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<document id="584ADD2668F0B3CAD6B015153F946BF3" ID-DOI="10.11646/zootaxa.5019.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222728" ID-ZooBank="241751BA-E759-49B8-A394-271B2A4650AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629385249404" checkinUser="plazi" docAuthor="Singh, Sudhir, Colombo, Wesley D., Shreevihar, Santhosh, Pandey, Vijay P. &amp; Tribull, Carly M." docDate="2021" docId="03ED87EE3B04445D31EDFC55FE43C779" docLanguage="en" docName="zootaxa.5019.1.1.pdf" docOrigin="Zootaxa 5019 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5019.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Holepyris patnae Singh &amp; Colombo &amp; Shreevihar &amp; Pandey &amp; Tribull 2021, comb. nov." docType="treatment" docVersion="5" lastPageNumber="60" masterDocId="FFD4FF963B3C4466317AFFAFFFCDC638" masterDocTitle="Rediscovery of Kurians types of Hymenoptera at Forest Research Institute, Dehradun, India with notes on Indian bethylid fauna" masterLastPageNumber="90" masterPageNumber="1" pageNumber="57" updateTime="1699070948635" updateUser="plazi">
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<mods:title id="FCF02F383744A0F0854AE6354C4672D3">Rediscovery of Kurians types of Hymenoptera at Forest Research Institute, Dehradun, India with notes on Indian bethylid fauna</mods:title>
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<mods:namePart id="EF94C4563298D7DA6AFE445F132BBFD8">Colombo, Wesley D.</mods:namePart>
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<mods:namePart id="07174F392BDB4C33D89B7E697F9A43E3">Shreevihar, Santhosh</mods:namePart>
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<mods:namePart id="5183DCAE6458BA3E63ADF1201A607B3F">Pandey, Vijay P.</mods:namePart>
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<treatment id="03ED87EE3B04445D31EDFC55FE43C779" ID-DOI="http://doi.org/10.5281/zenodo.5222835" ID-Zenodo-Dep="5222835" LSID="urn:lsid:plazi:treatment:03ED87EE3B04445D31EDFC55FE43C779" httpUri="http://treatment.plazi.org/id/03ED87EE3B04445D31EDFC55FE43C779" lastPageId="59" lastPageNumber="60" pageId="56" pageNumber="57">
<subSubSection id="C35E65733B04445E31EDFC55FD13C22D" box="[151,734,1018,1045]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="8BFB36F83B04445E31EDFC55FD13C22D" blockId="56.[151,734,1018,1081]" box="[151,734,1018,1045]" pageId="56" pageNumber="57">
<heading id="D0B381943B04445E31EDFC55FD13C22D" bold="true" box="[151,734,1018,1045]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="3">
<emphasis id="B930EAEA3B04445E31EDFC55FD13C22D" bold="true" box="[151,734,1018,1045]" pageId="56" pageNumber="57">
34.
