treatments-xml/data/2A/2B/87/2A2B87933E12BF0055947B44FEB2E19A.xml
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<document id="59A4BFF7804EF6108109FC82F4792B08" ID-DOI="10.11646/zootaxa.2531.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5302621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629683319273" checkinUser="felipe" docAuthor="Yang, Zhuo &amp; Zhou, Hong-Zhang" docDate="2010" docId="2A2B87933E12BF0055947B44FEB2E19A" docLanguage="en" docName="zt02531p014.pdf" docOrigin="Zootaxa 2531 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2531.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Naddia nanlingensis Yang &amp; Zhou 2010, sp. n." docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="D612FFEB3E10BF0455037949FFE9E300" masterDocTitle="Three new species of Naddia Fauvel (Coleoptera: Staphylinidae) from China" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1699095747488" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CF1DE6B0B2B91E676C2835ADC4975013">Three new species of Naddia Fauvel (Coleoptera: Staphylinidae) from China</mods:title>
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<mods:name id="EA7CDD27A0797098EE7D4D959AD1E15B" type="personal">
<mods:role id="6F5A210F0071EAD756A20053DD4CF14B">
<mods:roleTerm id="5AC7D271575FBF0C2A2E643163C5BBB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED44A4586D6B38F2DFE0AEC6AF9B064E">Yang, Zhuo</mods:namePart>
<mods:affiliation id="9BD9D025D4E4EDBC338638B479340687">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chao Yang District, 100101 Beijing, P. R. China &amp; Graduate University of the Chinese Academy of Sciences, 19 Yuquan Road, Shijingshan District, 100039 Beijing, P. R. China Corresponding author. E-mail: zhouhz @ ioz. ac. cn</mods:affiliation>
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<mods:namePart id="28FC51C876D04BEDFEF896928187344F">Zhou, Hong-Zhang</mods:namePart>
<mods:affiliation id="441E9D44B879D6EEDD72A04F5C40A2FB">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chao Yang District, 100101 Beijing, P. R. China</mods:affiliation>
<mods:nameIdentifier id="D5079DB7209ABFDD2A5E49E03386A060" type="email">zhouhz@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="7D4A02E25E60CE667E3C2B58F92DEB05">Zootaxa</mods:title>
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<mods:date id="63C8CC676EFBDC4C9240EF770AEDC0D2">2010</mods:date>
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<mods:number id="4892941807B938B114B8C31ACEC8E3DE">2010-07-09</mods:number>
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<mods:number id="4BBF3D5C465EA92E65157AAD59D11F3B">2531</mods:number>
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<mods:identifier id="C6964C01205B2E765C6DE43EAD908CB8" type="DOI">10.11646/zootaxa.2531.1.1</mods:identifier>
<mods:identifier id="0BC9031DCC052EA8ED8C50229466FF9A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6801160250CE33B0130EABEF9CC087D1" type="Zenodo-Dep">5302621</mods:identifier>
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<treatment id="2A2B87933E12BF0055947B44FEB2E19A" ID-DOI="http://doi.org/10.5281/zenodo.5309660" ID-Zenodo-Dep="5309660" LSID="urn:lsid:plazi:treatment:2A2B87933E12BF0055947B44FEB2E19A" httpUri="http://treatment.plazi.org/id/2A2B87933E12BF0055947B44FEB2E19A" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="EA98650E3E12BF0655947B44FD46E127" box="[151,687,525,551]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="A23D36853E12BF0655947B44FD46E127" blockId="2.[151,687,525,586]" box="[151,687,525,551]" pageId="2" pageNumber="3">
<heading id="F97581E93E12BF0655947B44FD46E127" bold="true" box="[151,687,525,551]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="90F6EA973E12BF0655947B44FD46E127" bold="true" box="[151,687,525,551]" pageId="2" pageNumber="3">
1.
