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<document id="FA1732B539CD5EB4B14FBEE014922889" ID-DOI="10.1080/00222930903094621" ID-ISSN="1464-5262" ID-Zenodo-Dep="5217087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629293244201" checkinUser="felipe" docAuthor="Yanes, Yurena, Martín, Javier, Moro, Leopoldo, Alonso, María R. &amp; Ibáñez, Miguel" docDate="2009" docId="4014878B4560793C40A1817EFD0BFEAB" docLanguage="en" docName="JNatHist.43.35-36.2179-2207.pdf" docOrigin="Journal of Natural History 43 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222930903094621" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Napaeus ornamentatus Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2202" masterDocId="BC2DFFF345777925413C8541FFAFFF94" masterDocTitle="On the relationships of the genus Napaeus (Gastropoda: Pulmonata: Enidae) with the descriptions of four new species from the Canary Islands" masterLastPageNumber="2207" masterPageNumber="2179" pageNumber="2200" updateTime="1699067505050" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0F9DBF97E7709CF8BEB5790DE7889E63">On the relationships of the genus Napaeus (Gastropoda: Pulmonata: Enidae) with the descriptions of four new species from the Canary Islands</mods:title>
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<mods:namePart id="0366978BE83EA16EB51AC3C786974223">Yanes, Yurena</mods:namePart>
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<mods:namePart id="B56DB99BB058EB86D76ED2EC607C67E3">Martín, Javier</mods:namePart>
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<mods:namePart id="3CE81D788CDBABD90D7DEF0B09BC54FA">Moro, Leopoldo</mods:namePart>
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<mods:namePart id="2F1D86577A85429D2CF394ADE4A2CBB5">Alonso, María R.</mods:namePart>
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<mods:namePart id="02A54576C3111AF707459CF3FA3F09D1">Ibáñez, Miguel</mods:namePart>
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<mods:typeOfResource id="2D8F0EF0B36A64B26AF2C5A0D3F71E28">text</mods:typeOfResource>
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<mods:title id="DFBA659B4F95974FB49A5F3AD85EE7E8">Journal of Natural History</mods:title>
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<mods:date id="0627F99E152575A52E4B09314E8B3D78">2009</mods:date>
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<mods:number id="5D82BF992FD68EA29BBA4BBBF48299E3">2009-08-26</mods:number>
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<mods:detail id="492467B242F3109AB1930853AB6D11CC" type="volume">
<mods:number id="D114250E3EB361559B788E9ACF575D84">43</mods:number>
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<mods:detail id="37AF7122090773B085E29199C9322D40" type="issue">
<mods:number id="AC5677CFD6A4DAD1B6F27F3113DE906E">35 - 36</mods:number>
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<mods:start id="CB342CA92336F6882CEF9DBE19341E39">2179</mods:start>
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<mods:identifier id="754BDE328F85B4B69D06D13192606D5F" type="ISSN">1464-5262</mods:identifier>
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<treatment id="4014878B4560793C40A1817EFD0BFEAB" LSID="urn:lsid:plazi:treatment:4014878B4560793C40A1817EFD0BFEAB" httpUri="http://treatment.plazi.org/id/4014878B4560793C40A1817EFD0BFEAB" lastPageId="25" lastPageNumber="2202" pageId="23" pageNumber="2200">
<subSubSection id="80A765164560793240A1817EFCECFBC3" box="[413,835,1087,1111]" pageId="23" pageNumber="2200" type="nomenclature">
<paragraph id="C802369D4560793240A1817EFCECFBC3" blockId="23.[413,835,1087,1111]" box="[413,835,1087,1111]" pageId="23" pageNumber="2200">
<heading id="934A81F14560793240A1817EFCECFBC3" box="[413,835,1087,1111]" centered="true" fontSize="36" level="2" pageId="23" pageNumber="2200" reason="7">
<taxonomicName id="0FBD4D1E4560793240A1817EFD4CFBC3" ID-CoL="45NQK" authority="Moro" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[413,739,1087,1111]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="2200" phylum="Mollusca" rank="species" species="ornamentatus" status="sp. nov.">
<emphasis id="FAC9EA8F4560793240A1817EFD35FBC3" bold="true" box="[413,666,1087,1111]" italics="true" pageId="23" pageNumber="2200">Napaeus ornamentatus</emphasis>
Moro
</taxonomicName>
<taxonomicNameLabel id="E1FA57F44560793243D58101FCECFBC3" box="[745,835,1088,1111]" pageId="23" pageNumber="2200" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="80A765164560793C41AC81C6FD0BFEAB" lastPageId="25" lastPageNumber="2202" pageId="23" pageNumber="2200" type="materials_examined">