<taxonomicName id="4C444D7B3B04445E31B9FC55FD9CC22C" authority="(Kurian, 1955)" authorityName="Singh &amp; Colombo &amp; Shreevihar &amp; Pandey &amp; Tribull" authorityYear="2021" baseAuthorityName="Kurian" baseAuthorityYear="1955" box="[195,593,1018,1045]" class="Insecta" family="Bethylidae" genus="Holepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="patnae" status="comb. nov.">
<emphasis id="B930EAEA3B04445E31B9FC55FE5CC22D" bold="true" box="[195,401,1018,1045]" italics="true" pageId="56" pageNumber="57">Holepyris patnae</emphasis>
(
<bibRefCitation id="EFD54B093B04445E30DBFC55FD85C22C" author="Kurian, C." box="[417,584,1018,1044]" pageId="56" pageNumber="57" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357913B04445E3322FC54FD13C22D" box="[600,734,1019,1045]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35E65733B04445E31EDFBB0FE53C201" box="[151,414,1054,1081]" pageId="56" pageNumber="57" type="description">
<paragraph id="8BFB36F83B04445E31EDFBB0FE53C201" blockId="56.[151,734,1018,1081]" box="[151,414,1054,1081]" pageId="56" pageNumber="57">
<figureCitation id="137F2A7D3B04445E31EDFBB0FF21C201" box="[151,236,1055,1081]" captionStart="FIGURE 34" captionStartId="57.[151,250,1909,1933]" captionTargetBox="[206,1379,181,1876]" captionTargetId="figure-19@57.[206,1381,181,1885]" captionTargetPageId="57" captionText="FIGURE 34. AL. Holepyris patnae (Kurian, 1955) comb. nov. Holotype. A. Mesosoma, lateral view; B. Habitus after remounting, lateral view; C. Head, frontal view; D. Mesosoma, dorsal view; E, Lower margin of head showing dentition of mandibles; F. Mesosoma, lateral view; G. Antenna; H. Forewing; I. Legs, left pro- and right metaleg; J. Metasoma, dorsal view; K. Three slides, upper two of paratypes and lower of holotype; L. Labels." figureDoi="http://doi.org/10.5281/zenodo.5222798" httpUri="https://zenodo.org/record/5222798/files/figure.png" pageId="56" pageNumber="57">Figs 34</figureCitation>
, AL;
<figureCitation id="137F2A7D3B04445E3043FBB1FE9AC201" box="[313,343,1054,1081]" captionStart="FIGURE 51" captionStartId="82.[151,250,1824,1848]" captionTargetBox="[329,1259,181,1800]" captionTargetId="figure-19@82.[329,1259,181,1800]" captionTargetPageId="82" captionText="FIGURE 51. AN. Male genitalia. AB. Goniozus madrassippattanami Kurian, 1955, paratype. A. dorsal view; B. ventral view; CD. Goniozus salvadorae Kurian, 1954b, paratype; C. dorsal view; D. ventral view; EF. Goniozus thalasodi Kurian, 1955, holotype; E. dorsal view; F. ventral view; GH. Epyris indicae (Kurian, 1955), holotype; G. dorsal view; H. ventral view; IJ. Epyris manii Kurian, 1955, holotype; I. dorsal view; J. ventral view; KL. Holepyris patnae (Kurian, 1955), holotype; K. dorsal view; L. ventral view; MN. Pristocera agraensis (Kurian, 1955), holotype; M. dorsal view; N. ventral view." figureDoi="http://doi.org/10.5281/zenodo.5222843" httpUri="https://zenodo.org/record/5222843/files/figure.png" pageId="56" pageNumber="57">51</figureCitation>
, KL.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B04445E31EDFBCBFD2DC243" box="[151,736,1123,1147]" pageId="56" pageNumber="57" type="reference_group">
<paragraph id="8BFB36F83B04445E31EDFBCBFD2DC243" blockId="56.[151,736,1123,1178]" box="[151,736,1123,1147]" pageId="56" pageNumber="57">
<taxonomicName id="4C444D7B3B04445E31EDFBCBFE7EC243" authority="Kurian, 1955" authorityName="Kurian" authorityYear="1955" box="[151,435,1123,1147]" class="Insecta" family="Bethylidae" genus="Epyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="patnae">
<emphasis id="B930EAEA3B04445E31EDFBCBFEE8C243" box="[151,293,1124,1147]" italics="true" pageId="56" pageNumber="57">Epyris patnae</emphasis>
<bibRefCitation id="EFD54B093B04445E3051FBCCFE7EC243" author="Kurian, C." box="[299,435,1123,1147]" pageId="56" pageNumber="57" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
</taxonomicName>
, 4: 89, 97, 98, Figs: 107111.
