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<document id="884A8038A9FC4F453291FA36EF18DF0C" ID-CLB-Dataset="298741" ID-DOI="10.11646/zootaxa.5471.4.2" ID-GBIF-Dataset="2370cfa6-ebc7-4962-ae93-ade18759c5cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190291" ID-ZooBank="A625B488-6A82-44F4-99AD-A66CAB9FC996" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718893826382" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="E56187A31F12FFE56C9EFF0FFD8B4B21" docLanguage="en" docName="zootaxa.5471.4.2.pdf" docOrigin="Zootaxa 5471 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loeblites umphangensis Jałoszyński 2024, sp. n." docType="treatment" docVersion="6" lastPageNumber="426" masterDocId="1958FFDB1F16FFE16C09FF98FF914C6C" masterDocTitle="Three new species, one subspecies, and new records of the Oriental genus Loeblites Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="432" masterPageNumber="422" pageNumber="426" updateTime="1719007836080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6AC56D84E1AA69CDD8A1F9D723FFE86D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2C5EFE5B6C7E7CB604C7D8302AB143EB">Three new species, one subspecies, and new records of the Oriental genus Loeblites Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="0CC004ABD36ED15675D49FE06842FD16">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="D48351F5421C476F27D964329C171496">Zootaxa</mods:title>
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<mods:date id="B28CD4F0C5F3E68426D3537836AE6AA9">2024</mods:date>
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<mods:number id="A3764569604533C8A7D8A178D4C7FAE9">2024-06-20</mods:number>
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<mods:number id="16E4A79698A5490616B67142272EA720">4</mods:number>
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<mods:classification id="B833A7BE3D219021D40BD051C4E3770E">journal article</mods:classification>
<mods:identifier id="766EC012E0E02974C5EB5A4BAB790A7E" type="CLB-Dataset">298741</mods:identifier>
<mods:identifier id="C8EED4E619199621C3FAAC69B7B1BC32" type="DOI">10.11646/zootaxa.5471.4.2</mods:identifier>
<mods:identifier id="E4CF01CEF49078CB3325A4EF7362A903" type="GBIF-Dataset">2370cfa6-ebc7-4962-ae93-ade18759c5cc</mods:identifier>
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<treatment id="E56187A31F12FFE56C9EFF0FFD8B4B21" ID-DOI="http://doi.org/10.5281/zenodo.12190846" ID-GBIF-Taxon="233317987" ID-Zenodo-Dep="12190846" LSID="urn:lsid:plazi:treatment:E56187A31F12FFE56C9EFF0FFD8B4B21" httpUri="http://treatment.plazi.org/id/E56187A31F12FFE56C9EFF0FFD8B4B21" lastPageNumber="426" pageId="4" pageNumber="426">
<subSubSection id="25D2653E1F12FFE56C9EFF0FFD944CDD" box="[151,517,151,178]" pageId="4" pageNumber="426" type="nomenclature">
<paragraph id="6D7736B51F12FFE56C9EFF0FFD944CDD" blockId="4.[151,517,151,213]" box="[151,517,151,178]" pageId="4" pageNumber="426">
<heading id="363F81D91F12FFE56C9EFF0FFD944CDD" bold="true" box="[151,517,151,178]" fontSize="11" level="1" pageId="4" pageNumber="426" reason="1">
<emphasis id="5FBCEAA71F12FFE56C9EFF0FFD944CDD" bold="true" box="[151,517,151,178]" pageId="4" pageNumber="426">
<taxonomicName id="AAC84D361F12FFE56C9EFF0FFE2D4CDE" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[151,444,151,178]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="umphangensis" status="sp. nov.">
<emphasis id="5FBCEAA71F12FFE56C9EFF0FFE2D4CDE" bold="true" box="[151,444,151,178]" italics="true" pageId="4" pageNumber="426">Loeblites umphangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="448F57DC1F12FFE56DCAFF0FFD944CDD" box="[451,517,151,177]" pageId="4" pageNumber="426" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56C9EFF23FE224CB9" box="[151,435,187,213]" pageId="4" pageNumber="426" type="description">