<paragraph id="C802369D4560793241AC81C6FE85FB0B" blockId="23.[144,298,1159,1183]" box="[144,298,1159,1183]" pageId="23" pageNumber="2200">
<emphasis id="FAC9EA8F4560793241AC81C6FE85FB0B" box="[144,298,1159,1183]" italics="true" pageId="23" pageNumber="2200">Type material</emphasis>
</paragraph>
<paragraph id="C802369D4560793241AC81EEFE56FA93" blockId="23.[144,1104,1199,1415]" pageId="23" pageNumber="2200">
<materialsCitation id="78D53CC04560793241AC81EEFE56FA93" collectingDate="2008-03-05" collectionCode="UTM" collectorName="de Taguluche &amp; La Gomera" elevation="800" location="Mirador de Taguluche" pageId="23" pageNumber="2200" specimenCode="MT 0400, RBS7113" specimenCount="1" typeStatus="holotype">
<typeStatus id="1706883F4560793241AC81EEFF54FB53" box="[144,251,1199,1223]" pageId="23" pageNumber="2200" type="holotype">Holotype</typeStatus>
(
<figureCitation id="50862A1845607932403581EEFE26FB53" box="[265,393,1199,1223]" captionStart="Figure 4" captionStartId="14.[212,279,1195,1217]" captionTargetBox="[220,1162,176,1153]" captionTargetId="figure-214@14.[212,1172,168,1165]" captionTargetPageId="14" captionText="Figure 4. Napaeus shell drawings. (A) N. tenoensis (holotype, AIT); (B) N. bajamarensis sp. nov. (holotype, TFMC MT 0397); (C) N. boucheti (holotype, AIT); (D) N. obesatus; (E) N. propinquus; (F) N. ornamentatus sp. nov. (holotype, TFMC MT 0400); (G) N. orientalis (holotype, AIT); (H) N. ocellatus (syntype, ZMZ 513767); (J) N. rupicola; (K) N. avaloensis (holotype, TFMC MT 0382); (L) N. maculatus (holotype, TFMC MT 0386); (M) N. helvolus; (N) N. voggenreiteri (holotype, TFMC MT 0384); (O) N. isletae (holotype, AIT); (P) N. variatus; (Q) N. encaustus; (R) N. moquinianus; (S) N. teobaldoi sp. nov. (holotype, TFMC MT 0399); (T) N. lajaensis (holotype, TFMC MT 0381); (U) N. procerus (holotype, TFMC MT 0385); (V) N. rufobrunneus." figureDoi="http://doi.org/10.5281/zenodo.5217097" httpUri="https://zenodo.org/record/5217097/files/figure.png" pageId="23" pageNumber="2200">Figures 4F</figureCitation>
,
<figureCitation id="50862A184560793240A881EEFE17FB53" box="[404,440,1199,1223]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (AD) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="23" pageNumber="2200">7C</figureCitation>
): TFMC (
<specimenCode id="981B9EE645607932431081EEFD35FB53" box="[556,666,1199,1223]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="23" pageNumber="2200">MT 0400</specimenCode>
). Type locality:
<location id="CD62604645607932427381EEFBFFFB53" LSID="urn:lsid:plazi:treatment:4014878B4560793C40A1817EFD0BFEAB:CD62604645607932427381EEFBFFFB53" box="[847,1104,1199,1223]" name="Mirador de Taguluche" pageId="23" pageNumber="2200">
Mirador
<collectorName id="6548534B45607932428B81EEFBFFFB53" box="[951,1104,1199,1223]" pageId="23" pageNumber="2200">de Taguluche</collectorName>
</location>
(
<collectorName id="6548534B4560793241A4818EFEB0FB73" box="[152,287,1231,1255]" pageId="23" pageNumber="2200">La Gomera</collectorName>
;
<collectionCode id="AEACAE584560793240158191FEC4FB73" box="[297,363,1232,1255]" pageId="23" pageNumber="2200">UTM</collectionCode>
coordinates: 28
<specimenCode id="981B9EE6456079324322818EFD24FB73" box="[542,651,1231,1255]" collectionCode="RBS" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="23" pageNumber="2200" type="Herbarium">RBS7113</specimenCode>
;
<quantity id="0F459B784560793243A9818EFD75FB73" box="[661,730,1231,1255]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="2200" unit="m" value="800.0">
<elevation id="4390D1AE4560793243A9818EFD75FB73" box="[661,730,1231,1255]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="2200" unit="m" value="800.0">800 m</elevation>
</quantity>
altitude). Leg. Javier Martín and Leopoldo Moro,
<date id="BC03105D45607932406881AEFE5BFA93" box="[340,500,1263,1287]" pageId="23" pageNumber="2200" value="2008-03-05">
<collectingDate id="AC47E9B545607932406881AEFE5BFA93" box="[340,500,1263,1287]" pageId="23" pageNumber="2200" value="2008-03-05">5 March 2008</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="C802369D4560793241868051FC18FA13" blockId="23.[144,1104,1199,1415]" pageId="23" pageNumber="2200">
<materialsCitation id="78D53CC04560793241868051FC1CFA13" collectionCode="UTM, CML" location="western La Gomera" pageId="23" pageNumber="2200" specimenCode="RBS7011, 28, RBS7508, RBS7609, MT 0136" specimenCount="15" typeStatus="paratype">
<typeStatus id="1706883F4560793241868051FE86FABC" box="[186,297,1296,1320]" pageId="23" pageNumber="2200" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="DEBBFD1445607932400E804EFE10FAB3" box="[306,447,1295,1319]" pageId="23" pageNumber="2200" type="generic">15 specimens</specimenCount>