</paragraph>
</subSubSection>
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<paragraph id="8BFB36F83B04445E31EDFB2DFD64C2A2" blockId="56.[151,736,1123,1178]" box="[151,681,1154,1178]" pageId="56" pageNumber="57">
<materialsCitation id="3B2C3CA53B04445E31EDFB2DFD68C2A2" box="[151,677,1154,1178]" collectionCode="NFIC-FRI" country="India" location="India" pageId="56" pageNumber="57" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
Type locality:
<taxonomicName id="4C444D7B3B04445E3053FB2DFEA9C2A1" authorityName="Ragonot" authorityYear="1888" box="[297,356,1154,1177]" class="Insecta" family="Pyralidae" genus="Patna" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">Patna</taxonomicName>
,
<collectingCountry id="F35376683B04445E3014FB2DFE6FC2A2" box="[366,418,1154,1178]" name="India" pageId="56" pageNumber="57">India</collectingCountry>
(
<specimenCount id="9D42FD713B04445E30CBFB2DFE09C2A2" box="[433,452,1154,1178]" pageId="56" pageNumber="57" type="male"></specimenCount>
<typeStatus id="54FF885A3B04445E30B0FB2DFDE8C2A2" box="[458,549,1154,1178]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED55AE3D3B04445E334AFB2DFD52C2A2" box="[560,671,1154,1178]" pageId="56" pageNumber="57">NFIC-FRI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B04445E31EDFB67FD67C072" pageId="56" pageNumber="57" type="diagnosis">
<paragraph id="8BFB36F83B04445E31EDFB67FD67C072" blockId="56.[151,1437,1224,2043]" pageId="56" pageNumber="57">
<emphasis id="B930EAEA3B04445E31EDFB67FEC7C2DA" bold="true" box="[151,266,1224,1250]" pageId="56" pageNumber="57">Diagnosis</emphasis>
. Male. Length about 2.8 mm. General colour dark reddish-brown to black. Head as long as wide; frons strongly coriaceus; ocelli distinct, posterior to supra-ocular line; eye glabrous; median clypeal lobe conspicuous, anterior margin rounded, as long as lateral ones; mandible tridentate; antenna with sparse pubescence, scape 0.2 × longer than pedicel, pedicel shorter than first flagellomere. Pronotum strongly coriaceous; mesonotum shorter than pronotal dorsal area; notauli indistinct, shorter than parapsidal signum; mesoscuto-mesoscutellar suture sulcate, trabeculate, deep. Metapectal-propodeal disc longer than wide; metapostnotal median carina present; first abdominal spiracle laterally placed. Tarsal claw bifid. Wings hyaline; forewing with pterostigma light brown; hind wing with four hamuli. Metasoma polished. Genitalia: Harpe simple, longer than gonostipes in dorsal view, apex rounded, not arched inward; gonostipes wide, not covering aedeagal base; cuspis elongate, apex anterior to apex of harpe; digitus with apex posterior to apex of cuspis, its posterior margin smooth with three teeth; aedeagus subtrapezoidal, its apex posterior to apex of digitus. Female. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B04445E31BDF9FBFC50C153" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="8BFB36F83B04445E31BDF9FBFEE7C0AA" blockId="56.[151,1437,1224,2043]" pageId="56" pageNumber="57">
<emphasis id="B930EAEA3B04445E31BDF9FBFE7CC056" bold="true" box="[199,433,1620,1646]" pageId="56" pageNumber="57">Material examined.</emphasis>
<materialsCitation id="3B2C3CA53B04445E30C3F9FBFB92C056" box="[441,1119,1619,1647]" collectingDate="1911-05-26" country="India" location="INDIA" pageId="56" pageNumber="57" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FF885A3B04445E30C3F9FBFDEBC056" box="[441,550,1620,1646]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
: [
<collectingCountry id="F35376683B04445E334DF9FCFD43C056" box="[567,654,1619,1646]" name="India" pageId="56" pageNumber="57">INDIA</collectingCountry>
,]
<taxonomicName id="4C444D7B3B04445E33DBF9FAFD29C056" authorityName="Ragonot" authorityYear="1888" box="[673,740,1621,1646]" class="Insecta" family="Pyralidae" genus="Patna" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAEA3B04445E33DBF9FAFD29C056" box="[673,740,1621,1646]" italics="true" pageId="56" pageNumber="57">Patna</emphasis>
</taxonomicName>
,
<date id="FFFA10383B04445E3388F9FBFCA1C056" box="[754,876,1620,1646]" pageId="56" pageNumber="57" value="1911-05-26">
<collectingDate id="EFBEE9D03B04445E3388F9FBFCA1C056" box="[754,876,1620,1646]" pageId="56" pageNumber="57" value="1911-05-26">26-V-1911</collectingDate>
</date>
, parasitic, C. Nº. 881
</materialsCitation>
.
