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<document id="678BB04474E6FB8E94EAEEDAC055158B" ID="10.11646/zootaxa.4117.1.10" ID-CLB-Dataset="38970" ID-DOI="10.11646/zootaxa.4117.1.10" ID-GBIF-Dataset="07aea016-04f8-4440-8f83-13ed74c2b0bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="255511" ID-ZooBank="9E563E76-E83B-45A2-BE87-2A552F95FD14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464592098733" checkinUser="plazi" docAuthor="Hibino, Yusuke &amp; Kimura, Seishi" docDate="2016" docId="DF525071CE06FFA8C3ABE9DCFDBFFDA9" docLanguage="en" docName="data.AD6B2A2E27B6520A7B419389A1ED8602.cached" docOrigin="Zootaxa 4117 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apterichtus succinus Hibino &amp; Kimura, 2016, sp. nov." docType="treatment" docUuid="734F8D4B-17C2-4E1E-ADE4-77FA3409619E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="150" masterDocId="236B2809CE06FFACC33CEF47FF94FF94" masterDocTitle="A new species of Apterichtus (Anguilliformes: Ophichthidae) from the Marquesas Islands" masterLastPageNumber="150" masterPageNumber="146" pageNumber="146" updateTime="1698662648725" updateUser="ExternalLinkService">
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<mods:title id="B295A1ED7CD15D6A961E596137E9F1F9">A new species of Apterichtus (Anguilliformes: Ophichthidae) from the Marquesas Islands</mods:title>
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<mods:namePart id="5D20423CB9A3B231585F3AD5335B85EE">Hibino, Yusuke</mods:namePart>
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<mods:namePart id="D23304280D40195146356593EF22A76F">Kimura, Seishi</mods:namePart>
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<mods:typeOfResource id="33023D84BB425043C7C30CB887D4EBEF">text</mods:typeOfResource>
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<mods:title id="A9578954047CA68E82D542EFF08A86E0">Zootaxa</mods:title>
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<mods:part id="AA150ABA6CC75C0FEC7B1E8C2B218B2A">
<mods:date id="59601E902501C59A0745B032EEB4EF0C">2016</mods:date>
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<mods:number id="C9FBC7EA6A67D0E5AD81A0FF9FBA7C9E">4117</mods:number>
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<treatment id="DF525071CE06FFA8C3ABE9DCFDBFFDA9" ID-DOI="http://doi.org/10.5281/zenodo.6055264" ID-GBIF-Taxon="120733371" ID-Zenodo-Dep="6055264" LSID="urn:lsid:zoobank.org:act:734F8D4B-17C2-4E1E-ADE4-77FA3409619E" httpUri="http://treatment.plazi.org/id/DF525071CE06FFA8C3ABE9DCFDBFFDA9" lastPageId="4" lastPageNumber="150" pageId="0" pageNumber="146">
<subSubSection id="1FE1B2ECCE06FFACC3ABE9DCFC12F888" pageId="0" pageNumber="146" type="nomenclature">
<paragraph id="5744E167CE06FFACC3ABE9DCFE6DF921" blockId="0.[151,505,1691,1750]" box="[151,505,1691,1717]" pageId="0" pageNumber="146">
<heading id="0C0C560BCE06FFACC3ABE9DCFE6DF921" bold="true" box="[151,505,1691,1717]" fontSize="11" level="1" pageId="0" pageNumber="146" reason="1">
<emphasis id="658F3D75CE06FFACC3ABE9DCFE6DF921" bold="true" box="[151,505,1691,1717]" pageId="0" pageNumber="146">
<taxonomicName id="90FB9AE4CE06FFACC3ABE9DCFE00F921" ID-CoL="FYHN" box="[151,404,1691,1717]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="0" pageNumber="146" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE06FFACC3ABE9DCFE00F921" bold="true" box="[151,404,1691,1717]" italics="true" pageId="0" pageNumber="146">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE06FFACC2A7E9DCFE6DF921" box="[411,505,1691,1717]" pageId="0" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5744E167CE06FFACC3ABE9FAFE32F942" blockId="0.[151,505,1691,1750]" box="[151,422,1725,1750]" pageId="0" pageNumber="146">
<figureCitation id="CFC0FDE2CE06FFACC3ABE9FAFE94F942" box="[151,256,1725,1750]" captionStart="FIGURE 1" captionStartId="1.[151,250,1231,1253]" captionTargetBox="[183,1388,490,1202]" captionTargetId="figure@1.[173,1414,481,1210]" captionTargetPageId="1" captionText="FIGURE 1. Apterichtus succinus sp. nov., holotype, USNM 409323, 175 mm TL, Marquesas Islands (fresh condition, photographed by Jeffery T. Williams)." httpUri="https://zenodo.org/record/255512/files/figure.png" pageId="0" pageNumber="146">Figures 1</figureCitation>
,
<figureCitation id="CFC0FDE2CE06FFACC232E9F9FE8AF942" box="[270,286,1726,1750]" captionStart="FIGURE 2" captionStartId="2.[151,250,1174,1196]" captionTargetBox="[167,1427,412,1147]" captionTargetId="figure@2.[159,1427,409,1153]" captionTargetPageId="2" captionText="FIGURE 2. Sensory pores on head (a), teeth on upper jaw and palatal area (b), and those of mandible (c) of Apterichtus succinus sp. nov., holotype, USNM 409323. IMT, intermaxillary teeth; IO, infraorbital pores; LL, lateral-line pores; MD, mandibular pores; PO, preopercular pores; SO, supraorbital pores; ST, supratemporal pores; UJT, upper-jaw teeth; VT, vomerine tooth. Arrows indicate interorbital (left) and mid-temporal (right) pores." httpUri="https://zenodo.org/record/255513/files/figure.png" pageId="0" pageNumber="146">2</figureCitation>
,
<figureCitation id="CFC0FDE2CE06FFACC216E9F9FEACF942" box="[298,312,1726,1750]" captionStart="FIGURE 3" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[168,1436,781,1906]" captionTargetId="figure@3.[151,1436,772,1915]" captionTargetPageId="3" captionText="FIGURE 3. Marking patterns of head and nape, traced from fresh color photographs: (a) Apterichtus succinus sp. nov., holotype, USNM 409323; (b) A. klazingai, USNM 375002, 247 mm TL, Ile Uvea, Wallis Island; (c) A. klazingai, BPBM 36764, 298 mm TL, Maumere Bay, Flores, Indonesia." httpUri="https://zenodo.org/record/255514/files/figure.png" pageId="0" pageNumber="146">3</figureCitation>
a;
<tableCitation id="1A79D4DCCE06FFACC26FE9FAFE32F942" box="[339,422,1725,1750]" captionStart="TABLE 1" captionStartId="3.[151,239,296,319]" captionText="TABLE 1. Frequency distribution of vertebral counts in Apterichtus succinus sp. nov. and A. klazingai." pageId="0" pageNumber="146">Table 1</tableCitation>