and 26 shells, collected between 1981 and 2009, from
<location id="CD6260464560793242C18051FEB9FAD3" LSID="urn:lsid:plazi:treatment:4014878B4560793C40A1817EFD0BFEAB:CD6260464560793242C18051FEB9FAD3" name="western La Gomera" pageId="23" pageNumber="2200">western La Gomera</location>
(
<collectionCode id="AEACAE58456079324017806EFEC2FAD3" box="[299,365,1327,1351]" pageId="23" pageNumber="2200">UTM</collectionCode>
coordinates: 28
<specimenCode id="981B9EE645607932431A806EFD20FAD3" box="[550,655,1327,1351]" collectionCode="RBS" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="23" pageNumber="2200" type="Herbarium">RBS7011</specimenCode>
,
<specimenCode id="981B9EE645607932439C806EFD15FAD3" box="[672,698,1327,1351]" collectionCode="RBS" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="23" pageNumber="2200" type="Herbarium">28</specimenCode>
RBS7016, 28RBS7113, 28RBS7115, 28RBS7116, 28RBS7212, 28RBS7214, 28RBS7319, 28RBS7405, 28
<specimenCode id="981B9EE6456079324291800EFBBBFAF3" box="[941,1044,1359,1383]" collectionCode="RBS" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="23" pageNumber="2200" type="Herbarium">RBS7508</specimenCode>
and 28
<specimenCode id="981B9EE6456079324196802EFEBBFA13" box="[170,276,1391,1415]" collectionCode="RBS" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="23" pageNumber="2200" type="Herbarium">RBS7609</specimenCode>
), deposited in TFMC (
<specimenCode id="981B9EE6456079324337802EFDD9FA13" box="[523,630,1391,1415]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="23" pageNumber="2200">MT 0136</specimenCode>
/1),
<collectionCode id="AEACAE584560793243A5802EFD78FA13" box="[665,727,1391,1415]" country="0" httpUri="http://grbio.org/cool/ib7f-dbfe" name="Universidad Nacional de Tucuman, Coleccion de Mamiferos Lillo (Argentina)" pageId="23" pageNumber="2200">CML</collectionCode>
(six shells) and AIT
</materialsCitation>
.
</paragraph>
<paragraph id="C802369D4560793241AC80F4FEA6FA59" blockId="23.[144,265,1461,1485]" box="[144,265,1461,1485]" pageId="23" pageNumber="2200">
<emphasis id="FAC9EA8F4560793241AC80F4FEA6FA59" box="[144,265,1461,1485]" italics="true" pageId="23" pageNumber="2200">Etymology</emphasis>
</paragraph>
<paragraph id="C802369D4560793241AC809CFC2BFA61" blockId="23.[144,900,1501,1525]" box="[144,900,1501,1525]" pageId="23" pageNumber="2200">The specific name derives from the distinctive shell ornamentation.</paragraph>
<paragraph id="C802369D4560793241AC8364FBFFF9F0" blockId="23.[144,470,1573,1595]" lastBlockId="23.[144,1104,1609,1701]" pageId="23" pageNumber="2200">
<emphasis id="FAC9EA8F4560793241AC8364FE79F9A9" box="[144,470,1573,1597]" italics="true" pageId="23" pageNumber="2200">Geographic range and habitat</emphasis>
Ground dweller endemic to La Gomera, with a distribution area of about
<quantity id="0F459B784560793242E6830CFB8FF9F1" box="[986,1056,1613,1637]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="2200" unit="km" value="60.0">60 km</quantity>
2 on
</paragraph>
<paragraph id="C802369D4560793241AC832CFB8FF931" blockId="23.[144,1104,1609,1701]" pageId="23" pageNumber="2200">
the western side of the island, mainly related to lowland vegetation and semi-arid subtropical scrub and palm grove, ranging in altitude from
<quantity id="0F459B784560793243C483CCFC3BF930" box="[760,916,1677,1701]" metricMagnitude="2" metricUnit="m" metricValue="5.875" metricValueMax="10.0" metricValueMin="1.75" pageId="23" pageNumber="2200" unit="m" value="587.5" valueMax="1000.0" valueMin="175.0">175 to 1000 m</quantity>
(
<figureCitation id="50862A1845607932429D83CCFBBCF931" box="[929,1043,1677,1701]" captionStart="Figure 1" captionStartId="11.[144,211,1524,1546]" captionTargetBox="[144,1104,173,1488]" captionTargetId="figure-84@11.[144,1104,173,1488]" captionTargetPageId="11" captionText="Figure 1. (A) Geographical location of the Azores, Madeira and the Canaries; arrows indicate possible colonization patterns used by land snails; position of sea mounts comes from Van den Broeck et al. 2008; (B) geographical distribution of the Napaeus species; symbols represent 1×1 km Universal Transverse Mercator squares; arrow points to the Pleistocene deposit from Mancha de La Laja-Bajamar, where the oldest Napaeus species (N. lajaensis) has been collected (Castillo et al., 2006)." figureDoi="http://doi.org/10.5281/zenodo.5217089" httpUri="https://zenodo.org/record/5217089/files/figure.png" pageId="23" pageNumber="2200">Figure 1B</figureCitation>
).
