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<document id="EC142CAE235100E6E8906E081A4E777A" ID-DOI="10.11646/zootaxa.4811.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3944031" ID-ZooBank="EB3E2796-D738-405B-AAB7-4D8209214114" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594719006450" checkinUser="plazi" docAuthor="Weglarz, Kathryn M. &amp; Bartlett, Charles R" docDate="2020" docId="9C5C87C4AB41FFE0FF2EF99AFDFEFA9F" docLanguage="en" docName="zootaxa.4811.1.1.pdf" docOrigin="Zootaxa 4811 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chionomus bellicosus Weglarz &amp; Bartlett 2020, new combination" docType="treatment" docVersion="3" lastPageNumber="18" masterDocId="6065FFBCAB4EFFF1FFB9FFA3FF90FF90" masterDocTitle="A revision of the planthopper genus Chionomus Fennah (Hemiptera: Fulgoroidea: Delphacidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="16" updateTime="1698781408566" updateUser="plazi">
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<mods:title id="2A620F8E8E7F4AB75F9E905CD57C3A4D">A revision of the planthopper genus Chionomus Fennah (Hemiptera: Fulgoroidea: Delphacidae)</mods:title>
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<mods:name id="901C8966DE83AEC1AF2CB3C720CFA009" type="personal">
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<mods:namePart id="879411C4F8BFE350D917DAFD6940B275">Weglarz, Kathryn M.</mods:namePart>
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<mods:namePart id="B0DE39064309EC644C60E78B39236DBC">Bartlett, Charles R</mods:namePart>
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<mods:title id="3855862922284212127CD86637B10A87">Zootaxa</mods:title>
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<mods:date id="52FEDE9297D29D95A5C1C53F8464F84B">2020</mods:date>
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<mods:number id="33A886F2BA4264AA50788E45136A4B50">2020-07-14</mods:number>
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<mods:number id="6B8AD10F9D37D3008FF05AEA7EF54370">4811</mods:number>
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<treatment id="9C5C87C4AB41FFE0FF2EF99AFDFEFA9F" LSID="urn:lsid:plazi:treatment:9C5C87C4AB41FFE0FF2EF99AFDFEFA9F" httpUri="http://treatment.plazi.org/id/9C5C87C4AB41FFE0FF2EF99AFDFEFA9F" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="5CEF6559AB41FFFEFF2EF99AFC21F9C3" box="[151,945,1593,1620]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="144A36D2AB41FFFEFF2EF99AFC21F9C3" blockId="15.[151,945,1593,1655]" box="[151,945,1593,1620]" pageId="15" pageNumber="16">
<heading id="4F0281BEAB41FFFEFF2EF99AFC21F9C3" bold="true" box="[151,945,1593,1620]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="2681EAC0AB41FFFEFF2EF99AFC21F9C3" bold="true" box="[151,945,1593,1620]" pageId="15" pageNumber="16">
<taxonomicName id="D3F54D51AB41FFFEFF2EF99AFD5DF9C3" ID-CoL="9FNJX" authority="(Muir &amp; Giffard, 1924)" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Muir &amp; Giffard" baseAuthorityYear="1924" box="[151,717,1593,1620]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bellicosus" status="comb. nov.">
<emphasis id="2681EAC0AB41FFFEFF2EF99AFE32F9C4" bold="true" box="[151,418,1593,1620]" italics="true" pageId="15" pageNumber="16">Chionomus bellicosus</emphasis>
(
<bibRefCitation id="70644B23AB41FFFEFE08F99AFD55F9C3" author="Muir, F. A. G. &amp; Giffard, W. M." box="[433,709,1593,1619]" pageId="15" pageNumber="16" pagination="1 - 53" refId="ref29217" refString="Muir, F. A. G. &amp; Giffard, W. M. (1924) Studies in North American Delphacidae. Bulletin of the Experiment Station of the Hawaiian Sugar Planters' Association, Entomological Series, 15, 1 - 53." type="journal article" year="1924">Muir &amp; Giffard, 1924</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="3DB257BBAB41FFFEFD62F99AFC21F9C3" box="[731,945,1593,1619]" pageId="15" pageNumber="16" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB41FFFEFF2EF9FEFEADF9E7" box="[151,317,1629,1655]" pageId="15" pageNumber="16" type="description">
