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<mods:title id="39FEB4E98D82807BDFBF842EF6F40346">New taxonomic twists in shore flies as revealed in discovery of Neozealides, a new genus from New Zealand (Diptera: Ephydridae)</mods:title>
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<mods:namePart id="D3CC829EB35261BBE3BEAA586CA75F61">Mathis, Wayne N.</mods:namePart>
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<paragraph id="E37936B6FFB1FFEE63AD7205FDB9F828" blockId="1.[151,536,1917,1944]" box="[151,536,1917,1944]" pageId="1" pageNumber="488">
<heading id="B83181DAFFB1FFEE63AD7205FDB9F828" bold="true" box="[151,536,1917,1944]" fontSize="11" level="1" pageId="1" pageNumber="488" reason="1">
<emphasis id="D1B2EAA4FFB1FFEE63AD7205FDB9F828" bold="true" box="[151,536,1917,1944]" pageId="1" pageNumber="488">
Tribe
<taxonomicName id="24C64D35FFB1FFEE63D87205FE67F827" ID-CoL="87CTY" authority="Phillips" authorityName="Phillips" box="[226,454,1917,1943]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="488" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini Phillips</taxonomicName>
<emphasis id="D1B2EAA4FFB1FFEE62F77205FDB9F828" bold="true" box="[461,536,1917,1944]" italics="true" pageId="1" pageNumber="488">et alia</emphasis>
</emphasis>
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</paragraph>
<paragraph id="E37936B6FFB1FFEE63AD72B8FDCAF848" blockId="1.[151,1436,1984,2040]" pageId="1" pageNumber="488">
Hydrinini
<bibRefCitation id="87574B47FFB1FFEE623B72B9FE62F869" author="Cresson, E. T. Jr." box="[257,451,1984,2009]" pageId="1" pageNumber="488" pagination="159 - 180" refId="ref8208" refString="Cresson, E. T. Jr. (1944) Synopses of North American Ephydridae (Diptera). Parts IA and II. Transactions of the American Entomological Society, 70, 159 - 180." type="journal article" year="1944">Cresson 1944: 175</bibRefCitation>
.
<typeStatus id="3C7D8814FFB1FFEE62F672BAFE5FF86A" box="[460,510,1986,2010]" pageId="1" pageNumber="488">Type</typeStatus>
genus:
<taxonomicName id="24C64D35FFB1FFEE617472B9FD03F869" authorityName="Rafinesque" authorityYear="1815" box="[590,674,1985,2009]" class="Insecta" family="Ephydridae" genus="Hydrina" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB1FFEE617472B9FD03F869" box="[590,674,1985,2009]" italics="true" pageId="1" pageNumber="488">Hydrina</emphasis>
</taxonomicName>
of authors, not Robineau-Desvoidy 1830 (=
<taxonomicName id="24C64D35FFB1FFEE675772B9FA30F869" authority="Stenhammar 1844" authorityName="Stenhammar" authorityYear="1844" box="[1133,1425,1984,2009]" class="Insecta" family="Ephydridae" genus="Philygria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB1FFEE675772B9FB6CF869" box="[1133,1229,1985,2009]" italics="true" pageId="1" pageNumber="488">Philygria</emphasis>
Stenhammar 1844
</taxonomicName>
), unavailable, based on a junior homonym.
</paragraph>
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<subSubSection id="ABDC653DFFB2FFED63AD75E0FC5EFEFA" pageId="2" pageNumber="489" type="reference_group">
<paragraph id="E37936B6FFB2FFED63AD75E0FE47FF5E" blockId="2.[151,1437,152,330]" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED63AD75E0FEF0FF01" ID-CoL="87CTY" authority="Phillips" authorityName="Phillips" box="[151,337,152,177]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini Phillips</taxonomicName>
<emphasis id="D1B2EAA4FFB2FFED626375E1FE3BFF01" box="[345,410,153,177]" italics="true" pageId="2" pageNumber="489">et alia</emphasis>
in
<bibRefCitation id="87574B47FFB2FFED628775E1FD21FF01" author="Cresson, E. T. Jr." box="[445,640,152,177]" pageId="2" pageNumber="489" pagination="225 - 260" refId="ref8246" refString="Cresson, E. T. Jr. (1949) A systematic annotated arrangement of the genera and species of the North American Ephydridae (Diptera). IV. The subfamily Napaeinae. Transactions of the American Entomological Society, 74, 225 - 260." type="journal article" year="1949">Cresson 1949: 251</bibRefCitation>
.
