329 lines
37 KiB
XML
329 lines
37 KiB
XML
<document id="1B96A3E595DC7FF49A7241ED50333BB6" ID-CLB-Dataset="5001" ID-DOI="10.11646/zootaxa.4984.1.10" ID-GBIF-Dataset="327074ac-0102-495e-92f7-d89c9441ef2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4926688" ID-ZooBank="A10DA1E6-FAA6-4DD9-9A58-CD6C1CD7ECCA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623423665944" checkinUser="plazi" docAuthor="Shear, William A." docDate="2021" docId="03FCF141AD2AFFACFF0CFEDAFA10F9AF" docLanguage="en" docName="zootaxa.4984.1.10.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caseyidae Verhoeff 1909" docType="treatment" docVersion="4" lastPageNumber="110" masterDocId="FFC58939AD2BFFAEFF9BFF86FF8DFFF6" masterDocTitle="Additions to the millipede family Caseyidae. II. Martenseya, a new genus of miniature, blind millipedes from California (Diplopoda, Chordeumatida Striariidea, Striarioidea, Caseyidae)" masterLastPageNumber="113" masterPageNumber="108" pageNumber="109" updateTime="1699010113355" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5848BC812AFE035DD6054AB71E47B9D5">Additions to the millipede family Caseyidae. II. Martenseya, a new genus of miniature, blind millipedes from California (Diplopoda, Chordeumatida Striariidea, Striarioidea, Caseyidae)</mods:title>
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<mods:namePart id="E6F71A24B7270D65AEE07DED17F6FC21">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="CC2558997252056174E0DE1FEA11FE66">text</mods:typeOfResource>
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<mods:title id="0C8D78B44D363ACC955B96191D5EA528">Zootaxa</mods:title>
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<mods:date id="8FBA434C14B477D5305984EE2478271F">2021</mods:date>
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<mods:number id="9A176149AA0ACBF5E0CAA46BBA76EB12">2021-06-10</mods:number>
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<mods:number id="6F84A2BD66938C4BB98B69F3CE63BE18">4984</mods:number>
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<treatment id="03FCF141AD2AFFACFF0CFEDAFA10F9AF" ID-DOI="http://doi.org/10.5281/zenodo.4926690" ID-GBIF-Taxon="184605100" ID-Zenodo-Dep="4926690" LSID="urn:lsid:plazi:treatment:03FCF141AD2AFFACFF0CFEDAFA10F9AF" httpUri="http://treatment.plazi.org/id/03FCF141AD2AFFACFF0CFEDAFA10F9AF" lastPageId="2" lastPageNumber="110" pageId="1" pageNumber="109">
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<subSubSection id="C34F13DCAD2AFFAFFF0CFEDAFDBFFE80" box="[151,562,348,374]" pageId="1" pageNumber="109" type="nomenclature">
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<paragraph id="8BEA4057AD2AFFAFFF0CFEDAFDBFFE80" blockId="1.[151,562,348,374]" box="[151,562,348,374]" pageId="1" pageNumber="109">
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<heading id="D0A2F73BAD2AFFAFFF0CFEDAFDBFFE80" bold="true" box="[151,562,348,374]" fontSize="11" level="1" pageId="1" pageNumber="109" reason="1">
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<emphasis id="B9219C45AD2AFFAFFF0CFEDAFDBFFE80" bold="true" box="[151,562,348,374]" pageId="1" pageNumber="109">
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Family
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<taxonomicName id="4C553BD4AD2AFFAFFF6CFEDAFDBFFE80" ID-CoL="7NGD9" authority="Verhoeff, 1909" authorityName="Verhoeff" authorityYear="1909" box="[247,562,348,374]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">
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Caseyidae
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<bibRefCitation id="EFC43DA6AD2AFFAFFEE7FEDAFDBFFE80" author="Verhoeff, K. W." box="[380,562,348,374]" pageId="1" pageNumber="109" pagination="566 - 572" refId="ref5320" refString="Verhoeff, K. W. (1909) Neues System der Diplopoda - Ascospermophora. Zoologische Anzeiger, 34, 566 - 572." type="journal article" year="1909">Verhoeff, 1909</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F13DCAD2AFFAFFF0CFE26FC75FE21" pageId="1" pageNumber="109" type="reference_group">
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<paragraph id="8BEA4057AD2AFFAFFF0CFE26FC52FE4E" blockId="1.[151,1016,416,471]" box="[151,991,416,441]" pageId="1" pageNumber="109">