<paragraph id="144A36D2AB41FFFEFF2EF9FEFEADF9E7" blockId="15.[151,945,1593,1655]" box="[151,317,1629,1655]" pageId="15" pageNumber="16">
(
<figureCitation id="8CCE2A57AB41FFFEFF26F9FEFE9CF9E7" box="[159,268,1629,1655]" captionStart="FIGURE 6" captionStartId="16.[151,250,1694,1719]" captionTargetBox="[152,1432,181,1668]" captionTargetId="figure@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of Chionomus bellicosus (holotype; A, C, E-I; holotype terminalia in balsam); A. dorsal view, habitus brachypterous holotype; B. dorsal view, habitus macropter; C. lateral view, habitus brachypterous holotype; D. lateral view, habitus macropter; E. labels of holotype; F. front habitus of holotype; G. aedeagus and anal tube of holotype, left lateral view; H. parameres of holotype, caudal view; I. pygofer of holotype, caudal view, J. lateral view of male terminalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3944043" httpUri="https://zenodo.org/record/3944043/files/figure.png" pageId="15" pageNumber="16">Figures 6</figureCitation>
,
<figureCitation id="8CCE2A57AB41FFFEFEA1F9FEFEA5F9E7" box="[280,309,1629,1655]" captionStart="FIGURE 22" captionStartId="40.[151,250,1104,1129]" captionTargetBox="[256,1332,181,1078]" captionTargetId="figure@40.[256,1332,181,1078]" captionTargetPageId="40" captionText="FIGURE 22. Line art of Chionomus bellicosus, terminalia, A. pygofer, caudal view, B. anal tube, left lateral view, C. aedeagus, left lateral view; D. left paramere, widest view." figureDoi="http://doi.org/10.5281/zenodo.3944076" httpUri="https://zenodo.org/record/3944076/files/figure.png" pageId="15" pageNumber="16">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB41FFFEFF2EF902FD1EF929" box="[151,654,1696,1721]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="144A36D2AB41FFFEFF2EF902FD1EF929" blockId="15.[151,654,1696,1721]" box="[151,654,1696,1721]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="34E511FCAB41FFFEFF2EF902FD1EF929" box="[151,654,1696,1721]" pageId="15" pageNumber="16">
<treatmentCitation id="955410C3AB41FFFEFF2EF902FD19F929" author="Muir, F. A. G. &amp; Giffard, W. M." box="[151,649,1696,1721]" page="34" pageId="15" pageNumber="16" year="1924">
<taxonomicName id="D3F54D51AB41FFFEFF2EF902FD19F929" ID-CoL="34HH3" authority="Muir &amp; Giffard, 1924: 34" authorityName="Muir &amp; Giffard" authorityPageNumber="34" authorityYear="1924" box="[151,649,1696,1721]" class="Insecta" family="Delphacidae" genus="Delphacodes" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bellicosa">
<emphasis id="2681EAC0AB41FFFEFF2EF902FEEDF929" box="[151,381,1697,1721]" italics="true" pageId="15" pageNumber="16">Delphacodes bellicosa</emphasis>
<bibRefCitation id="70644B23AB41FFFEFE3DF903FD19F929" author="Muir, F. A. G. &amp; Giffard, W. M." box="[388,649,1696,1721]" pageId="15" pageNumber="16" pagination="1 - 53" refId="ref29217" refString="Muir, F. A. G. &amp; Giffard, W. M. (1924) Studies in North American Delphacidae. Bulletin of the Experiment Station of the Hawaiian Sugar Planters' Association, Entomological Series, 15, 1 - 53." type="journal article" year="1924">Muir &amp; Giffard, 1924: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB41FFFEFF2EF94BFCC2F891" box="[151,850,1767,1794]" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="144A36D2AB41FFFEFF2EF94BFCC2F891" blockId="15.[151,1437,1767,2010]" box="[151,850,1767,1794]" pageId="15" pageNumber="16">
<materialsCitation id="A49D3C8FAB41FFFEFF2EF94BFCDFF891" box="[151,847,1767,1794]" country="United States of America" county="Tulare County" location="Three Rivers" pageId="15" pageNumber="16" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="2681EAC0AB41FFFEFF2EF94BFED0F892" bold="true" box="[151,320,1767,1794]" pageId="15" pageNumber="16">