<taxonomicName id="65824D063E12BF0655B77B44FDB6E127" ID-CoL="45JWJ" authority="Yang &amp; Zhou" authorityName="Yang &amp; Zhou" authorityYear="2010" box="[180,607,525,551]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="90F6EA973E12BF0655B77B44FE47E127" bold="true" box="[180,430,525,551]" italics="true" pageId="2" pageNumber="3">Naddia nanlingensis</emphasis>
Yang &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="8BC557EC3E12BF06576E7B44FD46E127" box="[621,687,525,551]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EA98650E3E12BF0655947B79FEDAE14A" box="[151,307,560,586]" pageId="2" pageNumber="3" type="description">
<paragraph id="A23D36853E12BF0655947B79FEDAE14A" blockId="2.[151,687,525,586]" box="[151,307,560,586]" pageId="2" pageNumber="3">
(
<figureCitation id="3AB92A003E12BF06559C7B79FEC3E14A" box="[159,298,560,586]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Figs. 1AK</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EA98650E3E12BF0655947B33FDD6E02A" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="A23D36853E12BF0655947B33FDD6E02A" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">
<emphasis id="90F6EA973E12BF0655947B33FE89E194" bold="true" box="[151,352,634,660]" pageId="2" pageNumber="3">Type material.</emphasis>
<materialsCitation id="12EA3CD83E12BF0654697B33FC40E1BA" collectingDate="2008-07-03" collectingDateMax="2008-07-09" collectingDateMin="2008-07-03" collectionCode="IZCAS" country="China" elevation="1500" latitude="24.89881" location="Xiaohuang Mountain" longLatPrecision="1" longitude="113.01946" municipality="Nanling" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="90F6EA973E12BF0654697B33FCCAE194" bold="true" box="[362,803,634,660]" pageId="2" pageNumber="3">
<typeStatus id="7D3988273E12BF0654697B33FE02E194" box="[362,491,634,660]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="DA9576153E12BF0654FF7B33FD80E194" box="[508,617,634,660]" name="China" pageId="2" pageNumber="3">CHINA</collectingCountry>
:
<collectingRegion id="6046F8673E12BF0657797B33FCF7E194" box="[634,798,634,660]" country="China" name="Guangdong" pageId="2" pageNumber="3">Guangdong</collectingRegion>
:
</emphasis>
<specimenCount id="B484FD0C3E12BF0656337B33FCADE18E" box="[816,836,634,654]" pageId="2" pageNumber="3" type="male"></specimenCount>
,
<collectingMunicipality id="4259ACFF3E12BF0656567B33FC29E194" box="[853,960,634,660]" pageId="2" pageNumber="3">Nanling</collectingMunicipality>
,
<location id="A75D605E3E12BF0656CC7B33FB0BE194" LSID="urn:lsid:plazi:treatment:2A2B87933E12BF0055947B44FEB2E19A:A75D605E3E12BF0656CC7B33FB0BE194" box="[975,1250,634,660]" country="China" latitude="24.89881" longLatPrecision="1" longitude="113.01946" municipality="Nanling" name="Xiaohuang Mountain" pageId="2" pageNumber="3" stateProvince="Guangdong">Xiaohuang Mountain</location>
[
<geoCoordinate id="C7B650423E12BF0651F77B33FA7EE194" box="[1268,1431,634,660]" degrees="24.89881" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="24.89881">N24.89881°</geoCoordinate>
,
<geoCoordinate id="C7B650423E12BF0655947BE9FEDFE1BA" box="[151,310,672,698]" degrees="113.01946" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="113.01946">E113.01946°</geoCoordinate>
],
<quantity id="657A9B603E12BF0654497BE9FE4EE1BA" box="[330,423,672,698]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="1500.0">
<elevation id="29AFD1B63E12BF0654497BE9FE4EE1BA" box="[330,423,672,698]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="D63C10453E12BF0654B77BE9FDBCE1BA" box="[436,597,672,698]" pageId="2" pageNumber="3" value="2008-07-03" valueMax="2008-07-09" valueMin="2008-07-03">