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<treatmentCitationGroup id="AB456779AD2AFFAFFF0CFE26FC52FE4E" box="[151,991,416,441]" pageId="1" pageNumber="109">
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<treatmentCitation id="0AF46646AD2AFFAFFF0CFE26FE40FE4E" author="Verhoeff, K. W." box="[151,461,416,441]" page="567" pageId="1" pageNumber="109" year="1909">
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<taxonomicName id="4C553BD4AD2AFFAFFF0CFE26FE40FE4E" ID-CoL="7NGD9" authority="Verhoeff 1909: 567" authorityName="Verhoeff" authorityPageNumber="567" authorityYear="1909" box="[151,461,416,441]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">
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Caseyidae
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<bibRefCitation id="EFC43DA6AD2AFFAFFE9EFE26FE40FE4E" author="Verhoeff, K. W." box="[261,461,416,441]" pageId="1" pageNumber="109" pagination="566 - 572" refId="ref5320" refString="Verhoeff, K. W. (1909) Neues System der Diplopoda - Ascospermophora. Zoologische Anzeiger, 34, 566 - 572." type="journal article" year="1909">Verhoeff 1909: 567</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Garner & Shelley 1989: 185 (complete references).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEA4057AD2AFFAFFF0CFE39FC75FE21" blockId="1.[151,1016,416,471]" box="[151,1016,447,471]" pageId="1" pageNumber="109">
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<treatmentCitationGroup id="AB456779AD2AFFAFFF0CFE39FC75FE21" box="[151,1016,447,471]" pageId="1" pageNumber="109">
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<treatmentCitation id="0AF46646AD2AFFAFFF0CFE39FD80FE21" author="Verhoeff, K. W." box="[151,525,447,471]" page="568" pageId="1" pageNumber="109" year="1909">
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<taxonomicName id="4C553BD4AD2AFFAFFF0CFE39FD80FE21" authority="Verhoeff 1909: 568" authorityName="Verhoeff" authorityPageNumber="568" authorityYear="1909" box="[151,525,447,471]" class="Diplopoda" family="Underwoodiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">
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Underwoodiidae
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<bibRefCitation id="EFC43DA6AD2AFFAFFEDDFE39FD80FE21" author="Verhoeff, K. W." box="[326,525,447,471]" pageId="1" pageNumber="109" pagination="566 - 572" refId="ref5320" refString="Verhoeff, K. W. (1909) Neues System der Diplopoda - Ascospermophora. Zoologische Anzeiger, 34, 566 - 572." type="journal article" year="1909">Verhoeff 1909: 568</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="0AF46646AD2AFFAFFD8CFE39FD43FE21" author="Causey, N. B." box="[535,718,447,471]" page="193" pageId="1" pageNumber="109" year="1963">
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<bibRefCitation id="EFC43DA6AD2AFFAFFD8CFE39FD43FE21" author="Causey, N. B." box="[535,718,447,471]" pageId="1" pageNumber="109" pagination="193 - 198" refId="ref4757" refString="Causey, N. B. (1963) Two new caseyid millipeds from California caves (Chordeumida: Chordeumidea). The Wassman Journal of Biology, 21, 193 - 198." type="journal article" year="1963">Causey 1963: 193</bibRefCitation>
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</treatmentCitation>
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(synonymy with
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<taxonomicName id="4C553BD4AD2AFFAFFC1BFE39FC60FE21" ID-CoL="7NGD9" authorityName="Verhoeff" authorityYear="1909" box="[896,1005,447,471]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F13DCAD2AFFACFF0CFD85FDFCFD3A" lastPageId="2" lastPageNumber="110" pageId="1" pageNumber="109" type="discussion">
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<paragraph id="8BEA4057AD2AFFAFFF0CFD85FCBDFD51" blockId="1.[151,1436,514,680]" pageId="1" pageNumber="109">
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<emphasis id="B9219C45AD2AFFAFFF0CFD85FF57FDEB" bold="true" box="[151,218,515,541]" pageId="1" pageNumber="109">Notes</emphasis>
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.