<typeStatus id="CB4E8870AB41FFFEFF2EF94BFF42F892" box="[151,210,1768,1794]" pageId="15" pageNumber="16">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="6CE27642AB41FFFEFEFEF94BFE12F892" box="[327,386,1768,1794]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
,
<collectingRegion id="D631F830AB41FFFEFE34F944FD91F891" box="[397,513,1767,1793]" country="United States of America" name="California" pageId="15" pageNumber="16">California</collectingRegion>
,
<collectingCounty id="FD2B4E5EAB41FFFEFDB5F944FD3FF892" box="[524,687,1767,1794]" pageId="15" pageNumber="16">Tulare County</collectingCounty>
,
<location id="112A6009AB41FFFEFD00F944FCDFF891" LSID="urn:lsid:plazi:treatment:9C5C87C4AB41FFE0FF2EF99AFDFEFA9F:112A6009AB41FFFEFD00F944FCDFF891" box="[697,847,1767,1793]" country="United States of America" county="Tulare County" name="Three Rivers" pageId="15" pageNumber="16" stateProvince="California">Three Rivers</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB41FFFEFF7EF8A8FC5AF849" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="144A36D2AB41FFFEFF7EF8A8FC5AF849" blockId="15.[151,1437,1767,2010]" pageId="15" pageNumber="16">
<emphasis id="2681EAC0AB41FFFEFF7EF8A8FED2F8B6" bold="true" box="[199,322,1803,1830]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Body brown to light brown, with ivory to cream markings. Vertex quadrate; foveae of frons and vertex dark, posterior compartments of vertex light with fuscous markings. Carinae distinct, off-white in color; antennae light brown. Pronotum light, white to cream in color, infuscate directly behind eyes, paranota white. Mesonotum brown, lateral and median carinae cream; wing with faint fuscate mark at apex of clavus. Armature of diaphragm bifurcate, hooked at apex; parameres sinuate, outer angles enlarged. Aedeagus tubular, directed dorsoposteriorly, large flange with teeth on apical ½ of left side, 2 subapical teeth on right.
</paragraph>
</subSubSection>
<caption id="408A665AAB5EFFE1FF2EF93DFA0BF8B3" ID-DOI="http://doi.org/10.5281/zenodo.3944043" ID-Zenodo-Dep="3944043" httpUri="https://zenodo.org/record/3944043/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1694,1719]" targetBox="[152,1432,181,1668]" targetPageId="16">
<paragraph id="144A36D2AB5EFFE1FF2EF93DFA0BF8B3" blockId="16.[151,1436,1694,1827]" pageId="16" pageNumber="17">
<emphasis id="2681EAC0AB5EFFE1FF2EF93DFE85F927" bold="true" box="[151,277,1694,1719]" pageId="16" pageNumber="17">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="D3F54D51AB5EFFE1FE2DF93CFDE5F927" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Muir &amp; Giffard" baseAuthorityYear="1924" box="[404,629,1695,1719]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="2681EAC0AB5EFFE1FE2DF93CFDE5F927" box="[404,629,1695,1719]" italics="true" pageId="16" pageNumber="17">Chionomus bellicosus</emphasis>
</taxonomicName>
(holotype; A, C, E-I; holotype terminalia in balsam); A. dorsal view, habitus brachypterous holotype; B. dorsal view, habitus macropter; C. lateral view, habitus brachypterous holotype; D. lateral view, habitus macropter; E. labels of holotype; F. front habitus of holotype; G. aedeagus and anal tube of holotype, left lateral view; H. parameres of holotype, caudal view; I. pygofer of holotype, caudal view, J. lateral view of male terminalia, left lateral view.
</paragraph>
</caption>
<subSubSection id="5CEF6559AB5EFFE0FF7EF8D7FBDBFC37" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="144A36D2AB5EFFE0FF7EF8D7FCA2FE67" blockId="16.[151,1436,1908,2044]" lastBlockId="17.[151,1437,153,1295]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="2681EAC0AB5EFFE1FF7EF8D7FEC9F81F" bold="true" box="[199,345,1908,1935]" pageId="16" pageNumber="17">Description.</emphasis>
<emphasis id="2681EAC0AB5EFFE1FED8F8D6FE32F81F" box="[353,418,1909,1935]" italics="true" pageId="16" pageNumber="17">Color</emphasis>
.