</paragraph>
<paragraph id="C802369D456F793D41E885E6FEEEFF2B" blockId="24.[212,321,167,191]" box="[212,321,167,191]" pageId="24" pageNumber="2201">
<emphasis id="FAC9EA8F456F793D41E885E6FEEEFF2B" box="[212,321,167,191]" italics="true" pageId="24" pageNumber="2201">Diagnosis</emphasis>
</paragraph>
<paragraph id="C802369D456F793D41E8858EFB24FED3" blockId="24.[211,1173,207,327]" pageId="24" pageNumber="2201">Shell elongated ovatecylindrical, cone-shaped above. Protoconch very fragile. Teleoconch heavily ornamented with strong, laminar, prominent and irregularly undulating radial oblique ribs. Shell colour tan brown, with first whorls slightly darker than the rest. Epiphallus without epiphallar caecum. Bursa copulatrix duct without diverticulum.</paragraph>
<paragraph id="C802369D456F793D41E8843EFEFBFE03" blockId="24.[212,340,383,407]" box="[212,340,383,407]" pageId="24" pageNumber="2201">
<emphasis id="FAC9EA8F456F793D41E8843EFEFBFE03" box="[212,340,383,407]" italics="true" pageId="24" pageNumber="2201">Description</emphasis>
</paragraph>
<paragraph id="C802369D456F793D41E884E6FBF0FD4B" blockId="24.[212,1172,423,1247]" pageId="24" pageNumber="2201">
Shell (
<figureCitation id="50862A18456F793D402684E6FE16FE2B" box="[282,441,423,447]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (AD) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="24" pageNumber="2201">Figures 7C, E</figureCitation>
) obese (SB/SH index), elongated ovatecylindrical, cone-shaped above, 77½ convex whorls, deeply marked suture. Body whorl intermediate (BH/SH index), protoconch smooth, shiny, with 1½1¾ whorls. Protoconch very fragile, broken in many shells and living specimens. Aperture short, wide (AH/SH and AB/SB indices, respectively), with elliptical section at palatal side, rounded at union of columellar and palatal edges. Upper palatal side forms angle slightly less than 90º with parietal side, aperture juts out only slightly from body whorl beginning. Whitish discontinuous peristome expanded as lip, more developed in lower part of palatal edge, reflected in columellar edge, where it partially covers umbilical slit. Older specimens with callosity between peristome edges, nodule at union of parietalpalatal area.
</paragraph>
<paragraph id="C802369D456F793D41C287A6FDCBFCCB" blockId="24.[212,1172,423,1247]" pageId="24" pageNumber="2201">
Shell colour dark brown, first teleoconch whorls slightly darker than rest. Heavily ornamented teleoconch characterized by strong, laminar, prominent, irregularly undulating, sometimes interrupted radial oblique ribs (
<figureCitation id="50862A18456F793D427B8666FC46FCAB" box="[839,1001,807,831]" captionStart="Figure 7" captionStartId="17.[144,211,1168,1190]" captionTargetBox="[144,1104,168,1139]" captionTargetId="figure-197@17.[144,1104,168,1139]" captionTargetPageId="17" captionText="Figure 7. Holotype shells (AD) and a specimen (E). (A) N. bajamarensis sp. nov.; (B) N. tafadaensis sp. nov.; (C) N. ornamentatus sp. nov.; (D) N. teobaldoi sp. nov.; (E) N. ornamentatus, paratype from Mirador de Taguluche (L. Moro photograph)." figureDoi="http://doi.org/10.5281/zenodo.5217105" httpUri="https://zenodo.org/record/5217105/files/figure.png" pageId="24" pageNumber="2201">Figures 7C, E</figureCitation>
,
<figureCitation id="50862A18456F793D42CA8666FB84FCAB" box="[1014,1067,807,831]" captionStart="Figure 10" captionStartId="20.[212,279,1133,1155]" captionTargetBox="[212,1172,168,1103]" captionTargetId="figure-150@20.[212,1172,167,1103]" captionTargetPageId="20" captionText="Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A) N. bajamarensis sp. nov.; (BC) N. tafadaensis sp. nov.; (D) N. ornamentatus sp. nov.; (E) N. teobaldoi sp. nov.; (F) shell ornamentation detail of the penultimate whorl of N. badiosus, from Barranco de Valle Seco (Tenerife); pt, peristome." figureDoi="http://doi.org/10.5281/zenodo.5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="24" pageNumber="2201">10D</figureCitation>
). Ribs of body whorl more regularly located.