<typeStatus id="3C7D8814FFB2FFED61B175E1FD1CFF01" box="[651,701,153,177]" pageId="2" pageNumber="489">Type</typeStatus>
genus:
<taxonomicName id="24C64D35FFB2FFED603475E1FBF6FF01" ID-CoL="636KN" authority="Haliday in Curtis 1837" authorityName="Haliday in Curtis" authorityYear="1837" box="[782,1111,152,177]" class="Insecta" family="Ephydridae" genus="Hyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED603475E1FCC4FF01" box="[782,869,153,177]" italics="true" pageId="2" pageNumber="489">Hyadina</emphasis>
Haliday in Curtis 1837
</taxonomicName>
.—
<bibRefCitation id="87574B47FFB2FFED674F75E0FE81FF7F" author="Mathis, W. N. &amp; Zatwarnicki, T." pageId="2" pageNumber="489" refId="ref8830" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A World Catalog of the Shore Flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, 423 pp." type="book" year="1995">Mathis &amp; Zatwarnicki 1995: 200214</bibRefCitation>
[world catalog].—
<bibRefCitation id="87574B47FFB2FFED62DA75CFFCE5FF7F" author="Hollmann-Schirrmacher, V." box="[480,836,183,208]" pageId="2" pageNumber="489" pagination="1 - 144" refId="ref8550" refString="Hollmann-Schirrmacher, V. (1998) Phylogeny of the subfamily Ilytheinae (Diptera, Ephydridae) with species reference to the genus Philygria. Studia dipterologica. Supplement, 5, 1 - 144." type="journal article" year="1998">Hollmann-Schirrmacher 1998: 29</bibRefCitation>
, 4556 [discussion].—
<bibRefCitation id="87574B47FFB2FFED671775CFFF4CFF5E" author="Zatwarnicki, T. &amp; Ryczko, I." pageId="2" pageNumber="489" pagination="495 - 515" refId="ref9182" refString="Zatwarnicki, T. &amp; Ryczko, I. (2014) Phylogeny of Hyadinini (Diptera: Ephydridae) with an emphasis on structures of the proboscis. Annales Zoologici, 64 (3), 495 - 515. https: // doi. org / 10.3161 / 000345414 X 68482 1" type="journal article" year="2014">Zatwarnicki &amp; Ryczko 2014: 495 515</bibRefCitation>
[phylogenetic analysis].
</paragraph>
<paragraph id="E37936B6FFB2FFED63AD758CFD75FEBD" blockId="2.[151,1437,152,330]" box="[151,724,244,269]" pageId="2" pageNumber="489">
<treatmentCitationGroup id="C3D61198FFB2FFED63AD758CFD75FEBD" box="[151,724,244,269]" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED63AD758CFEA2FEBD" authorityName="Lizarralde de Grosso" authorityYear="1989" box="[151,259,244,269]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Philygriini">Philygriini</taxonomicName>
[nomen nudum].—
<treatmentCitation id="626710A7FFB2FFED62F3758CFD71FEBD" author="Wirth, W. W. &amp; Stone, A." box="[457,720,244,269]" page="469" pageId="2" pageNumber="489" year="1956">
<bibRefCitation id="87574B47FFB2FFED62F3758CFD71FEBD" author="Wirth, W. W. &amp; Stone, A." box="[457,720,244,269]" pageId="2" pageNumber="489" pagination="372 - 482" refId="ref9058" refString="Wirth, W. W. &amp; Stone, A. (1956) Chapter 14. Aquatic Diptera. In: Usinger, R. L. (Ed.), Aquatic Insects of California. 508 + x pp. University of California Press, Berkeley, pp. 372 - 482, 64 figs." type="book chapter" year="1956">Wirth &amp; Stone 1956: 469</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E37936B6FFB2FFED63AD746BFC5EFEFA" blockId="2.[151,1437,152,330]" pageId="2" pageNumber="489">