<emphasis id="2681EAC0AB5EFFE1FE08F8D5FDBCF81F" box="[433,556,1910,1935]" italics="true" pageId="16" pageNumber="17">Macropter.</emphasis>
Body brown to light brown, matte, with cream or ivory markings. Carinae of head (including genal carinae) distinct, ochraceous to off-white in color; median carinae of vertex evident. Foveae of frons and vertex brown, posterior compartments of vertex light brown.Antennae cream to stramineous. Pronotum white to cream, fuscate markings directly posterior to eyes. Mesonotum dark brown to brown, median and lateral carinae light brown to cream. Legs stramineous to brown, with fuscous markings on anterior face of femora; apex of tarsi brown. Forewings hyaline, faint fuscous spot just before apex of clavus. Abdomen brown.
<emphasis id="2681EAC0AB5FFFE0FB07FF1EFAACFF47" box="[1214,1340,189,215]" italics="true" pageId="17" pageNumber="18">Brachypter</emphasis>
. Similar to above, tegmina infuscate, white stripe along apex, darkened spot near apex of clavus.
<emphasis id="2681EAC0AB5FFFE0FBD5FF42FB43FF6B" box="[1132,1235,225,251]" italics="true" pageId="17" pageNumber="18">Structure</emphasis>
.
<emphasis id="2681EAC0AB5FFFE0FB58FF42FA8BFF6B" box="[1249,1307,225,251]" italics="true" pageId="17" pageNumber="18">Body</emphasis>
. Length (in mm) [All specimens previously dissected]; width
<specimenCount id="02F3FD5BAB5FFFE0FD68FEA6FD76FE8F" box="[721,742,261,287]" pageId="17" pageNumber="18" type="male"></specimenCount>
macropter 0.82±0.09 (n=3);
<specimenCount id="02F3FD5BAB5FFFE0FB8BFEA6FBD7FE8F" box="[1074,1095,261,287]" pageId="17" pageNumber="18" type="male"></specimenCount>
brachypter 0.75 (n=1).
<emphasis id="2681EAC0AB5FFFE0FAE0FEA6FA0DFE8F" box="[1369,1437,261,287]" italics="true" pageId="17" pageNumber="18">Head.</emphasis>
Vertex length 0.19±0.02 (n=4); vertex width 0.19±0.01 (n=4); frons length 0.47±0.02 (n=4); frons width 0.26±0.02 (n=4).
<emphasis id="2681EAC0AB5FFFE0FF5BFEEDFEC9FEF7" box="[226,345,334,359]" italics="true" pageId="17" pageNumber="18">Macropter</emphasis>
. Carinae of vertex distinct, stem of Y-shaped carina faint but evident. In lateral view (
<figureCitation id="8CCE2A57AB5FFFE0FABFFEEEFA1EFEF8" box="[1286,1422,333,360]" captionStart="FIGURE 6" captionStartId="16.[151,250,1694,1719]" captionTargetBox="[152,1432,181,1668]" captionTargetId="figure@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of Chionomus bellicosus (holotype; A, C, E-I; holotype terminalia in balsam); A. dorsal view, habitus brachypterous holotype; B. dorsal view, habitus macropter; C. lateral view, habitus brachypterous holotype; D. lateral view, habitus macropter; E. labels of holotype; F. front habitus of holotype; G. aedeagus and anal tube of holotype, left lateral view; H. parameres of holotype, caudal view; I. pygofer of holotype, caudal view, J. lateral view of male terminalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3944043" httpUri="https://zenodo.org/record/3944043/files/figure.png" pageId="17" pageNumber="18">Figure 6c, d</figureCitation>
), fastigium rounded; projecting in front of the eye about 1/6th eye length. Carinae of frons and clypeus distinct (
<figureCitation id="8CCE2A57AB5FFFE0FAD5FED2FF4DFE20" captionStart="FIGURE 6" captionStartId="16.[151,250,1694,1719]" captionTargetBox="[152,1432,181,1668]" captionTargetId="figure@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of Chionomus bellicosus (holotype; A, C, E-I; holotype terminalia in balsam); A. dorsal view, habitus brachypterous holotype; B. dorsal view, habitus macropter; C. lateral view, habitus brachypterous holotype; D. lateral view, habitus macropter; E. labels of holotype; F. front habitus of holotype; G. aedeagus and anal tube of holotype, left lateral view; H. parameres of holotype, caudal view; I. pygofer of holotype, caudal view, J. lateral view of male terminalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3944043" httpUri="https://zenodo.org/record/3944043/files/figure.png" pageId="17" pageNumber="18">Figure 6f</figureCitation>
), frons bowed, widest just below compound eyes. Antennal segments subequal in length, segment II wider; 2nd segment sparsely setaceous, bearing sensory pits arranged 3, 4, 4, 21, evenly spaced around segment. Sensory pits surrounded by small setae.