<collectingDate id="C678E9AD3E12BF0654B77BE9FDBCE1BA" box="[436,597,672,698]" pageId="2" pageNumber="3" value="2008-07-03" valueMax="2008-07-09" valueMin="2008-07-03">39.VII.2008</collectingDate>
</date>
, Lei Gao collected (
<collectionCode id="C493AE403E12BF0656467BE9FC48E1BA" box="[837,929,672,698]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="3">IZCAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="12EA3CD83E12BF0656B67BD6FDE6E1DF" collectingDate="2008-07-03" collectingDateMax="2008-07-09" collectingDateMin="2008-07-03" collectionCode="IZCAS" country="China" elevation="1500" latitude="24.89881" location="Xiaohuang Mountain" longLatPrecision="1" longitude="113.01946" municipality="Nanling" pageId="2" pageNumber="3" specimenCount="2" specimenCount-female="2" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="90F6EA973E12BF0656B67BD6FABFE1B9" bold="true" box="[949,1366,671,697]" pageId="2" pageNumber="3">
<typeStatus id="7D3988273E12BF0656B67BD6FBD1E1B9" box="[949,1080,671,697]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="DA9576153E12BF0651467BD6FB42E1B9" box="[1093,1195,671,697]" name="China" pageId="2" pageNumber="3">CHINA</collectingCountry>
:
<collectingRegion id="6046F8673E12BF0651BA7BD6FAB8E1B9" box="[1209,1361,671,697]" country="China" name="Guangdong" pageId="2" pageNumber="3">Guangdong</collectingRegion>
:
</emphasis>
<specimenCount id="B484FD0C3E12BF06505D7BE9FA7DE1B3" box="[1374,1428,671,698]" pageId="2" pageNumber="3" type="female">2♀♀</specimenCount>
, same data as holotype (
<collectionCode id="C493AE403E12BF0654AF7B8CFDEEE1DF" box="[428,519,709,735]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="3">IZCAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="12EA3CD83E12BF06571E7B8DFE11E004" collectingDate="2008-07-03" collectingDateMax="2008-07-09" collectingDateMin="2008-07-03" collectionCode="IZCAS" country="China" elevation="1500" latitude="24.5547" longLatPrecision="7" longitude="112.5737" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong" typeStatus="paratype">
<specimenCount id="B484FD0C3E12BF06571E7B8DFDD8E1D8" box="[541,561,708,728]" pageId="2" pageNumber="3" type="female"></specimenCount>
, Nanling, Shikengkong [
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,
<geoCoordinate id="C7B650423E12BF0656EA7B8CFB90E1DF" box="[1001,1145,709,735]" degrees="112.5737" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="5" value="112.5737">E112.5737°</geoCoordinate>
],
<quantity id="657A9B603E12BF06518E7B8CFB00E1DF" box="[1165,1257,709,735]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="1500.0">
<elevation id="29AFD1B63E12BF06518E7B8CFB00E1DF" box="[1165,1257,709,735]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="D63C10453E12BF0651F57B8CFA7EE1DF" box="[1270,1431,709,735]" pageId="2" pageNumber="3" value="2008-07-03" valueMax="2008-07-09" valueMin="2008-07-03">
<collectingDate id="C678E9AD3E12BF0651F57B8CFA7EE1DF" box="[1270,1431,709,735]" pageId="2" pageNumber="3" value="2008-07-03" valueMax="2008-07-09" valueMin="2008-07-03">39.VII.2008</collectingDate>
</date>
, Lei Gao collected (
<collectionCode id="C493AE403E12BF06548E7BA3FE07E004" box="[397,494,746,772]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="3">IZCAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="12EA3CD83E12BF06570A7BA3FDD2E02A" collectingDate="2008-07-19" collectionCode="IZCAS" country="China" elevation="1030" latitude="24.92579" longLatPrecision="1" longitude="113.01638" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong" typeStatus="paratype">