<treatmentCitationGroup id="C3D61198FFB2FFED63AD746BFC5EFEFA" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED63AD746BFDE6FE9C" authority="Lizarralde de Grosso 1989: 51" authorityName="Lizarralde de Grosso" authorityPageNumber="51" authorityYear="1989" box="[151,583,275,300]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Philygriini">
Philygriini
<treatmentCitation id="626710A7FFB2FFED6231746BFDE6FE9C" author="Lizarralde de Grosso, M. S." box="[267,583,275,300]" page="51" pageId="2" pageNumber="489" year="1989">
<bibRefCitation id="87574B47FFB2FFED6231746BFDE6FE9C" author="Lizarralde de Grosso, M. S." box="[267,583,275,300]" pageId="2" pageNumber="489" refId="ref8586" refString="Lizarralde de Grosso, M. S. (1989) Ephydridae de la Republica Argentina (Insecta-Diptera). Serie Monografica y Didactica No. 3. Facultad de Ciencias Naturales y Instituto Miguel Lillo Universidad Nacional de Tucuman, 93 pp." type="book" year="1989">Lizarralde de Grosso 1989: 51</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3C7D8814FFB2FFED6168746CFD25FE9C" box="[594,644,276,300]" pageId="2" pageNumber="489">Type</typeStatus>
genus:
<taxonomicName id="24C64D35FFB2FFED61EC746BFC56FE9C" ID-CoL="63MVM" authority="Stenhammar 1844" authorityName="Stenhammar" authorityYear="1844" box="[726,1015,275,300]" class="Insecta" family="Ephydridae" genus="Philygria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED61EC746BFC97FE9B" box="[726,822,275,299]" italics="true" pageId="2" pageNumber="489">Philygria</emphasis>
Stenhammar 1844
</taxonomicName>
.—
<bibRefCitation id="87574B47FFB2FFED672F746BFA3CFE9B" author="Mathis, W. N. &amp; Zatwarnicki, T." box="[1045,1437,275,300]" pageId="2" pageNumber="489" refId="ref8830" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A World Catalog of the Shore Flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, 423 pp." type="book" year="1995">Mathis &amp; Zatwarnicki 1995: 190200</bibRefCitation>
[world catalog].—
<treatmentCitation id="626710A7FFB2FFED62477449FD79FEFA" author="Hollmann-Schirrmacher, V." box="[381,728,305,330]" page="50" pageId="2" pageNumber="489" year="1998">
<bibRefCitation id="87574B47FFB2FFED62477449FD79FEFA" author="Hollmann-Schirrmacher, V." box="[381,728,305,330]" pageId="2" pageNumber="489" pagination="1 - 144" refId="ref8550" refString="Hollmann-Schirrmacher, V. (1998) Phylogeny of the subfamily Ilytheinae (Diptera, Ephydridae) with species reference to the genus Philygria. Studia dipterologica. Supplement, 5, 1 - 144." type="journal article" year="1998">Hollmann-Schirrmacher 1998: 50</bibRefCitation>
</treatmentCitation>
[synonymy with
<taxonomicName id="24C64D35FFB2FFED60B07449FC54FEFA" ID-CoL="87CTY" authority="Phillips" authorityName="Phillips" box="[906,1013,305,330]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
].
