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<document id="84B48895EF2EA8F42ECEC996BF1DC681" ID-DOI="10.5281/zenodo.5170505" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170505" ID-ZooBank="B39DE284-15C2-4ADE-B1B9-9146D3DFE764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628285093964" checkinUser="felipe" docAuthor="Ferro, Michael L." docDate="2016" docId="CB3843409163FFB16CA2BA852F245563" docLanguage="en" docName="InsectaMundi.0472.1-57.pdf" docOrigin="Insecta Mundi 2016 (472)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Sonoma cobra Ferro 2016, new species" docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="37013B38916EFFBF6C62B9072D725479" masterDocTitle="Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America" masterLastPageNumber="57" masterPageNumber="1" pageNumber="12" updateTime="1699040133542" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="14AE556742C40488D63A5DCAA1C66251">Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America</mods:title>
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<mods:namePart id="C8BF53F0FC2A32D565A184C11A8005B8">Ferro, Michael L.</mods:namePart>
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<mods:title id="BC26FD728E6C7989A5DE77B3763D5A55">Insecta Mundi</mods:title>
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<mods:date id="AA2933E187D6AC8D5D8299C557D3D2AC">2016</mods:date>
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<mods:number id="46155CAC4ACB5E299C19ADD203AB1221">2016-03-11</mods:number>
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<mods:number id="05E817805949FE7D877DFBE7585CAEAC">472</mods:number>
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<mods:identifier id="F30CD5287017C5BC87D2286691BAF179" type="ISSN">1942-1354</mods:identifier>
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<treatment id="CB3843409163FFB16CA2BA852F245563" ID-DOI="http://doi.org/10.5281/zenodo.5185119" ID-Zenodo-Dep="5185119" LSID="urn:lsid:plazi:treatment:CB3843409163FFB16CA2BA852F245563" httpUri="http://treatment.plazi.org/id/CB3843409163FFB16CA2BA852F245563" lastPageId="14" lastPageNumber="13" pageId="13" pageNumber="12">
<subSubSection id="0B8BA1DD9163FFB26CA2BA852F3C57E0" box="[192,590,897,922]" pageId="13" pageNumber="12" type="nomenclature">
<paragraph id="432EF2569163FFB26CA2BA852F3C57E0" blockId="13.[192,590,897,954]" box="[192,590,897,922]" pageId="13" pageNumber="12">
<heading id="1866453A9163FFB26CA2BA852F3C57E0" bold="true" box="[192,590,897,922]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
<emphasis id="71E52E449163FFB26CA2BA852F3C57E0" bold="true" box="[192,590,897,922]" pageId="13" pageNumber="12">
7.
<taxonomicName id="849189D59163FFB26CBCBA862CD057E0" ID-CoL="6Z6Q4" authority="Ferro, 2016" authorityName="Ferro" authorityYear="2016" box="[222,418,897,921]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cobra" status="sp. nov.">
<emphasis id="71E52E449163FFB26CBCBA862CD057E0" bold="true" box="[222,418,897,921]" italics="true" pageId="13" pageNumber="12">Sonoma cobra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AD6933F9163FFB26DCBBA852F3C57E0" box="[425,590,897,922]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9163FFB26CA2BAA528255103" pageId="13" pageNumber="12" type="description">
<paragraph id="432EF2569163FFB26CA2BAA52CD757C3" blockId="13.[192,590,897,954]" box="[192,421,929,954]" pageId="13" pageNumber="12">
<figureCitation id="DBAAEED39163FFB26CA2BAA52C7A57C0" box="[192,264,930,954]" captionStart="Figures 7–11" captionStartId="23.[192,288,1808,1830]" captionTargetBox="[240,1378,224,1735]" captionTargetId="figure-13@23.[240,1378,192,1841]" captionTargetPageId="23" captionText="Figures 7–11. Sonoma spp. holotype habiti. 7) S. cobra. 8) S. colberti. 9) S. maryae. 10) S. quellazaire. 11) S. rossellinae." figureDoi="http://doi.org/10.5281/zenodo.5170509" httpUri="https://zenodo.org/record/5170509/files/figure.png" pageId="13" pageNumber="12">Fig. 7</figureCitation>
,
<figureCitation id="DBAAEED39163FFB26D77BAA62C4757C0" box="[277,309,929,953]" captionStart="Figures 50–52" captionStartId="34.[192,288,1783,1805]" captionTargetBox="[240,1326,286,1701]" captionTargetId="figure-17@34.[192,1442,192,1836]" captionTargetPageId="34" captionText="Figures 50–52. Aedeagi of Sonoma spp. 50) S. repanda Marsh and Schuster. 51) S. quercicola Chandler. Modified from Chandler (1986). 52) S. cobra new species. Right side of figures is anatomical left." figureDoi="http://doi.org/10.5281/zenodo.5170535" httpUri="https://zenodo.org/record/5170535/files/figure.png" pageId="13" pageNumber="12">52</figureCitation>
; Map 10.
