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<mods:title id="A48F4BC92F9B7D76E8EB8A6075F1E1D7">Taxonomy and distribution of the genus Trichomyrmex Mayr, 1865 (Hymenoptera: Formicidae) in the Arabian Peninsula, with the description of two new species</mods:title>
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<treatment id="03F0F7622C6C974BFD9153D78E1BFC06" ID-DOI="http://doi.org/10.5281/zenodo.6060810" ID-GBIF-Taxon="124955331" ID-Zenodo-Dep="6060810" LSID="urn:lsid:plazi:treatment:03F0F7622C6C974BFD9153D78E1BFC06" httpUri="http://treatment.plazi.org/id/03F0F7622C6C974BFD9153D78E1BFC06" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<paragraph id="8BE646742C6C974AFD9153D78CA2FD04" blockId="18.[569,1019,691,753]" box="[569,1019,691,718]" pageId="18" pageNumber="19">
<heading id="D0AEF1182C6C974AFD9153D78CA2FD04" box="[569,1019,691,718]" centered="true" fontSize="11" level="4" pageId="18" pageNumber="19" reason="4">
<taxonomicName id="4C593DF72C6C974AFD9153D78CA2FD04" authority="Forel, 1902" authorityName="Forel" authorityYear="1902" box="[569,1019,691,718]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6C974AFD9153D78C04FD07" bold="true" box="[569,861,691,718]" italics="true" pageId="18" pageNumber="19">Trichomyrmex lameerei</emphasis>
(
<bibRefCitation id="EFC83B852C6C974AFCC453D78CAAFD04" author="Forel A." box="[876,1011,691,717]" journalOrPublisher="A. Diehl. Annales de la Societe Entomologique de Belgique" pageId="18" pageNumber="19" pagination="147 - 158" part="46" refString="Forel A. 1902 a. Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr. A. Diehl. Annales de la Societe Entomologique de Belgique 46: 147 - 158." title="Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr" type="journal article" year="1902">Forel, 1902</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6C974AFD7553B28C0FFD38" box="[733,854,726,753]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BE646742C6C974AFD7553B28C0FFD38" blockId="18.[569,1019,691,753]" box="[733,854,726,753]" pageId="18" pageNumber="19">
<figureCitation id="13625AF12C6C974AFD7553B28C47FD38" box="[733,798,726,753]" captionStart="Fig. 9" captionStartId="20.[189,231,1962,1988]" captionTargetBox="[433,1155,340,1923]" captionTargetId="figure@20.[433,1155,340,1923]" captionTargetPageId="20" captionText="Fig. 9. Trichomyrmex lameerei (Forel, 1902), lectotype, worker (CASENT 0249871). A. Body in profile. B. Body in dorsal view. C. Head in full-face view (photos: Shannon Hartman)." httpUri="https://zenodo.org/record/193054/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
AC
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6C974AFF15527C8BE6FB8E" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BE646742C6C974AFF15527C8A77FCB0" blockId="18.[189,1399,791,889]" pageId="18" pageNumber="19">
<taxonomicName id="4C593DF72C6C974AFF15527C8DF7FCFA" authority="Forel, 1902a: 150" authorityName="Forel" authorityPageNumber="150" authorityYear="1902" box="[189,686,792,819]" class="Insecta" family="Formicidae" genus="Holcomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6C974AFF15527C8E8AFCFB" box="[189,467,792,818]" italics="true" pageId="18" pageNumber="19">Holcomyrmex lameerei</emphasis>
<bibRefCitation id="EFC83B852C6C974AFE76527C8D2FFCFB" author="Forel A." box="[478,630,792,818]" journalOrPublisher="A. Diehl. Annales de la Societe Entomologique de Belgique" pageId="18" pageNumber="19" pagination="147 - 158" part="46" refString="Forel A. 1902 a. Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr. A. Diehl. Annales de la Societe Entomologique de Belgique 46: 147 - 158." title="Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr" type="journal article" year="1902" yearSuffix="a">Forel, 1902a</bibRefCitation>
: 150
</taxonomicName>
(workers). Algeria. Palearctic, syntype worker [examined], (Lectotype here designated; Hamada between Houderat, Sahara Algeria, A. Forel (CASENT0249871) (MHNG); paralectotype worker with same data as the lectotype (CASENT0249870) [examined].