</paragraph>
<paragraph id="5744E167CE06FFACC3ABE841FC12F888" blockId="0.[151,902,1798,1820]" box="[151,902,1798,1820]" pageId="0" pageNumber="146">
<treatmentCitationGroup id="77EBC649CE06FFACC3ABE841FC12F888" box="[151,902,1798,1820]" pageId="0" pageNumber="146">
<taxonomicName id="90FB9AE4CE06FFACC3ABE841FE98F888" ID-CoL="ZCD" box="[151,268,1798,1820]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="0" pageNumber="146" phylum="Chordata" rank="genus">
<emphasis id="658F3D75CE06FFACC3ABE841FE98F888" box="[151,268,1798,1820]" italics="true" pageId="0" pageNumber="146">Apterichtus</emphasis>
</taxonomicName>
sp.:
<treatmentCitation id="D65AC776CE06FFACC200E841FDFCF888" author="Delrieu-Trottin" box="[316,616,1798,1820]" page="4" pageId="0" pageNumber="146" year="2015">
<bibRefCitation id="336A9C96CE06FFACC200E841FDC5F888" author="Delrieu-Trottin" box="[316,593,1798,1820]" pageId="0" pageNumber="146" refString="Delrieu-Trottin, E., Williams, J. T., Bacchet, P., Kulbicki, M., Mourier, J., Galzin, R., Lison de Loma, T., Mou-Tham, G., Siu, G. &amp; Planes, S. (2015) Shore fishes of the Marquesas Islands, an updated checklist with new records and new percentage of endemic species. Check List, 11, 1 - 13. http: // dx. doi. org / 10.15560 / 11.5.1758" type="journal article" year="2015">
Delrieu-Trottin
<emphasis id="658F3D75CE06FFACC2E0E840FD9DF888" box="[476,521,1798,1820]" italics="true" pageId="0" pageNumber="146">et al</emphasis>
., 2015
</bibRefCitation>
: 4
</treatmentCitation>
(listed; Marquesas Islands).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1FE1B2ECCE06FFACC3ABE80FFEF9F83D" pageId="0" pageNumber="146" type="materials_examined">
<paragraph id="5744E167CE06FFACC3ABE80FFEF9F83D" blockId="0.[151,1436,1864,2033]" pageId="0" pageNumber="146">
<emphasis id="658F3D75CE06FFACC3ABE80FFE9FF8F5" bold="true" box="[151,267,1864,1889]" pageId="0" pageNumber="146">
<typeStatus id="88405FC5CE06FFACC3ABE80FFE92F8F5" box="[151,262,1864,1889]" pageId="0" pageNumber="146" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="31EA79A2CE06FFACC22EE80EFEF7F8F5" LSID="urn:lsid:biocol.org:col:34871" box="[274,355,1865,1889]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="146">USNM</collectionCode>
<quantity id="90034C82CE06FFACC255E80EFDBCF8F4" box="[361,552,1864,1889]" metricMagnitude="5" metricUnit="m" metricValue="4.093231749999999" pageId="0" pageNumber="146" unit="mm" value="4.09323175E8">409323, 175 mm</quantity>
TL, female with mature eggs (
<quantity id="90034C82CE06FFACC0BDE80EFB9EF8F4" box="[897,1034,1865,1889]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="0" pageNumber="146" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70.8 mm</quantity>
in diameter), 9.386 S, 140.119 W, Bay of Vaiehu, Ura Pou Island, Marquesas Islands, depth
<quantity id="90034C82CE06FFACC019E82BFCF4F810" box="[805,864,1900,1925]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="146" unit="m" value="25.0">25 m</quantity>
, flat sand bottom, collected by Jeffery T. Williams
<emphasis id="658F3D75CE06FFACC3ABE8D6FF5BF83D" box="[151,207,1937,1961]" italics="true" pageId="0" pageNumber="146">et al.</emphasis>
,
<date id="2345C7A7CE06FFACC3E1E8D6FEFDF83D" box="[221,361,1937,1961]" pageId="0" pageNumber="146" value="2011-11-04">4 Nov. 2011</date>
.
</paragraph>
</subSubSection>
<subSubSection id="1FE1B2ECCE06FFADC3FBE8F3FAB9FE39" lastPageId="1" lastPageNumber="147" pageId="0" pageNumber="146" type="diagnosis">
<paragraph id="5744E167CE06FFADC3FBE8F3FC5DFF6C" blockId="0.[151,1436,1864,2033]" lastBlockId="1.[151,1436,151,429]" lastPageId="1" lastPageNumber="147" pageId="0" pageNumber="146">
<emphasis id="658F3D75CE06FFACC3FBE8F3FED5F859" bold="true" box="[199,321,1972,1997]" pageId="0" pageNumber="146">Diagnosis.</emphasis>
An elongate species of
<taxonomicName id="90FB9AE4CE06FFACC162E8F2FD4BF859" box="[606,735,1973,1997]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="0" pageNumber="146" phylum="Chordata" rank="genus">
<emphasis id="658F3D75CE06FFACC162E8F2FD4BF859" box="[606,735,1973,1997]" italics="true" pageId="0" pageNumber="146">Apterichtus</emphasis>
</taxonomicName>
with the following combination of characters: tail 1.8, head 14.8, and body depth
<quantity id="90034C82CE06FFACC2AEE89FFE76F865" box="[402,482,2008,2033]" metricMagnitude="0" metricUnit="m" metricValue="1.7729199999999998" pageId="0" pageNumber="146" unit="in" value="69.8">69.8 in</quantity>
total length; tip of lower jaw between verticals through anterior margin of eye and mid-eye; four preopercular and five supratemporal pores; teeth conical, uniserial on jaws, a single vomerine tooth; head and breast white, body yellowish brown with numerous small orange-brown spots mainly larger than eye on head and nape but a few spots smaller than a half of eye; and VF 54-133.
</paragraph>
<paragraph id="5744E167CE07FFADC3FBEE43FAB9FE39" blockId="1.[151,1436,151,429]" pageId="1" pageNumber="147">
<emphasis id="658F3D75CE07FFADC3FBEE43FCDFFE89" bold="true" box="[199,843,260,285]" pageId="1" pageNumber="147">
Counts and measurements (in mm) of the
<typeStatus id="88405FC5CE07FFADC1E1EE43FCD2FE89" box="[733,838,260,285]" pageId="1" pageNumber="147" type="holotype">holotype</typeStatus>
.
</emphasis>
Preanal vertebrae 54; total vertebrae 133; preanal lateral-line pores 55, 6 pores in branchial region. Total length 175; head 11.8; trunk 62.8; tail 99.4; body depth at gill openings 2.5; body width at gill openings 1.9; body depth at anus 2.2; body width at anus 2.0; head depth at branchial basket 2.9; head width at branchial basket 2.3; snout length 1.4; tip of snout to tip of lower jaw 2.1, tip of snout to rictus 4.5; eye diameter 1.1; interorbital distance 0.4; gill-opening length 1.6; isthmus width 0.5.
