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<document id="8A7339FFF9CA190BF48CFA468A7C453D" ID-DOI="10.5281/zenodo.182666" ID-GBIF-Dataset="d1698b4a-ec6b-4a5f-96cf-956535545e85" ID-ISSN="1175-5326" ID-Zenodo-Dep="182666" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459872547299" checkinUser="plazi" docAuthor="Ellenrieder, Natalia Von &amp; Garrison, Rosser W." docDate="2008" docId="03ED87D9FF94BE62A9D1A4ECFBA667AB" docLanguage="en" docName="zt01805p051.pdf" docOrigin="Zootaxa 1805" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Oreiallagma oreas Ris 1918, comb. nov." docType="treatment" docVersion="11" lastPageNumber="46" masterDocId="FFD4FFA1FFB8BE4FA946A043FFC86549" masterDocTitle="Oreiallagma gen. nov. with a redefinition of Cyanallagma Kennedy 1920 and Mesamphiagrion Kennedy 1920, and the description of M. dunklei sp. nov. and M. ecuatoriale sp. nov. from Ecuador (Odonata: Coenagrionidae)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="45" updateTime="1698226780440" updateUser="plazi">
<mods:mods id="DCE92103EFBC03AE5ADB10569F10F4FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4E23B0409640D9D5027B1F092447E26A">
<mods:title id="914ADBD886A28ADD29EF8BE1373D3153">Oreiallagma gen. nov. with a redefinition of Cyanallagma Kennedy 1920 and Mesamphiagrion Kennedy 1920, and the description of M. dunklei sp. nov. and M. ecuatoriale sp. nov. from Ecuador (Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="966F923136BD2BBE82E5AC88B40B67DE" type="personal">
<mods:role id="A33F8D438F7D50B201E826FFB5E2784B">
<mods:roleTerm id="D40BC2E7B85625D4C2897A9F594393D4">Author</mods:roleTerm>
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<mods:namePart id="DAC7C2CFB7D034A055441794F91970AF">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:name id="2AD2D0ECA495DE6D7993AED3AECB2B7E" type="personal">
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</mods:role>
<mods:namePart id="4636E6F261784366C9E11FE3B5793E1D">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DFC1A19587E87BFA844E0985DA356BDC">text</mods:typeOfResource>
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<mods:title id="5ACAA06ADB28AB21406FD0E427E37275">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="23DD391657BE6E32EB4222D7E18F84B2">
<mods:date id="0CB2DCC4C48BFE7A3E508788107336AD">2008</mods:date>
<mods:detail id="96C637CA29246FDFD3D7F1CBC7494A6E" type="volume">
<mods:number id="BDC8031A20A42DA083E2E497EE6EA311">1805</mods:number>
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<mods:extent id="725ABAF2F238662ED0CF230D513D5E33" unit="page">
<mods:start id="F464A162865022734DF374F69AAE2CE4">1</mods:start>
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<mods:classification id="DC640C76005659B29EEE9BD3F4CCB113">journal article</mods:classification>
<mods:identifier id="9E168F556E3E5DB7D43C59A99906E290" type="DOI">10.5281/zenodo.182666</mods:identifier>
<mods:identifier id="DDDBBDF1EDAB00151A0967BCBCAFB1B4" type="GBIF-Dataset">d1698b4a-ec6b-4a5f-96cf-956535545e85</mods:identifier>
<mods:identifier id="D4541CE9ABD8A19AEB3AF0E1634752F7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8A9C224EC6B2C3464E8A183B2C5021A0" type="Zenodo-Dep">182666</mods:identifier>
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<treatment id="03ED87D9FF94BE62A9D1A4ECFBA667AB" ID-DOI="http://doi.org/10.5281/zenodo.5691621" ID-GBIF-Taxon="119369805" ID-Zenodo-Dep="5691621" LSID="urn:lsid:plazi:treatment:03ED87D9FF94BE62A9D1A4ECFBA667AB" httpUri="http://treatment.plazi.org/id/03ED87D9FF94BE62A9D1A4ECFBA667AB" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C35E6544FF94BE63A9D1A4ECFE1D61A5" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BFB36CFFF94BE63A9D1A4ECFD416180" blockId="44.[151,649,1199,1260]" box="[151,649,1199,1225]" pageId="44" pageNumber="45">