<emphasis id="2681EAC0AB5FFFE0FE41FE7EFDE6FE67" box="[504,630,477,503]" italics="true" pageId="17" pageNumber="18">Brachypter</emphasis>
. Same as above.
</paragraph>
<paragraph id="144A36D2AB5FFFE0FF7EFDA2FACBFDF3" blockId="17.[151,1437,153,1295]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFDA2FE0CFD8B" box="[199,412,513,539]" italics="true" pageId="17" pageNumber="18">Thorax. Macropter</emphasis>
. Carinae of mesonotum evident, median carina ending at scutellum, lateral carinae faint, diverging posteriorly to reach hind margin. Legs with 2 rows of small setae on ventral margin of femora, mostly bare. Calcar bearing 17 23 small teeth (
<typeStatus id="CB4E8870AB5FFFE0FDA6FDEAFD13FDF3" box="[543,643,585,611]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
with 17).
<emphasis id="2681EAC0AB5FFFE0FD4FFDEAFCE4FDF3" box="[758,884,585,611]" italics="true" pageId="17" pageNumber="18">Brachypter</emphasis>
. Same as above; tegmina apically rounded.
</paragraph>
<paragraph id="144A36D2AB5FFFE0FF7EFDCEFC54FD17" blockId="17.[151,1437,153,1295]" box="[199,964,621,647]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFDCEFE2DFD17" box="[199,445,621,647]" italics="true" pageId="17" pageNumber="18">Abdomen. Brachypter</emphasis>
. Compressed dorsoventrally, tapering caudad.
</paragraph>
<paragraph id="144A36D2AB5FFFE0FF7EFD32FB70FC13" blockId="17.[151,1437,153,1295]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFD32FEA1FD3B" box="[199,305,657,683]" italics="true" pageId="17" pageNumber="18">Genitalia</emphasis>
. Pygofer in caudal view (
<figureCitation id="8CCE2A57AB5FFFE0FDF0FD32FD2CFD3B" box="[585,700,657,683]" captionStart="FIGURE 6" captionStartId="16.[151,250,1694,1719]" captionTargetBox="[152,1432,181,1668]" captionTargetId="figure@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of Chionomus bellicosus (holotype; A, C, E-I; holotype terminalia in balsam); A. dorsal view, habitus brachypterous holotype; B. dorsal view, habitus macropter; C. lateral view, habitus brachypterous holotype; D. lateral view, habitus macropter; E. labels of holotype; F. front habitus of holotype; G. aedeagus and anal tube of holotype, left lateral view; H. parameres of holotype, caudal view; I. pygofer of holotype, caudal view, J. lateral view of male terminalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3944043" httpUri="https://zenodo.org/record/3944043/files/figure.png" pageId="17" pageNumber="18">Figures 6i</figureCitation>
,
<figureCitation id="8CCE2A57AB5FFFE0FD7EFD32FD63FD3B" box="[711,755,657,683]" captionStart="FIGURE 22" captionStartId="40.[151,250,1104,1129]" captionTargetBox="[256,1332,181,1078]" captionTargetId="figure@40.[256,1332,181,1078]" captionTargetPageId="40" captionText="FIGURE 22. Line art of Chionomus bellicosus, terminalia, A. pygofer, caudal view, B. anal tube, left lateral view, C. aedeagus, left lateral view; D. left paramere, widest view." figureDoi="http://doi.org/10.5281/zenodo.3944076" httpUri="https://zenodo.org/record/3944076/files/figure.png" pageId="17" pageNumber="18">22a</figureCitation>
), about as tall as wide, rounded; margins rounded. Opening to inner chamber triangular, pointed ventrad. Armature of diaphragm bifurcate, hooked apically, distinctly projecting caudad. Parameres, widest in basal half, basal angle strong, projecting, quadrate; dorsolaterally diverging to quadrate apices, lateral margins concave, inner angles weak, pointed; outer angles produced to rounded apices. Aedeagus circular in cross-section, slightly curved ventrad, directed dorsoposteriorly, widest near base, slightly tapering for most of length. Aedeagus bearing toothed flange on left distal half, to subapical teeth on right. Segment X quadrate; processes distinct, projected posteriorly. Segment XI produced, about 2/3rds length of segment X.