</paragraph>
<paragraph id="C802369D456F793D41C28626FD5CFB4B" blockId="24.[212,1172,423,1247]" pageId="24" pageNumber="2201">
Genital system (
<figureCitation id="50862A18456F793D40858626FD91FCEB" box="[441,574,871,895]" captionStart="Figure 11" captionStartId="21.[144,211,1117,1139]" captionTargetBox="[144,1104,168,1087]" captionTargetId="figure-219@21.[144,1104,168,1087]" captionTargetPageId="21" captionText="Figure 11. Genital system. (A) N. tafadaensis sp. nov., paratype from Montaña Tafada; (B) N. ornamentatus sp. nov., paratype from Mirador de Taguluche; (C) N. teobaldoi sp. nov., paratype from Barranco Hondo. A1, A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; di, diaphragm; dp, distal penis; e, epiphallus; ec, epiphallar caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; pp, proximal penis; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5217117" httpUri="https://zenodo.org/record/5217117/files/figure.png" pageId="24" pageNumber="2201">Figure 11B</figureCitation>
;
<specimenCount id="DEBBFD14456F793D43778626FD40FCEB" box="[587,751,871,895]" pageId="24" pageNumber="2201" type="generic">five specimens</specimenCount>
dissected). Atrium very short. Penis mainly tubular, slightly longer than A
<subScript id="543934D8456F793D43B486D3FD3DFC30" attach="left" box="[648,658,914,932]" fontSize="8" pageId="24" pageNumber="2201">1</subScript>
portion of penial appendix. Penis with three portions, distal portion located between atrium and insertion of the penial retractor muscle; proximal portion small, slightly widened, with incipient penial papilla; intermediate portion slender, tubular, slightly longer than distal portion. Epiphallus long, tubular, without epiphallar caecum, opening distally on penis. Flagellum short. Vas deferens opens laterally on proximal end of epiphallus. Penial appendix arises near distal end of penis, is longer than penis and epiphallus together. A
<subScript id="543934D8456F793D42F08113FC79FBF0" attach="left" box="[972,982,1106,1124]" fontSize="8" pageId="24" pageNumber="2201">1</subScript>
penial appendix portion similar in length to penis. Appendicular retractor muscle inserted laterally, at certain distance of proximal top of A
<subScript id="543934D8456F793D43B781D3FD3AFB30" attach="left" box="[651,661,1170,1188]" fontSize="8" pageId="24" pageNumber="2201">1</subScript>
portion; insertion on diaphragm walls very close, but not fused to penial retractor. Free oviduct slightly longer than vagina. Bursa copulatrix complex without diverticulum.
</paragraph>
<paragraph id="C802369D456F793D41E88059FE98FAA4" blockId="24.[212,311,1304,1328]" box="[212,311,1304,1328]" pageId="24" pageNumber="2201">
<emphasis id="FAC9EA8F456F793D41E88059FE98FAA4" box="[212,311,1304,1328]" italics="true" pageId="24" pageNumber="2201">Remarks</emphasis>
</paragraph>
<paragraph id="C802369D456F793D41E88001FD56F903" blockId="24.[212,1172,1343,1688]" pageId="24" pageNumber="2201">
<emphasis id="FAC9EA8F456F793D41E88001FE32FACC" box="[212,413,1344,1368]" italics="true" pageId="24" pageNumber="2201">Shell morphology.</emphasis>
Within the conchological group of
<taxonomicName id="0FBD4D1E456F793D42598001FC45FACC" baseAuthorityName="Webb and Berthelot" baseAuthorityYear="1833" box="[869,1002,1344,1368]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="variatus">
<emphasis id="FAC9EA8F456F793D42598001FC45FACC" box="[869,1002,1344,1368]" italics="true" pageId="24" pageNumber="2201">N. variatus</emphasis>
</taxonomicName>
, the shell of
<taxonomicName id="0FBD4D1E456F793D41E88021FE23FAEC" authority="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez, 2009" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[212,396,1376,1400]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus" status="sp. nov.">
<emphasis id="FAC9EA8F456F793D41E88021FE23FAEC" box="[212,396,1376,1400]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1FA57F4456F793D40A48021FE59FAE3" box="[408,502,1376,1399]" pageId="24" pageNumber="2201" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="50862A18456F793D4335801EFD2EFAE3" box="[521,641,1375,1399]" captionStart="Figure 4" captionStartId="14.[212,279,1195,1217]" captionTargetBox="[220,1162,176,1153]" captionTargetId="figure-214@14.