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<taxonomicName id="4C553BD4AD2AFFAFFF72FD85FE2AFDEB" authorityName="Verhoeff" authorityYear="1909" box="[233,423,515,541]" class="Diplopoda" family="Underwoodiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Underwoodiidae</taxonomicName>
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, with only the genus
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<taxonomicName id="4C553BD4AD2AFFAFFD02FD85FBB5FDEB" authority="Cook & Collins, 1895" authorityName="Cook & Collins" authorityYear="1895" box="[665,1080,514,541]" class="Diplopoda" family="Caseyidae" genus="Underwoodia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis id="B9219C45AD2AFFAFFD02FD85FCBEFDEB" box="[665,819,515,541]" italics="true" pageId="1" pageNumber="109">Underwoodia</emphasis>
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<bibRefCitation id="EFC43DA6AD2AFFAFFCA0FD85FBB5FDEB" author="Cook, O. F. & Collins, G. N." box="[827,1080,514,541]" pageId="1" pageNumber="109" pagination="1 - 100" refId="ref4852" refString="Cook, O. F. & Collins, G. N. (1895) The Craspedosomatidae of North America. Annals of the New York Academy of Science, 9, 1 - 100. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55430. x" type="journal article" year="1895">Cook & Collins, 1895</bibRefCitation>
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</taxonomicName>
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, was synonymized with
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<taxonomicName id="4C553BD4AD2AFFAFFAD5FD85FF4BFDC9" authorityName="Verhoeff" authorityYear="1909" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
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by
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<bibRefCitation id="EFC43DA6AD2AFFAFFF76FDA3FE19FDB6" author="Causey, N. B." box="[237,404,549,576]" pageId="1" pageNumber="109" pagination="193 - 198" refId="ref4757" refString="Causey, N. B. (1963) Two new caseyid millipeds from California caves (Chordeumida: Chordeumidea). The Wassman Journal of Biology, 21, 193 - 198." type="journal article" year="1963">Causey (1963)</bibRefCitation>
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. The synonymy was supported by
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<bibRefCitation id="EFC43DA6AD2AFFAFFC94FDA3FBBEFDC9" author="Gardner, M. R. & Shelley, R. M." box="[783,1075,549,575]" pageId="1" pageNumber="109" pagination="177 - 268" refId="ref4908" refString="Gardner, M. R. & Shelley, R. M. (1989) New records, species, and genera of caseyid millipds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan - Pacific Entomologist, 65, 177 - 268." type="journal article" year="1989">Gardner & Shelley (1989)</bibRefCitation>
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.
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<emphasis id="B9219C45AD2AFFAFFBA6FDA3FB53FDC9" box="[1085,1246,549,575]" italics="true" pageId="1" pageNumber="109">
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<taxonomicName id="4C553BD4AD2AFFAFFBA6FDA3FB57FDC9" authorityName="Cook & Collins" authorityYear="1895" box="[1085,1242,549,575]" class="Diplopoda" family="Caseyidae" genus="Underwoodia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">Underwoodia</taxonomicName>
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,
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</emphasis>
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with two species distributed transcontinentally in northern North America and one in the Russian Far East, is the only trans-Beringian genus in the family (
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<bibRefCitation id="EFC43DA6AD2AFFAFFEE1FDEDFD9FFD73" author="Shelley, R. M." box="[378,530,619,645]" pageId="1" pageNumber="109" pagination="168 - 176" refId="ref5180" refString="Shelley, R. M. (1993) The milliped genus Underwoodia (Chordeumatida: Caseyidae). Canadian Journal of Zoology, 71, 168 - 176. https: // doi. org / 10.1139 / z 93 - 023" type="journal article" year="1993">Shelley 1993</bibRefCitation>
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). Species of
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<taxonomicName id="4C553BD4AD2AFFAFFD04FDEDFC9FFD73" authorityName="Verhoeff" authorityYear="1909" box="[671,786,619,645]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
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may be distinguished from other chordeumatidan families in northwestern North America according to the key below.
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</paragraph>
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<paragraph id="8BEA4057AD2AFFAFFF5CF881FDC4F80F" blockId="1.[151,1437,1799,2041]" pageId="1" pageNumber="109">
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The present classification of the
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<taxonomicName id="4C553BD4AD2AFFAFFDB6F881FD2DF8D7" authorityName="Verhoeff" authorityYear="1909" box="[557,672,1799,1825]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
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is not analytical, without subfamilial or tribal taxa established.