<heading id="D0B381A3FF94BE63A9D1A4ECFD416180" bold="true" box="[151,649,1199,1225]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B930EADDFF94BE63A9D1A4ECFD416180" bold="true" box="[151,649,1199,1225]" pageId="44" pageNumber="45">
<taxonomicName id="4C444D4CFF94BE63A9D1A4ECFE336180" ID-CoL="6T4P5" authority="Ris 1918" authorityName="Ris" authorityYear="1918" box="[151,507,1199,1225]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="oreas" status="comb. nov.">
<emphasis id="B930EADDFF94BE63A9D1A4ECFEBE6180" bold="true" box="[151,374,1199,1225]" italics="true" pageId="44" pageNumber="45">Oreiallagma oreas</emphasis>
(
<bibRefCitation id="EFD54B3EFF94BE63A8C3A4ECFE3B6180" author="Ris" box="[389,499,1199,1225]" pageId="44" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris 1918</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357A6FF94BE63AB45A4ECFD416180" box="[515,649,1199,1225]" pageId="44" pageNumber="45">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A491FE1D61A5" blockId="44.[151,649,1199,1260]" box="[151,469,1234,1260]" pageId="44" pageNumber="45">
<figureCitation id="137F2A4AFF94BE63A9D1A491FEDF61A5" box="[151,279,1234,1260]" captionStart="FIGURES 25 – 29" captionStartId="20.[151,269,1543,1567]" captionTargetBox="[188,1403,280,1513]" captionTargetId="figure@20.[183,1415,274,1525]" captionTargetPageId="20" captionText="FIGURES 25 – 29. Posterior lobe of prothorax. (25) Oreiallagma prothoracicum, male paratype, Ecuador, Intaj, (a) dorsal view, (b) medio-dorsal view; (26), O. oreas, male holotype, Colombia, Monte Socorro, dorsal view; (27) O. acutum, male lectotype, Bolivia, Río Zongo, dorsal view; (28) O. quadricolor, Peru, (a – b) posterior lobe of prothorax, male, holotype, Santa Ana, (a) dorsal view, (b) antero-lateral view, (c) female, Morro Leguía, dorsal view; (29) O. thelkterion, Venezuela, dorsal view, (a) male, Monte Zerpa, (b) female, Guaramacal." httpUri="https://zenodo.org/record/182676/files/figure.png" pageId="44" pageNumber="45">Figures 26</figureCitation>
; 52; 89; 94; 104
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF94BE63A9D1A561FDA960F0" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="8BFB36CFFF94BE63A9D1A561FDED6030" blockId="44.[151,1436,1314,1625]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="AB5411E1FF94BE63A9D1A561FDED6030" pageId="44" pageNumber="45">
<taxonomicName id="4C444D4CFF94BE63A9D1A561FE146070" ID-CoL="9WYDW" authority="Ris 1918: 136" authorityName="Ris" authorityPageNumber="136" authorityYear="1918" box="[151,476,1314,1337]" class="Insecta" family="Coenagrionidae" genus="Telagrion" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="oreas">
<emphasis id="B930EADDFF94BE63A9D1A561FEF66070" box="[151,318,1314,1337]" italics="true" pageId="44" pageNumber="45">Telagrion oreas</emphasis>
<treatmentCitation id="0AE510DEFF94BE63A803A561FE146070" author="Ris" box="[325,476,1314,1337]" page="136" pageId="44" pageNumber="45" year="1918">
<bibRefCitation id="EFD54B3EFF94BE63A803A561FE606070" author="Ris" box="[325,424,1314,1337]" pageId="44" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris 1918</bibRefCitation>
: 136
</treatmentCitation>
</taxonomicName>
, 138–139, fig. 78 (diagnosis from
<taxonomicName id="4C444D4CFF94BE63AA0AA561FC2C6070" ID-CoL="754P5" box="[844,996,1314,1337]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF94BE63AA0AA561FC2C6070" box="[844,996,1314,1337]" italics="true" pageId="44" pageNumber="45">T. quadricolor</emphasis>
</taxonomicName>
, description of male, illustration of S10).
<treatmentCitation id="0AE510DEFF94BE63A9D1A501FEA46010" author="Kimmins" box="[151,364,1346,1369]" page="189" pageId="44" pageNumber="45" year="1945">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A501FEF26010" author="Kimmins" box="[151,314,1346,1369]" pageId="44" pageNumber="50" refString="Kimmins, D. E. (1945) A new species of Odonata from Ecuador. Annals of the Magazine on Natural History, (11) 12, 187 - 189." type="journal article" year="1945">Kimmins 1945</bibRefCitation>
: 189
</treatmentCitation>
(comparison with
<taxonomicName id="4C444D4CFF94BE63AB72A501FD246010" ID-CoL="74SP5" box="[564,748,1346,1369]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="prothoracicum">
<emphasis id="B930EADDFF94BE63AB72A501FD246010" box="[564,748,1346,1369]" italics="true" pageId="44" pageNumber="45">T. prothoracicum</emphasis>
</taxonomicName>
). St.
<treatmentCitation id="0AE510DEFF94BE63A9FCA521FEBD6030" author="Quentin" box="[186,373,1378,1401]" page="48" pageId="44" pageNumber="45" year="1960">
<bibRefCitation id="EFD54B3EFF94BE63A9FCA521FE986030" author="Quentin" box="[186,336,1378,1401]" pageId="44" pageNumber="51" refString="St. Quentin, D. (1960) Zur Kenntnis der Agrioninae (Coenagrioninae) Sudamerikas (Odonata). Beitrage Neotropische Fauna, 2 (1), 45 - 64." type="journal article" year="1960">Quentin 1960</bibRefCitation>
: 48
</treatmentCitation>
(key for males).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A5C1FDA960F0" blockId="44.[151,1436,1314,1625]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A5C1FED560D0" author="Santos" box="[151,285,1410,1433]" pageId="44" pageNumber="51" refString="Santos, N. D. (1965) Contribuicao ao conhecimento dos &quot; Odonata &quot; da regiao de Pocos de Caldas, MG. &quot; Minagrion &quot; gen. n., para &quot; Telagrion mecistogastrum &quot; Selys Longchamps 1865, com a descricao de uma nova especie. Atas da Sociedade de Biologia do Rio de Janeiro, 9 (1), 8 - 12." type="journal article" year="1965">Santos 1965</bibRefCitation>
: 9 (possible placement in
<taxonomicName id="4C444D4CFF94BE63AB74A5C1FD6060D0" ID-CoL="5CR8" box="[562,680,1410,1433]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B930EADDFF94BE63AB74A5C1FD6060D0" box="[562,680,1410,1433]" italics="true" pageId="44" pageNumber="45">Leptagrion</emphasis>
</taxonomicName>
).
<bibRefCitation id="EFD54B3EFF94BE63A9D1A5E1FE4860F0" author="Davies" box="[151,384,1442,1465]" pageId="44" pageNumber="50" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. Volume 1. Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communications (Supplement), Utrecht, 3: ix + 127 pp." type="book" year="1984">Davies &amp; Tobin 1984</bibRefCitation>
: 94 (synonymic list).