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</paragraph>
</subSubSection>
<subSubSection id="ABDC653DFFB2FFED63AD7400FE94FB37" pageId="2" pageNumber="489" type="materials_examined">
<paragraph id="E37936B6FFB2FFED63AD7400FB73FDF6" blockId="2.[151,1437,376,1159]" pageId="2" pageNumber="489">
<emphasis id="D1B2EAA4FFB2FFED63AD7400FEB3FE23" bold="true" box="[151,274,376,403]" pageId="2" pageNumber="489">Diagnosis.</emphasis>
The tribe
<taxonomicName id="24C64D35FFB2FFED62B67400FE5DFE22" box="[396,508,376,402]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
is similar to
<taxonomicName id="24C64D35FFB2FFED61AD7400FD57FE22" box="[663,758,376,402]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Ilytheini">Ilytheini</taxonomicName>
but is distinguished by the following combination of characters: Aristal hairs tending to be reduced (sometimes secondarily moderately well developed); acrostichal setae reduced, prescutellar pair lacking; dorsocentral setae frequently reduced in size and/or number (1+2, 0+2, or sutural+1); subepandrial plate fused with gonite dorsally, forming a rounded projection, sometimes joined medially over aedeagus to form a gonal arch (= subepandrial plate); gonites and hypandrium usually fused; gonite produced posteriorly as a long triangular to almost parallel sided projection, with apex variously modified.
</paragraph>
<paragraph id="E37936B6FFB2FFED63FD7728FD91FD66" blockId="2.[151,1437,376,1159]" pageId="2" pageNumber="489">
<emphasis id="D1B2EAA4FFB2FFED63FD7728FEEDFDDB" bold="true" box="[199,332,592,619]" pageId="2" pageNumber="489">Discussion.</emphasis>
The monophyly of the tribe
<taxonomicName id="24C64D35FFB2FFED61A07728FCABFDDA" box="[666,778,592,618]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
is established by the following synapomorphies (
<bibRefCitation id="87574B47FFB2FFED66787729FE9FFD3E" author="Zatwarnicki, T." pageId="2" pageNumber="489" pagination="65 - 119" refId="ref9117" refString="Zatwarnicki, T. (1992) A new classification of Ephydridae based on phylogenetic reconstruction (Diptera: Cyclorrhapha). Genus, 3, 65 - 119." type="journal article" year="1992">Zatwarnicki 1992: 76</bibRefCitation>
;
<bibRefCitation id="87574B47FFB2FFED6271770CFD6BFD3F" author="Hollmann-Schirrmacher, V." box="[331,714,628,655]" pageId="2" pageNumber="489" pagination="1 - 144" refId="ref8550" refString="Hollmann-Schirrmacher, V. (1998) Phylogeny of the subfamily Ilytheinae (Diptera, Ephydridae) with species reference to the genus Philygria. Studia dipterologica. Supplement, 5, 1 - 144." type="journal article" year="1998">Hollmann-Schirrmacher 1998: 50</bibRefCitation>
): (1) reduction of scutellar/prescutellar acrostichal setae; (2) fusion of gonite with hypandrium; (3) reduction of dorsocentral setae; (4) lack of long branches on the arista; and (5) ventral elongation of the epandrium.
</paragraph>
<paragraph id="E37936B6FFB2FFED63FD7798FCE9FC3A" blockId="2.[151,1437,376,1159]" pageId="2" pageNumber="489">
Only four described genera of the tribe
<taxonomicName id="24C64D35FFB2FFED61BC7798FD57FD4A" box="[646,758,736,762]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
occur in
<collectingCountry id="9BD17626FFB2FFED605B7799FC5AFD4A" box="[865,1019,736,763]" name="New Zealand" pageId="2" pageNumber="489">New Zealand</collectingCountry>
, and one,
<emphasis id="D1B2EAA4FFB2FFED67547798FB5DFD4A" box="[1134,1276,736,762]" italics="true" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED67547798FB59FD4A" authority="Mathis &amp; Zatwarnicki, 2019" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2019" box="[1134,1272,736,762]" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus" status="gen. nov.">Neozealides</taxonomicName>
,
</emphasis>
the subject of this paper, and
<taxonomicName id="24C64D35FFB2FFED6279767CFD68FCAE" authority="Tonnoir and Malloch" authorityName="Tonnoir and Malloch" box="[323,713,772,798]" class="Insecta" family="Ephydridae" genus="Parahyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED6279767CFE74FCAE" box="[323,469,772,798]" italics="true" pageId="2" pageNumber="489">Parahyadina</emphasis>
Tonnoir and Malloch
</taxonomicName>
are apparently endemic to these islands. The four genera can be identified by the key that follows.