</paragraph>
<paragraph id="144A36D2AB5FFFE0FF7EFC2EFBDBFC37" blockId="17.[151,1437,153,1295]" box="[199,1099,908,935]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFC2EFE33FC37" bold="true" box="[199,419,908,935]" pageId="17" pageNumber="18">Plant associations.</emphasis>
<taxonomicName id="D3F54D51AB5FFFE0FE13FC2EFD1EFC37" box="[426,654,909,935]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="distichum">
<emphasis id="2681EAC0AB5FFFE0FE13FC2EFD1EFC37" box="[426,654,909,935]" italics="true" pageId="17" pageNumber="18">Paspalum distichum</emphasis>
</taxonomicName>
L. (knotgrass,
<taxonomicName id="D3F54D51AB5FFFE0FC8FFC2EFC04FC37" authorityName=", Muir" authorityYear="1926" box="[822,916,909,935]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) (
<bibRefCitation id="70644B23AB5FFFE0FC13FC2EFBAEFC37" author="Wilson, S. W." box="[938,1086,908,935]" pageId="17" pageNumber="18" pagination="72 - 78" refId="ref30386" refString="Wilson, S. W. (1985) Descriptions of the immature stages of Delphacodes bellicosa (Homoptera: Fulgoroidea: Delphacidae). Pan-Pacific Entomologist, 61, 72 - 78." type="journal article" year="1985">Wilson 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB5FFFE0FF7EFC13FABDFC7F" pageId="17" pageNumber="18" type="distribution">
<paragraph id="144A36D2AB5FFFE0FF7EFC13FE4DFC5B" blockId="17.[151,1437,153,1295]" box="[199,477,944,971]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFC13FEF1FC5B" bold="true" box="[199,353,944,971]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="6CE27642AB5FFFE0FEDEFC12FE33FC5B" box="[359,419,945,971]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
: CA.
</paragraph>
<paragraph id="144A36D2AB5FFFE0FF7EFC77FABDFC7F" blockId="17.[151,1437,153,1295]" box="[199,1325,980,1007]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFC77FD89FC7F" bold="true" box="[199,537,980,1007]" pageId="17" pageNumber="18">GenBank accession number.</emphasis>
Material for molecular work was unavailable at the time of this study.
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB5FFFE0FF7EFC5AFB59FBEF" pageId="17" pageNumber="18" type="discussion">
<paragraph id="144A36D2AB5FFFE0FF7EFC5AFB59FBEF" blockId="17.[151,1437,153,1295]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFC5AFEABFB83" bold="true" box="[199,315,1017,1043]" pageId="17" pageNumber="18">Remarks.</emphasis>
This species is uncommon in collections. It is closely allied with
<taxonomicName id="D3F54D51AB5FFFE0FB86FC5AFB58FB83" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Crawford" baseAuthorityYear="1914" box="[1087,1224,1017,1043]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="2681EAC0AB5FFFE0FB86FC5AFB58FB83" box="[1087,1224,1017,1043]" italics="true" pageId="17" pageNumber="18">C. pacificus</emphasis>
</taxonomicName>
but can be distinguished by the shape of the aedeagus and the hook of the bifurcate armature of the diaphragm being only apical. This bifurcate armature is also mirrored in
<taxonomicName id="D3F54D51AB5FFFE0FDFBFBE2FD23FBCB" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Van Duzee" baseAuthorityYear="1897" box="[578,691,1089,1115]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="puellus">
<emphasis id="2681EAC0AB5FFFE0FDFBFBE2FD23FBCB" box="[578,691,1089,1115]" italics="true" pageId="17" pageNumber="18">C. puellus</emphasis>
</taxonomicName>
but the processes are
<taxonomicName id="D3F54D51AB5FFFE0FC12FBE2FC5FFBCB" box="[939,975,1089,1115]" class="Not" higherTaxonomySource="CoL" kingdom="Viruses" pageId="17" pageNumber="56" rank="class">not</taxonomicName>