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<bibRefCitation id="EFC43DA6AD2AFFAFFAC0F881FE18F8B3" author="Gardner, M. R. & Shelley, R. M." pageId="1" pageNumber="109" pagination="177 - 268" refId="ref4908" refString="Gardner, M. R. & Shelley, R. M. (1989) New records, species, and genera of caseyid millipds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan - Pacific Entomologist, 65, 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
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had little to say about the possible relationships of the genera, excepting that
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<taxonomicName id="4C553BD4AD2AFFAFFB6AF8ADFAE0F8B2" authorityName="Chamberlin" authorityYear="1941" box="[1265,1389,1835,1860]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis id="B9219C45AD2AFFAFFB6AF8ADFAE0F8B2" box="[1265,1389,1835,1860]" italics="true" pageId="1" pageNumber="109">Vasingtona</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C553BD4AD2AFFAFFF0CF8C9FEB6F89F" authorityName="Gardner & Shelley" authorityYear="1989" box="[151,315,1871,1897]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis id="B9219C45AD2AFFAFFF0CF8C9FEB6F89F" box="[151,315,1871,1897]" italics="true" pageId="1" pageNumber="109">Ochrogramma</emphasis>
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</taxonomicName>
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shared two synapomorphies in the males, a ventral lamella extending from the mandible and long gonapophyses on the second coxae. Material available to me includes at least three additional distinctive genera in addition to
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<taxonomicName id="4C553BD4AD2AFFAFFEADF811FE37F846" authority="There" authorityName="Shear" authorityYear="2021" box="[310,442,1943,1968]" class="Diplopoda" family="Caseyidae" genus="Martenseya" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9219C45AD2AFFAFFEADF811FE37F846" box="[310,442,1943,1968]" italics="true" pageId="1" pageNumber="109">Martenseya</emphasis>
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</taxonomicName>
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<emphasis id="B9219C45AD2AFFAFFE59F811FD9CF847" bold="true" box="[450,529,1943,1969]" pageId="1" pageNumber="109">
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<taxonomicNameLabel id="A212213EAD2AFFAFFE59F811FD9CF847" box="[450,529,1943,1969]" pageId="1" pageNumber="109" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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There are also many new species of
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<emphasis id="B9219C45AD2AFFAFFC2EF811FB82F847" box="[949,1039,1943,1969]" italics="true" pageId="1" pageNumber="109">
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<taxonomicName id="4C553BD4AD2AFFAFFC2EF811FB81F847" authorityName="Chamberlin" authorityYear="1951" box="[949,1036,1943,1969]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">Opiona</taxonomicName>
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,
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</emphasis>
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which may turn out to be the most speciose genus in the family. Until this additional diversity is described, it would be unwise to suggest any new taxa between the level of family and genus.
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</paragraph>
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<paragraph id="8BEA4057AD29FFACFF5CFF11FBA1FD97" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="110">
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Species of the family are rich in characters associated with secondary sexual modifications of the males. The second legpair has processes from the coxae which
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<bibRefCitation id="EFC43DA6AD29FFACFD49FF3DFB8AFF23" author="Gardner, M. R. & Shelley, R. M." box="[722,1031,187,213]" pageId="2" pageNumber="110" pagination="177 - 268" refId="ref4908" refString="Gardner, M. R. & Shelley, R. M. (1989) New records, species, and genera of caseyid millipds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan - Pacific Entomologist, 65, 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
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called gonapophyses, which may be long or relatively short. The third legs are often strikingly modified, with long, flattened coxae and reduced telopodites which sometimes articulate laterally near the midpoint of the extended coxae. The gonopods of most species are complex, and the terminology used by
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<bibRefCitation id="EFC43DA6AD29FFACFD16FEA1FC42FEB7" author="Gardner, M. R. & Shelley, R. M." box="[653,975,295,321]" pageId="2" pageNumber="110" pagination="177 - 268" refId="ref4908" refString="Gardner, M. R. & Shelley, R. M. (1989) New records, species, and genera of caseyid millipds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan - Pacific Entomologist, 65, 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
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was reevaluated by