</paragraph>
</subSubSection>
<caption id="0384B1EFCE07FFADC3ABEB88FD8EFA90" httpUri="https://zenodo.org/record/255512/files/figure.png" pageId="1" pageNumber="147" targetBox="[183,1388,490,1202]" targetPageId="1">
<paragraph id="5744E167CE07FFADC3ABEB88FD8EFA90" blockId="1.[151,1436,1231,1284]" pageId="1" pageNumber="147">
<emphasis id="658F3D75CE07FFADC3ABEB88FE8CFB70" bold="true" box="[151,280,1231,1253]" pageId="1" pageNumber="147">FIGURE 1.</emphasis>
<taxonomicName id="90FB9AE4CE07FFADC218EB88FE6FFB72" box="[292,507,1231,1254]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="1" pageNumber="147" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE07FFADC218EB88FE6FFB72" box="[292,507,1231,1254]" italics="true" pageId="1" pageNumber="147">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE07FFADC13BEB97FDC8FB71" box="[519,604,1232,1253]" pageId="1" pageNumber="147" rank="species">
<emphasis id="658F3D75CE07FFADC13BEB97FDC8FB71" bold="true" box="[519,604,1232,1253]" pageId="1" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, USNM 409323, 175 mm TL, Marquesas Islands (fresh condition, photographed by Jeffery T. Williams).
</paragraph>
</caption>
<subSubSection id="1FE1B2ECCE07FFAEC3FBEA6CFBC4FF40" lastPageId="2" lastPageNumber="148" pageId="1" pageNumber="147" type="description">
<paragraph id="5744E167CE07FFADC3FBEA6CFB80F83C" blockId="1.[151,1437,1323,2032]" pageId="1" pageNumber="147">
<emphasis id="658F3D75CE07FFADC3FBEA6CFECDFAD0" bold="true" box="[199,345,1323,1348]" pageId="1" pageNumber="147">Description.</emphasis>
Body elongate, subcylindrical, tip of tail strongly laterally compressed and extremely pointed (
<figureCitation id="CFC0FDE2CE07FFADC3A3EA08FF74FAFC" box="[159,224,1359,1384]" captionStart="FIGURE 1" captionStartId="1.[151,250,1231,1253]" captionTargetBox="[183,1388,490,1202]" captionTargetId="figure@1.[173,1414,481,1210]" captionTargetPageId="1" captionText="FIGURE 1. Apterichtus succinus sp. nov., holotype, USNM 409323, 175 mm TL, Marquesas Islands (fresh condition, photographed by Jeffery T. Williams)." httpUri="https://zenodo.org/record/255512/files/figure.png" pageId="1" pageNumber="147">Fig. 1</figureCitation>
). Head short; preanal length much shorter than tail,
<quantity id="90034C82CE07FFADC01AEA17FCF1FAFC" box="[806,869,1359,1384]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="1" pageNumber="147" unit="in" value="2.3">2.3 in</quantity>
TL. Snout short, its length slightly longer than eye diameter; its tip prominently pointed; short median groove ventrally on snout, its anterior tip below anteriormost margin of second infraorbital pore; slope of dorsal surface of snout approximately 40° relative to underside of snout. Anterior nostril tubular but extremely short; posterior nostril oval in shape with an inner valve, located on anteroventral margin of eye, opening ventrally. Eyes moderate in size, covered by a transparent skin; center of eye anterior to mid-jaw. Mouth inferior, distance from tip of snout to anterior tip of lower jaw almost equal to twice eye diameter; lower jaw short, its tip pointed and located between verticals through anterior margin of eye and center of eye; rictus short, posterior end well behind a vertical through posterior margin of eye; lips smooth, with distinct folds, fold on upper lip extending from a vertical through anterior margin of posterior nostril to posterior end of rictus. Gill openings entirely ventral, slit-like in shape, with its diameter less than twice eye diameter; anterior margins of gill openings close-set but not connected. Sensory pores on head developed, arrangement of those pores as follows (
<figureCitation id="CFC0FDE2CE07FFADC225E99CFECAF960" box="[281,350,1755,1780]" captionStart="FIGURE 2" captionStartId="2.[151,250,1174,1196]" captionTargetBox="[167,1427,412,1147]" captionTargetId="figure@2.[159,1427,409,1153]" captionTargetPageId="2" captionText="FIGURE 2. Sensory pores on head (a), teeth on upper jaw and palatal area (b), and those of mandible (c) of Apterichtus succinus sp. nov., holotype, USNM 409323. IMT, intermaxillary teeth; IO, infraorbital pores; LL, lateral-line pores; MD, mandibular pores; PO, preopercular pores; SO, supraorbital pores; ST, supratemporal pores; UJT, upper-jaw teeth; VT, vomerine tooth. Arrows indicate interorbital (left) and mid-temporal (right) pores." httpUri="https://zenodo.org/record/255513/files/figure.png" pageId="1" pageNumber="147">Fig. 2</figureCitation>
a): 1 + 6 on supraorbital, 3 + 4 on infraorbital, five on lower jaw, four on preopercle, and five on supratemporal, one of those on mid-temporal; a single median interorbital pore. Lateral-line pores small but obvious. Lateral line almost complete except for near tip of tail. Teeth (
<figureCitation id="CFC0FDE2CE07FFADC0D8E864FBA2F8A8" box="[996,1078,1827,1852]" captionStart="FIGURE 2" captionStartId="2.[151,250,1174,1196]" captionTargetBox="[167,1427,412,1147]" captionTargetId="figure@2.[159,1427,409,1153]" captionTargetPageId="2" captionText="FIGURE 2. Sensory pores on head (a), teeth on upper jaw and palatal area (b), and those of mandible (c) of Apterichtus succinus sp. nov., holotype, USNM 409323. IMT, intermaxillary teeth; IO, infraorbital pores; LL, lateral-line pores; MD, mandibular pores; PO, preopercular pores; SO, supraorbital pores; ST, supratemporal pores; UJT, upper-jaw teeth; VT, vomerine tooth. Arrows indicate interorbital (left) and mid-temporal (right) pores." httpUri="https://zenodo.org/record/255513/files/figure.png" pageId="1" pageNumber="147">Figs. 2</figureCitation>
b, c) pointed, slightly recurved posteriorly; teeth on jaws uniserial; a single vomerine tooth on anterormost vomer, smoothly connected with upperjaw teeth; intermaxillary teeth larger and more slender than jaw teeth, arranged in a chevron shape along with edge of pre-ethmoid; intermaxillary teeth visible when mouth closed. All fins absent.
</paragraph>
<paragraph id="5744E167CE07FFAEC3FBE8F3FBC4FF40" blockId="1.[151,1437,1323,2032]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="148" pageId="1" pageNumber="147">
<emphasis id="658F3D75CE07FFADC3FBE8F3FEDEF858" box="[199,330,1972,1996]" italics="true" pageId="1" pageNumber="147">Coloration.</emphasis>
Ground color of body light reddish brown except white head and nape; head and nape with vivid reddish oval or circular spots, mostly equal to or slightly larger than eye diameter, only a few spots smaller than half of eye diameter (
<figureCitation id="CFC0FDE2CE04FFAEC2A5EFD0FE4BFF24" box="[409,479,151,176]" captionStart="FIGURE 3" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[168,1436,781,1906]" captionTargetId="figure@3.[151,1436,772,1915]" captionTargetPageId="3" captionText="FIGURE 3. Marking patterns of head and nape, traced from fresh color photographs: (a) Apterichtus succinus sp. nov., holotype, USNM 409323; (b) A. klazingai, USNM 375002, 247 mm TL, Ile Uvea, Wallis Island; (c) A. klazingai, BPBM 36764, 298 mm TL, Maumere Bay, Flores, Indonesia." httpUri="https://zenodo.org/record/255514/files/figure.png" pageId="2" pageNumber="148">Fig. 3</figureCitation>
a); remainder of body with numerous faint brown dots; iris light blue. In alcohol preservative, body mostly pale yellow, with faint pale brown spots on head and nape.