</paragraph>
<paragraph id="432EF2569163FFB26CA2BAE628D25043" blockId="13.[192,1440,993,1402]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26CA2BAE62C185780" bold="true" box="[192,362,993,1017]" pageId="13" pageNumber="12">Description.</emphasis>
<typeStatus id="9C2A4CF49163FFB26D0DBAE52C925783" box="[367,480,994,1018]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
, male. Measurements: head 0.27 long, 0.36 wide; pronotum 0.41 long, 0.43 wide; elytra 0.40 long, 0.28 wide; antennomeres 1–11 total 0.67; total length 1.80. Body brown, distal portion of antennae, maxillary palpi, and legs paler. Body covered in short setae (less than 1/2 length of the eye).
</paragraph>
<paragraph id="432EF2569163FFB26C92BD4529995003" blockId="13.[192,1440,993,1402]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26C92BD452C405023" box="[240,306,1090,1114]" italics="true" pageId="13" pageNumber="12">Head</emphasis>
. Eyes reduced, maximum length in dorsal view 0.75x length of first antennal segment, with approximately 9–10 facets. Antennomere 2 approximately 0.78x width of 1; 3 smallest.
</paragraph>
<paragraph id="432EF2569163FFB26C92BD85291550C3" blockId="13.[192,1440,993,1402]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26C92BD852C3A50E3" box="[240,328,1154,1178]" italics="true" pageId="13" pageNumber="12">Thorax</emphasis>
. Elytra with indistinct sutural foveae; 2–3 foveae lateral to sutural fovea; single elongate central fovea in basal 1/3. Apparently brachypterous. Metatibia unmodified.
</paragraph>
<paragraph id="432EF2569163FFB26C92BDC5283150A0" blockId="13.[192,1440,993,1402]" box="[240,1347,1218,1242]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26C92BDC52C1150A3" box="[240,355,1218,1242]" italics="true" pageId="13" pageNumber="12">Abdomen</emphasis>
. Tergite IV without microtrichia. Basal lateral foveae obscured, weak if present.
</paragraph>
<paragraph id="432EF2569163FFB26CA2BDE52E6F5083" blockId="13.[192,1440,993,1402]" box="[192,797,1250,1274]" pageId="13" pageNumber="12">Basal pubescence present on all visible ventrites.</paragraph>
<paragraph id="432EF2569163FFB26C92BC0528255103" blockId="13.[192,1440,993,1402]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26C92BC052C105163" box="[240,354,1282,1306]" italics="true" pageId="13" pageNumber="12">Aedeagus</emphasis>
. Left paramere: longer than endophallus; blade-like apical projection with 4 setae arranged palmately directed posteriorly to mesad. Endophallus: wide as base, narrowing to apex; s-shaped; apical 1/5 perpendicular to aedeagus; subapical wart with two stout setae. Right paramere: elongate apical hook-like process approximately 2/3 length of paramere, directed mesad with 4–5 setae at base.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9163FFB26CA2BCA52C365243" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="432EF2569163FFB26CA2BCA52E6D51A3" blockId="13.[192,1440,1441,1498]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26CA2BCA52CFE51C0" bold="true" box="[192,396,1441,1466]" pageId="13" pageNumber="12">Type Material.</emphasis>
<materialsCitation id="F3F9F80B9163FFB26DF6BCA52E6D51A3" collectionCode="CALIF, T, R, CSAC" collectorName="Shasta Co. Buckhorn Summit IV" county="Shasta Co." location="Shasta Co." pageId="13" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9C2A4CF49163FFB26DF6BCA52F7451C3" box="[404,518,1442,1466]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
, male: *
<collectionCode id="25806A939163FFB26E0BBCA52FB651C3" box="[617,708,1442,1466]" pageId="13" pageNumber="12">CALIF</collectionCode>
:
<collectorName id="EE6497809163FFB26EB2BCA5291851C0" box="[720,1130,1441,1466]" pageId="13" pageNumber="12">
<collectingCounty id="AA4F8ADA9163FFB26EB2BCA52E2451C3" box="[720,854,1442,1466]" pageId="13" pageNumber="12">Shasta Co.</collectingCounty>
Buckhorn Summit IV
</collectorName>
.15.1981
<collectionCode id="25806A939163FFB268BABCA5299E51C3" box="[1240,1260,1442,1466]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="12">T</collectionCode>
.