hooked; additionally, the ventral edge of the parameres is distinctly S-shaped in
<taxonomicName id="D3F54D51AB5FFFE0FDF7FBC6FD2FFBEF" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Van Duzee" baseAuthorityYear="1897" box="[590,703,1125,1151]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="puellus">
<emphasis id="2681EAC0AB5FFFE0FDF7FBC6FD2FFBEF" box="[590,703,1125,1151]" italics="true" pageId="17" pageNumber="18">C. puellus</emphasis>
</taxonomicName>
while they are nearly straight in
<emphasis id="2681EAC0AB5FFFE0FB88FBC6FB59FBEF" box="[1073,1225,1125,1151]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="D3F54D51AB5FFFE0FB88FBC6FB54FBEF" authorityName="Weglarz &amp; Bartlett" authorityYear="2020" baseAuthorityName="Muir &amp; Giffard" baseAuthorityYear="1924" box="[1073,1220,1125,1151]" class="Insecta" family="Delphacidae" genus="Chionomus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellicosus">C. bellicosus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="5CEF6559AB5FFFE0FF7EFB2AFDFEFA9F" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="144A36D2AB5FFFE0FF7EFB2AFDFEFA9F" blockId="17.[151,1437,153,1295]" pageId="17" pageNumber="18">
<emphasis id="2681EAC0AB5FFFE0FF7EFB2AFE7DFB33" bold="true" box="[199,493,1160,1187]" pageId="17" pageNumber="18">Type material examined.</emphasis>
<materialsCitation id="A49D3C8FAB5FFFE0FE4AFB2AFDFEFA9F" collectionCode="BPBM" collectorName="Brachypterous" pageId="17" pageNumber="18" specimenCount="3" specimenCount-male="3" typeStatus="holotype">
<typeStatus id="CB4E8870AB5FFFE0FE4AFB2AFDCBFB33" box="[499,603,1161,1187]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
[
<collectionCode id="72E4AE17AB5FFFE0FDD1FB2BFD2FFB33" box="[616,703,1160,1187]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="17" pageNumber="18" type="Museum">BPBM</collectionCode>
]:
<collectorName id="B9005304AB5FFFE0FD69FB2AFCE3FB33" box="[720,883,1161,1187]" pageId="17" pageNumber="18">Brachypterous</collectorName>
<specimenCount id="02F3FD5BAB5FFFE0FCC0FB2AFC00FB33" box="[889,912,1161,1187]" pageId="17" pageNumber="18" type="male"></specimenCount>
, “
<taxonomicName id="D3F54D51AB5FFFE0FC1CFB2AFBA9FB33" authorityName="Fieber" authorityYear="1866" box="[933,1081,1161,1187]" class="Insecta" family="Delphacidae" genus="Delphacodes" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Delphacodes</taxonomicName>
/ bellicosa /
<specimenCount id="02F3FD5BAB5FFFE0FB7DFB2AFB49FB33" box="[1220,1241,1161,1187]" pageId="17" pageNumber="18" type="male"></specimenCount>
M.G. [Handwrit- ten, white card with hole punch containing genitalia embedded in balsam] // Three Rivers / Cal. Clortson //
<typeStatus id="CB4E8870AB5FFFE0FA8DFB0EFA0CFB57" box="[1332,1436,1197,1223]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
[Red paper, vertically oriented, affixed to following label] //
<taxonomicName id="D3F54D51AB5FFFE0FC83FB72FC5BFB7B" authorityName="Fieber" authorityYear="1866" box="[826,971,1233,1259]" class="Insecta" family="Delphacidae" genus="Delphacodes" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Delphacodes</taxonomicName>
/ bellicosa / M.G. /
<specimenCount id="02F3FD5BAB5FFFE0FB1EFB72FB2CFB7B" box="[1191,1212,1233,1259]" pageId="17" pageNumber="18" type="male"></specimenCount>
[Handwritten, label with black border] // 1074 [Handwritten]”
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>