</paragraph>
</subSubSection>
<subSubSection id="1FE1B2ECCE04FFAEC3FBEF98FE3AFE89" pageId="2" pageNumber="148" type="etymology">
<paragraph id="5744E167CE04FFAEC3FBEF98FE3AFE89" blockId="2.[151,1436,151,357]" pageId="2" pageNumber="148">
<emphasis id="658F3D75CE04FFAEC3FBEF98FEDAFF6C" bold="true" box="[199,334,223,248]" pageId="2" pageNumber="148">Etymology.</emphasis>
The scientific name
<taxonomicName id="90FB9AE4CE04FFAEC104EFA7FD0CFF63" box="[568,664,224,247]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="succinus">
<emphasis id="658F3D75CE04FFAEC104EFA7FD0CFF63" box="[568,664,224,247]" italics="true" pageId="2" pageNumber="148">succinus</emphasis>
</taxonomicName>
, the Latin name of amber, refers to its coloration and the shape of its head and nape markings.
</paragraph>
</subSubSection>
<subSubSection id="1FE1B2ECCE04FFAEC3FBEE60FEFAFEF1" pageId="2" pageNumber="148" type="distribution">
<paragraph id="5744E167CE04FFAEC3FBEE60FEFAFEF1" blockId="2.[151,1436,151,357]" pageId="2" pageNumber="148">
<emphasis id="658F3D75CE04FFAEC3FBEE60FEF4FED4" bold="true" box="[199,352,295,320]" pageId="2" pageNumber="148">Distribution.</emphasis>
Known only a single specimen collected from shallow sandy bottom (
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depth) of the Marquesas Islands.
</paragraph>
</subSubSection>
<caption id="0384B1EFCE04FFAEC3ABEBD1FC42FA9C" httpUri="https://zenodo.org/record/255513/files/figure.png" pageId="2" pageNumber="148" targetBox="[167,1427,412,1147]" targetPageId="2">
<paragraph id="5744E167CE04FFAEC3ABEBD1FC42FA9C" blockId="2.[151,1436,1174,1288]" pageId="2" pageNumber="148">
<emphasis id="658F3D75CE04FFAEC3ABEBD1FE82FB38" bold="true" box="[151,278,1174,1196]" pageId="2" pageNumber="148">FIGURE 2.</emphasis>
Sensory pores on head (
<emphasis id="658F3D75CE04FFAEC123EBD0FDBFFB38" bold="true" box="[543,555,1175,1196]" pageId="2" pageNumber="148">a</emphasis>
), teeth on upper jaw and palatal area (
<emphasis id="658F3D75CE04FFAEC0F6EBD0FC4CFB38" bold="true" box="[970,984,1175,1196]" pageId="2" pageNumber="148">b</emphasis>
), and those of mandible (
<emphasis id="658F3D75CE04FFAEC7D0EBD0FB63FB38" bold="true" box="[1260,1271,1175,1196]" pageId="2" pageNumber="148">c</emphasis>
) of
<taxonomicName id="90FB9AE4CE04FFAEC61BEBD1FF7AFB5F" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE04FFAEC61BEBD1FF7AFB5F" italics="true" pageId="2" pageNumber="148">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE04FFAEC3C4EBF1FED8FB5F" box="[248,332,1206,1227]" pageId="2" pageNumber="148" rank="species">
<emphasis id="658F3D75CE04FFAEC3C4EBF1FED8FB5F" bold="true" box="[248,332,1206,1227]" pageId="2" pageNumber="148">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, USNM 409323. IMT, intermaxillary teeth; IO, infraorbital pores; LL, lateral-line pores; MD, mandibular pores; PO, preopercular pores; SO, supraorbital pores; ST, supratemporal pores; UJT, upper-jaw teeth; VT, vomerine tooth. Arrows indicate interorbital (left) and mid-temporal (right) pores.
</paragraph>
</caption>
<subSubSection id="1FE1B2ECCE04FFA8C3FBEA68FD5EFE61" lastPageId="4" lastPageNumber="150" pageId="2" pageNumber="148" type="discussion">
<paragraph id="5744E167CE04FFAFC3FBEA68FBB3FF6C" blockId="2.[151,1436,1327,2037]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148">
<emphasis id="658F3D75CE04FFAEC3FBEA68FEAFFADC" bold="true" box="[199,315,1327,1352]" pageId="2" pageNumber="148">Remarks.</emphasis>
The combination of preopercular and supratemporal pore counts is one of the most important diagnostic characters in the genus (
<bibRefCitation id="336A9C96CE04FFAEC10EEA13FCE4FAF9" author="McCosker" box="[562,880,1364,1389]" pageId="2" pageNumber="148" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino, 2015</bibRefCitation>
). Although the present new species agrees with
<taxonomicName id="90FB9AE4CE04FFAEC3ABEA3FFD80FA04" authority="Bohlke, 1968" authorityName="Bohlke" authorityYear="1968" box="[151,532,1399,1424]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="ansp">
<emphasis id="658F3D75CE04FFAEC3ABEA3FFECDFA04" box="[151,345,1400,1424]" italics="true" pageId="2" pageNumber="148">Apterichtus ansp</emphasis>
(
<bibRefCitation id="336A9C96CE04FFAEC250EA30FD98FA04" author="Bohlke" box="[364,524,1399,1424]" pageId="2" pageNumber="148" refString="Bohlke, J. E. (1968) A new species of the ophichthid eel genus Verma from the west Atlantic, with comments on related species. Notulae Naturae, 415, 1 - 12." type="journal article" year="1968">Böhlke, 1968</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="90FB9AE4CE04FFAEC119EA3FFC5BFA04" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[549,975,1399,1424]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="caecus">
<emphasis id="658F3D75CE04FFAEC119EA3FFD6AFA04" box="[549,766,1400,1424]" italics="true" pageId="2" pageNumber="148">Apterichtus caecus</emphasis>
(
<bibRefCitation id="336A9C96CE04FFAEC02DEA30FC53FA04" author="Linnaeus" box="[785,967,1399,1424]" pageId="2" pageNumber="148" refString="Linnaeus, C. (1758) Systema Naturae, Ed. X. (Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.). Impensis Direct. Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="90FB9AE4CE04FFAEC0DCEA3FFAFCFA04" authority="Kaup, 1856" authorityName="Kaup" authorityYear="1856" box="[992,1384,1399,1424]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="658F3D75CE04FFAEC0DCEA3FFB56FA04" box="[992,1218,1400,1424]" italics="true" pageId="2" pageNumber="148">Apterichtus gracilis</emphasis>
(
<bibRefCitation id="336A9C96CE04FFAEC7EAEA3FFACBFA04" author="Kaup" box="[1238,1375,1399,1424]" pageId="2" pageNumber="148" refString="Kaup, J. J. (1856) Catalogue of apodal fish in the collection of the British Museum. Published by order of the Trustees, London, 160 pp." type="book" year="1856">Kaup, 1856</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="90FB9AE4CE04FFAEC3ABEADBFD71FA21" authority="McCosker &amp; Hibino, 2015" authorityName="McCosker &amp; Hibino" authorityYear="2015" box="[151,741,1436,1461]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="658F3D75CE04FFAEC3ABEADBFE3FFA20" box="[151,427,1436,1460]" italics="true" pageId="2" pageNumber="148">Apterichtus jeffwilliamsi</emphasis>
<bibRefCitation id="336A9C96CE04FFAEC288EADBFD71FA21" author="McCosker" box="[436,741,1436,1461]" pageId="2" pageNumber="148" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">
McCosker &amp; Hibino,
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</bibRefCitation>
</taxonomicName>
in those counts, the new species differs from
<taxonomicName id="90FB9AE4CE04FFAEC7C8EADBFADEFA20" box="[1268,1354,1436,1460]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="ansp">
<emphasis id="658F3D75CE04FFAEC7C8EADBFADEFA20" box="[1268,1354,1436,1460]" italics="true" pageId="2" pageNumber="148">A. ansp</emphasis>
</taxonomicName>
and
<taxonomicName id="90FB9AE4CE04FFAEC6B8EADBFF79FA4C" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="658F3D75CE04FFAEC6B8EADBFF79FA4C" italics="true" pageId="2" pageNumber="148">A. gracilis</emphasis>
</taxonomicName>
in the presence of spots on the head (vs. absent) and the number of vomerine teeth (1 vs.
<quantity id="90034C82CE04FFAEC7F1EA87FA87FA4C" box="[1229,1299,1471,1496]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="2" pageNumber="148" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">34 in</quantity>
<taxonomicName id="90FB9AE4CE04FFAEC626EA87FAF9FA4C" box="[1306,1389,1472,1496]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="ansp">
<emphasis id="658F3D75CE04FFAEC626EA87FAF9FA4C" box="[1306,1389,1472,1496]" italics="true" pageId="2" pageNumber="148">A. ansp</emphasis>
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and
<quantity id="90034C82CE04FFAEC3ABEAA3FF4AFA69" box="[151,222,1508,1533]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="2" pageNumber="148" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
<taxonomicName id="90FB9AE4CE04FFAEC3D8EAA3FECDFA68" box="[228,345,1508,1532]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="658F3D75CE04FFAEC3D8EAA3FECDFA68" box="[228,345,1508,1532]" italics="true" pageId="2" pageNumber="148">A. gracilis</emphasis>
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). It differs from
<taxonomicName id="90FB9AE4CE04FFAEC129EAA3FD16FA68" box="[533,642,1508,1532]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="caucus">
<emphasis id="658F3D75CE04FFAEC129EAA3FD16FA68" box="[533,642,1508,1532]" italics="true" pageId="2" pageNumber="148">A. caucus</emphasis>
</taxonomicName>
in the size of spots on the head (large spots equal to or longer than eye vs. smaller than eye), the number of vomerine teeth (1 vs. 814), and the location of lower-jaw tip (between verticals through anterior margin of eye and mid-eye vs. anterior to anteriormost margin of eye).
<taxonomicName id="90FB9AE4CE04FFAEC627E96BFF63F9F3" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="succinus">
<emphasis id="658F3D75CE04FFAEC627E96BFF63F9F3" italics="true" pageId="2" pageNumber="148">Apterichtus succinus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="90FB9AE4CE04FFAEC2F9E917FDE5F9FC" box="[453,625,1616,1640]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="658F3D75CE04FFAEC2F9E917FDE5F9FC" box="[453,625,1616,1640]" italics="true" pageId="2" pageNumber="148">A. jeffwilliamsi</emphasis>
</taxonomicName>
in its snout length and profile (snout length 11.9% HL vs. 13.417.8% HL; profile 40° vs. 30°), size of spots (generally equal or larger than eye vs. smaller than eye diameter), and the shape of the anterior nostril (short tube vs. simple hole without tube). The present new species closely resembles
<taxonomicName id="90FB9AE4CE04FFAEC3ABE9FBFD89F940" authority="Weber, 1913" authorityName="Weber" authorityYear="1913" box="[151,541,1724,1749]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE04FFAEC3ABE9FBFE13F940" box="[151,391,1724,1748]" italics="true" pageId="2" pageNumber="148">Apterichtus klazingai</emphasis>
<bibRefCitation id="336A9C96CE04FFAEC2B2E9FBFD89F940" author="Weber" box="[398,541,1724,1749]" pageId="2" pageNumber="148" refString="Weber, M. (1913) Die Fische der Siboga-Expedition. E. J. Brill, Leiden, 710 pp." type="book" year="1913">
Weber,
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</bibRefCitation>
</taxonomicName>
in the location of its lower-jaw tip, the presence of head spots, snout profile of 40°, and in total vertebral count. The former species differs from the latter in its count of preopercular pores (4 vs. 3).
<taxonomicName id="90FB9AE4CE04FFAEC380E843FE38F888" box="[188,428,1796,1820]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE04FFAEC380E843FE38F888" box="[188,428,1796,1820]" italics="true" pageId="2" pageNumber="148">Apterichtus klazingai</emphasis>
</taxonomicName>
is the most widespread species, known from
<collectingCountry id="2FECA1F7CE04FFAEC097E843FBAFF889" box="[939,1083,1796,1821]" name="South Africa" pageId="2" pageNumber="148">South Africa</collectingCountry>
to
<collectingCountry id="2FECA1F7CE04FFAEC75CE843FAB1F889" box="[1120,1317,1796,1821]" name="French Polynesia" pageId="2" pageNumber="148">French Polynesia</collectingCountry>
, and it has high variation in its coloration or vertebral counts. Nevertheless, there is no variation in its counts of preopercular and supratemporal pores as far as our materials of
<specimenCount id="41FD2AEECE04FFAEC1F1E80BFCF2F8F1" box="[717,870,1868,1893]" pageId="2" pageNumber="148" type="generic">66 specimens</specimenCount>
collected from various localities. As stated above, these counts are very important to identify species of
<taxonomicName id="90FB9AE4CE04FFAEC1D0E837FCF9F81C" box="[748,877,1904,1928]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="genus">
<emphasis id="658F3D75CE04FFAEC1D0E837FCF9F81C" box="[748,877,1904,1928]" italics="true" pageId="2" pageNumber="148">Apterichtus</emphasis>
</taxonomicName>
because all congeners have a unique combination of counts with minor exceptions (
<bibRefCitation id="336A9C96CE04FFAEC117E8D3FC87F838" author="Hibino" box="[555,787,1940,1965]" pageId="2" pageNumber="148" refString="Hibino, Y., Shibata, J. &amp; Kimura, S. (2014) Description of a new snake eel, Apterichtus hatookai sp. nov. (Anguilliformes: Ophichthidae), from the Pacific coast of Japan. Ichthyological Research, 61, 317 - 321. http: // dx. doi. org / 10.1007 s / 10228 - 014 - 0398 - 7" type="journal article" year="2014">
Hibino
<emphasis id="658F3D75CE04FFAEC1BAE8D2FD50F838" box="[646,708,1940,1964]" italics="true" pageId="2" pageNumber="148">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation id="336A9C96CE04FFAEC018E8D3FBF2F839" author="McCosker" box="[804,1126,1940,1965]" pageId="2" pageNumber="148" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino, 2015</bibRefCitation>
). Consequently, the above difference is regarded as an interspecific difference. In addition, the new species can be distinguished from
<taxonomicName id="90FB9AE4CE04FFAEC6B8E8FFFF6AF860" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="148" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE04FFAEC6B8E8FFFF6AF860" italics="true" pageId="2" pageNumber="148">A. klazingai</emphasis>
</taxonomicName>
by having fewer preanal vertebrae (54 vs. 5663, mean 58) (
<tableCitation id="1A79D4DCCE04FFAEC0D8E89BFBA9F860" box="[996,1085,2012,2037]" captionStart="TABLE 1" captionStartId="3.[151,239,296,319]" captionText="TABLE 1. Frequency distribution of vertebral counts in Apterichtus succinus sp. nov. and A. klazingai." pageId="2" pageNumber="148">Table 1</tableCitation>
), a single tooth on prevomer (generally absent); shorter head (
<quantity id="90034C82CE05FFAFC137EFDFFDCFFF24" box="[523,603,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.7592" pageId="3" pageNumber="149" unit="in" value="14.8">14.8 in</quantity>
TL vs.
<quantity id="90034C82CE05FFAFC189EFDFFC8DFF24" box="[693,793,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="3" pageNumber="149" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">1214 in</quantity>
TL), and larger spots on head and nape (mostly equal or larger than eye with a few spots smaller than a half of eye, vs. various but generally smaller than eye, exceptionally with larger spots with numerous smaller spots smaller than a half of eye) (
<figureCitation id="CFC0FDE2CE05FFAFC0E9EF98FB8CFF6C" box="[981,1048,223,248]" captionStart="FIGURE 3" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[168,1436,781,1906]" captionTargetId="figure@3.[151,1436,772,1915]" captionTargetPageId="3" captionText="FIGURE 3. Marking patterns of head and nape, traced from fresh color photographs: (a) Apterichtus succinus sp. nov., holotype, USNM 409323; (b) A. klazingai, USNM 375002, 247 mm TL, Ile Uvea, Wallis Island; (c) A. klazingai, BPBM 36764, 298 mm TL, Maumere Bay, Flores, Indonesia." httpUri="https://zenodo.org/record/255514/files/figure.png" pageId="3" pageNumber="149">Fig. 3</figureCitation>
).
</paragraph>
<caption id="0384B1EFCE05FFAFC3ABEE6FFB72FEAB" box="[151,1254,295,319]" pageId="3" pageNumber="149">
<paragraph id="5744E167CE05FFAFC3ABEE6FFB72FEAB" blockId="3.[151,1254,295,319]" box="[151,1254,295,319]" pageId="3" pageNumber="149">
<emphasis id="658F3D75CE05FFAFC3ABEE6FFE9EFEAB" bold="true" box="[151,266,296,319]" pageId="3" pageNumber="149">TABLE 1.</emphasis>
Frequency distribution of vertebral counts in
<taxonomicName id="90FB9AE4CE05FFAFC1CFEE60FC45FEAB" box="[755,977,295,319]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE05FFAFC1CFEE60FC45FEAB" box="[755,977,295,319]" italics="true" pageId="3" pageNumber="149">Apterichtus succinus</emphasis>
</taxonomicName>
<emphasis id="658F3D75CE05FFAFC0EBEE6FFBB9FEAB" bold="true" box="[983,1069,296,319]" pageId="3" pageNumber="149">
<taxonomicNameLabel id="7EBC800ECE05FFAFC0EBEE6FFBB9FEAB" box="[983,1069,296,319]" pageId="3" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="90FB9AE4CE05FFAFC75CEE60FB4BFEAB" box="[1120,1247,295,319]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE05FFAFC75CEE60FB4BFEAB" box="[1120,1247,295,319]" italics="true" pageId="3" pageNumber="149">A. klazingai</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="5744E167CE05FFAFC17FEE1DFD4EFEE4" blockId="3.[159,1399,346,666]" box="[579,730,346,368]" pageId="3" pageNumber="149">Total vertebrae</paragraph>
<paragraph id="5744E167CE05FFAFC17FEEC3FAE3FE0E" blockId="3.[159,1399,346,666]" box="[579,1399,388,410]" pageId="3" pageNumber="149">133 134 135 136 137 138 139 140 141 142</paragraph>
<paragraph id="5744E167CE05FFAFC3A3EEEBFDC4FE57" blockId="3.[159,1399,346,666]" box="[159,592,428,454]" pageId="3" pageNumber="149">
<taxonomicName id="90FB9AE4CE05FFAFC3A3EEEBFE06FE52" box="[159,402,428,454]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE05FFAFC3A3EEEBFE06FE52" box="[159,402,428,454]" italics="true" pageId="3" pageNumber="149">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE05FFAFC2A5EEF7FE7EFE51" box="[409,490,432,453]" pageId="3" pageNumber="149" rank="species">
<emphasis id="658F3D75CE05FFAFC2A5EEF7FE7EFE51" bold="true" box="[409,490,432,453]" pageId="3" pageNumber="149">
sp.
<date id="2345C7A7CE05FFAFC282EEF7FE7EFE51" box="[446,490,432,453]" pageId="3" pageNumber="149">nov.</date>
</emphasis>
</taxonomicNameLabel>
1
</paragraph>
<paragraph id="5744E167CE05FFAFC3A3EE9DFD67FD8E" blockId="3.[159,1399,346,666]" pageId="3" pageNumber="149">
<taxonomicName id="90FB9AE4CE05FFAFC3A3EE9DFEECFE64" box="[159,376,474,496]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="klazingai">Apterichtus klazingai</taxonomicName>
3 2 2 5 9 6 4* 2 1? Preanal vertebrae
</paragraph>
<paragraph id="5744E167CE05FFAFC3A3ED6AFDC4FDF8" blockId="3.[159,1399,346,666]" pageId="3" pageNumber="149">
? 5 4 5 5 5 6 5 7 5 8 5 9 6 0 6 1 6 2 6 3
<taxonomicName id="90FB9AE4CE05FFAFC3A3ED11FE06FDFB" box="[159,402,598,624]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE05FFAFC3A3ED11FE06FDFB" box="[159,402,598,624]" italics="true" pageId="3" pageNumber="149">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE05FFAFC2A5ED1EFE7EFDFA" box="[409,490,601,622]" pageId="3" pageNumber="149" rank="species">
<emphasis id="658F3D75CE05FFAFC2A5ED1EFE7EFDFA" bold="true" box="[409,490,601,622]" pageId="3" pageNumber="149">
sp.
<date id="2345C7A7CE05FFAFC282ED1EFE7EFDFA" box="[446,490,601,622]" pageId="3" pageNumber="149">nov.</date>
</emphasis>
</taxonomicNameLabel>
1
</paragraph>
<paragraph id="5744E167CE05FFAFC3A3EDC3FAFEFD0E" blockId="3.[159,1399,346,666]" box="[159,1386,644,666]" pageId="3" pageNumber="149">
<taxonomicName id="90FB9AE4CE05FFAFC3A3EDC3FEECFD0E" box="[159,376,644,666]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="klazingai">Apterichtus klazingai</taxonomicName>
<date id="2345C7A7CE05FFAFC1CCEDC3FC23FD0E" box="[752,951,644,666]" pageId="3" pageNumber="149" value="2012-05-05">5 5 12</date>
8 2 1 1*
</paragraph>
<paragraph id="5744E167CE05FFAFC3ABEDFDFEE1FD45" blockId="3.[151,373,698,721]" box="[151,373,698,721]" pageId="3" pageNumber="149">
*including
<typeStatus id="88405FC5CE05FFAFC233EDFDFEE4FD45" box="[271,368,698,721]" pageId="3" pageNumber="149" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="0384B1EFCE05FFAFC3ABE8D7FD2DF877" httpUri="https://zenodo.org/record/255514/files/figure.png" pageId="3" pageNumber="149" targetBox="[168,1436,781,1906]" targetPageId="3">
<paragraph id="5744E167CE05FFAFC3ABE8D7FD2DF877" blockId="3.[151,1435,1935,2019]" pageId="3" pageNumber="149">
<emphasis id="658F3D75CE05FFAFC3ABE8D7FE83F831" bold="true" box="[151,279,1936,1958]" pageId="3" pageNumber="149">FIGURE 3.</emphasis>
Marking patterns of head and nape, traced from fresh color photographs: (
<emphasis id="658F3D75CE05FFAFC703E8C8FBC1F832" bold="true" box="[1087,1109,1935,1958]" pageId="3" pageNumber="149">a)</emphasis>
<taxonomicName id="90FB9AE4CE05FFAFC763E8D7FAA1F832" box="[1119,1333,1936,1958]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="succinus" status="sp. nov.">
<emphasis id="658F3D75CE05FFAFC763E8D7FAA1F832" box="[1119,1333,1936,1958]" italics="true" pageId="3" pageNumber="149">Apterichtus succinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EBC800ECE05FFAFC67CE8D7FA01F831" box="[1344,1429,1936,1957]" pageId="3" pageNumber="149" rank="species">
<emphasis id="658F3D75CE05FFAFC67CE8D7FA01F831" bold="true" box="[1344,1429,1936,1957]" pageId="3" pageNumber="149">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, USNM 409323; (
<emphasis id="658F3D75CE05FFAFC28BE8E9FE5AF851" bold="true" box="[439,462,1966,1989]" pageId="3" pageNumber="149">b)</emphasis>
<taxonomicName id="90FB9AE4CE05FFAFC2EBE8F7FDC0F851" box="[471,596,1967,1989]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE05FFAFC2EBE8F7FDC0F851" box="[471,596,1967,1989]" italics="true" pageId="3" pageNumber="149">A. klazingai</emphasis>
</taxonomicName>
, USNM 375002, 247 mm TL, Ile Uvea, Wallis Island; (
<emphasis id="658F3D75CE05FFAFC790E8E9FB54F851" bold="true" box="[1196,1216,1966,1989]" pageId="3" pageNumber="149">c)</emphasis>
<taxonomicName id="90FB9AE4CE05FFAFC7F5E8F7FAD2F851" box="[1225,1350,1967,1989]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="149" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE05FFAFC7F5E8F7FAD2F851" box="[1225,1350,1967,1989]" italics="true" pageId="3" pageNumber="149">A. klazingai</emphasis>
</taxonomicName>
, BPBM 36764, 298 mm TL, Maumere Bay, Flores, Indonesia.
</paragraph>
</caption>
<paragraph id="5744E167CE02FFA8C3FBEFDFFD5EFE61" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="150">
In the Marquesas Islands, numerous endemic fishes were found including
<taxonomicName id="90FB9AE4CE02FFA8C705EFDFFEB8FF41" authority="McCosker &amp; Hibino, 2015" authorityName="McCosker &amp; Hibino" authorityYear="2015" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="mysi">
<emphasis id="658F3D75CE02FFA8C705EFDFFB6DFF3B" box="[1081,1273,152,176]" italics="true" pageId="4" pageNumber="150">Apterichtus mysi</emphasis>
<bibRefCitation id="336A9C96CE02FFA8C639EFD0FEB8FF41" author="McCosker" pageId="4" pageNumber="150" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino, 2015</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="336A9C96CE02FFA8C207EFFBFDE4FF41" author="McCosker" box="[315,624,188,213]" pageId="4" pageNumber="150" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino, 2015</bibRefCitation>
;
<bibRefCitation id="336A9C96CE02FFA8C147EFFBFC26FF41" author="Delrieu-Trottin" box="[635,946,188,213]" pageId="4" pageNumber="150" refString="Delrieu-Trottin, E., Williams, J. T., Bacchet, P., Kulbicki, M., Mourier, J., Galzin, R., Lison de Loma, T., Mou-Tham, G., Siu, G. &amp; Planes, S. (2015) Shore fishes of the Marquesas Islands, an updated checklist with new records and new percentage of endemic species. Check List, 11, 1 - 13. http: // dx. doi. org / 10.15560 / 11.5.1758" type="journal article" year="2015">
Delrieu-Trottin
<emphasis id="658F3D75CE02FFA8C011EFFAFCF2FF40" box="[813,870,188,212]" italics="true" pageId="4" pageNumber="150">et al.</emphasis>
, 2015
</bibRefCitation>
). The new species is also possibly endemic in that area. Although
<taxonomicName id="90FB9AE4CE02FFA8C2A2EFA7FCCBFF6C" authority="Snyder, 1904" authorityName="Snyder" authorityYear="1904" box="[414,863,223,248]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="flavicaudus">
<emphasis id="658F3D75CE02FFA8C2A2EFA7FD3FFF6C" box="[414,683,224,248]" italics="true" pageId="4" pageNumber="150">Apterichtus flavicaudus</emphasis>
(
<bibRefCitation id="336A9C96CE02FFA8C181EF98FCC3FF6C" author="Snyder" box="[701,855,223,248]" pageId="4" pageNumber="150" refString="Snyder, J. O. (1904) A catalogue of the shore fishes collected by the steamer &quot; Albatross &quot; about the Hawaiian Islands in 1902. Bulletin of the United States Fish Commission, 22, 513 - 538." type="journal article" year="1904">Snyder, 1904</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="90FB9AE4CE02FFA8C0A0EFA7FBE7FF63" box="[924,1139,224,248]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="moseri">
<emphasis id="658F3D75CE02FFA8C0A0EFA7FBE7FF63" box="[924,1139,224,248]" italics="true" pageId="4" pageNumber="150">Apterichtus moseri</emphasis>
</taxonomicName>
(