<paragraph id="6D7736B51F12FFE56C9EFF23FE224CB9" blockId="4.[151,517,151,213]" box="[151,435,187,213]" pageId="4" pageNumber="426">
(
<figureCitation id="F5F32A301F12FFE56C96FF23FE954CB9" box="[159,260,187,213]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="4" pageNumber="426">Figs 2, 6</figureCitation>
,
<figureCitation id="F5F32A301F12FFE56D19FF23FEC84CB9" box="[272,345,187,213]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">14–18</figureCitation>
,
<figureCitation id="F5F32A301F12FFE56D6DFF23FE134CB9" box="[356,386,187,213]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="4" pageNumber="426">30</figureCitation>
,
<figureCitation id="F5F32A301F12FFE56D87FF23FE3A4CB9" box="[398,427,187,213]" captionStart="FIGURE 32" captionStartId="7.[151,250,1999,2023]" captionTargetBox="[177,1411,849,1976]" captionTargetId="figure-234@7.[177,1411,849,1976]" captionTargetPageId="7" captionText="FIGURE 32. Distribution of nominal species of Loeblites." figureDoi="http://doi.org/10.5281/zenodo.12190305" httpUri="https://zenodo.org/record/12190305/files/figure.png" pageId="4" pageNumber="426">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56C9EFE9BFD1B4DE5" pageId="4" pageNumber="426" type="materials_examined">
<paragraph id="6D7736B51F12FFE56C9EFE9BFD1B4DE5" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56C9EFE9BFEF54D71" bold="true" box="[151,356,259,285]" pageId="4" pageNumber="426">Material studied.</emphasis>
<materialsCitation id="DDA03CE81F12FFE56D63FE9BFDFA4D09" ID-GBIF-Occurrence="4898138306" collectingDate="1993-10-02" collectionCode="MHNG" collectorName="Schwendinger" country="Thailand" determinerName="P. Jaloszynski" elevation="1250" location="Mae sot Umphang" pageId="4" pageNumber="426" specimenCount="1" specimenCount-male="1" stateProvince="Mts. n Umphang" typeStatus="holotype">
<emphasis id="5FBCEAA71F12FFE56D63FE9BFCAC4D71" bold="true" box="[362,829,259,285]" pageId="4" pageNumber="426">
<typeStatus id="B27388171F12FFE56D63FE9BFE4A4D71" box="[362,475,259,285]" pageId="4" pageNumber="426" type="holotype">Holotype</typeStatus>
: THAILAND (Tak Province):
</emphasis>
<specimenCount id="7BCEFD3C1F12FFE56F4DFE9CFCCA4D72" box="[836,859,260,286]" count="1" pageId="4" pageNumber="426" type="male">♂</specimenCount>
, two labels: “
<collectingCountry id="15DF76251F12FFE56FF9FE9BFB194D71" box="[1008,1160,259,285]" name="Thailand" pageId="4" pageNumber="426">THAILAND</collectingCountry>
: /
<collectingRegion id="AF0CF8571F12FFE568A9FE9CFAC44D71" box="[1184,1365,259,286]" pageId="4" pageNumber="426">Mts.n Umphang</collectingRegion>
/
<location id="6817606E1F12FFE56963FE9CFEBD4D2D" LSID="urn:lsid:plazi:treatment:E56187A31F12FFE56C9EFF0FFD8B4B21:6817606E1F12FFE56963FE9CFEBD4D2D" country="Thailand" name="Mae sot Umphang" pageId="4" pageNumber="426" stateProvince="Mts. n Umphang">Mae sot Umphang</location>
/ 1250m 10.2.93 /
<collectorName id="C03D53631F12FFE56DF3FEBFFD0C4D2D" box="[506,669,295,321]" pageId="4" pageNumber="426">Schwendinger</collectorName>
” [white], “
<taxonomicName id="AAC84D361F12FFE56F1EFEBFFC344D2D" authorityName="Franz" authorityYear="1986" box="[791,933,295,321]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="5FBCEAA71F12FFE56F1EFEBFFC344D2D" box="[791,933,295,321]" italics="true" pageId="4" pageNumber="426">LOEBLITES</emphasis>
</taxonomicName>
/
<taxonomicName id="AAC84D361F12FFE56FB1FEBFFBC94D2D" authorityName="Jałoszyński" authorityYear="2024" box="[952,1112,295,321]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="umphangensis">
<emphasis id="5FBCEAA71F12FFE56FB1FEBFFBC94D2D" box="[952,1112,295,321]" italics="true" pageId="4" pageNumber="426">umphangensis</emphasis>
</taxonomicName>
m. / det.
