736,475,501]" pageId="3" pageNumber="425">
|
||
<emphasis id="5FBCEAA71F15FFE26CCEFE43FEDF4D99" bold="true" box="[199,334,475,501]" pageId="3" pageNumber="425">Etymology.</emphasis>
|
||
After Anai Valley, the
|
||
<typeStatus id="B27388171F15FFE26E46FE43FDEE4D99" box="[591,639,475,501]" pageId="3" pageNumber="425">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25D2653E1F15FFE26CCEFE67FCB14EC5" pageId="3" pageNumber="425" type="discussion">
|
||
<paragraph id="6D7736B51F15FFE26CCEFE67FCB14EC5" blockId="3.[151,1437,151,681]" pageId="3" pageNumber="425">
|
||
<emphasis id="5FBCEAA71F15FFE26CCEFE67FEAA4E75" bold="true" box="[199,315,511,537]" pageId="3" pageNumber="425">Remarks.</emphasis>
|
||
<taxonomicName id="AAC84D361F15FFE26D4AFE67FD8C4E74" authorityName="Jałoszyński" authorityYear="2024" box="[323,541,511,537]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="anaiensis">
|
||
<emphasis id="5FBCEAA71F15FFE26D4AFE67FD8C4E74" box="[323,541,511,537]" italics="true" pageId="3" pageNumber="425">Loeblites anaiensis</emphasis>
|
||
</taxonomicName>
|
||
is externally unremarkable and the aedeagus must be examined to identify the species. The conspicuously long parameres and the uniquely shaped and subquadrate distal median sclerotized endophallic structure allow for unambiguous identification. The spermatheca is most similar to that of
|
||
<taxonomicName id="AAC84D361F15FFE2693AFDDFFA044E0C" authorityName="Jaloszynski" authorityYear="2005" box="[1331,1429,583,608]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="5FBCEAA71F15FFE2693AFDDFFA044E0C" box="[1331,1429,583,608]" italics="true" pageId="3" pageNumber="425">L. minor</emphasis>
|
||
</taxonomicName>
|
||
, except that the distal margin (i.e., the one opposite the ‘base’, where the ductus spermathecae and accessory gland are inserted) is broadly rounded, and in
|
||
<taxonomicName id="AAC84D361F15FFE26E5CFD17FD274EC4" authorityName="Jaloszynski" authorityYear="2005" box="[597,694,655,680]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="5FBCEAA71F15FFE26E5CFD17FD274EC4" box="[597,694,655,680]" italics="true" pageId="3" pageNumber="425">L. minor</emphasis>
|
||
</taxonomicName>
|
||
concave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |