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<document id="E25E30F66A529FCE9F9DDA628F9EFDC6" ID-DOI="10.26107/RBZ-2019-0046" ID-ISSN="2345-7600" ID-Zenodo-Dep="5343729" ID-ZooBank="051B2EC9-50B0-421A-BD4A-9679F9DD3A9F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630197588800" checkinUser="diego" docAuthor="Samoh, Abdulloh, Satasook, Chutamas &amp; Grootaert, Patrick" docDate="2019" docId="53495D02F25BFFCDFCFAFA2A2A68F6DA" docLanguage="en" docName="RBZ-2019-0046.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Ngirhaphium meieri Samoh &amp; Satasook &amp; Grootaert 2019, new species" docType="treatment" docUuid="2374B56D-1B45-431A-953F-58F531022B29" docUuidSource="ZooBank" docVersion="4" lastPageNumber="651" masterDocId="AF70257AF25DFFC6FFCDFFEF2E7BF650" masterDocTitle="NGS-barcodes, haplotype networks combined to external morphology help to identify new species in the mangrove genus Ngirhaphium Evenhuis &amp; Grootaert, 2002 (Diptera: Dolichopodidae: Rhaphiinae) in Southeast Asia" masterLastPageNumber="659" masterPageNumber="640" pageNumber="646" updateTime="1699183134106" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6AA515CDCEB3AA0E93A8BA899B45B61B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BDD7FBE446DF61C70AC128CECFAEEE86">NGS-barcodes, haplotype networks combined to external morphology help to identify new species in the mangrove genus Ngirhaphium Evenhuis &amp; Grootaert, 2002 (Diptera: Dolichopodidae: Rhaphiinae) in Southeast Asia</mods:title>
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<mods:name id="F0DFE438E7D933FEFF58C2818FD9C51C" type="personal">
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<mods:namePart id="4BA1866844871A50DA0C39D5B72FF703">Samoh, Abdulloh</mods:namePart>
</mods:name>
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<mods:namePart id="0B2CD28989D931E789583BCE9EF16781">Satasook, Chutamas</mods:namePart>
</mods:name>
<mods:name id="BD824CF5E7CD3C7878BD0672A176EA90" type="personal">
<mods:role id="093EBDE51BA84384584176A28EE7F240">
<mods:roleTerm id="F8DAA0BACD2A861DD4D76A94E3692CD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46C36A9BF1042197BBA9AC1A7E92AEBE">Grootaert, Patrick</mods:namePart>
</mods:name>
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<mods:title id="59E552F70D2E32D830DB554A184939EC">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="0B09997EB13BBD8026EC005C97B45ED3">
<mods:date id="ED210724378CAF314AC40E0398312DC3">2019</mods:date>
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<mods:number id="638408B18842EA94217C46F129584DCC">2019-11-21</mods:number>
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<mods:number id="1CE34331160107CEC553875FF7353AEA">67</mods:number>
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<mods:classification id="7F6F1D2E6157D1302FD126E0172320DC">journal article</mods:classification>
<mods:identifier id="95A42A851220F50B8C8C8211302214D6" type="DOI">10.26107/RBZ-2019-0046</mods:identifier>
<mods:identifier id="D7926B51225D2B5B0CE020DC9B187E5B" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F0EAD805CD88FEB166CC05349D12ECC2" type="Zenodo-Dep">5343729</mods:identifier>
<mods:identifier id="11DE27D63D02C911D9353B62B7EAB64E" type="ZooBank">051B2EC9-50B0-421A-BD4A-9679F9DD3A9F</mods:identifier>
</mods:mods>
<treatment id="53495D02F25BFFCDFCFAFA2A2A68F6DA" LSID="urn:lsid:zoobank.org:act:2374B56D-1B45-431A-953F-58F531022B29" httpUri="http://treatment.plazi.org/id/53495D02F25BFFCDFCFAFA2A2A68F6DA" lastPageId="11" lastPageNumber="651" pageId="6" pageNumber="646">
<subSubSection id="93FABF9FF25BFFC0FCFAFA2A2BD8F38E" box="[823,1443,1477,1502]" pageId="6" pageNumber="646" type="nomenclature">
<paragraph id="DB5FEC14F25BFFC0FCFAFA2A2BD8F38E" blockId="6.[823,1443,1477,1534]" box="[823,1443,1477,1502]" pageId="6" pageNumber="646">
<heading id="80175B78F25BFFC0FCFAFA2A2BD8F38E" bold="true" box="[823,1443,1477,1502]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="646" reason="2">
<emphasis id="E9943006F25BFFC0FCFAFA2A2BD8F38E" bold="true" box="[823,1443,1477,1502]" pageId="6" pageNumber="646">
<taxonomicName id="1CE09797F25BFFC0FCFAFA2A2B6BF38E" ID-CoL="9HTMT" authority="Samoh &amp; Grootaert" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[823,1296,1477,1502]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="646" phylum="Arthropoda" rank="species" species="meieri" status="sp. nov.">
<emphasis id="E9943006F25BFFC0FCFAFA2A2A67F38E" bold="true" box="[823,1052,1477,1502]" italics="true" pageId="6" pageNumber="646">Ngirhaphium meieri</emphasis>
Samoh &amp; Grootaert
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF25BFFC0FAD0FA292BD8F38E" box="[1309,1443,1478,1502]" pageId="6" pageNumber="646" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF25BFFC0FBDFFA092ABDF3AE" box="[1042,1222,1510,1534]" pageId="6" pageNumber="646" type="description">
<paragraph id="DB5FEC14F25BFFC0FBDFFA092ABDF3AE" blockId="6.[823,1443,1477,1534]" box="[1042,1222,1510,1534]" pageId="6" pageNumber="646">
(
<figureCitation id="43DBF091F25BFFC0FBD7FA092AF9F3AE" box="[1050,1154,1510,1534]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="7.[836,871,982,1004]" captionStartId-1="8.[113,148,1995,2017]" captionStartId-2="9.[137,172,1968,1990]" captionTargetBox-0="[832,1478,193,968]" captionTargetBox-1="[112,1457,228,1982]" captionTargetBox-2="[142,1476,317,1952]" captionTargetId-0="figure-456@7.[819,1482,164,981]" captionTargetId-1="figure-11@8.[99,1467,209,1988]" captionTargetId-2="figure-8@9.[123,1490,297,1961]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Fig. 7. Ngirhaphium meieri, new species, female habitus. Scale = 1 mm." captionText-1="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." captionText-2="Fig. 9. Distribution map of Ngirhaphium meieri, new species and Ngirhaphium thaicum, new species." figureDoi-0="http://doi.org/10.5281/zenodo.5343745" figureDoi-1="http://doi.org/10.5281/zenodo.5343751" figureDoi-2="http://doi.org/10.5281/zenodo.5343753" httpUri-0="https://zenodo.org/record/5343745/files/figure.png" httpUri-1="https://zenodo.org/record/5343751/files/figure.png" httpUri-2="https://zenodo.org/record/5343753/files/figure.png" pageId="6" pageNumber="646">Figs. 7–9</figureCitation>
,
<figureCitation id="43DBF091F25BFFC0FB42FA092AC5F3AE" box="[1167,1214,1510,1534]" captionStart="Fig" captionStartId="10.[812,847,838,860]" captionTargetBox="[813,1451,191,825]" captionTargetId="figure-789@10.[796,1466,181,765]" captionTargetPageId="10" captionText="Fig. 10. Haplotype network of Ngirhaphium murphyi. Singapore: Pulau Ubin, Mandai Sungei Buloh; Thailand: Satun and Krabi." figureDoi="http://doi.org/10.5281/zenodo.5343757" httpUri="https://zenodo.org/record/5343757/files/figure.png" pageId="6" pageNumber="646">10D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF25BFFC0FCE0F9C92ACEF1EE" pageId="6" pageNumber="646" type="diagnosis">
<paragraph id="DB5FEC14F25BFFC0FCE0F9C92A8EF14E" blockId="6.[811,1454,1574,1822]" pageId="6" pageNumber="646">
<emphasis id="E9943006F25BFFC0FCE0F9C92DE4F06E" bold="true" box="[813,927,1574,1598]" pageId="6" pageNumber="646">Diagnosis.</emphasis>
A large species with apical segment of arista twice as long as basal aristal segment. Prongs on fore tarsomere 4 with exterior (posterior) prong sharply pointed longer than interior (anterior) prong. Apical tarsomere of fore leg with a black somewhat twisted prong below the posterior claw. Apical tarsomere of mid leg with a double dorsal comb of black squamiform bristles forming a flag. Mesonotum and abdominal tergites shining metallic green.
</paragraph>
<paragraph id="DB5FEC14F25BFFC0FCE0F8A92ACEF1EE" blockId="6.[811,1452,1862,1982]" pageId="6" pageNumber="646">Cercus in lateral view slightly shorter than dorsal surstylus. Cercus brown, tip pointed bearing a single long, yellow bristle. Dorsal surstylus mushroom-shaped, yellowish brown, bordered with short, yellow bristles.</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF25BFFC1FCE0F8092FFFF029" lastPageId="7" lastPageNumber="647" pageId="6" pageNumber="646" type="materials_examined">
<paragraph id="DB5FEC14F25BFFC0FCE0F8092BD0FE6E" blockId="6.[812,1452,2022,2110]" pageId="6" pageNumber="646">
<emphasis id="E9943006F25BFFC0FCE0F8092A7CF1AE" bold="true" box="[813,1031,2022,2046]" pageId="6" pageNumber="646">Material examined.</emphasis>
<materialsCitation id="6B88E649F25BFFC0FBC1F8092BD0FE6E" collectingDate="2015-02-09" collectorName="Samoh A" country="Thailand" latitude="8.907639" location="Bang Yai" longLatPrecision="1" longitude="98.39989" municipality="Takuapa" pageId="6" pageNumber="646" specimenCount="1" specimenCount-male="1" stateProvince="Phangnga" typeStatus="holotype">
<typeStatus id="045B52B6F25BFFC0FBC1F8092AE3F1AE" box="[1036,1176,2022,2046]" pageId="6" pageNumber="646" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="CDE6279DF25BFFC0FB51F8092AC9F1AD" box="[1180,1202,2022,2045]" pageId="6" pageNumber="646" type="male">♂</specimenCount>
, labelled:
<collectingCountry id="A3F7AC84F25BFFC0FAD6F8092BD3F1AE" box="[1307,1448,2022,2046]" name="Thailand" pageId="6" pageNumber="646">THAILAND</collectingCountry>
:
<collectingRegion id="192422F6F25BFFC0FCE1F7E92A6DFE4E" box="[812,1046,2054,2078]" country="Thailand" name="Phangnga" pageId="6" pageNumber="646">Phang Nga province</collectingRegion>
,
<collectingMunicipality id="3B3B766EF25BFFC0FBE8F7E92AFDFE4E" box="[1061,1158,2054,2078]" pageId="6" pageNumber="646">Takuapa</collectingMunicipality>
,
<location id="DE3FBACFF25BFFC0FB59F7E92B79FE4E" LSID="urn:lsid:plazi:treatment:53495D02F25BFFCDFCFAFA2A2A68F6DA:DE3FBACFF25BFFC0FB59F7E92B79FE4E" box="[1172,1282,2054,2078]" country="Thailand" latitude="8.907639" longLatPrecision="1" longitude="98.39989" municipality="Takuapa" name="Bang Yai" pageId="6" pageNumber="646" stateProvince="Phangnga">Bang Yai</location>
, (
<geoCoordinate id="BED48AD3F25BFFC0FAD4F7EB2BDDFE4D" box="[1305,1446,2052,2077]" degrees="8" direction="north" minutes="54" orientation="latitude" pageId="6" pageNumber="646" precision="1" seconds="27.5" value="8.907639">8°54′27.5″N</geoCoordinate>
,
<geoCoordinate id="BED48AD3F25BFFC0FCE0F7CB2DBBFE6D" box="[813,960,2084,2109]" degrees="98" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="646" precision="1" seconds="59.6" value="98.39989">98°23′59.6″E</geoCoordinate>
),
<collectorName id="761589C2F25BFFC0FC1BF7C92A3BFE6E" box="[982,1088,2086,2110]" pageId="6" pageNumber="646">Samoh A</collectorName>
, sweep netting,
<date id="AF5ECAD4F25BFFC0FB38F7C92BD0FE6E" box="[1269,1451,2086,2110]" pageId="6" pageNumber="646" value="2015-02-09">
<collectingDate id="BF1A333CF25BFFC0FB38F7C92BD0FE6E" box="[1269,1451,2086,2110]" pageId="6" pageNumber="646" value="2015-02-09">9 February 2015</collectingDate>
</date>
</materialsCitation>
</paragraph>
<caption id="8F9FBC9CF25AFFC1FC89FC392DE1F258" ID-DOI="http://doi.org/10.5281/zenodo.5343745" ID-Zenodo-Dep="5343745" httpUri="https://zenodo.org/record/5343745/files/figure.png" pageId="7" pageNumber="647" startId="7.[836,871,982,1004]" targetBox="[832,1478,193,968]" targetPageId="7">
<paragraph id="DB5FEC14F25AFFC1FC89FC392DE1F258" blockId="7.[836,1474,982,1032]" pageId="7" pageNumber="647">
Fig. 7.
<taxonomicName id="1CE09797F25AFFC1FC42FC392A23F5BC" authority="Samoh &amp; Satasook &amp; Grootaert, 2019" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[911,1112,982,1004]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="647" phylum="Arthropoda" rank="species" species="meieri" status="sp. nov.">
<emphasis id="E9943006F25AFFC1FC42FC392A23F5BC" box="[911,1112,982,1004]" italics="true" pageId="7" pageNumber="647">Ngirhaphium meieri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF25AFFC1FBAAFC382A9AF5BC" box="[1127,1249,982,1004]" pageId="7" pageNumber="647" rank="species">new species</taxonomicNameLabel>
, female habitus. Scale = 1 mm.
</paragraph>
</caption>
<caption id="8F9FBC9CF25AFFC1FF45FB582FAFF2B9" ID-DOI="http://doi.org/10.5281/zenodo.5343749" ID-Zenodo-Dep="5343749" httpUri="https://zenodo.org/record/5343749/files/figure.png" pageId="7" pageNumber="647" startId="7.[136,172,1207,1229]" targetBox="[137,774,190,1193]" targetPageId="7">
<paragraph id="DB5FEC14F25AFFC1FF45FB582FAFF2B9" blockId="7.[136,775,1207,1258]" pageId="7" pageNumber="647">
Fig. 6.
<taxonomicName id="1CE09797F25AFFC1FF19FB582FE4F29D" authority="Samoh &amp; Satasook &amp; Grootaert, 2019" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[212,415,1207,1229]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="647" phylum="Arthropoda" rank="species" species="meieri" status="sp. nov.">
<emphasis id="E9943006F25AFFC1FF19FB582FE4F29D" box="[212,415,1207,1229]" italics="true" pageId="7" pageNumber="647">Ngirhaphium meieri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF25AFFC1FE63FB582C52F29D" box="[430,553,1207,1229]" pageId="7" pageNumber="647" rank="species">new species</taxonomicNameLabel>
, male habitus (photo: Abdulloh Samoh). Scale = 1 mm.
</paragraph>
</caption>
<paragraph id="DB5FEC14F25AFFC1FF45FACE2F72F309" blockId="7.[136,775,1313,1369]" pageId="7" pageNumber="647">
(
<collectionCode id="BDF174D1F25AFFC1FF42FACE2F7EF369" box="[143,261,1313,1337]" pageId="7" pageNumber="647">PSUNHM</collectionCode>
). (Doli3 _26_005E, collected Abdulloh Samoh,
<collectionCode id="BDF174D1F25AFFC1FF44FAAE2E86F309" box="[137,253,1345,1369]" pageId="7" pageNumber="647">PSUNHM</collectionCode>
).
</paragraph>
<paragraph id="DB5FEC14F25AFFC1FF45FA6E2FFFF029" blockId="7.[134,776,1409,1657]" pageId="7" pageNumber="647">
<emphasis id="E9943006F25AFFC1FF45FA6E2E85F3C9" bold="true" box="[136,254,1409,1433]" pageId="7" pageNumber="647">
<typeStatus id="045B52B6F25AFFC1FF45FA6E2E82F3C9" box="[136,249,1409,1433]" pageId="7" pageNumber="647" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="CDE6279DF25AFFC1FECFFA6E2F2FF3C9" box="[258,340,1409,1433]" pageId="7" pageNumber="647" type="male">2 males</specimenCount>
;
<materialsCitation id="6B88E649F25AFFC1FE90FA6E2C6DF389" collectingDate="2015-02-09" collectionCode="PSUNHM" collectorName="Samoh A" latitude="8.907639" location="Bang Yai" longLatPrecision="1" longitude="98.39989" municipality="Takuapa" pageId="7" pageNumber="647" specimenCount="2" specimenCount-female="2" stateProvince="Phangnga" typeStatus="paratype">
<specimenCount id="CDE6279DF25AFFC1FE90FA6E2FB9F3C9" box="[349,450,1409,1433]" pageId="7" pageNumber="647" type="female">2 females</specimenCount>
,
<collectingRegion id="192422F6F25AFFC1FE07FA6E2CE5F3C9" box="[458,670,1409,1433]" country="Thailand" name="Phangnga" pageId="7" pageNumber="647">Phang Nga province</collectingRegion>
,
<collectingMunicipality id="3B3B766EF25AFFC1FD6BFA6E2D78F3C9" box="[678,771,1409,1433]" pageId="7" pageNumber="647">Takuapa</collectingMunicipality>
,
<location id="DE3FBACFF25AFFC1FF4BFA4E2E89F3E9" LSID="urn:lsid:plazi:treatment:53495D02F25BFFCDFCFAFA2A2A68F6DA:DE3FBACFF25AFFC1FF4BFA4E2E89F3E9" box="[134,242,1441,1465]" latitude="8.907639" longLatPrecision="1" longitude="98.39989" municipality="Takuapa" name="Bang Yai" pageId="7" pageNumber="647" stateProvince="Phangnga">Bang Yai</location>
, (
<geoCoordinate id="BED48AD3F25AFFC1FEC5FA702FE9F3E8" box="[264,402,1439,1464]" degrees="8" direction="north" minutes="54" orientation="latitude" pageId="7" pageNumber="647" precision="1" seconds="27.5" value="8.907639">8°54′27.5″N</geoCoordinate>
,
<geoCoordinate id="BED48AD3F25AFFC1FE6DFA702C4FF3E8" box="[416,564,1439,1464]" degrees="98" direction="east" minutes="23" orientation="longitude" pageId="7" pageNumber="647" precision="1" seconds="59.6" value="98.39989">98°23′59.6″E</geoCoordinate>
),
<collectorName id="761589C2F25AFFC1FD87FA4E2CCFF3E9" box="[586,692,1441,1465]" pageId="7" pageNumber="647">Samoh A</collectorName>
, sweep netting,
<date id="AF5ECAD4F25AFFC1FF13FA2E2FF7F389" box="[222,396,1473,1497]" pageId="7" pageNumber="647" value="2015-02-09">
<collectingDate id="BF1A333CF25AFFC1FF13FA2E2FF7F389" box="[222,396,1473,1497]" pageId="7" pageNumber="647" value="2015-02-09">9 February 2015</collectingDate>
</date>
(
<collectionCode id="BDF174D1F25AFFC1FE57FA2E2C6BF389" box="[410,528,1473,1497]" pageId="7" pageNumber="647">PSUNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6B88E649F25AFFC1FDEDFA2E2CD5F049" collectingDate="2018-07-13" collectionCode="PSUNHM" collectorName="Samoh A" latitude="8.907639" location="Bang Yai" longLatPrecision="1" longitude="98.39989" municipality="Takuapa" pageId="7" pageNumber="647" specimenCount="3" specimenCount-female="3" stateProvince="Phangnga" typeStatus="paratype">
<specimenCount id="CDE6279DF25AFFC1FDEDFA2E2CF2F389" box="[544,649,1473,1497]" pageId="7" pageNumber="647" type="female">3 females</specimenCount>
,
<collectingRegion id="192422F6F25AFFC1FD5FFA2E2E9CF3A9" country="Thailand" name="Phangnga" pageId="7" pageNumber="647">Phang Nga province</collectingRegion>
,
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,
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, (
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,
<geoCoordinate id="BED48AD3F25AFFC1FDABFA302C81F3A8" box="[614,762,1503,1528]" degrees="98" direction="east" minutes="23" orientation="longitude" pageId="7" pageNumber="647" precision="1" seconds="59.6" value="98.39989">98°23′59.6″E</geoCoordinate>
),
<collectorName id="761589C2F25AFFC1FF45F9EE2E94F049" box="[136,239,1537,1561]" pageId="7" pageNumber="647">Samoh A</collectorName>
, sweep netting,
<date id="AF5ECAD4F25AFFC1FE57F9EE2C58F049" box="[410,547,1537,1561]" pageId="7" pageNumber="647" value="2018-07-13">
<collectingDate id="BF1A333CF25AFFC1FE57F9EE2C58F049" box="[410,547,1537,1561]" pageId="7" pageNumber="647" value="2018-07-13">13 July 2018</collectingDate>
</date>
(
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)
</materialsCitation>
;
<materialsCitation id="6B88E649F25AFFC1FD75F9EE2F04F029" collectingDate="2018-07-10" collectionCode="PSUNHM" collectorName="Samoh A." county="Kraburi" latitude="10.32375" location="Phra Khayang cave" longLatPrecision="1" longitude="98.76611" municipality="Lam Liang" pageId="7" pageNumber="647" specimenCount="1" specimenCount-male="1" stateProvince="Ranong" typeStatus="paratype">
<specimenCount id="CDE6279DF25AFFC1FD75F9EE2D79F049" box="[696,770,1537,1561]" pageId="7" pageNumber="647" type="male">1 male</specimenCount>
,
<collectingRegion id="192422F6F25AFFC1FF45F9CE2F38F069" box="[136,323,1569,1593]" country="Thailand" name="Ranong" pageId="7" pageNumber="647">Ranong province</collectingRegion>
,
<collectingCounty id="323E9498F25AFFC1FE83F9CE2FDFF069" box="[334,420,1569,1593]" pageId="7" pageNumber="647">Kraburi</collectingCounty>
,
<collectingMunicipality id="3B3B766EF25AFFC1FE7DF9CE2C52F069" box="[432,553,1569,1593]" pageId="7" pageNumber="647">Lam Liang</collectingMunicipality>
,
<location id="DE3FBACFF25AFFC1FDF8F9CE2D7CF068" LSID="urn:lsid:plazi:treatment:53495D02F25BFFCDFCFAFA2A2A68F6DA:DE3FBACFF25AFFC1FDF8F9CE2D7CF068" box="[565,775,1569,1593]" county="Kraburi" latitude="10.32375" longLatPrecision="1" longitude="98.76611" municipality="Lam Liang" name="Phra Khayang cave" pageId="7" pageNumber="647" stateProvince="Ranong">Phra Khayang cave</location>
(
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<geoCoordinate id="BED48AD3F25AFFC1FEE1F9D02FBBF008" box="[300,448,1599,1624]" degrees="98" direction="east" minutes="45" orientation="longitude" pageId="7" pageNumber="647" precision="1" seconds="58.0" value="98.76611">98°45′58.0″E</geoCoordinate>
),
<collectorName id="761589C2F25AFFC1FE1EF9AE2C3BF009" box="[467,576,1601,1625]" pageId="7" pageNumber="647">Samoh A.</collectorName>
, sweep netting,
<date id="AF5ECAD4F25AFFC1FD20F9AE2E89F029" pageId="7" pageNumber="647" value="2018-07-10">
<collectingDate id="BF1A333CF25AFFC1FD20F9AE2E89F029" pageId="7" pageNumber="647" value="2018-07-10">10 July 2018</collectingDate>
</date>
(
<collectionCode id="BDF174D1F25AFFC1FECEF98E2F02F029" box="[259,377,1633,1657]" pageId="7" pageNumber="647">PSUNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF25AFFC1FF47F94E2CD9F169" pageId="7" pageNumber="647" type="etymology">
<paragraph id="DB5FEC14F25AFFC1FF47F94E2CD9F169" blockId="7.[136,775,1697,1849]" pageId="7" pageNumber="647">
<emphasis id="E9943006F25AFFC1FF47F94E2F69F0E9" bold="true" box="[138,274,1697,1721]" pageId="7" pageNumber="647">Etymology.</emphasis>
The species is dedicated to Rudolf Meier, head of Evolutionary Biology Laboratory, Department of Biological Sciences at the National University of
<collectingCountry id="A3F7AC84F25AFFC1FD56F90E2D7CF0A9" box="[667,775,1761,1785]" name="Singapore" pageId="7" pageNumber="647">Singapore</collectingCountry>
(NUS), who provided us an opportunity to learn and do a molecular study of the marine dolichopodid flies.
</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF25AFFCCFF45F88E2DF9F118" lastPageId="10" lastPageNumber="650" pageId="7" pageNumber="647" type="description">
<paragraph id="DB5FEC14F25AFFC1FF45F88E2BF9F0C8" blockId="7.[135,776,1889,2073]" lastBlockId="7.[834,1476,1088,1688]" pageId="7" pageNumber="647">
<emphasis id="E9943006F25AFFC1FF45F88E2F69F129" bold="true" box="[136,274,1889,1913]" pageId="7" pageNumber="647">Description.</emphasis>
Male. (
<figureCitation id="43DBF091F25AFFC1FEACF88E2FDAF129" box="[353,417,1889,1913]" captionStart="Fig" captionStartId="7.[136,172,1207,1229]" captionTargetBox="[137,774,190,1193]" captionTargetId="figure-438@7.[127,787,175,1202]" captionTargetPageId="7" captionText="Fig. 6. Ngirhaphium meieri, new species, male habitus (photo: Abdulloh Samoh). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343749" httpUri="https://zenodo.org/record/5343749/files/figure.png" pageId="7" pageNumber="647">Fig. 6</figureCitation>
). Length body:
<quantity id="1C1841F1F25AFFC1FD89F88E2CB9F128" box="[580,706,1889,1913]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="8.8" metricValueMin="8.6" pageId="7" pageNumber="647" unit="mm" value="8.7" valueMax="8.8" valueMin="8.6">8.6-8.8 mm</quantity>
; wing:
<quantity id="1C1841F1F25AFFC1FF4AF86E2F7FF1C8" box="[135,260,1921,1945]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.6" metricValueMin="8.4" pageId="7" pageNumber="647" unit="mm" value="8.5" valueMax="8.6" valueMin="8.4">8.4-8.6 mm</quantity>
. Head. Frons shining metallic green (not dusted), deeply excavated. Face greenish brown in ground-colour, silvery grey dusted; apex of face and clypeus yellowish brown in ground-colour, parallel-sided. Eyes pass beyond border of face; eyes densely set with minute white hairs. Ocellar callus globular protruding from frons with two very long ocellars, directed backward, divergent. Vertical bristles long, half as long as ocellars, rather anteriad on frons at level of ocellar callus, close to eye border, long, black, directed forward and cruciate. Pair of long black postverticals directed backward and crossing. Postoculars above strong, black in single row, below white and mixed with very long white hairs below mouth; postcranium greenish in ground-colour but grey dusted. Palpus long, strap-shaped, yellowish with few short black hairs, no apical bristle. Labella brown with black hairs. Antenna long, completely black. First segment long, 3 times as long as second segment; second segment short, apically with crown of short black bristles. Third segment very long strap-shaped, laterally flattened, with a lateral rim, about 6.5 times as long as width at base. Arista apical, apical segment twice as long as basal article, gradually tapering towards tip. Total length antenna:
<quantity id="1C1841F1F25AFFC1FAD9F9AF2B11F007" box="[1300,1386,1600,1624]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="647" unit="mm" value="2.1">2.1 mm</quantity>
. Length of scape: 0.39; pedicel: 0.13; postpedicel: 0.87; basal aristal article: 0.23; apical aristal article: 0.468 (all in mm).
</paragraph>
<paragraph id="DB5FEC14F25AFFC1FC8EF92F2BE9FE48" blockId="7.[834,1477,1728,2072]" pageId="7" pageNumber="647">Thorax and scutellum dark metallic green in ground-colour (bluish when seen from in front), covered with fine grey dusting. All hairs and bristles black. Pleura more densely grey dusted than mesonotum. Acrostichals biseriate, about 7 pairs, rows widening slightly behind. Presutural dorsocentrals multiseriate; 6 postsutural dorsocentrals: 4 short and 2 long prescutellars; 1 pair of strong scutellars. One long humeral with shorter bristle in front; 1 strong posthumeral, 2 strong notopleurals, 1 postsutural, 1 supraalar, 1 very strong postalar. Propleural bristles black, 6 short, fine upper and 2 lower propleural bristles (lower one twice as long as upper).</paragraph>
<caption id="8F9FBC9CF255FFCEFFBCF8242DB2FE4A" ID-DOI="http://doi.org/10.5281/zenodo.5343751" ID-Zenodo-Dep="5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="8" pageNumber="648" startId="8.[113,148,1995,2017]" targetBox="[112,1457,228,1982]" targetPageId="8">
<paragraph id="DB5FEC14F255FFCEFFBCF8242DB2FE4A" blockId="8.[113,1452,1995,2075]" pageId="8" pageNumber="648">
Fig. 8.
<taxonomicName id="1CE09797F255FFCEFF7BF8242F01F1B1" authority="Samoh &amp; Satasook &amp; Grootaert, 2019" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[182,378,1995,2017]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="648" phylum="Arthropoda" rank="species" species="meieri" status="sp. nov.">
<emphasis id="E9943006F255FFCEFF7BF8242F01F1B1" box="[182,378,1995,2017]" italics="true" pageId="8" pageNumber="648">Ngirhaphium meieri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF255FFCEFE4BF8232F86F1B1" box="[390,509,1995,2017]" pageId="8" pageNumber="648" rank="species">new species</taxonomicNameLabel>
, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm.
</paragraph>
</caption>
<caption id="8F9FBC9CF254FFCFFF44F85F2A09F196" ID-DOI="http://doi.org/10.5281/zenodo.5343753" ID-Zenodo-Dep="5343753" box="[137,1138,1968,1990]" httpUri="https://zenodo.org/record/5343753/files/figure.png" pageId="9" pageNumber="649" startId="9.[137,172,1968,1990]" targetBox="[142,1476,317,1952]" targetPageId="9">
<paragraph id="DB5FEC14F254FFCFFF44F85F2A09F196" blockId="9.[137,1138,1968,1990]" box="[137,1138,1968,1990]" pageId="9" pageNumber="649">
Fig. 9. Distribution map of
<taxonomicName id="1CE09797F254FFCFFE56F85F2C24F195" authority="Samoh &amp; Satasook &amp; Grootaert, 2019" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[411,607,1968,1989]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="649" phylum="Arthropoda" rank="species" species="meieri" status="sp. nov.">
<emphasis id="E9943006F254FFCFFE56F85F2C24F195" box="[411,607,1968,1989]" italics="true" pageId="9" pageNumber="649">Ngirhaphium meieri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF254FFCFFDA1F85F2C9AF196" box="[620,737,1968,1990]" pageId="9" pageNumber="649" rank="species">new species</taxonomicNameLabel>
and
<taxonomicName id="1CE09797F254FFCFFCD8F85F2D92F195" authority="Samoh &amp; Satasook &amp; Grootaert, 2019" authorityName="Samoh &amp; Satasook &amp; Grootaert" authorityYear="2019" box="[789,1001,1968,1989]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="649" phylum="Arthropoda" rank="species" species="thaicum" status="sp. nov.">
<emphasis id="E9943006F254FFCFFCD8F85F2D92F195" box="[789,1001,1968,1989]" italics="true" pageId="9" pageNumber="649">Ngirhaphium thaicum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F2A78D7DF254FFCFFC3BF85F2A15F196" box="[1014,1134,1968,1990]" pageId="9" pageNumber="649" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="DB5FEC14F257FFCCFFBDFF512F01F7E5" blockId="10.[111,753,189,437]" pageId="10" pageNumber="650">
Legs (
<figureCitation id="43DBF091F257FFCCFF79FF522E8EF685" box="[180,245,189,213]" captionStart="Fig" captionStartId="6.[113,148,1372,1394]" captionTargetBox="[99,1465,178,1370]" captionTargetId="figure-243@6.[99,1465,178,1370]" captionTargetPageId="6" captionText="Fig. 5. Ngirhaphium thaicum, new species male terminalia (27_009) A, epandrium left side; B, cerci dorsal view; C, left surstylus inside view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343741" httpUri="https://zenodo.org/record/5343741/files/figure.png" pageId="10" pageNumber="650">Fig. 5</figureCitation>
) yellow, all bristles black. All coxae brownish black in ground-colour, covered with fine greyish dusting. All trochanters brown. Tip of hind tibia annulated brown at tip. Fore tarsus with tip of tarsomere 1 brownish, tarsomeres 2–4 almost entirely brown (especially dorsally), tarsomere 5 entirely black. Mid tarsus more yellowish than fore tarsus, with apical tarsomere black; hind tarsus black but base of tarsomere 1 and 2 paler.
</paragraph>
<paragraph id="DB5FEC14F257FFCCFFBDFE312C95F565" blockId="10.[111,753,477,822]" pageId="10" pageNumber="650">Fore leg. Coxa with short black bristles. Fore femur slightly swollen on basal half; row of minute posteroventrals in apical half. Short preapical posterior bristle and 1 stronger anterior preapical bristle directed forward. Fore tibia with 4 strong ad, 4 strong pd and crown of 4 apicals. All tarsal segments densely set with black hairs and short black bristles. Tarsomere 4 with dorsal asymmetrical fork, extending over tarsomere 5; exterior (posterior) prong longer than interior (anterior) prong, both sharply pointed when seen in lateral view. Apical tarsomere with a black, blunt, somewhat twisted prong below the posterior claw, as long as apical tarsomere.</paragraph>
<caption id="8F9FBC9CF257FFCCFCE1FCA92BECF529" ID-DOI="http://doi.org/10.5281/zenodo.5343757" ID-Zenodo-Dep="5343757" httpUri="https://zenodo.org/record/5343757/files/figure.png" pageId="10" pageNumber="650" startId="10.[812,847,838,860]" targetBox="[813,1451,191,825]" targetPageId="10">
<paragraph id="DB5FEC14F257FFCCFCE1FCA92BECF529" blockId="10.[812,1453,838,889]" pageId="10" pageNumber="650">
Fig. 10. Haplotype network of
<emphasis id="E9943006F257FFCCFB92FCA92B45F50B" box="[1119,1342,838,859]" italics="true" pageId="10" pageNumber="650">
<taxonomicName id="1CE09797F257FFCCFB92FCA92B41F50B" authorityName="Evenhuis &amp; Grootaert" authorityYear="2002" box="[1119,1338,838,859]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="650" phylum="Arthropoda" rank="species" species="murphyi">Ngirhaphium murphyi</taxonomicName>
.
</emphasis>
Singapore: Pulau Ubin, Mandai Sungei Buloh; Thailand: Satun and Krabi.
</paragraph>
</caption>
<paragraph id="DB5FEC14F257FFCCFFBDFCB12F09F5E5" blockId="10.[112,751,861,949]" pageId="10" pageNumber="650">Two well-developed pulvilli and empodium present. Length of femur, tibia and tarsal segments (in mm): 1.6: 1.62: 0.8: 0.26: 0.2: 0.16: 0.2.</paragraph>
<paragraph id="DB5FEC14F257FFCCFFBDFC322F1AF365" blockId="10.[111,752,989,1333]" pageId="10" pageNumber="650">
Mid leg. Coxa with short bristles anteriorly; no exterior bristle. Mid femur as wide as fore femur; ventrally with inconspicuous bristles; 1 strong anterior preapical and 2 tiny posterior preapicals. Tibia with 6 ad, 6 pd (might be considered as dorsal), 8 longer av and crown of long apicals. Tarsomeres 1–4 ventrally at tip with pair of short spine-like bristles as well as comb of shorter black spinules. Apical tarsomere of mid leg with a double, dorsal comb of black squamiform bristles forming a flag (
<figureCitation id="43DBF091F257FFCCFE38FB322C4FF2A5" box="[501,564,1245,1269]" captionStart="Fig" captionStartId="6.[113,148,1372,1394]" captionTargetBox="[99,1465,178,1370]" captionTargetId="figure-243@6.[99,1465,178,1370]" captionTargetPageId="6" captionText="Fig. 5. Ngirhaphium thaicum, new species male terminalia (27_009) A, epandrium left side; B, cerci dorsal view; C, left surstylus inside view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343741" httpUri="https://zenodo.org/record/5343741/files/figure.png" pageId="10" pageNumber="650">Fig. 5</figureCitation>
inset). Length of femur, tibia and tarsal segments (in mm): 1.9: 2.4: 1.3: 0.74: 0.54: 0.3: 0.3.
</paragraph>
<paragraph id="DB5FEC14F257FFCCFFBDFAB22C9CF005" blockId="10.[111,753,1373,1622]" pageId="10" pageNumber="650">Hind leg. Coxa bare. Trochanter with a long dorsal bristle. Hind femur thicker than fore and mid femur, ventrally almost bare; 1 strong anterior preapical, 1 fine posterior preapical. Tibia stronger bristled than mid tibia with 7 long av, 7 ad, 7 pd and crown of long apicals. Tarsomeres 1–4 ventrally at tip with pair of short spine-like bristles as well as comb of shorter black spinules. Length of femur, tibia and tarsal segments (in mm): 1.8: 3.3: 1.4: 0.8: 0.64: 0.36: 0.4.</paragraph>
<paragraph id="DB5FEC14F257FFCCFFBDF9922FDFF125" blockId="10.[111,752,1661,1909]" pageId="10" pageNumber="650">
Wing (
<figureCitation id="43DBF091F257FFCCFF77F9922E80F0C5" box="[186,251,1661,1685]" captionStart="Fig" captionStartId="7.[136,172,1207,1229]" captionTargetBox="[137,774,190,1193]" captionTargetId="figure-438@7.[127,787,175,1202]" captionTargetPageId="7" captionText="Fig. 6. Ngirhaphium meieri, new species, male habitus (photo: Abdulloh Samoh). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343749" httpUri="https://zenodo.org/record/5343749/files/figure.png" pageId="10" pageNumber="650">Fig. 6</figureCitation>
) tinged brownish (paler on imaged specimen), anteriorly between costa and R
<subScript id="4764EE51F257FFCCFE04F9462F9AF0E7" attach="left" box="[457,481,1705,1719]" fontSize="6" pageId="10" pageNumber="650">4+5</subScript>
a little darker yellowish brown. Tp (crossvein r-m) brown seamed. Veins dark brown, yellowish at base. M
<subScript id="4764EE51F257FFCCFE96F9062F0FF0A7" attach="left" box="[347,372,1769,1783]" fontSize="6" pageId="10" pageNumber="650">1+2</subScript>
sharply bent upwards and ending in costa closely near tip of R
<subScript id="4764EE51F257FFCCFE63F8E62FBDF147" attach="left" box="[430,454,1801,1815]" fontSize="6" pageId="10" pageNumber="650">4+5</subScript>
. Tp straight, a little longer than apical part of M
<subScript id="4764EE51F257FFCCFE93F8C62F0DF167" attach="left" box="[350,374,1833,1847]" fontSize="6" pageId="10" pageNumber="650">3+4</subScript>
. Anal vein reaching wing border. Haltere with white knob, stem a little infuscate. Squama white, with long white cilia.
</paragraph>
<paragraph id="DB5FEC14F257FFCCFFBDF8722C49FE65" blockId="10.[111,752,1949,2101]" pageId="10" pageNumber="650">Abdomen subshining dark metallic green. Tergites densely set with quite long black bristles; hind-marginal bristles slightly longer than other bristles. Only tergites 1 and 5 with much longer marginal bristles. Sternites with very short hairs except for longer marginals on sternite 4.</paragraph>
<paragraph id="DB5FEC14F257FFCCFCE6FC5F2BEFF318" blockId="10.[810,1452,944,1352]" pageId="10" pageNumber="650">
Terminalia (
<figureCitation id="43DBF091F257FFCCFC74FC5F2A7BF598" box="[953,1024,944,968]" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8</figureCitation>
). Cercus yellowish brown to brown, slightly shorter than dorsal surstylus (
<figureCitation id="43DBF091F257FFCCFB06FC3F2B58F5B8" box="[1227,1315,976,1000]" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8A</figureCitation>
). Both cerci look like fused for almost entire length, only tips free (
<figureCitation id="43DBF091F257FFCCFAB2FC1F2D35F278" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8B</figureCitation>
). Tip of cercus pointed, with a single yellow apical bristle, dorsally set with long blackish brown bristles, sometimes pale at tip. Dorsal surstylus brown, paler than cercus, with much widened tip in the shape of a mushroom, forming a transverse clasper on cercus, bordered with short but stout yellow bristles (
<figureCitation id="43DBF091F257FFCCFC2CFB5F2A40F298" box="[993,1083,1200,1224]" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8A</figureCitation>
). Ventral and dorsal surstyli not fused. Ventral surstylus yellowish, oval (
<figureCitation id="43DBF091F257FFCCFB27FB3F2B44F2B8" box="[1258,1343,1232,1256]" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8C</figureCitation>
), tip with 3 long bristles on apical border and a few short hair-like bristles at inside. Hypandrium dorsally split, with a large rounded black protuberance set with denticles (
<figureCitation id="43DBF091F257FFCCFAE2FADF2BFCF318" box="[1327,1415,1328,1352]" captionStart="Fig" captionStartId="8.[113,148,1995,2017]" captionTargetBox="[112,1457,228,1982]" captionTargetId="figure-11@8.[99,1467,209,1988]" captionTargetPageId="8" captionText="Fig. 8. Ngirhaphium meieri, new species, holotype male terminalia: A, Lateral view of genital capsule with left ventral surstylus removed; B, Cerci dorsally; C. Left ventral surstylus; D, Ventral view of genital capsule. Abbreviations: ae = aedeagus; c = cercus; ds = dorsal surstylus; hy = hypandrium; sp = sperm pump; vs = ventral surstylus. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343751" httpUri="https://zenodo.org/record/5343751/files/figure.png" pageId="10" pageNumber="650">Fig. 8A</figureCitation>
).
</paragraph>
<paragraph id="DB5FEC14F257FFCCFCE6FA9F2BE0F3D8" blockId="10.[811,1435,1392,1416]" box="[811,1435,1392,1416]" pageId="10" pageNumber="650">
Female. (
<figureCitation id="43DBF091F257FFCCFC5CFA9F2DAEF3D8" box="[913,981,1392,1416]" captionStart="Fig" captionStartId="7.[836,871,982,1004]" captionTargetBox="[832,1478,193,968]" captionTargetId="figure-456@7.[819,1482,164,981]" captionTargetPageId="7" captionText="Fig. 7. Ngirhaphium meieri, new species, female habitus. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5343745" httpUri="https://zenodo.org/record/5343745/files/figure.png" pageId="10" pageNumber="650">Fig. 7</figureCitation>
). Body: 8.9–9.0 mm; Wing:
<quantity id="1C1841F1F257FFCCFADFFA9F2BE3F3D8" box="[1298,1432,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="8.8" metricValueMin="8.6" pageId="10" pageNumber="650" unit="mm" value="8.7" valueMax="8.8" valueMin="8.6">8.6–8.8 mm</quantity>
.
</paragraph>
<paragraph id="DB5FEC14F257FFCCFCE6FA5F2A2BF3B9" blockId="10.[811,1450,1456,1513]" pageId="10" pageNumber="650">The female has larger labellae than the male, which is a characteristic of the genus.</paragraph>
<paragraph id="DB5FEC14F257FFCCFCE6F9FF2DA4F0F8" blockId="10.[811,1450,1552,1704]" pageId="10" pageNumber="650">
The antenna is slightly shorter than in male with apical aristal segment twice as long as basal segment. Length of scape: 0.377; pedicel: 0.13; postpedicel: 0.741; basal aristal article: 0.234; apical aristal article: 0.455 (all in mm). Total length
<quantity id="1C1841F1F257FFCCFCBAF97F2DA0F0F8" box="[887,987,1680,1704]" metricMagnitude="-3" metricUnit="m" metricValue="1.94" pageId="10" pageNumber="650" unit="mm" value="1.94">1.94 mm</quantity>
.
</paragraph>
<paragraph id="DB5FEC14F257FFCCFCE6F93E2DF9F118" blockId="10.[811,1452,1744,1864]" pageId="10" pageNumber="650">Fore tarsomere 4 without dorsal fork. Tarsomere 5 with an apical dorsal projection and without ventral prong below the exterior claw. Mid tarsomere 5 without dorsal squamiform bristles.</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF257FFCCFCE6F89F2A0DF1F8" pageId="10" pageNumber="650" type="distribution">
<paragraph id="DB5FEC14F257FFCCFCE6F89F2A0DF1F8" blockId="10.[811,1450,1904,1960]" pageId="10" pageNumber="650">
<emphasis id="E9943006F257FFCCFCE6F89F2DC5F1D8" bold="true" box="[811,958,1904,1928]" pageId="10" pageNumber="650">Distribution.</emphasis>
<collectingRegion id="192422F6F257FFCCFC08F89F2A47F1D9" box="[965,1084,1904,1929]" country="Thailand" name="Phangnga" pageId="10" pageNumber="650">Phang Nga</collectingRegion>
and
<collectingRegion id="192422F6F257FFCCFBBEF89E2ABDF1D9" box="[1139,1222,1905,1929]" country="Thailand" name="Ranong" pageId="10" pageNumber="650">Ranong</collectingRegion>
provinces,
<collectingCountry id="A3F7AC84F257FFCCFA88F89F2BDCF1D8" box="[1349,1447,1904,1928]" name="Thailand" pageId="10" pageNumber="650">Thailand</collectingCountry>
, Andaman Sea (Indian Ocean).
</paragraph>
</subSubSection>
<subSubSection id="93FABF9FF257FFCDFCE6F83F2A68F6DA" lastPageId="11" lastPageNumber="651" pageId="10" pageNumber="650" type="discussion">
<paragraph id="DB5FEC14F257FFCCFCE6F83F2BD0FE78" blockId="10.[811,1451,2000,2088]" pageId="10" pageNumber="650">
<emphasis id="E9943006F257FFCCFCE6F83F2DE1F1B8" bold="true" box="[811,922,2000,2024]" pageId="10" pageNumber="650">Remarks.</emphasis>
The new species is morphologically similar to
<taxonomicName id="1CE09797F257FFCCFA5CF83E2DEAFE58" authorityName="Grootaert &amp; Puniamoorthy" authorityYear="2014" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="650" phylum="Arthropoda" rank="species" species="sivasothii">
<emphasis id="E9943006F257FFCCFA5CF83E2DEAFE58" italics="true" pageId="10" pageNumber="650">N. sivasothii</emphasis>
</taxonomicName>
and even more to
<taxonomicName id="1CE09797F257FFCCFB9BF81E2A8EFE58" authorityName="Samoh, Boonrotpongi &amp; Grootaert" authorityYear="2015" box="[1110,1269,2032,2056]" class="Insecta" family="Dolichopodidae" genus="Ngirhaphium" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="650" phylum="Arthropoda" rank="species" species="chutamasae">
<emphasis id="E9943006F257FFCCFB9BF81E2A8EFE58" box="[1110,1269,2032,2056]" italics="true" pageId="10" pageNumber="650">N. chutamasae</emphasis>
</taxonomicName>
. Differences can be found in the key to the Southeast Asian species here below.
</paragraph>
<paragraph id="DB5FEC14F256FFCDFDFBFF9D2A68F6DA" blockId="11.[566,1043,114,138]" box="[566,1043,114,138]" pageId="11" pageNumber="651">
<emphasis id="E9943006F256FFCDFDFBFF9D2DAEF6D9" box="[566,981,114,137]" italics="true" pageId="11" pageNumber="651">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2019
</paragraph>
</subSubSection>
</treatment>
</document>
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