</paragraph>
<paragraph id="8BE646742C6C974AFF1552C48C45FC73" blockId="18.[189,1215,928,1095]" box="[189,796,928,954]" pageId="18" pageNumber="19">
<taxonomicName id="4C593DF72C6C974AFF1552C48E88FC73" box="[189,465,928,954]" class="Insecta" family="Formicidae" genus="Holcomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6C974AFF1552C48E88FC73" box="[189,465,928,954]" italics="true" pageId="18" pageNumber="19">Holcomyrmex lameerei</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6C974AFE4652C48DCFFC73" author="Santschi F." box="[494,662,928,954]" journalOrPublisher="Revue Suisse de Zoologie" pageId="18" pageNumber="19" pagination="305 - 334" part="15" refString="Santschi F. 1907. Fourmis de Tunisie capturees en 1906. Revue Suisse de Zoologie 15: 305 - 334." title="Fourmis de Tunisie capturees en 1906" type="journal article" year="1907">Santschi 1907</bibRefCitation>
: 327 (q.m.).
</paragraph>
<paragraph id="8BE646742C6C974AFF1552A08DE0FC14" blockId="18.[189,1215,928,1095]" box="[189,697,963,989]" pageId="18" pageNumber="19">
<taxonomicName id="4C593DF72C6C974AFF1552A08E95FC14" ID-CoL="6RP7L" box="[189,460,963,989]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6C974AFF1552A08E95FC14" box="[189,460,963,989]" italics="true" pageId="18" pageNumber="19">Monomorium lameerei</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6C974AFE4252A78D24FC14" author="Emery C." box="[490,637,963,989]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="18" pageNumber="19" pagination="663 - 686" part="1908" refString="Emery C. 1908. Beitrage zur Monographie der Formiciden des palaarktischen Faunengebietes. (Hym.) Teil V. Deutsche Entomologische Zeitschrift 1908: 663 - 686." title="Beitrage zur Monographie der Formiciden des palaarktischen Faunengebietes. (Hym.) Teil V" type="journal article" year="1908">Emery 1908</bibRefCitation>
: 673.
</paragraph>
<paragraph id="8BE646742C6C974AFF1552838C3FFBEA" blockId="18.[189,1215,928,1095]" pageId="18" pageNumber="19">
<taxonomicName id="4C593DF72C6C974AFF1552838DC4FBC9" ID-CoL="6RP7L" box="[189,669,998,1025]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei" subGenus="Equestrimessor">
<emphasis id="B92D9A662C6C974AFF1552838E07FBC9" box="[189,350,999,1024]" italics="true" pageId="18" pageNumber="19">Monomorium</emphasis>
(
<emphasis id="B92D9A662C6C974AFEC652838D7CFBC9" box="[366,549,999,1024]" italics="true" pageId="18" pageNumber="19">Equestrimessor</emphasis>
)
<emphasis id="B92D9A662C6C974AFD9E52828DC4FBC9" box="[566,669,998,1024]" italics="true" pageId="18" pageNumber="19">lameerei</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6C974AFD1352828C39FBC9" author="Santschi F." box="[699,864,998,1024]" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="18" pageNumber="19" pagination="90 - 92" part="1919" refString="Santschi F. 1919. Nouveaux genre et sous-genre de fourmis barbaresques (Hym.). Bulletin de la Societe Entomologique de France 1919: 90 - 92." title="Nouveaux genre et sous-genre de fourmis barbaresques (Hym.)" type="journal article" year="1919">Santschi 1919</bibRefCitation>
: 92.
<taxonomicName id="4C593DF72C6C974AFF15556E8D25FBEA" ID-CoL="6RP7L" box="[189,636,1033,1060]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei" subGenus="Xeromyrmex">
<emphasis id="B92D9A662C6C974AFF15556E8E07FBEA" box="[189,350,1034,1059]" italics="true" pageId="18" pageNumber="19">Monomorium</emphasis>
(
<emphasis id="B92D9A662C6C974AFEC7556E8D5DFBEA" box="[367,516,1034,1059]" italics="true" pageId="18" pageNumber="19">Xeromyrmex</emphasis>
)
<emphasis id="B92D9A662C6C974AFDBD556D8D25FBEA" box="[533,636,1033,1059]" italics="true" pageId="18" pageNumber="19">lameerei</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6C974AFD32556E8C74FBEA" author="Emery C." box="[666,813,1033,1060]" journalOrPublisher="Genera Insectorum" pageId="18" pageNumber="19" pagination="95 - 206" part="174" refString="Emery C. 1922.: Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae [part]. Genera Insectorum 174 B: 95 - 206, Brussels." title="Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae [part]" type="journal article" year="1922">Emery 1922</bibRefCitation>
: 177.
</paragraph>
<paragraph id="8BE646742C6C974AFF1555488BE6FB8E" blockId="18.[189,1215,928,1095]" box="[189,1215,1068,1095]" pageId="18" pageNumber="19">
<taxonomicName id="4C593DF72C6C974AFF1555488E8FFB8F" box="[189,470,1068,1094]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6C974AFF1555488E8FFB8F" box="[189,470,1068,1094]" italics="true" pageId="18" pageNumber="19">Trichomyrmex lameerei</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6C974AFE5C55488CFEFB8E" author="Ward P. S. &amp; Brady S. G. &amp; Fisher B. L. &amp; Schultz T. R." box="[500,935,1068,1095]" journalOrPublisher="Systematic Entomology" pageId="18" pageNumber="19" pagination="61 - 81" part="40" publicationUrl="http://dx.doi.org/10.1111/syen.12090" refString="Ward P. S., Brady S. G., Fisher B. L. &amp; Schultz T. R. 2015. The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology 40: 61 - 81. http: // dx. doi. org / 10.1111 / syen. 12090" title="The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae)" type="journal article" year="2015">Ward, Brady, Fisher &amp; Schultz, 2015</bibRefCitation>
: 16.
<emphasis id="B92D9A662C6C974AFC7355498CB3FB8F" box="[987,1002,1069,1094]" italics="true" pageId="18" pageNumber="19"></emphasis>
<bibRefCitation id="EFC83B852C6C974AFC5955488BDAFB8E" author="Bolton B." box="[1009,1155,1068,1095]" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology" pageId="18" pageNumber="19" pagination="263 - 452" part="54" publicationUrl="http://dx.doi.org/10.5281/zenodo.26850" refString="Bolton B. 1987. A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History), Entomology 54: 263 - 452. http: // dx. doi. org / 10.5281 / zenodo. 26850" title="A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae)" type="journal article" year="1987">Bolton 1987</bibRefCitation>
: 297.