<determinerName id="98ABDA711F12FFE568B4FEB0FACB4D2D" box="[1213,1370,295,322]" pageId="4" pageNumber="426">P. Jałoszyński</determinerName>
, 2024 / HOLOTYPUS” [red, printed] (
<collectionCode id="0BD9AE701F12FFE56E0EFED3FDF24D09" box="[519,611,331,357]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="426" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="DDA03CE81F12FFE56E7EFED3FD174DE5" ID-GBIF-Occurrence="4898138302" collectingDate="1993-10-02" collectionCode="MHNG" collectorName="Schwendinger" country="Thailand" determinerName="P. Jaloszynski" elevation="1250" location="Tak Province" pageId="4" pageNumber="426" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Tak" typeStatus="paratype">
<emphasis id="5FBCEAA71F12FFE56E7EFED3FD7E4D09" bold="true" box="[631,751,331,357]" pageId="4" pageNumber="426">
<typeStatus id="B27388171F12FFE56E7EFED3FD7E4D09" box="[631,751,331,357]" pageId="4" pageNumber="426" type="paratype">Paratypes</typeStatus>
</emphasis>
(9 exx.):
<specimenCount id="7BCEFD3C1F12FFE56F56FED3FC304D0A" box="[863,929,331,358]" count="5" pageId="4" pageNumber="426" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="7BCEFD3C1F12FFE56FA7FED3FC7E4D0A" box="[942,1007,331,358]" count="4" pageId="4" pageNumber="426" type="female">4 ♀♀</specimenCount>
, same data as for holotype, each with yellow “PARATYPUS” label (
<collectionCode id="0BD9AE701F12FFE56DE4FEF7FDD64DE5" box="[493,583,367,393]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="426" type="Museum">MHNG</collectionCode>
, cPJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56CCEFE0BFAC34E75" pageId="4" pageNumber="426" type="diagnosis">
<paragraph id="6D7736B51F12FFE56CCEFE0BFAC34E75" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56CCEFE0BFED34DC1" bold="true" box="[199,322,403,429]" pageId="4" pageNumber="426">Diagnosis.</emphasis>
Pronotum with fine, inconspicuous punctures; aedeagus (
<figureCitation id="F5F32A301F12FFE56FE5FE0BFBD34DC1" box="[1004,1090,403,429]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">Fig. 14</figureCitation>
) stout, with short parameres, endophallus (
<figureCitation id="F5F32A301F12FFE56D38FE2FFE174DBD" box="[305,390,439,465]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">Fig. 18</figureCitation>
) symmetrical, with strongly oblique lateral sclerites and with sclerotized median structure distal to lateral sclerites strongly transverse, much lighter than lateral sclerites, with anterior margin straight at middle and laterally strongly bent lateroproximally; spermatheca in lateral view (
<figureCitation id="F5F32A301F12FFE5682DFE67FBE44E75" box="[1060,1141,511,537]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="4" pageNumber="426">Fig. 30</figureCitation>
) oval and elongate.
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56CCEFDBBFF6D4A3D" pageId="4" pageNumber="426" type="description">
<paragraph id="6D7736B51F12FFE56CCEFDBBFEFA4E0E" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56CCEFDBBFEC84E51" bold="true" box="[199,345,547,573]" pageId="4" pageNumber="426">Description.</emphasis>
Body of male (
<figureCitation id="F5F32A301F12FFE56DF6FDBBFDD04E51" box="[511,577,547,573]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="4" pageNumber="426">Fig. 2</figureCitation>
) slender and strongly convex, light brown, covered with light brown vestiture. BL 2.45–2.48 mm.