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<bibRefCitation id="EFC43DA6AD29FFACFB59FEA1FF68FE93" author="Shear, W. A. & Leonard, W. P." pageId="2" pageNumber="110" pagination="23 - 34" refId="ref5101" refString="Shear, W. A. & Leonard, W. P. (2007) Additions to the millipede family Caseyidae. I. Caseya richarti, n. sp., and new records of previously described species in the genus Caseya Cook & Collins 1895 (Diplopoda, Chordeumatida, Caseyidae). Zootaxa, 1524 (1), 23 - 34. https: // doi. org / 10.11646 / zootaxa. 1524.1.2" type="journal article" year="2007">Shear and Leonard (2007)</bibRefCitation>
|
|
. For example, the structure they called a telopodite is in fact the colpocoxite, resembling that found in the related family
|
|
<taxonomicName id="4C553BD4AD29FFACFEA1FEE9FE3EFE7F" box="[314,435,367,393]" class="Diplopoda" family="Striariidae" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="family">Striariidae</taxonomicName>
|
|
: a sclerotized structrure derived from the coxal gland of the eighth leg. What they refer to as colpocoxites are in fact angiocoxites, modifications of the rims of the coxal glands. The ninth legs are reduced to a single telopodite podomere that is large and usually button-like, protruding laterally beyond the pleurotergites of the seventh ring and allowing the identification of males as such in the field. Curiously,
|
|
<bibRefCitation id="EFC43DA6AD29FFACFBFDFE5DFA11FE03" author="Gardner, M. R. & Shelley, R. M." box="[1126,1436,475,501]" pageId="2" pageNumber="110" pagination="177 - 268" refId="ref4908" refString="Gardner, M. R. & Shelley, R. M. (1989) New records, species, and genera of caseyid millipds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan - Pacific Entomologist, 65, 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
|
|
never described or even mentioned the highly modified coxae of the tenth legpair, simply stating that the tenth coxae are enlarged and bear an eversible gland. In fact in the species of
|
|
<taxonomicName id="4C553BD4AD29FFACFC1DFDA5FC55FDCB" authorityName="Cook & Collins" authorityYear="1895" box="[902,984,547,573]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFC1DFDA5FC55FDCB" box="[902,984,547,573]" italics="true" pageId="2" pageNumber="110">Caseya</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C553BD4AD29FFACFB89FDA5FBEBFDCB" authorityName="Chamberlin" authorityYear="1951" box="[1042,1126,547,573]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFB89FDA5FBEBFDCB" box="[1042,1126,547,573]" italics="true" pageId="2" pageNumber="110">Opiona</emphasis>
|
|
</taxonomicName>
|
|
I have examined, the tenth coxae have an array of lobes and processes that are definitely of taxonomic value.
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFD35FDFCFD3A" blockId="2.[151,625,690,717]" box="[151,625,690,717]" pageId="2" pageNumber="110">
|
|
<heading id="D0A2F73BAD29FFACFF0CFD35FDFCFD3A" bold="true" box="[151,625,690,717]" fontSize="11" level="1" pageId="2" pageNumber="110" reason="1">
|
|
<emphasis id="B9219C45AD29FFACFF0CFD35FDFCFD3A" bold="true" box="[151,625,690,717]" pageId="2" pageNumber="110">
|
|
Key to genera of the family
|
|
<taxonomicName id="4C553BD4AD29FFACFE69FD34FDFCFD3A" authorityName="Verhoeff" authorityYear="1909" box="[498,625,690,716]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C34F13DCAD29FFACFF0CFD77FA10F9AF" pageId="2" pageNumber="110" type="key">
|
|
<keyStep id="30A158F2AD29FFACFF0CFD77FA11FCD5" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFD77FA11FCF0" blockId="2.[151,1436,753,804]" box="[151,1436,753,775]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFD77FA11FCF0" box="[151,1436,753,775]" pageId="2" pageNumber="110">
|
|
1a. Eyes lacking; usually less than
|
|
<quantity id="4CADEDB2AD29FFACFD95FD77FDDCFCF0" box="[526,593,753,774]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="110" unit="mm" value="10.0">10 mm</quantity>
|
|
long................................................................ 2
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFC88FA11FCD5" blockId="2.[151,1436,753,804]" box="[151,1436,782,804]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFC88FA11FCD5" box="[151,1436,782,804]" pageId="2" pageNumber="110">
|
|
1b. Eyes present; usually
|
|
<quantity id="4CADEDB2AD29FFACFE2FFC88FE7AFCD2" box="[436,503,782,804]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="110" unit="mm" value="10.0">10 mm</quantity>
|
|
long or more................................................................. 3
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFCCFFA11FC40" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFCCFFA11FC8D" blockId="2.[151,1437,841,951]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFCCFFA11FC8D" pageId="2" pageNumber="110">
|
|
2a. Length of mature males
|
|
<quantity id="4CADEDB2AD29FFACFE57FCCFFD97FCA8" box="[460,538,841,862]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="110" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
|
|
; legpair 3 unmodified; gonopods without flagellocoxites; Marin Co.,
|
|
<collectingRegion id="49918EB5AD29FFACFB2DFCCFFA9AFCA9" box="[1206,1303,841,863]" country="United States of America" name="California" pageId="2" pageNumber="110">California</collectingRegion>
|
|
..............................................................................................