[212,1172,168,1165]" captionTargetPageId="14" captionText="Figure 4. Napaeus shell drawings. (A) N. tenoensis (holotype, AIT); (B) N. bajamarensis sp. nov. (holotype, TFMC MT 0397); (C) N. boucheti (holotype, AIT); (D) N. obesatus; (E) N. propinquus; (F) N. ornamentatus sp. nov. (holotype, TFMC MT 0400); (G) N. orientalis (holotype, AIT); (H) N. ocellatus (syntype, ZMZ 513767); (J) N. rupicola; (K) N. avaloensis (holotype, TFMC MT 0382); (L) N. maculatus (holotype, TFMC MT 0386); (M) N. helvolus; (N) N. voggenreiteri (holotype, TFMC MT 0384); (O) N. isletae (holotype, AIT); (P) N. variatus; (Q) N. encaustus; (R) N. moquinianus; (S) N. teobaldoi sp. nov. (holotype, TFMC MT 0399); (T) N. lajaensis (holotype, TFMC MT 0381); (U) N. procerus (holotype, TFMC MT 0385); (V) N. rufobrunneus." figureDoi="http://doi.org/10.5281/zenodo.5217097" httpUri="https://zenodo.org/record/5217097/files/figure.png" pageId="24" pageNumber="2201">Figure 4F</figureCitation>
) is quite similar in form and size to that of
<taxonomicName id="0FBD4D1E456F793D41E880C1FEC7FA0C" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[212,360,1408,1432]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456F793D41E880C1FEC7FA0C" box="[212,360,1408,1432]" italics="true" pageId="24" pageNumber="2201">N. propinquus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D4049803EFE4BFA03" box="[373,484,1407,1431]" captionStart="Figure 4" captionStartId="14.[212,279,1195,1217]" captionTargetBox="[220,1162,176,1153]" captionTargetId="figure-214@14.[212,1172,168,1165]" captionTargetPageId="14" captionText="Figure 4. Napaeus shell drawings. (A) N. tenoensis (holotype, AIT); (B) N. bajamarensis sp. nov. (holotype, TFMC MT 0397); (C) N. boucheti (holotype, AIT); (D) N. obesatus; (E) N. propinquus; (F) N. ornamentatus sp. nov. (holotype, TFMC MT 0400); (G) N. orientalis (holotype, AIT); (H) N. ocellatus (syntype, ZMZ 513767); (J) N. rupicola; (K) N. avaloensis (holotype, TFMC MT 0382); (L) N. maculatus (holotype, TFMC MT 0386); (M) N. helvolus; (N) N. voggenreiteri (holotype, TFMC MT 0384); (O) N. isletae (holotype, AIT); (P) N. variatus; (Q) N. encaustus; (R) N. moquinianus; (S) N. teobaldoi sp. nov. (holotype, TFMC MT 0399); (T) N. lajaensis (holotype, TFMC MT 0381); (U) N. procerus (holotype, TFMC MT 0385); (V) N. rufobrunneus." figureDoi="http://doi.org/10.5281/zenodo.5217097" httpUri="https://zenodo.org/record/5217097/files/figure.png" pageId="24" pageNumber="2201">Figure 4E</figureCitation>
), and also comparable with that of
<taxonomicName id="0FBD4D1E456F793D426380C1FBA6FA0C" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[863,1033,1408,1432]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="maffioteanus">
<emphasis id="FAC9EA8F456F793D426380C1FBA6FA0C" box="[863,1033,1408,1432]" italics="true" pageId="24" pageNumber="2201">N. maffioteanus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D4529803EFB28FA03" box="[1045,1159,1407,1431]" captionStart="Figure 5" captionStartId="15.[144,211,920,942]" captionTargetBox="[152,1091,172,885]" captionTargetId="figure-303@15.[144,1104,168,891]" captionTargetPageId="15" captionText="Figure 5. Napaeus shell drawings. (A) N. maffioteanus; (B) N. chrysaloides; (C) N. indifferens; (D) N. huttereri (holotype, AIT); (E) N. elegans (holotype, AIT); (F) N. esbeltus (holotype, AIT); (G) N. exilis (holotype, AIT); (H) N. subgracilior; (J) N. osoriensis; (K) N. palmaensis (syntype, NHM 1895.2.2.219); (L) N. myosotis; (M) N. lichenicola (holotype, TFMC MT 389); (N) N. gruereanus; (O) N. barquini (holotype, TFMC MT 0383); (P) N. roccellicola; (Q) N. beguirae (holotype, AIT); (R) N. nanodes; (S) N. doliolum (paratype, AIT); (T) N. bechi (holotype, AIT); (U) N. texturatus; (V) N. taguluchensis (holotype, AIT); (X) N. tagamichensis (paratype, AIT); (Y) N. pygmaeus (holotype, AIT)." figureDoi="http://doi.org/10.5281/zenodo.5217101" httpUri="https://zenodo.org/record/5217101/files/figure.png" pageId="24" pageNumber="2201">Figure 5A</figureCitation>
), although the latter species is more cylindrical. The first whorls of
<taxonomicName id="0FBD4D1E456F793D42E480E1FB3CFA2C" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[984,1171,1440,1464]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456F793D42E480E1FB3CFA2C" box="[984,1171,1440,1464]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="0FBD4D1E456F793D403B8081FE0BFA4C" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[263,420,1472,1496]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456F793D403B8081FE0BFA4C" box="[263,420,1472,1496]" italics="true" pageId="24" pageNumber="2201">N. propinquus</emphasis>
</taxonomicName>
are smaller than those of
<taxonomicName id="0FBD4D1E456F793D43EE8081FC29FA4C" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[722,902,1472,1496]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="maffioteanus">