</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFDF3FE8B4F55" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
Head in dorsal view transverse, broadest at eyes, HL
<quantity id="AA309B501F12FFE56F4BFDF3FC7A4EEA" box="[834,1003,619,646]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="4" pageNumber="426" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
, HW
<quantity id="AA309B501F12FFE5683CFDF3FB4E4EEA" box="[1077,1247,619,646]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="4" pageNumber="426" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
; vertex strongly transverse and evenly convex, anteriorly confluent with frons; frons flattened between prominent supraantennal tubercles; tempora in dorsal view about as long as eyes; eyes small, oval, with slightly concave posteroventral margin, finely faceted. Vertex and frons covered with fine, inconspicuous punctures and sparse, moderately long, suberect setae. Antennae long and slender, slightly shorter than BL, AnL
<quantity id="AA309B501F12FFE56FCAFD63FBF64F7A" box="[963,1127,763,790]" metricMagnitude="-3" metricUnit="m" metricValue="2.005" metricValueMax="2.08" metricValueMin="1.93" pageId="4" pageNumber="426" unit="mm" value="2.005" valueMax="2.08" valueMin="1.93">1.93–2.08 mm</quantity>
, all antennomeres elongate, as in
<figureCitation id="F5F32A301F12FFE56CDBFC87FE864F55" box="[210,279,799,825]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1606,1630]" captionTargetBox="[156,1431,276,1573]" captionTargetId="figure-54@2.[151,1435,270,1579]" captionTargetPageId="2" captionText="FIGURES 1–8. Males of Loeblites anaiensis sp. n. (1, 5); L. umphangensis sp. n. (2, 6); L. pseudominor pseudominor sp. n. (3, 7); and L. pseudominor sumatrensis subsp. n. (4, 8). Dorsal habitus (1–4) and antenna (5–8)." figureDoi="http://doi.org/10.5281/zenodo.12190293" httpUri="https://zenodo.org/record/12190293/files/figure.png" pageId="4" pageNumber="426">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFCDCFE664F81" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
Pronotum elongate, as broad as head, broadest slightly in front of anterior third; PL
<quantity id="AA309B501F12FFE5688FFCDBFB7C4F32" box="[1158,1261,835,862]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="426" unit="mm" value="0.7">0.70 mm</quantity>
, PW
<quantity id="AA309B501F12FFE56925FCDBFF574FEE" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="4" pageNumber="426" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
. Anterior margin broadly rounded, sides strongly rounded in anterior third, posteriorly broadly and weakly constricted; posterior margin nearly straight. Four dorsal antebasal pits large and deep, two lateral pits at each side distinct. Pronotum with fine, inconspicuous punctures, those behind antebasal pits not differing from those on disc; setae long, sparse and suberect.
</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFC6FFD414812" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
Elytra more convex than pronotum, oval, broadest distinctly in front of middle; EL
<quantity id="AA309B501F12FFE56871FC6FFA8F487E" box="[1144,1310,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" metricValueMax="1.35" metricValueMin="1.33" pageId="4" pageNumber="426" unit="mm" value="1.34" valueMax="1.35" valueMin="1.33">1.33–1.35 mm</quantity>
, EW
<quantity id="AA309B501F12FFE56954FC6FFF6D485A" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="8.8" metricValueMin="8.299999999999999" pageId="4" pageNumber="426" unit="mm" value="0.855" valueMax="0.88" valueMin="0.83">0.83– 0.88 mm</quantity>
, EI 1.56–1.61. Humeral calli weakly elevated, elongate; apex of each elytron subtriangular, sharply-angled but with blunt tip. Punctures slightly more distinct than those on head and pronotum, but superficial, diffuse and inconspicuous; setae similar to those on pronotum.