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6C974AFF15550A8B12FAE9" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BE646742C6C974AFF15550A8EF2FB41" blockId="18.[189,427,1134,1160]" box="[189,427,1134,1160]" pageId="18" pageNumber="19">
<heading id="D0AEF1182C6C974AFF15550A8EF2FB41" bold="true" box="[189,427,1134,1160]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B92D9A662C6C974AFF15550A8EF2FB41" bold="true" box="[189,427,1134,1160]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6C974AFF4255F88EE5FB7E" blockId="18.[189,1399,1180,1313]" box="[234,444,1180,1207]" pageId="18" pageNumber="19">
<heading id="D0AEF1182C6C974AFF4255F88EE5FB7E" bold="true" box="[234,444,1180,1207]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="6">
<emphasis id="B92D9A662C6C974AFF4255F88EE5FB7E" bold="true" box="[234,444,1180,1207]" pageId="18" pageNumber="19">Previous records</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6C974AFF1555A48B12FAE9" blockId="18.[189,1399,1180,1313]" pageId="18" pageNumber="19">UNITED ARAB EMIRATES: w, Dibba, 25.62˚ N, 56.27˚ E, alt. 19 m, Mar. 1995, Collingwood leg.; w, Lahbab-al-Madam, 24.97˚ N, 55.77˚ E, alt. 173 m, 12 Mar. 2005, A.V. Harten leg.; w, Sharjah, 25.32˚ N, 55.51˚ E, alt. 24 m, 1117 Oct. 2004, A.V. Harten leg. (all material in KSMA).</paragraph>
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<subSubSection id="C34315FF2C6C974BFF1554238E20FDF0" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8BE646742C6C974AFF1554238E96FAAB" blockId="18.[189,463,1351,1378]" box="[189,463,1351,1378]" pageId="18" pageNumber="19">
<heading id="D0AEF1182C6C974AFF1554238E96FAAB" bold="true" box="[189,463,1351,1378]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B92D9A662C6C974AFF1554238E96FAAB" bold="true" box="[189,463,1351,1378]" pageId="18" pageNumber="19">Description of worker</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6C974AFF1554138A2FFA7A" blockId="18.[189,1398,1398,1495]" pageId="18" pageNumber="19">
<heading id="D0AEF1182C6C974AFF1554138A2FFA7A" allCaps="true" centered="true" fontSize="11" level="3" pageId="18" pageNumber="19" reason="5">MEASUREMENTS. TL 4.064.50; HL 0.951.09; HW 1.001.15; SL 0.530.75; EL 0.160.18; ML 1.251.40; PW 0.600.61; PTL 0.300.31; PTW 0.210.25; PPL 0.170.23; PPW 0.270.30; CI 105106; EI 16;</heading>
</paragraph>
<paragraph id="8BE646742C6C974AFF1554D88DA6FA1F" blockId="18.[189,1398,1398,1495]" box="[189,767,1468,1495]" pageId="18" pageNumber="19">SI 5365 (CASENT0249870, CASENT0249871).</paragraph>
<paragraph id="8BE646742C6C974AFF15549A8A63F997" blockId="18.[189,1399,1534,1631]" pageId="18" pageNumber="19">HEAD. Broader than long, with emarginated posterior margin, rounded corners and feebly convex sides; anterior clypeal margin feebly concave; eyes of moderate size (EL 0.150.16 × HW) with 12 ommatidia in longest row; scapes when laid back from their insertions fail to reach posterior margin of head.</paragraph>
<paragraph id="8BE646742C6C974AFF1557E28E48F90A" blockId="18.[189,1398,1669,1731]" pageId="18" pageNumber="19">MESOSOMA. Metanotal groove impressed; propodeal dorsum 1.5 × longer than propodeal declivity in profile.</paragraph>
<paragraph id="8BE646742C6C974AFF15578F8C67F8EE" blockId="18.[189,1398,1770,1831]" pageId="18" pageNumber="19">PETIOLE. Peduncle long; two small lateral projections at junction between node and peduncle seen in dorsal view; petiolar node high and rounded in profile.</paragraph>
<paragraph id="8BE646742C6C974BFF15562B8CA1FE85" blockId="18.[189,1399,1871,2003]" lastBlockId="19.[189,1398,271,333]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">PILOSITY. Underside of head with abundant long J-shaped hairs forming a distinct psammophore; cephalic surface with sparse short hairs directed inward to midline of head; anterior clypeal margin with abundant long hairs; scapes with sparse appressed pubescence; funiculus with dense appressed pubescence; mesosoma with abundant short hairs, few longer pairs on pronotum and mesonotum; petiole with abundant short hairs; postpetiole with about seven pairs of long hairs directed backward; gaster with few sparse long hairs and abundant appressed pubescence.</paragraph>