<specimenCount id="B484FD0C3E12BF06570A7BA3FDF4E1FE" box="[521,541,746,766]" pageId="2" pageNumber="3" type="female"></specimenCount>
, Nanling, Ruyang [
<geoCoordinate id="C7B650423E12BF0656147BA3FC5EE004" box="[791,951,746,772]" degrees="24.92579" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="24.92579">N24.92579°</geoCoordinate>
,
<geoCoordinate id="C7B650423E12BF0656C57BA3FB98E004" box="[966,1137,746,772]" degrees="113.01638" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="113.01638">E113.01638°</geoCoordinate>
],
<quantity id="657A9B603E12BF0651847BA3FB00E004" box="[1159,1257,746,772]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="3" unit="m" value="1030.0">
<elevation id="29AFD1B63E12BF0651847BA3FB00E004" box="[1159,1257,746,772]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="3" unit="m" value="1030.0">1030 m</elevation>
</quantity>
,
<date id="D63C10453E12BF0651FA7BA3FA7EE004" box="[1273,1431,746,772]" pageId="2" pageNumber="3" value="2008-07-19">
<collectingDate id="C678E9AD3E12BF0651FA7BA3FA7EE004" box="[1273,1431,746,772]" pageId="2" pageNumber="3" value="2008-07-19">19.VII.2008</collectingDate>
</date>
, Rongrong Wang collected (
<collectionCode id="C493AE403E12BF0654DB7A59FDDAE02A" box="[472,563,784,810]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="3">IZCAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA98650E3E12BF0055C57A7CFF18E2BA" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="A23D36853E12BF0655C57A7CFD5EE04F" blockId="2.[151,1436,634,1982]" box="[198,695,821,847]" pageId="2" pageNumber="3">
<emphasis id="90F6EA973E12BF0655C57A7CFEB7E04F" bold="true" box="[198,350,821,847]" pageId="2" pageNumber="3">Description.</emphasis>
Body length:
<quantity id="657A9B603E12BF0657067A7CFD5AE04F" box="[517,691,821,847]" metricMagnitude="-2" metricUnit="m" metricValue="2.025" metricValueMax="2.09" metricValueMin="1.96" pageId="2" pageNumber="3" unit="mm" value="20.25" valueMax="20.9" valueMin="19.6">19.620.9 mm</quantity>
.
</paragraph>
<paragraph id="A23D36853E12BF0655C57A13FE3EE09A" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">Female measurements (mean, n = 4): HL: 3.54; HW: 3.82; CL: 0.98; PO: 2.40; PL: 3.75; PW: 3.38; EL: 3.35; EW: 3.74; ELS: 1.57.</paragraph>
<paragraph id="A23D36853E12BF0655C57AECFEB1E0E4" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">Male measurements (n = 1): HL: 3.43; HW: 3.53; CL: 0.98; PO: 2.25; PL: 3.35; PW: 3.08; EL: 3.23; EW: 3.32; ELS: 1.67.</paragraph>
<paragraph id="A23D36853E12BF0655C57ABBFC05E75A" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">
Male. Body elongate, black, dull, pubescence on sides of 35
<superScript id="55F79BCD3E12BF0656D67AB9FC08E0FE" attach="left" box="[981,993,1008,1022]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
abdominal tergites and base of 7
<superScript id="55F79BCD3E12BF06508C7AB9FA72E0FE" attach="none" box="[1423,1435,1008,1022]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
abdominal tergite silver-yellow, pubescence of 6
<superScript id="55F79BCD3E12BF0657CF7D51FD31E726" attach="none" box="[716,728,1048,1062]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
, 89
<superScript id="55F79BCD3E12BF0656117D51FCF7E726" attach="left" box="[786,798,1048,1062]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
and remaining part of 7
<superScript id="55F79BCD3E12BF0651367D51FBA8E726" attach="left" box="[1077,1089,1048,1062]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergite black. Maxillary palpi brown, apex of last segment slightly paler, tibiae yellow and tarsi brown.