</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFB10FB8E48A9" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">Legs long and slender; femora abruptly clavate, dorsal longitudinal grooves on femoral clava complete and with distinct glandular opening at highest site of each femur; all tibiae nearly straight.</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFB57FAED49AE" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
Aedeagus (
<figureCitation id="F5F32A301F12FFE56D4CFB57FE544885" box="[325,453,1231,1257]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">Figs 14–18</figureCitation>
) elongate but relatively stout; AeL
<quantity id="AA309B501F12FFE56F5CFB57FC6A4886" box="[853,1019,1231,1258]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="4" pageNumber="426" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
; median lobe in ventral view equally broad near base and near middle, distal region rapidly narrowing towards broadly subtriangular apex, with its distal margin distinctly angulate; endophallus (
<figureCitation id="F5F32A301F12FFE56E55FA8FFD3D495D" box="[604,684,1303,1329]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">Fig. 18</figureCitation>
) with pair of strongly elongate and oblique lateral sclerites (
<figureCitation id="F5F32A301F12FFE5694CFA8FFA06495D" box="[1349,1431,1303,1329]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1960,1984]" captionTargetBox="[157,1436,711,1930]" captionTargetId="figure-225@3.[151,1436,705,1936]" captionTargetPageId="3" captionText="FIGURES 9–18. Loeblites anaiensis sp. n. (9–13); L. umphangensis sp. n. (14–18). Aedeagus in ventral (9, 11, 14, 16) and lateral (10, 12, 15, 17) views, and close-up of endophallic sclerites in ventral view (13, 18)." figureDoi="http://doi.org/10.5281/zenodo.12190295" httpUri="https://zenodo.org/record/12190295/files/figure.png" pageId="4" pageNumber="426">Fig. 18</figureCitation>
;
<emphasis id="5FBCEAA71F12FFE56C9EFAA3FF274939" box="[151,182,1339,1365]" italics="true" pageId="4" pageNumber="426">lsc</emphasis>
) distally connected with small and lightly pigmented median sclerotized structure, the latter strongly transverse and with distal margin straight at middle and laterally strongly bent lateroproximally, proximally lateral sclerites connected with sclerotized median proximal complex, with its proximal margin rounded and strongly convex; parameres shorter than median lobe and their apices only slightly exceeding apex of median lobe, lacking setae.
</paragraph>
<paragraph id="6D7736B51F12FFE56CCEFA53FF6D4A3D" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
Female. Externally similar to male, with antennae indistinctly shorter in relation to body; BL
<quantity id="AA309B501F12FFE568C0FA53FAFF498A" box="[1225,1390,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.65" metricValueMin="2.45" pageId="4" pageNumber="426" unit="mm" value="2.55" valueMax="2.65" valueMin="2.45">2.45–2.65 mm</quantity>
; HL
<quantity id="AA309B501F12FFE56C9EFA77FEAD4A66" box="[151,316,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="4" pageNumber="426" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
, HW
<quantity id="AA309B501F12FFE56D75FA77FDB04A66" box="[380,545,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="4" pageNumber="426" unit="mm" value="0.575" valueMax="0.6" valueMin="0.55">0.55–0.60 mm</quantity>
, AnL
<quantity id="AA309B501F12FFE56E6AFA77FC984A66" box="[611,777,1519,1546]" metricMagnitude="-3" metricUnit="m" metricValue="2.005" metricValueMax="2.08" metricValueMin="1.93" pageId="4" pageNumber="426" unit="mm" value="2.005" valueMax="2.08" valueMin="1.93">1.93–2.08 mm</quantity>
; PL
<quantity id="AA309B501F12FFE56F33FA77FC4E4A66" box="[826,991,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="4" pageNumber="426" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.70–0.75 mm</quantity>
, PW
<quantity id="AA309B501F12FFE56813FA77FB514A66" box="[1050,1216,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="4" pageNumber="426" unit="mm" value="0.575" valueMax="0.6" valueMin="0.55">0.55–0.60 mm</quantity>
; EL
<quantity id="AA309B501F12FFE568FAFA77FA094A66" box="[1267,1432,1519,1546]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.45" metricValueMin="1.33" pageId="4" pageNumber="426" unit="mm" value="1.39" valueMax="1.45" valueMin="1.33">1.33–1.45 mm</quantity>
, EW
<quantity id="AA309B501F12FFE56CC3F98BFEE04A42" box="[202,369,1555,1582]" metricMagnitude="-4" metricUnit="m" metricValue="9.05" metricValueMax="9.3" metricValueMin="8.8" pageId="4" pageNumber="426" unit="mm" value="0.905" valueMax="0.93" valueMin="0.88">0.88–0.93 mm</quantity>
, EI 1.51–1.57. Spermatheca (
<figureCitation id="F5F32A301F12FFE56EC9F98BFC824A41" box="[704,787,1555,1581]" captionStart="FIGURES 29–31" captionStartId="6.[151,264,1151,1175]" captionTargetBox="[157,1430,316,1121]" captionTargetId="figure-66@6.[151,1436,309,1127]" captionTargetPageId="6" captionText="FIGURES 29–31. Spermatheca of Loeblites anaiensis sp. n. (29); L. umphangensis sp. n. (30); and L. pseudominor pseudominor sp. n. (31)." figureDoi="http://doi.org/10.5281/zenodo.12190303" httpUri="https://zenodo.org/record/12190303/files/figure.png" pageId="4" pageNumber="426">Fig. 30</figureCitation>
) in lateral view
<quantity id="AA309B501F12FFE56FC7F98BFBD04A42" box="[974,1089,1555,1582]" metricMagnitude="-5" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="426" unit="mm" value="0.063">0.063 mm</quantity>
wide, oval, with narrow distal ‘pocket’.