<paragraph id="8BE646742C6D974BFF1550108BA4FE1D" blockId="19.[189,1398,372,469]" pageId="19" pageNumber="20">SCULPTURE. Cephalic surface finely and densely longitudinally costulate; mandibles longitudinally rugulose; mesosomal dorsum finely transversely costulate; petiolar node nearly smooth and slightly shining; postpetiolar dorsum irregularly longitudinally rugulose; gaster smooth and shining.</paragraph>
<paragraph id="8BE646742C6D974BFF1550988E20FDF0" blockId="19.[189,1398,507,569]" pageId="19" pageNumber="20">COLOUR. Head, mesosoma, petiole, postpetiole and appendages reddish brown, gaster dark brown or blackish brown.</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974BFF1553048D25FD60" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BE646742C6D974BFF1553048E95FDB3" blockId="19.[189,460,608,634]" box="[189,460,608,634]" pageId="19" pageNumber="20">
<heading id="D0AEF1182C6D974BFF1553048E95FDB3" bold="true" box="[189,460,608,634]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="B92D9A662C6D974BFF1553048E95FDB3" bold="true" box="[189,460,608,634]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6D974BFF1553EB8D25FD60" blockId="19.[189,636,655,681]" box="[189,636,655,681]" pageId="19" pageNumber="20">
Separation is given under
<taxonomicName id="4C593DF72C6D974BFE5953F48D2DFD60" box="[497,628,655,681]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="chobauti">
<emphasis id="B92D9A662C6D974BFE5953F48D2DFD60" box="[497,628,655,681]" italics="true" pageId="19" pageNumber="20">T. chobauti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974BFF1553B48D7BFCF5" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="8BE646742C6D974BFF1553B48D67FD23" blockId="19.[189,574,720,746]" box="[189,574,720,746]" pageId="19" pageNumber="20">
<heading id="D0AEF1182C6D974BFF1553B48D67FD23" bold="true" box="[189,574,720,746]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="B92D9A662C6D974BFF1553B48D67FD23" bold="true" box="[189,574,720,746]" pageId="19" pageNumber="20">Ecological and biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6D974BFF15539B8D7BFCF5" blockId="19.[189,1398,767,828]" pageId="19" pageNumber="20">One paralectotype specimen is mounted with a seed between mandibles which indicates a granivorous feeding habitat of this species.</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974BFF1552078E1BFC06" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BE646742C6D974BFF1552078E0EFCB4" blockId="19.[189,343,867,893]" box="[189,343,867,893]" pageId="19" pageNumber="20">
<heading id="D0AEF1182C6D974BFF1552078E0EFCB4" bold="true" box="[189,343,867,893]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="B92D9A662C6D974BFF1552078E0EFCB4" bold="true" box="[189,343,867,893]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6D974BFF1552F68E1BFC06" blockId="19.[189,1398,914,975]" pageId="19" pageNumber="20">
This species was described from Algeria and recently recorded from United Arab Emirates (
<bibRefCitation id="EFC83B852C6D974BFB7452F68E6FFC06" author="Collingwood C. A. &amp; Agosti D. &amp; Sharaf M. R. &amp; Harten A." journalOrPublisher="Arthropod Fauna of the United Arab Emirates" pageId="19" pageNumber="20" pagination="405 - 474" part="4" refString="Collingwood C. A., Agosti D., Sharaf M. R. &amp; Harten A. 2011. Order Hymenoptera, Family Formicidae. Arthropod Fauna of the United Arab Emirates 4: 405 - 474." title="Order Hymenoptera, Family Formicidae" type="journal article" year="2011">
Collingwood
<emphasis id="B92D9A662C6D974BFF1552D28FA1FC06" box="[189,248,949,975]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>