|
|
<taxonomicName id="4C553BD4AD29FFACFB79FCE1FADEFC8A" authority="Shear, 2021" authorityName="Shear" authorityYear="2021" box="[1250,1363,871,892]" class="Diplopoda" family="Caseyidae" genus="Martenseya" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B9219C45AD29FFACFB79FCE1FADEFC8A" box="[1250,1363,871,892]" italics="true" pageId="2" pageNumber="110">Martenseya</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B9219C45AD29FFACFAC2FCE0FA11FC8D" bold="true" box="[1369,1436,870,891]" pageId="2" pageNumber="110">
|
|
<taxonomicNameLabel id="A212213EAD29FFACFAC2FCE0FA11FC8D" box="[1369,1436,870,891]" pageId="2" pageNumber="110" rank="genus">n. gen.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFC02FA11FC40" blockId="2.[151,1437,841,951]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFC02FA11FC40" pageId="2" pageNumber="110">
|
|
2b. Length of mature males
|
|
<quantity id="4CADEDB2AD29FFACFE55FC02FDA4FC6F" box="[462,553,900,921]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="110" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 mm</quantity>
|
|
; legpair 3 with slightly elongate coxae, reduced telopodites; gonopods with flagellocoxites; caves in Calaveras Co.,
|
|
<collectingRegion id="49918EB5AD29FFACFE53FC27FDA4FC41" box="[456,553,929,951]" country="United States of America" name="California" pageId="2" pageNumber="110">California</collectingRegion>
|
|
.................................................................
|
|
<emphasis id="B9219C45AD29FFACFAA3FC27FA11FC40" box="[1336,1436,929,950]" italics="true" pageId="2" pageNumber="110">Speyoseya</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFC5AFA11FBBF" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFC5AFA11FBF8" blockId="2.[151,1437,988,1097]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFC5AFA11FBF8" pageId="2" pageNumber="110">3a. Male mandibular stipes with broad, flattened ventral process; gonapophyses of legpair 2 as long or longer than telopodites; gonopods with many tiny, much branched processes......................................................... 4</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFB90FA11FBBF" blockId="2.[151,1437,988,1097]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFB90FA11FBBF" pageId="2" pageNumber="110">3b. Male mandibular stipes unmodified; gonapophyses of legpair 2 shorter than telopodites; gonopods lacking tiny, much branched processes........................................................................................... 5</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFBE8FA11FB57" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFBE8FA10FB72" blockId="2.[151,1437,1134,1186]" box="[151,1437,1134,1156]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFBE8FA10FB72" box="[151,1437,1134,1156]" pageId="2" pageNumber="110">
|
|
4a. Length
|
|
<quantity id="4CADEDB2AD29FFACFEB7FBE8FE1FFB72" box="[300,402,1134,1156]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="2" pageNumber="110" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">18-22 mm</quantity>
|
|
; southwestern
|
|
<collectingRegion id="49918EB5AD29FFACFDBAFBE8FD4AFB72" box="[545,711,1134,1156]" country="Canada" name="British Columbia" pageId="2" pageNumber="110">British Columbia</collectingRegion>
|
|
to northwestern
|
|
<collectingRegion id="49918EB5AD29FFACFCF2FBE8FC3CFB75" box="[873,945,1134,1155]" country="United States of America" name="Oregon" pageId="2" pageNumber="110">Oregon</collectingRegion>
|
|
...............................
|
|
<taxonomicName id="4C553BD4AD29FFACFAA9FBE9FA10FB72" ID-CoL="7Q67B" authorityName="Chamberlin" authorityYear="1941" box="[1330,1437,1135,1156]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFAA9FBE9FA10FB72" box="[1330,1437,1135,1156]" italics="true" pageId="2" pageNumber="110">Vasingtona</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFB0AFA11FB57" blockId="2.[151,1437,1134,1186]" box="[151,1436,1163,1186]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFB0AFA11FB57" box="[151,1436,1163,1186]" pageId="2" pageNumber="110">
|
|
4b. Length less than
|
|
<quantity id="4CADEDB2AD29FFACFE1EFB0DFE46FB57" box="[389,459,1163,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="110" unit="mm" value="15.0">15 mm</quantity>
|
|
; northwestern
|
|
<collectingRegion id="49918EB5AD29FFACFDC3FB0AFD2DFB57" box="[600,672,1164,1185]" country="United States of America" name="Oregon" pageId="2" pageNumber="110">Oregon</collectingRegion>
|
|
to extreme northwestern
|
|
<collectingRegion id="49918EB5AD29FFACFC0DFB0AFC7AFB54" box="[918,1015,1164,1186]" country="United States of America" name="California" pageId="2" pageNumber="110">California</collectingRegion>
|
|
......................
|
|
<taxonomicName id="4C553BD4AD29FFACFA8BFB0AFA11FB57" ID-CoL="7PMVS" authorityName="Gardner & Shelley" authorityYear="1989" box="[1296,1436,1164,1185]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFA8BFB0AFA11FB57" box="[1296,1436,1164,1185]" italics="true" pageId="2" pageNumber="110">Ochrogramma</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFB40FA11FB0F" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFB40FA11FB2A" blockId="2.[151,1436,1222,1274]" box="[151,1436,1222,1244]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFB40FA11FB2A" box="[151,1436,1222,1244]" pageId="2" pageNumber="110">
|
|
5a. Length
|
|
<quantity id="4CADEDB2AD29FFACFEB7FB40FE0EFB2A" box="[300,387,1222,1244]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="110" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 mm</quantity>
|
|
...................................................................................… 6
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFB65FA11FB0F" blockId="2.[151,1436,1222,1274]" box="[151,1436,1251,1274]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFB65FA11FB0F" box="[151,1436,1251,1274]" pageId="2" pageNumber="110">
|
|
5b. Length
|
|
<quantity id="4CADEDB2AD29FFACFEB7FB65FEE2FB0F" box="[300,367,1251,1273]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="110" unit="mm" value="15.0">15 mm</quantity>
|
|
or greater............................................................................... 7
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFA98FA11FA5F" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFA98FA11FAA7" blockId="2.[151,1437,1310,1450]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFA98FA11FAA7" pageId="2" pageNumber="110">
|
|
6a. Legpair 3 coxae much elongated, with apical tuft of setae, telopodites reduced; gonopods relatively simple, consisting of two coxites, flagellocoxite absent;
|
|
<collectingRegion id="49918EB5AD29FFACFD9EFABAFDFAFAA4" box="[517,631,1340,1362]" country="United States of America" name="Washington" pageId="2" pageNumber="110">Washington</collectingRegion>
|
|
Co.,
|
|
<collectingRegion id="49918EB5AD29FFACFD30FABAFD7EFAA7" box="[683,755,1340,1361]" country="United States of America" name="Oregon" pageId="2" pageNumber="110">Oregon</collectingRegion>
|
|
...............................................
|
|
<taxonomicName id="4C553BD4AD29FFACFAAFFABAFA11FAA7" ID-CoL="7PHLY" authorityName="Gardner & Shelley" authorityYear="1989" box="[1332,1436,1340,1361]" class="Diplopoda" family="Caseyidae" genus="Metopiona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFAAFFABAFA11FAA7" box="[1332,1436,1340,1361]" italics="true" pageId="2" pageNumber="110">Metopiona</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CFADFFA11FA5F" blockId="2.[151,1437,1310,1450]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFADFFA11FA5F" pageId="2" pageNumber="110">
|
|
6b. Legpair 3 coxae only slightly elongated, lacking apical tuft of setae, telopodites of nearly normal size; gonopods with flagellocoxites; central to northern
|
|
<collectingRegion id="49918EB5AD29FFACFE61FAF0FD12FA7A" box="[506,671,1398,1420]" country="Canada" name="British Columbia" pageId="2" pageNumber="110">British Columbia</collectingRegion>
|
|
(but also
|
|
<location id="8E8A168CAD29FFACFD67FAF0FC73FA7A" LSID="urn:lsid:plazi:treatment:03FCF141AD2AFFACFF0CFEDAFA10F9AF:8E8A168CAD29FFACFD67FAF0FC73FA7A" box="[764,1022,1398,1420]" country="Canada" name="northern Rocky Mountains" pageId="2" pageNumber="110" stateProvince="British Columbia">northern Rocky Mountains</location>
|
|
from
|
|
<collectingRegion id="49918EB5AD29FFACFBAEFAF0FBF0FA7A" box="[1077,1149,1398,1420]" country="Canada" name="Alberta" pageId="2" pageNumber="110">Alberta</collectingRegion>
|
|
to Utah, southern
|
|
<collectingCountry id="F34200C7AD29FFACFAB7FAF0FAF9FA7A" box="[1324,1396,1398,1420]" name="Canada" pageId="2" pageNumber="110">Canada</collectingCountry>
|
|
and northern United States from Quebec and Vermont west to North Dakota and Manitoba; Russian Far East).....
|
|
<taxonomicName id="4C553BD4AD29FFACFA83FA12FA11FA5F" ID-CoL="7Q5RB" authorityName="Cook & Collins" authorityYear="1895" box="[1304,1436,1428,1449]" class="Diplopoda" family="Caseyidae" genus="Underwoodia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFA83FA12FA11FA5F" box="[1304,1436,1428,1449]" italics="true" pageId="2" pageNumber="110">Underwoodia</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30A158F2AD29FFACFF0CFA49FA10F9AF" pageId="2" pageNumber="110">
|
|
<paragraph id="8BEA4057AD29FFACFF0CFA49FA11F9E8" blockId="2.[151,1437,1486,1626]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CFA49FA11F9E8" pageId="2" pageNumber="110">
|
|
7a. Legpair 3 telopodites much reduced, prefemora enlarged, flattened; legpair 7 coxae without lobes or processes; gonopod coxites with multiple flagelliform branches in addition to flagellocoxites; San Francisco Bay area to Alexander Archipelago,
|
|
<collectingRegion id="49918EB5AD29FFACFF79F98FFEA8F9E9" box="[226,293,1545,1567]" country="United States of America" name="Alaska" pageId="2" pageNumber="110">Alaska</collectingRegion>
|
|
........................................................................................
|
|
<taxonomicName id="4C553BD4AD29FFACFACFF98FFA11F9E8" ID-CoL="7PNGK" authorityName="Chamberlin" authorityYear="1951" box="[1364,1436,1545,1566]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9219C45AD29FFACFACFF98FFA11F9E8" box="[1364,1436,1545,1566]" italics="true" pageId="2" pageNumber="110">Opiona</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BEA4057AD29FFACFF0CF9A0FA10F9AF" blockId="2.[151,1437,1486,1626]" pageId="2" pageNumber="110">
|
|
<keyLead id="30A4E362AD29FFACFF0CF9A0FA10F9AF" pageId="2" pageNumber="110">
|
|
7b. Legpair 3 telopodites not much reduced, prefemora not enlarged; legpair 7 coxae with lobes and/or processes; southern
|
|
<collectingRegion id="49918EB5AD29FFACFAF7F9A0FE96F9AC" country="United States of America" name="California" pageId="2" pageNumber="110">California</collectingRegion>
|
|
(
|
|
<collectingCounty id="628B38DBAD29FFACFEB3F9C2FE47F9AC" box="[296,458,1604,1626]" pageId="2" pageNumber="110">Los Angeles Co.</collectingCounty>
|
|
) and
|
|
<location id="8E8A168CAD29FFACFE64F9C2FDB5F9AC" LSID="urn:lsid:plazi:treatment:03FCF141AD2AFFACFF0CFEDAFA10F9AF:8E8A168CAD29FFACFE64F9C2FDB5F9AC" box="[511,568,1604,1626]" county="Los Angeles Co." name="Sierra" pageId="2" pageNumber="110" stateProvince="California">Sierra</location>
|
|
<collectingRegion id="49918EB5AD29FFACFDA5F9C2FD0AF9AC" box="[574,647,1604,1626]" country="United States of America" name="Nevada" pageId="2" pageNumber="110">Nevada</collectingRegion>
|
|
to northwestern
|
|
<collectingRegion id="49918EB5AD29FFACFCB2F9C2FC16F9AC" box="[809,923,1604,1626]" country="United States of America" name="Washington" pageId="2" pageNumber="110">Washington</collectingRegion>
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....................................
|
|
<taxonomicName id="4C553BD4AD29FFACFACDF9C2FA10F9AF" ID-CoL="7NRZD" authority="Shear, 2021" authorityName="Shear" authorityYear="2021" box="[1366,1437,1604,1625]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
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<emphasis id="B9219C45AD29FFACFACDF9C2FA10F9AF" box="[1366,1437,1604,1625]" italics="true" pageId="2" pageNumber="110">Caseya</emphasis>
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|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
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</treatment>
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</document> |