</paragraph>
<paragraph id="A23D36853E12BF0655C57D2FFDDBE660" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">
Head (
<figureCitation id="3AB92A003E12BF0654187D2FFE94E780" box="[283,381,1126,1152]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
) large, subquadrate, not widened backward, sides almost straight; eyes small, oval, distinctly protruding, but not from lateral outline of head, longitudinal diameter much shorter than postocular area; base broadly and shallowly emarginate, posterior angles obviously protruding backward with apices rounded; surface densely covered with umbilicate punctures, with short yellow pubescence. Antennae (
<figureCitation id="3AB92A003E12BF06506F7D9FFF53E615" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Fig. 1G</figureCitation>
) short and stout, hardly reaching midlength of pronotum, moderately thickened and more opaque apicad, last segment asymmetrical, sharply pointed; ratios of antennomere length (1 to 11) as follows: 20:17:15:14:15:16:13:15:22:13:30.
</paragraph>
<paragraph id="A23D36853E12BF0655C57C22FECFE540" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">Pronotum nearly horseshoe-shaped and convex, longer than wide, slightly narrower than head, widest at anterior margin, anterior margin raised medially, sides subparallel, gradually narrowed toward base; anterior angles pointed, posterior angles widely rounded; surface densely and regularly covered with umbilicate punctures, coarser anteriorly; with shiny impunctate midline, becoming indistinct and almost obsolete at about midlength; pubescence similar to that on head. Scutellum triangular, densely covered with black velvety pubescence.</paragraph>
<paragraph id="A23D36853E12BF0655C57F02FC6FE5B0" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">Elytra short, nearly quadrate, slightly dilated posteriad, wider than long, a little shorter and broader than pronotum; sides straight, posterior margin slightly emarginate medially; surface feebly and densely punctate, covered with dense black and sparse golden yellow pubescence.</paragraph>
<paragraph id="A23D36853E12BF0655C57FF7FB4BE498" blockId="2.[151,1436,634,1982]" pageId="2" pageNumber="3">
Abdomen elongate and gradually widened posteriad; surface finely and densely punctate, 35
<superScript id="55F79BCD3E12BF06502C7FF2FAD2E5C9" attach="left" box="[1327,1339,1723,1737]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergites with black pubescence in middle and patches of yellow pubescence laterally, patches gradually decreasing in size towards apex, 7
<superScript id="55F79BCD3E12BF0654847E41FE7AE416" attach="left" box="[391,403,1800,1814]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergite also with transverse patch of golden pubescence at base; posterior margin of 8
<superScript id="55F79BCD3E12BF06508C7E41FA72E416" attach="none" box="[1423,1435,1800,1814]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
sternite (
<figureCitation id="3AB92A003E12BF0655FD7E78FEB1E44B" box="[254,344,1841,1867]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Fig. 1B</figureCitation>
) broadly and deeply emarginate medially, surface of tergite bearing moderately long golden pubescence. 10
<superScript id="55F79BCD3E12BF0654517E1FFEB6E464" attach="left" box="[338,351,1878,1892]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergite (
<figureCitation id="3AB92A003E12BF0654C47E10FDCDE473" box="[455,548,1881,1907]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Fig. 1C</figureCitation>
) with golden pubescence in basal half, 9
<superScript id="55F79BCD3E12BF06511E7E1FFBC3E464" attach="left" box="[1053,1066,1878,1892]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
sternite (
<figureCitation id="3AB92A003E12BF06519C7E10FB12E473" box="[1183,1275,1881,1907]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="2" pageNumber="3">Fig. 1E</figureCitation>
) triangularly excised in middle of posterior margin and with pyramid-shaped deep depression at base.