<bibRefCitation id="336A9C96CE02FFA8C7B8EF98FA07FF6C" author="Jordan" box="[1156,1427,223,248]" pageId="4" pageNumber="150" refString="Jordan, D. S. &amp; Snyder, J. O. (1901) A review of the apodal fishes or eels of Japan, with descriptions of nineteen new species. Proceedings of the United States National Museum, 23, 837 - 890. http: // dx. doi. org / 10.5479 / si. 00963801.23 - 1239.837" type="journal article" year="1901">
<collectingCountry id="2FECA1F7CE02FFA8C7B8EF98FB45FF6C" box="[1156,1233,223,248]" name="Jordan" pageId="4" pageNumber="150">Jordan</collectingCountry>
&amp; Snyder, 1901
</bibRefCitation>
) were recorded from these islands by
<bibRefCitation id="336A9C96CE02FFA8C144EE43FC31FE88" author="Randall" box="[632,933,260,285]" pageId="4" pageNumber="150" refString="Randall, J. E. &amp; Earle, J. L. (2000) Annotated checklist of the shore fishes of the Marquesas Islands. Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History, 66, 1 - 39." type="journal article" year="2000">Randall &amp; Earle (2000)</bibRefCitation>
, those records were probably based on misidentification of
<taxonomicName id="90FB9AE4CE02FFA8C247EE6FFE59FEAB" box="[379,461,296,319]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="mysi">
<emphasis id="658F3D75CE02FFA8C247EE6FFE59FEAB" box="[379,461,296,319]" italics="true" pageId="4" pageNumber="150">A. mysi</emphasis>
</taxonomicName>
and
<taxonomicName id="90FB9AE4CE02FFA8C138EE6FFD1EFED4" box="[516,650,296,320]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE02FFA8C138EE6FFD1EFED4" box="[516,650,296,320]" italics="true" pageId="4" pageNumber="150">A. klazingai</emphasis>
</taxonomicName>
, respectively, and
<bibRefCitation id="336A9C96CE02FFA8C060EE60FB01FED4" author="McCosker" box="[860,1173,295,320]" pageId="4" pageNumber="150" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino (2015)</bibRefCitation>
excluded these records from each distribution without explanations. All specimens recorded in
<bibRefCitation id="336A9C96CE02FFA8C08BEE0BFB29FEF0" author="Randall" box="[951,1213,332,357]" pageId="4" pageNumber="150" refString="Randall, J. E. &amp; Earle, J. L. (2000) Annotated checklist of the shore fishes of the Marquesas Islands. Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History, 66, 1 - 39." type="journal article" year="2000">Randall &amp; Earle (2000)</bibRefCitation>
are deposited in the Bishop Museum, but there were no specimens of those species from Marquesas during YH and JMcs revisionary work. Most of these formerly identified as
<taxonomicName id="90FB9AE4CE02FFA8C1B5EED3FCBFFE38" box="[649,811,404,428]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="flavicaudus">
<emphasis id="658F3D75CE02FFA8C1B5EED3FCBFFE38" box="[649,811,404,428]" italics="true" pageId="4" pageNumber="150">A. flavicaudus</emphasis>
</taxonomicName>
were designated as
<typeStatus id="88405FC5CE02FFA8C72BEED2FB16FE39" box="[1047,1154,405,429]" pageId="4" pageNumber="150" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="90FB9AE4CE02FFA8C790EED3FA95FE38" box="[1196,1281,404,428]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="mysi">
<emphasis id="658F3D75CE02FFA8C790EED3FA95FE38" box="[1196,1281,404,428]" italics="true" pageId="4" pageNumber="150">A. mysi</emphasis>
</taxonomicName>
.
<taxonomicName id="90FB9AE4CE02FFA8C62EEED3FA14FE38" box="[1298,1408,404,428]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="moseri">
<emphasis id="658F3D75CE02FFA8C62EEED3FA14FE38" box="[1298,1408,404,428]" italics="true" pageId="4" pageNumber="150">A. moseri</emphasis>
</taxonomicName>
is endemic to Japanese waters (
<bibRefCitation id="336A9C96CE02FFA8C2E6EEF0FC98FE44" author="McCosker" box="[474,780,439,464]" pageId="4" pageNumber="150" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino, 2015</bibRefCitation>
). Consequently, only
<taxonomicName id="90FB9AE4CE02FFA8C0C7EEFFFBD9FE5B" box="[1019,1101,440,463]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="mysi">
<emphasis id="658F3D75CE02FFA8C0C7EEFFFBD9FE5B" box="[1019,1101,440,463]" italics="true" pageId="4" pageNumber="150">A. mysi</emphasis>
</taxonomicName>
,
<taxonomicName id="90FB9AE4CE02FFA8C767EEFFFB74FE44" box="[1115,1248,440,464]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE02FFA8C767EEFFFB74FE44" box="[1115,1248,440,464]" italics="true" pageId="4" pageNumber="150">A. klazingai</emphasis>
</taxonomicName>
, and
<taxonomicName id="90FB9AE4CE02FFA8C621EEFFFA08FE5B" box="[1309,1436,440,463]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="succinus">
<emphasis id="658F3D75CE02FFA8C621EEFFFA08FE5B" box="[1309,1436,440,463]" italics="true" pageId="4" pageNumber="150">A. succinus</emphasis>
</taxonomicName>
are now known to occur in the Marquesas Islands.
</paragraph>
</subSubSection>
<subSubSection id="1FE1B2ECCE02FFA8C3FBEEB8FDBFFDA9" pageId="4" pageNumber="150" type="materials_examined">
<paragraph id="5744E167CE02FFA8C3FBEEB8FDBFFDA9" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="150">
<emphasis id="658F3D75CE02FFA8C3FBEEB8FE49FD8C" bold="true" box="[199,477,511,536]" pageId="4" pageNumber="150">Comparative material.</emphasis>
Collection and registration data for
<taxonomicName id="90FB9AE4CE02FFA8C0A8ED47FB1CFD8C" box="[916,1160,512,536]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="150" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="658F3D75CE02FFA8C0A8ED47FB1CFD8C" box="[916,1160,512,536]" italics="true" pageId="4" pageNumber="150">Apterichtus klazingai</emphasis>
</taxonomicName>
(
<emphasis id="658F3D75CE02FFA8C7A2ED46FB38FD8C" box="[1182,1196,513,536]" italics="true" pageId="4" pageNumber="150">n</emphasis>
= 66) are given by
<bibRefCitation id="336A9C96CE02FFA8C3ABED63FDB3FDA9" author="McCosker" box="[151,551,548,573]" pageId="4" pageNumber="150" refString="McCosker, J. E. &amp; Hibino, Y. (2015) A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species. Zootaxa, 3941 (1), 49 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3941.1.2" type="journal article" year="2015">McCosker &amp; Hibino (2015: 6667)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>