<materialsCitation id="3B2C3CA53B04445E3511F9FAFEEBC0AA" collectingDate="1911-05-26" country="India" location="INDIA" pageId="56" pageNumber="57" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54FF885A3B04445E3511F9FAFB1CC057" box="[1131,1233,1621,1647]" pageId="56" pageNumber="57" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D42FD713B04445E35A6F9FBFACDC056" box="[1244,1280,1620,1646]" pageId="56" pageNumber="57" type="male">1♂</specimenCount>
, same data of the
<typeStatus id="54FF885A3B04445E31BAF9D7FEEBC0AA" box="[192,294,1656,1682]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFB36F83B04445E31BDF933FC50C153" blockId="56.[151,1437,1224,2043]" pageId="56" pageNumber="57">
<emphasis id="B930EAEA3B04445E31BDF933FE41C08E" bold="true" box="[199,396,1692,1718]" pageId="56" pageNumber="57">Type conditions.</emphasis>
<typeStatus id="54FF885A3B04445E30E9F933FE36C08E" box="[403,507,1692,1718]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
partly dissected
<taxonomicName id="4C444D7B3B04445E33C1F932FCDEC08E" box="[699,787,1692,1719]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="73" phylum="Arthropoda" rank="species" species="slide">on slide</taxonomicName>
Nº. 90/B and rest was mounted on the pin.
<emphasis id="B930EAEA3B04445E3478F933FA9CC08E" box="[1282,1361,1692,1718]" italics="true" pageId="56" pageNumber="57">Pinned</emphasis>
: Head and Metasoma both separately glued to the pith to which mesosoma is pinned; mesosoma, with both pro- and mesolegs attached.
<taxonomicName id="4C444D7B3B04445E3000F94BFE25C0C6" box="[378,488,1764,1790]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="slides">
<emphasis id="B930EAEA3B04445E3000F94BFE25C0C6" box="[378,488,1764,1790]" italics="true" pageId="56" pageNumber="57">On slides</emphasis>
</taxonomicName>
: S1 C1—one forewing and one hind wing. S1 C2—one proleg, one leg without coxa; one antenna complete.
<typeStatus id="54FF885A3B04445E309BF8A6FD8EC11B" box="[481,579,1801,1827]" pageId="56" pageNumber="57" type="paratype">Paratype</typeStatus>
partly dissected
<taxonomicName id="4C444D7B3B04445E3279F8A6FC91C11A" box="[771,860,1800,1827]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="73" phylum="Arthropoda" rank="species" species="slide">on slide</taxonomicName>
Nº. 68/B and remaining rested on the pin.
<emphasis id="B930EAEA3B04445E343FF8A7FA59C11A" box="[1349,1428,1800,1826]" italics="true" pageId="56" pageNumber="57">Pinned</emphasis>
: mesosoma without any appendages.