<collectionCode id="25806A939163FFB26892BCA5287451C3" box="[1264,1286,1442,1466]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="12">R</collectionCode>
.Haig, Coll. / Berlesed from Oak duff (
<specimenCount id="559739DF9163FFB26D91BCC52F6151A3" box="[499,531,1473,1498]" pageId="13" pageNumber="12" type="male">1♂</specimenCount>
). Deposited in
<collectionCode id="25806A939163FFB26EAFBCC52E6E51A3" box="[717,796,1474,1498]" pageId="13" pageNumber="12">CSAC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="432EF2569163FFB26CA2BF062C365243" blockId="13.[192,1440,1537,1594]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26CA2BF062F475260" bold="true" box="[192,565,1537,1561]" pageId="13" pageNumber="12">Geographical Distribution.</emphasis>
<taxonomicName id="849189D59163FFB26E5EBF052F9B5263" authorityName="Ferro" authorityYear="2016" box="[572,745,1538,1562]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cobra">
<emphasis id="71E52E449163FFB26E5EBF052F9B5263" box="[572,745,1538,1562]" italics="true" pageId="13" pageNumber="12">Sonoma cobra</emphasis>
</taxonomicName>
is known from a single specimen collected in Shasta Co.,
<collectingRegion id="81553CB49163FFB26CA2BF252C325243" box="[192,320,1570,1594]" country="United States of America" name="California" pageId="13" pageNumber="12">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9163FFB26CA2BF6628745363" pageId="13" pageNumber="12" type="discussion">
<paragraph id="432EF2569163FFB26CA2BF662F0552C3" blockId="13.[192,1440,1633,1818]" pageId="13" pageNumber="12">
<emphasis id="71E52E449163FFB26CA2BF662C295200" bold="true" box="[192,347,1633,1657]" pageId="13" pageNumber="12">Comments.</emphasis>
The specimen was collected during April from Berlesed oak duff. The reduced eyes, apparent brachyptery, and collection method indicate that the species may be associated with more subterranean habitats than its congeners.
</paragraph>
<paragraph id="432EF2569163FFB26C92BFC528745363" blockId="13.[192,1440,1633,1818]" pageId="13" pageNumber="12">
Aedeagal characters of
<taxonomicName id="849189D59163FFB26E6FBFC52FC952A3" authorityName="Ferro" authorityYear="2016" box="[525,699,1730,1754]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cobra">
<emphasis id="71E52E449163FFB26E6FBFC52FC952A3" box="[525,699,1730,1754]" italics="true" pageId="13" pageNumber="12">Sonoma cobra</emphasis>
</taxonomicName>
are unique. Lack of elongate latero-apical setae on the parameres separate it from
<taxonomicName id="849189D59163FFB26DBBBFE52F255283" box="[473,599,1762,1786]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="spadica">
<emphasis id="71E52E449163FFB26DBBBFE52F255283" box="[473,599,1762,1786]" italics="true" pageId="13" pageNumber="12">S. spadica</emphasis>
</taxonomicName>
,
<taxonomicName id="849189D59163FFB26E06BFE52F9B5283" box="[612,745,1762,1786]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="repanda">
<emphasis id="71E52E449163FFB26E06BFE52F9B5283" box="[612,745,1762,1786]" italics="true" pageId="13" pageNumber="12">S. repanda</emphasis>
</taxonomicName>
, and
<taxonomicName id="849189D59163FFB26F48BFE52EDB5283" authorityName="Marsh and Schuster. Right" authorityYear="1962" box="[810,937,1762,1786]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="dolabra">
<emphasis id="71E52E449163FFB26F48BFE52EDB5283" box="[810,937,1762,1786]" italics="true" pageId="13" pageNumber="12">S. dolabra</emphasis>
</taxonomicName>
. The s-shaped endophallus and hook-like apical process on the right paramere will separate
<taxonomicName id="849189D59163FFB26F53BE052EE75363" authorityName="Ferro" authorityYear="2016" box="[817,917,1794,1818]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cobra">
<emphasis id="71E52E449163FFB26F53BE052EE75363" box="[817,917,1794,1818]" italics="true" pageId="13" pageNumber="12">S. cobra</emphasis>
</taxonomicName>
from all other known species.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9160FFB16CA2B9C52F245563" pageId="14" pageNumber="13" type="etymology">
<paragraph id="432EF2569160FFB16CA2B9C52F245563" blockId="14.[192,1440,194,282]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2B9C52C2E54A3" bold="true" box="[192,348,194,218]" pageId="14" pageNumber="13">Etymology.</emphasis>
<taxonomicName id="849189D59160FFB16D01B9C52F6054A3" authorityName="Ferro" authorityYear="2016" box="[355,530,194,218]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="cobra">
<emphasis id="71E52E449160FFB16D01B9C52F6054A3" box="[355,530,194,218]" italics="true" pageId="14" pageNumber="13">Sonoma cobra</emphasis>
</taxonomicName>
is named for its unique serpentine endophallus complete with “fang-like” setae. “Cobra” is derived from
<emphasis id="71E52E449160FFB16E53B9E52F985483" box="[561,746,226,250]" italics="true" pageId="14" pageNumber="13">
<taxonomicName id="849189D59160FFB16E53B9E52F065483" authorityName="Ferro" authorityYear="2016" box="[561,628,226,250]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="cobra">cobra</taxonomicName>
de capelo
</emphasis>
, a Portuguese term, and today is commonly used to refer to snakes in the family
<taxonomicName id="849189D59160FFB16D83B8052F205563" box="[481,594,258,282]" class="Reptilia" family="Elapidae" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="13" phylum="Chordata" rank="family">Elapidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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