<bibRefCitation id="87574B47FFB2FFED611A7650FCF2FCF3" author="Edmiston, J. F. &amp; Mathis, W. N." box="[544,851,808,835]" pageId="2" pageNumber="489" pagination="1 - 16" refId="ref8411" refString="Edmiston, J. F. &amp; Mathis, W. N. (2007) New Zealand species of the shore-fly genus Nostima Coquillett (Diptera: Ephydridae). Zootaxa, 1661, 1 - 16." type="journal article" year="2007">Edmiston &amp; Mathis (2007)</bibRefCitation>
revised the three
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species of
<taxonomicName id="24C64D35FFB2FFED66027651FE49FCD7" authority=", Mathis &amp; Zatwarnicki (2019)" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2019" class="Insecta" family="Ephydridae" genus="Nostima" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED66027651FA3DFCF2" box="[1336,1436,809,834]" italics="true" pageId="2" pageNumber="489">Nostima,</emphasis>
<bibRefCitation id="87574B47FFB2FFED63AD7634FE49FCD7" author="Mathis, W. N. &amp; Zatwarnicki, T." box="[151,488,844,871]" pageId="2" pageNumber="489" pagination="401 - 440" refId="ref8868" refString="Mathis, W. N. &amp; Zatwarnicki, T. (2019) Revision of the shore-fly genus Parahyadina Tonnoir and Malloch and New Zealand Hyadina Haliday (Diptera: Ephydridae). Zootaxa, 4623 (3), 401 - 440." type="journal article" year="2019">Mathis &amp; Zatwarnicki (2019)</bibRefCitation>
</taxonomicName>
revised the single species of
<taxonomicName id="24C64D35FFB2FFED60047634FC3FFCD6" authorityName="Haliday in Curtis" authorityYear="1837" box="[830,926,844,870]" class="Insecta" family="Ephydridae" genus="Hyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED60047634FC3FFCD6" box="[830,926,844,870]" italics="true" pageId="2" pageNumber="489">Hyadina</emphasis>
</taxonomicName>
and the eleven species of
<emphasis id="D1B2EAA4FFB2FFED67EB7634FACAFCD6" box="[1233,1387,844,870]" italics="true" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED67EB7634FAC7FCD6" box="[1233,1382,844,870]" class="Insecta" family="Ephydridae" genus="Parahyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">Parahyadina</taxonomicName>
,
</emphasis>
and herein, we treat the two
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species of
<emphasis id="D1B2EAA4FFB2FFED61837608FCE9FC3A" box="[697,840,880,906]" italics="true" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED61837608FCE5FC3A" authority="Mathis &amp; Zatwarnicki, 2019" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2019" box="[697,836,880,906]" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus" status="gen. nov.">Neozealides</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="E37936B6FFB2FFED63FD76ECFE94FB37" blockId="2.[151,1437,376,1159]" pageId="2" pageNumber="489">
The
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fauna of
<taxonomicName id="24C64D35FFB2FFED613976ECFD2BFC1E" authority="Mathis &amp; Zatwarnicki, 2019" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2019" box="[515,650,916,942]" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D1B2EAA4FFB2FFED613976ECFD2BFC1E" box="[515,650,916,942]" italics="true" pageId="2" pageNumber="489">Neozealides</emphasis>
</taxonomicName>
and
<taxonomicName id="24C64D35FFB2FFED61FE76ECFCF7FC1E" box="[708,854,916,942]" class="Insecta" family="Ephydridae" genus="Parahyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED61FE76ECFCF7FC1E" box="[708,854,916,942]" italics="true" pageId="2" pageNumber="489">Parahyadina</emphasis>
</taxonomicName>
are anomalous within
<taxonomicName id="24C64D35FFB2FFED676676ECFB71FC1E" box="[1116,1232,916,942]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
. For example, the two
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species of