<emphasis id="FAC9EA8F456F793D43EE8081FC29FA4C" box="[722,902,1472,1496]" italics="true" pageId="24" pageNumber="2201">N. maffioteanus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D42A980FEFBA1FA43" box="[917,1038,1471,1495]" captionStart="Figure 6" captionStartId="16.[212,279,1357,1379]" captionTargetBox="[214,1168,170,1326]" captionTargetId="figure-114@16.[212,1172,168,1327]" captionTargetPageId="16" captionText="Figure 6. Scatter plots of some shell measurements for the new species and the nearest species of their respective conchological group. AH, aperture height; AS, aperture surface (plane view); FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface (plane view); SB, shell breadth; SH, shell height." figureDoi="http://doi.org/10.5281/zenodo.5217103" httpUri="https://zenodo.org/record/5217103/files/figure.png" pageId="24" pageNumber="2201">Figure 6G</figureCitation>
). The body whorl and the aperture of
<taxonomicName id="0FBD4D1E456F793D433B80A1FD6DFA6C" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[519,706,1504,1528]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456F793D433B80A1FD6DFA6C" box="[519,706,1504,1528]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="0FBD4D1E456F793D43C280A1FC1DFA6C" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[766,946,1504,1528]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="maffioteanus">
<emphasis id="FAC9EA8F456F793D43C280A1FC1DFA6C" box="[766,946,1504,1528]" italics="true" pageId="24" pageNumber="2201">N. maffioteanus</emphasis>
</taxonomicName>
are similar in form and size but smaller than those of
<taxonomicName id="0FBD4D1E456F793D43568341FCA7F98C" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[618,776,1536,1560]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456F793D43568341FCA7F98C" box="[618,776,1536,1560]" italics="true" pageId="24" pageNumber="2201">N. propinquus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D422580BEFC39F983" box="[793,918,1535,1559]" captionStart="Figure 6" captionStartId="16.[212,279,1357,1379]" captionTargetBox="[214,1168,170,1326]" captionTargetId="figure-114@16.[212,1172,168,1327]" captionTargetPageId="16" captionText="Figure 6. Scatter plots of some shell measurements for the new species and the nearest species of their respective conchological group. AH, aperture height; AS, aperture surface (plane view); FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface (plane view); SB, shell breadth; SH, shell height." figureDoi="http://doi.org/10.5281/zenodo.5217103" httpUri="https://zenodo.org/record/5217103/files/figure.png" pageId="24" pageNumber="2201">Figure 6H</figureCitation>
). The main difference between
<taxonomicName id="0FBD4D1E456F793D40078361FE58F9AC" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[315,503,1568,1592]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456F793D40078361FE58F9AC" box="[315,503,1568,1592]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
and the other two species is in the shell ornamentation, which is smooth and not strongly developed in
<taxonomicName id="0FBD4D1E456F793D423E8301FC18F9CC" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[770,951,1600,1624]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="maffioteanus">
<emphasis id="FAC9EA8F456F793D423E8301FC18F9CC" box="[770,951,1600,1624]" italics="true" pageId="24" pageNumber="2201">N. maffioteanus</emphasis>
</taxonomicName>
and
<taxonomicName id="0FBD4D1E456F793D42C98301FB3BF9CC" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[1013,1172,1600,1624]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456F793D42C98301FB3BF9CC" box="[1013,1172,1600,1624]" italics="true" pageId="24" pageNumber="2201">N. propinquus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D41E7831EFE6DF9E3" box="[219,450,1631,1655]" captionStart="Figure 12" captionStartId="22.[212,279,1181,1203]" captionTargetBox="[212,1172,168,1151]" captionTargetId="figure-208@22.[212,1172,167,1151]" captionTargetPageId="22" captionText="Figure 12. Shell ornamentation details of the penultimate whorl (AC, E, F) and that of the two last whorls (D). (A) N. tenoensis holotype, from Tamargo (Tenerife); (B) N. boucheti holotype, from Llano del Lance (La Palma); (C) N. obesatus, from La Milagrosa (Gran Canaria); (D) N. tabidus, from La Laguna (Tenerife); (E) N. maffioteanus, from Barranco de La Aldea (Gran Canaria); (F) N. propinquus, from Barranco de Las Galgas (Tenerife)." figureDoi="http://doi.org/10.5281/zenodo.5217119" httpUri="https://zenodo.org/record/5217119/files/figure.png" pageId="24" pageNumber="2201">Figures 12E and 12F</figureCitation>
