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<mods:title id="39CFBAD7F272385779F86BC8C2D5BFE2">Lethrus (Scelolethrus) golestanicus (Coleoptera: Geotrupidae: Lethrinae), a new species from Iran</mods:title>
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<treatment id="03EDFB556D18FF96FBF3C5DE1DDEFD96" ID-DOI="http://doi.org/10.5281/zenodo.5588331" ID-GBIF-Taxon="189134881" ID-Zenodo-Dep="5588331" LSID="urn:lsid:plazi:treatment:03EDFB556D18FF96FBF3C5DE1DDEFD96" httpUri="http://treatment.plazi.org/id/03EDFB556D18FF96FBF3C5DE1DDEFD96" lastPageNumber="590" pageId="5" pageNumber="590">
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<paragraph id="8BFB4A436D18FF96FBF3C5DE19FAFF2C" blockId="5.[151,1437,151,623]" pageId="5" pageNumber="590">
<heading id="D0B3FD2F6D18FF96FBF3C5DE19FAFF2C" bold="true" centered="true" fontSize="11" level="1" pageId="5" pageNumber="590" reason="1">
<emphasis id="B93096516D18FF96FBF3C5DE19FAFF2C" bold="true" pageId="5" pageNumber="590">
Identification key to males of
<taxonomicName id="4C4431C06D18FF96F977C5DE1BD2FF4B" ID-CoL="6PSGL" authorityName="Kraatz" authorityYear="1883" box="[531,912,151,178]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="sulcatus" subGenus="Scelolethrus">
<emphasis id="B93096516D18FF96F977C5DE1A30FF4B" bold="true" box="[531,626,151,178]" italics="true" pageId="5" pageNumber="590">Lethrus</emphasis>
(
<emphasis id="B93096516D18FF96F9E0C5DE1B58FF4B" bold="true" box="[644,794,151,178]" italics="true" pageId="5" pageNumber="590">Scelolethrus</emphasis>
)
<emphasis id="B93096516D18FF96F849C5DE1BD2FF4B" bold="true" box="[813,912,151,178]" italics="true" pageId="5" pageNumber="590">sulcatus</emphasis>
</taxonomicName>
species group with conspicuously conical pleurostomal processes
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35E19C86D18FF96FBF3C5951DDEFD96" pageId="5" pageNumber="590" type="key">
<key id="FFD51D726D18FF96FBF3C5951DDEFD96" pageId="5" pageNumber="590">
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1. Ventral mandibular process long (
<emphasis id="B93096516D18FF96F943C5951A7AFF08" box="[551,568,220,241]" italics="true" pageId="5" pageNumber="590">cf</emphasis>
.
<bibRefCitation id="EFD537B26D18FF96F920C5951B2AFF0B" author="Gusakov, A. A. &amp; Klimenko, A. A." box="[580,872,220,242]" pageId="5" pageNumber="590" pagination="3 - 8" refId="ref2795" refString="Gusakov, A. A. &amp; Klimenko, A. A. (2011) Novyy vid roda Lethrus Scopoli, 1777 (podroda Scelolethrus Semenov, 1892) iz Irana (Coleoptera: Scarabaeidae: Geotrupinae). A new species of the genus Lethrus Scopoli, 1777 (subgenus Scelolethrus Semenov, 1892) from Iran (Coleoptera: Scarabaeidae: Geotrupinae). Eversmannia, 25 - 26, 3 - 8. [in Russian, English abstract]" type="journal article" year="2011">Gusakov &amp; Klimenko 2011: 6</bibRefCitation>
, fig. 3 and 7, fig. 5).
<collectingCountry id="F3530AD36D18FF96FF56C5941C1EFF0B" box="[1074,1116,221,242]" name="Iran" pageId="5" pageNumber="590">Iran</collectingCountry>
:
<collectingRegion id="498084A16D18FF96FF01C5951CFDFF0B" box="[1125,1215,220,242]" country="Iran" name="Golestan" pageId="5" pageNumber="590">Golestan</collectingRegion>
..................................................................................
<emphasis id="B93096516D18FF96F88DC5B31BBCFEF6" box="[1001,1022,250,271]" italics="true" pageId="5" pageNumber="590">L.</emphasis>
<taxonomicName id="4C4431C06D18FF96FF67C5B31DDEFEF6" ID-CoL="724F4" authority="Gusakov &amp; Klimenko, 2011" authorityName="Gusakov &amp; Klimenko" authorityYear="2011" box="[1027,1436,249,271]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="percutens" subGenus="Scelolethrus">
(
<emphasis id="B93096516D18FF96FF6FC5B31C55FEF6" box="[1035,1047,250,271]" italics="true" pageId="5" pageNumber="590">S</emphasis>
.)
<emphasis id="B93096516D18FF96FF4FC5B31CC5FEF6" box="[1067,1159,250,271]" italics="true" pageId="5" pageNumber="590">percutens</emphasis>
<bibRefCitation id="EFD537B26D18FF96FFE9C5B01DDEFEF6" author="Gusakov, A. A. &amp; Klimenko, A. A." box="[1165,1436,249,271]" pageId="5" pageNumber="590" pagination="3 - 8" refId="ref2795" refString="Gusakov, A. A. &amp; Klimenko, A. A. (2011) Novyy vid roda Lethrus Scopoli, 1777 (podroda Scelolethrus Semenov, 1892) iz Irana (Coleoptera: Scarabaeidae: Geotrupinae). A new species of the genus Lethrus Scopoli, 1777 (subgenus Scelolethrus Semenov, 1892) from Iran (Coleoptera: Scarabaeidae: Geotrupinae). Eversmannia, 25 - 26, 3 - 8. [in Russian, English abstract]" type="journal article" year="2011">Gusakov &amp; Klimenko, 2011</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BFB4A436D18FF96FBF3C45E1DDEFED5" blockId="5.[151,1437,151,623]" box="[151,1436,279,301]" pageId="5" pageNumber="590">
<keyLead id="30B5E9766D18FF96FBF3C45E1DDEFED5" box="[151,1436,279,301]" pageId="5" pageNumber="590">
Ventral mandibular process short or almost absent (
<figureCitation id="137F56C66D18FF96F9ABC45E1B50FED4" box="[719,786,279,301]" captionStart="FIGURES 47" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 47. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="5" pageNumber="590">Figs. 5</figureCitation>
,
<figureCitation id="137F56C66D18FF96F879C45E1B0FFED4" box="[797,845,279,301]" captionStart="FIGURES 813" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 813. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 810, head, dorsal view; 1113, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="5" pageNumber="590">813</figureCitation>
)............................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B052E66D18FF96FBF3C47D1DDEFE46" pageId="5" pageNumber="590">
<paragraph id="8BFB4A436D18FF96FBF3C47D1DDEFE7D" blockId="5.[151,1437,151,623]" pageId="5" pageNumber="590">
<keyLead id="30B5E9766D18FF96FBF3C47D1DDEFE7D" pageId="5" pageNumber="590">
2. Profemora with distinctly prominent tooth anteroventrally (
<emphasis id="B93096516D18FF96F847C47D1B76FEB0" box="[803,820,308,329]" italics="true" pageId="5" pageNumber="590">cf</emphasis>
.
<bibRefCitation id="EFD537B26D18FF96F825C47D1C67FEB3" author="Kral, D. &amp; Olexa, A." box="[833,1061,308,330]" pageId="5" pageNumber="590" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa 1996: 63</bibRefCitation>
. fig. 37), in less developed males only with slight tubercle. Pronotal punctation double, consisting from coarse and very fine punctures. Inner edge of parameres broadly angulate subapically (
<figureCitation id="137F56C66D18FF96F967C4261A08FE7C" box="[515,586,367,389]" captionStart="FIGURES 1416" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 1416. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="5" pageNumber="590">Fig. 15</figureCitation>
).
<collectingCountry id="F3530AD36D18FF96F938C4261AC4FE7D" box="[604,646,367,388]" name="Iran" pageId="5" pageNumber="590">Iran</collectingCountry>
:
<collectingRegion id="498084A16D18FF96F9EBC4261AA9FE7C" box="[655,747,367,389]" country="Iran" name="Khorasan-e Shomali" pageId="5" pageNumber="590">Khorasan</collectingRegion>
Shamali Prov........................
<emphasis id="B93096516D18FF96FFEBC4261CE1FE7D" box="[1167,1187,367,388]" italics="true" pageId="5" pageNumber="590">L.</emphasis>
<taxonomicName id="4C4431C06D18FF96FFCDC4261DDEFE7D" ID-CoL="6Q4F3" authority="Reitter, 1904" authorityName="Reitter" authorityYear="1904" box="[1193,1436,367,389]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="mithras" subGenus="Scelolethrus">
(
<emphasis id="B93096516D18FF96FFD5C4261CFFFE7D" box="[1201,1213,367,388]" italics="true" pageId="5" pageNumber="590">S</emphasis>
.)
<emphasis id="B93096516D18FF96FFB5C4261D58FE7D" box="[1233,1306,367,388]" italics="true" pageId="5" pageNumber="590">mithras</emphasis>
Reitter, 1904
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFB4A436D18FF96FBF3C4C41DDEFE46" blockId="5.[151,1437,151,623]" pageId="5" pageNumber="590">
<keyLead id="30B5E9766D18FF96FBF3C4C41DDEFE46" pageId="5" pageNumber="590">
Profemora with transversal edge anteroventrally (
<emphasis id="B93096516D18FF96F9DEC4C51A89FE58" box="[698,715,396,417]" italics="true" pageId="5" pageNumber="590">cf</emphasis>
.
<bibRefCitation id="EFD537B26D18FF96F9B2C4C51BF0FE5B" author="Kral, D. &amp; Olexa, A." box="[726,946,396,418]" pageId="5" pageNumber="590" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa 1996: 63</bibRefCitation>
, fig. 35). Pronotal punctation simple and fine. Inner edge of parameres angulate or rounded subapically (
<figureCitation id="137F56C66D18FF96F9BCC4E01B0FFE46" box="[728,845,425,447]" captionStart="FIGURES 1416" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 1416. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="5" pageNumber="590">Figs. 14, 16</figureCitation>
)................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B052E66D18FF96FBF3C48E1DDEFD96" pageId="5" pageNumber="590">
<paragraph id="8BFB4A436D18FF96FBF3C48E1DDEFDEE" blockId="5.[151,1437,151,623]" pageId="5" pageNumber="590">
<keyLead id="30B5E9766D18FF96FBF3C48E1DDEFDEE" pageId="5" pageNumber="590">
3. Ventral mandibular processes weak and angulate in lateral aspect, invisible in dorsal aspect (
<figureCitation id="137F56C66D18FF96FF34C48E1C81FE24" box="[1104,1219,455,477]" captionStart="FIGURES 813" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 813. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 810, head, dorsal view; 1113, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="5" pageNumber="590">Figs. 10, 13</figureCitation>
). Inner edge of parameres broadly rounded subapically (
<figureCitation id="137F56C66D18FF96F949C4AD1A37FE03" box="[557,629,484,506]" captionStart="FIGURES 1416" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 1416. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="5" pageNumber="590">Fig. 16</figureCitation>
).
<collectingCountry id="F3530AD36D18FF96F9E1C4AD1B4FFE03" box="[645,781,484,506]" name="Turkmenistan" pageId="5" pageNumber="590">Turkmenistan</collectingCountry>
: north-western part of Kopet Dagh Mts., approximately between the regions of Kara Kala [= Garrygala or Maktymguly] and Bakharden [= Baharly]........
<emphasis id="B93096516D18FF96FF24C74B1C16FDEE" box="[1088,1108,514,535]" italics="true" pageId="5" pageNumber="590">L.</emphasis>
<taxonomicName id="4C4431C06D18FF96FF3DC74B1DDEFDEE" ID-CoL="6Q4GL" authority="Kral &amp; Olexa, 1996" authorityName="Kral &amp; Olexa" authorityYear="1996" box="[1113,1436,513,535]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="tekkensis" subGenus="Scelolethrus">
(
<emphasis id="B93096516D18FF96FF05C74B1C2FFDEE" box="[1121,1133,514,535]" italics="true" pageId="5" pageNumber="590">S</emphasis>
.)
<emphasis id="B93096516D18FF96FFE5C7481C9AFDEF" box="[1153,1240,513,534]" italics="true" pageId="5" pageNumber="590">tekkensis</emphasis>
<bibRefCitation id="EFD537B26D18FF96FFBAC7481DDEFDEE" author="Kral, D. &amp; Olexa, A." box="[1246,1436,513,535]" pageId="5" pageNumber="590" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa, 1996</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BFB4A436D18FF96FBF3C7561DDEFD96" blockId="5.[151,1437,151,623]" pageId="5" pageNumber="590">
<keyLead id="30B5E9766D18FF96FBF3C7561DDEFD96" pageId="5" pageNumber="590">
Ventral mandibular processes distinct, directed obliquely anteroventrally, pointed apically in lateral aspect, clearly visible in dorsal aspect (
<figureCitation id="137F56C66D18FF96FA09C77519F0FDA8" box="[365,434,572,594]" captionStart="FIGURES 47" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 47. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="5" pageNumber="590">Figs. 4</figureCitation>
,
<figureCitation id="137F56C66D18FF96FAD8C775198BFDAB" box="[444,457,572,594]" captionStart="FIGURES 813" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 813. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 810, head, dorsal view; 1113, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="5" pageNumber="590">8</figureCitation>
). Inner edge of parameres angulate subapically (
<figureCitation id="137F56C66D18FF96F8C5C7751BBDFDAB" box="[929,1023,572,594]" captionStart="FIGURES 47" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 47. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="5" pageNumber="590">Figs. 6, 7</figureCitation>
,
<figureCitation id="137F56C66D18FF96FF6EC7751C67FDAB" box="[1034,1061,572,594]" captionStart="FIGURES 1416" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 1416. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="5" pageNumber="590">14</figureCitation>
).
<collectingCountry id="F3530AD36D18FF96FF52C7741C1DFDAB" box="[1078,1119,573,594]" name="Iran" pageId="5" pageNumber="590">Iran</collectingCountry>
,
<collectingRegion id="498084A16D18FF96FF0EC7751C81FDAB" box="[1130,1219,572,594]" country="Iran" name="Golestan" pageId="5" pageNumber="590">Golestan</collectingRegion>
: region between Atrek and Gorgan river.................................................
<taxonomicName id="4C4431C06D18FF96F8B2C7131D5DFD96" ID-CoL="8RXWW" authority="Kral &amp; Hillert" authorityName="Kral &amp; Hillert" authorityYear="2021" box="[982,1311,601,623]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="golestanicus" status="sp. nov." subGenus="Scelolethrus">
<emphasis id="B93096516D18FF96F8B2C7131BA1FD96" box="[982,995,602,623]" italics="true" pageId="5" pageNumber="590">L</emphasis>
. (
<emphasis id="B93096516D18FF96F893C7131C41FD96" box="[1015,1027,602,623]" italics="true" pageId="5" pageNumber="590">S</emphasis>
.)
<emphasis id="B93096516D18FF96FF73C7101CCDFD97" box="[1047,1167,601,622]" italics="true" pageId="5" pageNumber="590">golestanicus</emphasis>
Král &amp; Hillert
</taxonomicName>
,
<taxonomicNameLabel id="A2032B2A6D18FF96FE4DC7131DDEFD96" box="[1321,1436,601,623]" pageId="5" pageNumber="590" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</treatment>
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