<emphasis id="D1B2EAA4FFB2FFED62D976C0FDD0FC62" box="[483,625,952,978]" italics="true" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED62D976C0FDCCFC62" authority="Mathis &amp; Zatwarnicki, 2019" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2019" box="[483,621,952,978]" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus" status="gen. nov.">Neozealides</taxonomicName>
,
</emphasis>
<taxonomicName id="24C64D35FFB2FFED614076C1FC4CFC63" authority="(Tonnoir &amp; Malloch)" baseAuthorityName="Tonnoir &amp; Malloch" box="[634,1005,952,979]" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="irrorata">
<emphasis id="D1B2EAA4FFB2FFED614076C1FD54FC62" box="[634,757,953,978]" italics="true" pageId="2" pageNumber="489">N. irrorata</emphasis>
(Tonnoir &amp; Malloch)
</taxonomicName>
and
<taxonomicName id="24C64D35FFB2FFED671C76C1FF72FC47" authority="(Tonnoir &amp; Malloch)" baseAuthorityName="Tonnoir &amp; Malloch" class="Insecta" family="Ephydridae" genus="Neozealides" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="obscurifrons">
<emphasis id="D1B2EAA4FFB2FFED671C76C1FB74FC62" box="[1062,1237,952,978]" italics="true" pageId="2" pageNumber="489">N. obscurifrons</emphasis>
(Tonnoir &amp; Malloch)
</taxonomicName>
, are endemic to the islands of
<collectingCountry id="9BD17626FFB2FFED611F76A5FD1AFC46" box="[549,699,988,1015]" name="New Zealand" pageId="2" pageNumber="489">New Zealand</collectingCountry>
and are more closely related to the
<taxonomicName id="24C64D35FFB2FFED677376A4FB08FC46" authorityName="Haliday in Curtis" authorityYear="1837" box="[1097,1193,988,1014]" class="Insecta" family="Ephydridae" genus="Hyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED677376A4FB08FC46" box="[1097,1193,988,1014]" italics="true" pageId="2" pageNumber="489">Hyadina</emphasis>
</taxonomicName>
and
<taxonomicName id="24C64D35FFB2FFED67E576A4FAE8FC46" authorityName="Stenhammar" authorityYear="1844" box="[1247,1353,988,1014]" class="Insecta" family="Ephydridae" genus="Philygria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="D1B2EAA4FFB2FFED67E576A4FAE8FC46" box="[1247,1353,988,1014]" italics="true" pageId="2" pageNumber="489">Philygria</emphasis>
</taxonomicName>
groups of genera that
<bibRefCitation id="87574B47FFB2FFED620F7178FD25FBAB" author="Zatwarnicki, T. &amp; Ryczko, I." box="[309,644,1024,1051]" pageId="2" pageNumber="489" pagination="495 - 515" refId="ref9182" refString="Zatwarnicki, T. &amp; Ryczko, I. (2014) Phylogeny of Hyadinini (Diptera: Ephydridae) with an emphasis on structures of the proboscis. Annales Zoologici, 64 (3), 495 - 515. https: // doi. org / 10.3161 / 000345414 X 68482 1" type="journal article" year="2014">Zatwarnicki &amp; Ryczko (2014)</bibRefCitation>
recently recognized rather than to species within
<emphasis id="D1B2EAA4FFB2FFED67937178FAB0FBAA" box="[1193,1297,1024,1050]" italics="true" pageId="2" pageNumber="489">
<taxonomicName id="24C64D35FFB2FFED67937178FAADFBAA" authorityName="Haliday in Curtis" authorityYear="1837" box="[1193,1292,1024,1050]" class="Insecta" family="Ephydridae" genus="Hyadina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">Hyadina</taxonomicName>
,
</emphasis>
the genus in which they were initially described. Thus, the
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fauna of
<taxonomicName id="24C64D35FFB2FFED60A6715CFBADFB8E" box="[924,1036,1060,1086]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="tribe" tribe="Hyadinini">Hyadinini</taxonomicName>
is structurally and phylogenetically diverse, which is undoubtedly a reflection of their varied evolution as well as the historical biogeography of the included taxa.
</paragraph>
</subSubSection>
</treatment>
</document>