, respectively), in contrast to that of
<taxonomicName id="0FBD4D1E456F793D427B8321FC56F9EC" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[839,1017,1632,1656]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456F793D427B8321FC56F9EC" box="[839,1017,1632,1656]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
(
<figureCitation id="50862A18456F793D453A831EFB27F9E3" box="[1030,1160,1631,1655]" captionStart="Figure 10" captionStartId="20.[212,279,1133,1155]" captionTargetBox="[212,1172,168,1103]" captionTargetId="figure-150@20.[212,1172,167,1103]" captionTargetPageId="20" captionText="Figure 10. Holotype shell ornamentation details of the penultimate whorl (A, B, D, E) and that of the body whorl columellar zone, near the peristome (C). (A) N. bajamarensis sp. nov.; (BC) N. tafadaensis sp. nov.; (D) N. ornamentatus sp. nov.; (E) N. teobaldoi sp. nov.; (F) shell ornamentation detail of the penultimate whorl of N. badiosus, from Barranco de Valle Seco (Tenerife); pt, peristome." figureDoi="http://doi.org/10.5281/zenodo.5217111" httpUri="https://zenodo.org/record/5217111/files/figure.png" pageId="24" pageNumber="2201">Figure 10D</figureCitation>
). Besides,
<taxonomicName id="0FBD4D1E456F793D400983C1FE41F90C" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[309,494,1664,1688]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="2201" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456F793D400983C1FE41F90C" box="[309,494,1664,1688]" italics="true" pageId="24" pageNumber="2201">N. ornamentatus</emphasis>
</taxonomicName>
has a high protoconch.
</paragraph>
<paragraph id="C802369D456E793C41AC85E6FD0BFEAB" blockId="25.[144,1104,167,319]" pageId="25" pageNumber="2202">
<emphasis id="FAC9EA8F456E793C41AC85E6FE0EFF2B" box="[144,417,167,191]" italics="true" pageId="25" pageNumber="2202">Genital system anatomy.</emphasis>
The genital systems of
<taxonomicName id="0FBD4D1E456E793C438A85E6FCDEFF2B" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[694,881,167,191]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456E793C438A85E6FCDEFF2B" box="[694,881,167,191]" italics="true" pageId="25" pageNumber="2202">N. ornamentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="0FBD4D1E456E793C428D85E6FBFFFF2B" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[945,1104,167,191]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456E793C428D85E6FBFFFF2B" box="[945,1104,167,191]" italics="true" pageId="25" pageNumber="2202">N. propinquus</emphasis>
</taxonomicName>
are similar in shape and dimensions but
<taxonomicName id="0FBD4D1E456E793C43658586FD5AFF4B" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[601,757,199,223]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="propinquus">
<emphasis id="FAC9EA8F456E793C43658586FD5AFF4B" box="[601,757,199,223]" italics="true" pageId="25" pageNumber="2202">N. propinquus</emphasis>
</taxonomicName>
has an epiphallar caecum that is absent in
<emphasis id="FAC9EA8F456E793C402985A6FE79FF6B" box="[277,470,231,255]" italics="true" pageId="25" pageNumber="2202">
<taxonomicName id="0FBD4D1E456E793C402985A6FE7DFF6B" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[277,466,231,255]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="ornamentatus">N. ornamentatus</taxonomicName>
.
</emphasis>
The genital system of
<taxonomicName id="0FBD4D1E456E793C43E885A6FC29FF6B" baseAuthorityName="Mousson" baseAuthorityYear="1872" box="[724,902,231,255]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="maffioteanus">
<emphasis id="FAC9EA8F456E793C43E885A6FC29FF6B" box="[724,902,231,255]" italics="true" pageId="25" pageNumber="2202">N. maffioteanus</emphasis>
</taxonomicName>
has an epiphallar caecum and also a well-developed diverticulum in the bursa copulatrix complex, whereas
<taxonomicName id="0FBD4D1E456E793C41CD8466FE05FEAB" authorityName="Yanes &amp; Martín &amp; Moro &amp; Alonso &amp; Ibáñez" authorityYear="2009" box="[241,426,295,319]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="2202" phylum="Mollusca" rank="species" species="ornamentatus">
<emphasis id="FAC9EA8F456E793C41CD8466FE05FEAB" box="[241,426,295,319]" italics="true" pageId="25" pageNumber="2202">N. ornamentatus</emphasis>
</taxonomicName>
lacks both structures.
</paragraph>
</subSubSection>
</treatment>
</document>