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56CCEF9C3FD364A19" box="[199,679,1627,1653]" pageId="4" pageNumber="426" type="distribution">
<paragraph id="6D7736B51F12FFE56CCEF9C3FD364A19" blockId="4.[151,1437,259,1869]" box="[199,679,1627,1653]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56CCEF9C3FEF04A19" bold="true" box="[199,353,1627,1653]" pageId="4" pageNumber="426">Distribution.</emphasis>
Northern
<collectingCountry id="15DF76251F12FFE56DDAF9C3FDA64A19" box="[467,567,1627,1653]" name="Thailand" pageId="4" pageNumber="426">Thailand</collectingCountry>
(
<figureCitation id="F5F32A301F12FFE56E4FF9C3FD0B4A19" box="[582,666,1627,1653]" captionStart="FIGURE 32" captionStartId="7.[151,250,1999,2023]" captionTargetBox="[177,1411,849,1976]" captionTargetId="figure-234@7.[177,1411,849,1976]" captionTargetPageId="7" captionText="FIGURE 32. Distribution of nominal species of Loeblites." figureDoi="http://doi.org/10.5281/zenodo.12190305" httpUri="https://zenodo.org/record/12190305/files/figure.png" pageId="4" pageNumber="426">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56CCEF9E7FCB84AF5" box="[199,809,1663,1690]" pageId="4" pageNumber="426" type="etymology">
<paragraph id="6D7736B51F12FFE56CCEF9E7FCB84AF5" blockId="4.[151,1437,259,1869]" box="[199,809,1663,1690]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56CCEF9E7FEDF4AF5" bold="true" box="[199,334,1663,1689]" pageId="4" pageNumber="426">Etymology.</emphasis>
After Umphang District, the
<typeStatus id="B27388171F12FFE56E9EF918FD564AF6" box="[663,711,1664,1690]" pageId="4" pageNumber="426">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="25D2653E1F12FFE56CCEF93BFD8B4B21" pageId="4" pageNumber="426" type="discussion">
<paragraph id="6D7736B51F12FFE56CCEF93BFD8B4B21" blockId="4.[151,1437,259,1869]" pageId="4" pageNumber="426">
<emphasis id="5FBCEAA71F12FFE56CCEF93BFEAA4AD1" bold="true" box="[199,315,1699,1725]" pageId="4" pageNumber="426">Remarks.</emphasis>
<taxonomicName id="AAC84D361F12FFE56D4DF93BFD8C4AD1" authorityName="Jałoszyński" authorityYear="2024" box="[324,541,1699,1725]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="anaiensis">
<emphasis id="5FBCEAA71F12FFE56D4DF93BFD8C4AD1" box="[324,541,1699,1725]" italics="true" pageId="4" pageNumber="426">Loeblites anaiensis</emphasis>
</taxonomicName>
is externally unremarkable (except that all studied specimens are light brown, while in all remaining species fully pigmented adults are dark brown), and the aedeagus must be examined to identify the species. The lightly pigmented and strongly transverse distal median sclerotized endophallic structure allows for unambiguous identification. The spermatheca is most similar to that of
<taxonomicName id="AAC84D361F12FFE5683AF888FB784B45" authorityName="Franz" authorityYear="1986" box="[1075,1257,1808,1833]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="mastigicornis">
<emphasis id="5FBCEAA71F12FFE5683AF888FB784B45" box="[1075,1257,1808,1833]" italics="true" pageId="4" pageNumber="426">L. mastigicornis</emphasis>
</taxonomicName>
, except that the distal ‘pocket’ is distinctly shorter.
</paragraph>
</subSubSection>
</treatment>
</document>
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