<taxonomicName id="4C444D7B3B04445E3343F883FD69C17E" box="[569,676,1836,1862]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="slides">
<emphasis id="B930EAEA3B04445E3343F883FD69C17E" box="[569,676,1836,1862]" italics="true" pageId="56" pageNumber="57">On slides</emphasis>
</taxonomicName>
: S2 C1—one hind wing only. S3 C1—one forewing and one hind wing. S3 C2—two legs, one with coxa; one antenna broken into two.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B04445D31BDF8DBFA41C725" lastPageId="59" lastPageNumber="60" pageId="56" pageNumber="57" type="discussion">
<paragraph id="8BFB36F83B04445E31BDF8DBFBEBC1C2" blockId="56.[151,1437,1224,2043]" pageId="56" pageNumber="57">
<emphasis id="B930EAEA3B04445E31BDF8DBFEF0C1B6" bold="true" box="[199,317,1908,1934]" pageId="56" pageNumber="57">Remarks:</emphasis>
The pinned
<typeStatus id="54FF885A3B04445E30B3F8DBFDE6C1B6" box="[457,555,1908,1934]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
was in poor shape, with a rusted pin that had damaged the mesosoma, with the remaining parts (head, legs, wings and metasoma) loosely attached and disintegrating. To preserve the type, it was remounted on a card. The metasoma of the
<typeStatus id="54FF885A3B04445E330CF813FD15C1EE" box="[630,728,1980,2006]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
was dissected to extract the genitalia, which are mounted under the rectangular cover on S1. The head and metasoma of the
<typeStatus id="54FF885A3B04445E324CF84EFC5BC1C3" box="[822,918,2017,2043]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
are missing.
</paragraph>
<caption id="DF3B66703B05445F31EDF8DAFC4EC1C1" ID-DOI="http://doi.org/10.5281/zenodo.5222798" ID-Zenodo-Dep="5222798" httpUri="https://zenodo.org/record/5222798/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1909,1933]" targetBox="[206,1379,181,1876]" targetPageId="57">
<paragraph id="8BFB36F83B05445F31EDF8DAFC4EC1C1" blockId="57.[151,1436,1909,2041]" pageId="57" pageNumber="58">
<emphasis id="B930EAEA3B05445F31EDF8DAFEAEC1B5" bold="true" box="[151,355,1909,1933]" pageId="57" pageNumber="58">FIGURE 34. AL.</emphasis>
<taxonomicName id="4C444D7B3B05445F3014F8DAFD0AC1B5" authority="(Kurian, 1955)" authorityName="Singh &amp; Colombo &amp; Shreevihar &amp; Pandey &amp; Tribull" authorityYear="2021" baseAuthorityName="Kurian" baseAuthorityYear="1955" box="[366,711,1909,1933]" class="Insecta" family="Bethylidae" genus="Holepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="patnae" status="comb. nov.">
<emphasis id="B930EAEA3B05445F3014F8DAFDEFC1B5" box="[366,546,1909,1933]" italics="true" pageId="57" pageNumber="58">Holepyris patnae</emphasis>
(
<bibRefCitation id="EFD54B093B05445F334EF8DAFD0DC1B5" author="Kurian, C." box="[564,704,1909,1933]" pageId="57" pageNumber="58" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
)
</taxonomicName>
<emphasis id="B930EAEA3B05445F33A8F8DAFC87C1B5" bold="true" box="[722,842,1909,1933]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A20357913B05445F33A8F8DAFC87C1B5" box="[722,842,1909,1933]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Holotype.
<emphasis id="B930EAEA3B05445F32B9F8DAFC11C1B5" bold="true" box="[963,988,1909,1933]" pageId="57" pageNumber="58">A.</emphasis>
Mesosoma, lateral view;
<emphasis id="B930EAEA3B05445F3588F8DAFAC4C1B5" bold="true" box="[1266,1289,1909,1933]" pageId="57" pageNumber="58">B.</emphasis>
Habitus after remounting, lateral view;
<emphasis id="B930EAEA3B05445F30D9F836FE76C189" bold="true" box="[419,443,1945,1969]" pageId="57" pageNumber="58">C.</emphasis>
Head, frontal view;
<emphasis id="B930EAEA3B05445F33EEF836FD60C189" bold="true" box="[660,685,1945,1969]" pageId="57" pageNumber="58">D.</emphasis>
Mesosoma, dorsal view; E, Lower margin of head showing dentition of mandibles;
<emphasis id="B930EAEA3B05445F3076F812FED2C1ED" bold="true" box="[268,287,1981,2005]" pageId="57" pageNumber="58">F.</emphasis>
Mesosoma, lateral view;
<emphasis id="B930EAEA3B05445F335AF812FDF6C1ED" bold="true" box="[544,571,1981,2005]" pageId="57" pageNumber="58">G.</emphasis>
Antenna;
<emphasis id="B930EAEA3B05445F33DBF812FD76C1ED" bold="true" box="[673,699,1981,2005]" pageId="57" pageNumber="58">H.</emphasis>
Forewing;
<emphasis id="B930EAEA3B05445F3254F812FCF3C1ED" bold="true" box="[814,830,1981,2005]" pageId="57" pageNumber="58">I.</emphasis>
Legs, left pro- and right metaleg;
<emphasis id="B930EAEA3B05445F35E9F812FB6BC1ED" bold="true" box="[1171,1190,1981,2005]" pageId="57" pageNumber="58">J.</emphasis>
Metasoma, dorsal view;
<emphasis id="B930EAEA3B05445F31EDF84EFF7CC1C1" bold="true" box="[151,177,2017,2041]" pageId="57" pageNumber="58">K.</emphasis>
Three slides, upper two of paratypes and lower of holotype;
<emphasis id="B930EAEA3B05445F3260F84EFCFCC1C1" bold="true" box="[794,817,2017,2041]" pageId="57" pageNumber="58">L.</emphasis>
Labels.
</paragraph>
</caption>
<caption id="DF3B66703B06445C31EDF838FDAEC1CF" ID-DOI="http://doi.org/10.5281/zenodo.5222800" ID-Zenodo-Dep="5222800" httpUri="https://zenodo.org/record/5222800/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[195,1392,181,1912]" targetPageId="58">
<paragraph id="8BFB36F83B06445C31EDF838FDAEC1CF" blockId="58.[151,1436,1943,2039]" pageId="58" pageNumber="59">
<emphasis id="B930EAEA3B06445C31EDF838FE97C197" bold="true" box="[151,346,1943,1967]" pageId="58" pageNumber="59">FIGURE 35. AJ.</emphasis>
<taxonomicName id="4C444D7B3B06445C301BF838FD60C197" authority="(Kurian, 1955)" authorityName="Singh &amp; Colombo &amp; Shreevihar &amp; Pandey &amp; Tribull" authorityYear="2021" baseAuthorityName="Kurian" baseAuthorityYear="1955" box="[353,685,1943,1967]" class="Insecta" family="Bethylidae" genus="Holepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="raonis" status="comb. nov.">
<emphasis id="B930EAEA3B06445C301BF838FDC3C197" box="[353,526,1943,1967]" italics="true" pageId="58" pageNumber="59">Holepyris raonis</emphasis>
(
<bibRefCitation id="EFD54B093B06445C3367F838FD6BC197" author="Kurian, C." box="[541,678,1943,1967]" pageId="58" pageNumber="59" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
)
</taxonomicName>
<emphasis id="B930EAEA3B06445C33CFF838FCE6C197" bold="true" box="[693,811,1943,1967]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="A20357913B06445C33CFF838FCE6C197" box="[693,811,1943,1967]" pageId="58" pageNumber="59" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Holotype.
<emphasis id="B930EAEA3B06445C32E4F838FC7BC197" bold="true" box="[926,950,1943,1967]" pageId="58" pageNumber="59">A.</emphasis>
Habitus, dorsal view;
<emphasis id="B930EAEA3B06445C35DBF838FB75C197" bold="true" box="[1185,1208,1943,1967]" pageId="58" pageNumber="59">B.</emphasis>
Habitus, lateral view;
<emphasis id="B930EAEA3B06445C31EDF814FF62C1EB" bold="true" box="[151,175,1979,2003]" pageId="58" pageNumber="59">C.</emphasis>
Head, frontal view;
<emphasis id="B930EAEA3B06445C30F9F814FE51C1EB" bold="true" box="[387,412,1979,2003]" pageId="58" pageNumber="59">D.</emphasis>
Mandibles;
<emphasis id="B930EAEA3B06445C3366F814FDFFC1EB" bold="true" box="[540,562,1979,2003]" pageId="58" pageNumber="59">E.</emphasis>
Mesosoma, dorso-lateral view;
<emphasis id="B930EAEA3B06445C3201F814FC43C1EB" bold="true" box="[891,910,1979,2003]" pageId="58" pageNumber="59">F.</emphasis>
Antenna;
<emphasis id="B930EAEA3B06445C3282F814FBDFC1EB" bold="true" box="[1016,1042,1979,2003]" pageId="58" pageNumber="59">G.</emphasis>
Forewing;
<emphasis id="B930EAEA3B06445C35F2F814FB6FC1EB" bold="true" box="[1160,1186,1979,2003]" pageId="58" pageNumber="59">H.</emphasis>
Legs, left pro- and right metaleg;
<emphasis id="B930EAEA3B06445C318FF870FEC8C1CF" bold="true" box="[245,261,2015,2039]" pageId="58" pageNumber="59">I.</emphasis>
Two slides of holotype;
<emphasis id="B930EAEA3B06445C3085F870FDDFC1CF" bold="true" box="[511,530,2015,2039]" pageId="58" pageNumber="59">J.</emphasis>
Labels.
</paragraph>
</caption>
<paragraph id="8BFB36F83B07445D31BDFF38FA41C725" blockId="59.[151,1436,151,321]" pageId="59" pageNumber="60">
We detected the clypeus trilobate, eyes mid-sized and glabrous, the dorsal pronotal area with posterior pronotal sulcus foveolate, and with humeral angle not projected, the mesoscuto-scutellar suture sulcate, the metanotum undeveloped medially, the wings are fully developed, the forewings with three closed cells (C
<subScript id="17C034BD3B07445D35E4FF5EFB6BC6C7" attach="both" box="[1182,1190,241,255]" fontSize="6" pageId="59" pageNumber="60">2</subScript>
c, R
<subScript id="17C034BD3B07445D35A8FF5EFB17C6C7" attach="both" box="[1234,1242,241,255]" fontSize="6" pageId="59" pageNumber="60">2</subScript>
c, 1Cu
<subScript id="17C034BD3B07445D3458FF5EFAE7C6C7" attach="right" box="[1314,1322,241,255]" fontSize="6" pageId="59" pageNumber="60">2</subScript>
c), and the metapostnotal median carina is long. All of these observations support the placement of this species in
<taxonomicName id="4C444D7B3B07445D346DFEACFA48C725" authorityName="Kieffer" authorityYear="1904" box="[1303,1413,259,285]" class="Insecta" family="Bethylidae" genus="Holepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAEA3B07445D346DFEACFA48C725" box="[1303,1413,259,285]" italics="true" pageId="59" pageNumber="60">Holepyris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B07445D31BDFE88FE43C779" box="[199,398,295,321]" pageId="59" pageNumber="60" type="biology_ecology">
<paragraph id="8BFB36F83B07445D31BDFE88FE43C779" blockId="59.[151,1436,151,321]" box="[199,398,295,321]" pageId="59" pageNumber="60">
<emphasis id="B930EAEA3B07445D31BDFE88FEDCC779" bold="true" box="[199,273,295,321]" pageId="59" pageNumber="60">Hosts.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>