</paragraph>
<paragraph id="A23D36853E12BF0655C57EEDFA7BE4BE" blockId="2.[151,1436,634,1982]" box="[198,1426,1956,1982]" pageId="2" pageNumber="3">Legs robust, first four protarsal segments markedly dilated, sub-bilobed, each covered with golden setae.</paragraph>
<caption id="F6FD660D3E13BF0755947E32FF14E4F5" ID-DOI="http://doi.org/10.5281/zenodo.5302623" ID-Zenodo-Dep="5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,269,1915,1939]" targetBox="[159,1436,201,1908]" targetPageId="3">
<paragraph id="A23D36853E13BF0755947E32FF14E4F5" blockId="3.[151,1437,1915,2037]" pageId="3" pageNumber="4">
<emphasis id="90F6EA973E13BF0755947E32FEB4E493" bold="true" box="[151,349,1915,1939]" pageId="3" pageNumber="4">FIGURES 1AK.</emphasis>
<taxonomicName id="65824D063E13BF0754607E35FD3FE493" authority="Yang &amp; Zhou" authorityName="Yang &amp; Zhou" authorityYear="2010" box="[355,726,1916,1939]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="90F6EA973E13BF0754607E35FDD6E493" box="[355,575,1916,1939]" italics="true" pageId="3" pageNumber="4">Naddia nanlingensis</emphasis>
Yang &amp; Zhou
</taxonomicName>
<emphasis id="90F6EA973E13BF0757DF7E32FCF1E493" bold="true" box="[732,792,1915,1939]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="8BC557EC3E13BF0757DF7E32FCF1E493" box="[732,792,1915,1939]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8
<superScript id="55F79BCD3E13BF0754437ED5FEA2E4A9" attach="left" box="[320,331,1948,1961]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
sternite; CD—10
<superScript id="55F79BCD3E13BF07571B7ED5FDCAE4A9" attach="left" box="[536,547,1948,1961]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
tergite; E—9
<superScript id="55F79BCD3E13BF0757B67ED5FD29E4A9" attach="left" box="[693,704,1948,1961]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm.
</paragraph>
</caption>
<paragraph id="A23D36853E14BF0055C579D1FEAAE222" blockId="4.[151,1436,152,666]" pageId="4" pageNumber="5">
Aedeagus (
<figureCitation id="3AB92A003E14BF00544D79D1FE37E3B2" box="[334,478,152,178]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="4" pageNumber="5">Figs. 1HK</figureCitation>
) symmetrical; median lobe elongate, with weakly sclerotized middle part and globular basal part, in ventral view median lobe feebly narrowed towards round apex, in lateral view moderately curved ventrad; paramere (
<figureCitation id="3AB92A003E14BF00577979ABFD33E3FC" box="[634,730,226,252]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="4" pageNumber="5">Fig. 1H</figureCitation>
) degenerate, very fine and slender, with four short setae medioapically.
</paragraph>
<paragraph id="A23D36853E14BF0055C57864FF18E2BA" blockId="4.[151,1436,152,666]" pageId="4" pageNumber="5">
Female. First four protarsal segments similar to those of male. Second gonocoxite (
<figureCitation id="3AB92A003E14BF0051A77864FB12E247" box="[1188,1275,301,327]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="4" pageNumber="5">Fig. 1F</figureCitation>
) with minute stylus bearing two equally long apical setae, basal half of second gonocoxite with long golden setae. 10
<superScript id="55F79BCD3E14BF00508C781BFA75E260" attach="right" box="[1423,1436,338,352]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
tergite (
<figureCitation id="3AB92A003E14BF0055F17833FEA4E294" box="[242,333,378,404]" captionStart="FIGURES 1" captionStartId="3.[151,269,1915,1939]" captionTargetBox="[159,1436,201,1908]" captionTargetId="figure-72@3.[151,1436,197,1908]" captionTargetPageId="3" captionText="FIGURES 1AK. Naddia nanlingensis Yang &amp; Zhou sp. n., male (AC, E, GK, holotype) and female (D, F, paratype). A—head; B—8th sternite; CD—10th tergite; E—9th sternite; F—detail of genital segment with second gonocoxite; G— antennae; H—detail of paramere; I—aedeagus, ventral view; J—aedeagus, lateral view; K—aedeagus, dorsal view. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.5302623" httpUri="https://zenodo.org/record/5302623/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
) triangular, moderately wide, apex broadly arcuate, with long golden setae at and near apical margin.
</paragraph>
</subSubSection>
<subSubSection id="EA98650E3E14BF0055C5788CFD0BE14F" pageId="4" pageNumber="5" type="distribution">
<paragraph id="A23D36853E14BF0055C5788CFDB6E2DF" blockId="4.[151,1436,152,666]" box="[198,607,453,479]" pageId="4" pageNumber="5">
<emphasis id="90F6EA973E14BF0055C5788CFE8EE2DF" bold="true" box="[198,359,453,479]" pageId="4" pageNumber="5">Distribution.</emphasis>
<collectingCountry id="DA9576153E14BF00546E788CFE5AE2DF" box="[365,435,453,479]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
(
<collectingRegion id="6046F8673E14BF0054C0788CFDBAE2DF" box="[451,595,453,479]" country="China" name="Guangdong" pageId="4" pageNumber="5">Guangdong</collectingRegion>
).
</paragraph>
<paragraph id="A23D36853E14BF0055C578A3FD0BE14F" blockId="4.[151,1436,152,666]" pageId="4" pageNumber="5">
<emphasis id="90F6EA973E14BF0055C578A3FEFAE104" bold="true" box="[198,275,490,516]" pageId="4" pageNumber="5">Notes.</emphasis>
In general appearance, the present new species is similar to
<taxonomicName id="65824D063E14BF00510378A2FA72E104" authority="Shibata, 1979" authorityName="Shibata" authorityYear="1979" box="[1024,1435,490,516]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="90F6EA973E14BF00510378A2FB05E104" box="[1024,1260,491,516]" italics="true" pageId="4" pageNumber="5">Naddia taiwanensis</emphasis>
<bibRefCitation id="C6134B743E14BF0051F478A3FA72E104" author="Shibata, Y." box="[1271,1435,490,516]" pageId="4" pageNumber="5" pagination="19 - 29" refId="ref7385" refString="Shibata, Y. (1979) New or little-known Staphylinidae (Coleoptera). The Entomological Review of Japan, 33, 19 - 29." type="journal article" year="1979">Shibata, 1979</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="DA9576153E14BF0055D97B59FEDCE12A" box="[218,309,528,554]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, but the latter has distinctly divergent posterior areas of head, orange-red pubescence on the elytra and shiny impunctate midline on pronotum.
</paragraph>
</subSubSection>
<subSubSection id="EA98650E3E14BF0055C57B13FEB2E19A" pageId="4" pageNumber="5" type="etymology">
<paragraph id="A23D36853E14BF0055C57B13FEB2E19A" blockId="4.[151,1436,152,666]" pageId="4" pageNumber="5">
<emphasis id="90F6EA973E14BF0055C57B13FEBAE174" bold="true" box="[198,339,602,628]" pageId="4" pageNumber="5">Etymology.</emphasis>
The species is named after the
<typeStatus id="7D3988273E14BF0057C57B13FD11E174" box="[710,760,602,628]" pageId="4" pageNumber="5">type</typeStatus>
locality, Nanling National Nature Reserve in
<collectingRegion id="6046F8673E14BF0050107B13FEEBE19A" country="China" name="Guangdong" pageId="4" pageNumber="5">Guangdong Province</collectingRegion>
,
<collectingCountry id="DA9576153E14BF00540D7BC9FEB1E19A" box="[270,344,640,666]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>