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF94BE62A9D1A581FCF0644B" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
<paragraph id="8BFB36CFFF94BE63A9D1A581FDDD6090" blockId="44.[151,1436,1314,1625]" box="[151,533,1474,1497]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A581FEFC6090" author="Garrison" box="[151,308,1474,1497]" pageId="44" pageNumber="50" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison 1991</bibRefCitation>
: 13 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A5A1FD8B60B0" blockId="44.[151,1436,1314,1625]" box="[151,579,1506,1529]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A5A1FEE160B0" author="Bridges" box="[151,297,1506,1529]" pageId="44" pageNumber="49" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group, and species-group names of the Odonata of the world (Third Edition). Urbana, xlvi + 905 pp." type="book" year="1994">Bridges 1994</bibRefCitation>
: VII.174 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A641FDFE6350" blockId="44.[151,1436,1314,1625]" box="[151,566,1538,1561]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A641FE806350" author="Steinmann" box="[151,328,1538,1561]" pageId="44" pageNumber="51" refString="Steinmann, H. (1997) World catalogue of Odonata. Volume I. Zygoptera. In: Wermuth, H. &amp; Fischer, M. (eds.). Das Tierreich. The Animal Kingdom. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen, Walter de Gruyter, Berlin, 110, xxi + 500 pp." type="book" year="1997">Steinmann 1997</bibRefCitation>
: 358 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A661FE3F6370" blockId="44.[151,1436,1314,1625]" box="[151,503,1570,1593]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A661FEDE6370" author="Tsuda" box="[151,278,1570,1593]" pageId="44" pageNumber="51" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately published, Osaka, vi + 430 pp." type="book" year="2000">Tsuda 2000</bibRefCitation>
: 50 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A601FE196310" blockId="44.[151,1436,1314,1625]" box="[151,465,1602,1625]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A601FEFD6310" author="Lencioni" box="[151,309,1602,1625]" pageId="44" pageNumber="50" refString="Lencioni, F. A. A. (2004) Telagrion nathaliae spec. nov. (Zygoptera: Coenagrionidae). Odonatologica, 33 (1), 91 - 98." type="journal article" year="2004">Lencioni 2004</bibRefCitation>
: 92 (mention).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A6C9FB8E6385" blockId="44.[151,1436,1674,2020]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A9D1A6C9FF2F63ED" bold="true" box="[151,231,1674,1700]" pageId="44" pageNumber="45">
<typeStatus id="54FF886DFF94BE63A9D1A6C9FF2A63ED" box="[151,226,1674,1700]" pageId="44" pageNumber="45">Types</typeStatus>
:
</emphasis>
ɗ
<typeStatus id="54FF886DFF94BE63A84AA6C9FEB063ED" box="[268,376,1674,1700]" pageId="44" pageNumber="45" type="holotype">
<emphasis id="B930EADDFF94BE63A84AA6C9FEB063ED" bold="true" box="[268,376,1674,1700]" pageId="44" pageNumber="45">holotype</emphasis>
</typeStatus>
in FNS (examined) with following data: &quot;Monte Socorro
<quantity id="4CBC9B2AFF94BE63AD66A6C9FBB063ED" box="[1056,1144,1674,1700]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="44" pageNumber="45" unit="m" value="2300.0">2300 m</quantity>
/ [Valle del Cauca Dept.]
<collectingCountry id="F353765FFF94BE63A9D1A6F1FEC66385" box="[151,270,1714,1740]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
, W. Cordill./ vii. 0 9 [A.H.] Fassl&quot;; (Senckenberg Register No. 10764).
</paragraph>
<paragraph id="8BFB36CFFF94BE62A980A699FCF0644B" blockId="44.[151,1436,1674,2020]" lastBlockId="45.[151,1436,152,738]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A980A699FE6B63BD" bold="true" box="[198,419,1754,1780]" pageId="44" pageNumber="45">Characterization.</emphasis>
Medial lobe of posterior lobe of prothorax is slightly constricted at base, subquadrate, with shallow medial concavity on posterior margin (
<figureCitation id="137F2A4AFF94BE63ABB8A741FC9E6255" box="[766,854,1794,1820]" captionStart="FIGURES 25 – 29" captionStartId="20.[151,269,1543,1567]" captionTargetBox="[188,1403,280,1513]" captionTargetId="figure@20.[183,1415,274,1525]" captionTargetPageId="20" captionText="FIGURES 25 – 29. Posterior lobe of prothorax. (25) Oreiallagma prothoracicum, male paratype, Ecuador, Intaj, (a) dorsal view, (b) medio-dorsal view; (26), O. oreas, male holotype, Colombia, Monte Socorro, dorsal view; (27) O. acutum, male lectotype, Bolivia, Río Zongo, dorsal view; (28) O. quadricolor, Peru, (a – b) posterior lobe of prothorax, male, holotype, Santa Ana, (a) dorsal view, (b) antero-lateral view, (c) female, Morro Leguía, dorsal view; (29) O. thelkterion, Venezuela, dorsal view, (a) male, Monte Zerpa, (b) female, Guaramacal." httpUri="https://zenodo.org/record/182676/files/figure.png" pageId="44" pageNumber="45">Fig. 26</figureCitation>
; as in
<taxonomicName id="4C444D4CFF94BE63AAE7A740FBD36255" box="[929,1051,1795,1820]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B930EADDFF94BE63AAE7A740FBD36255" box="[929,1051,1795,1820]" italics="true" pageId="44" pageNumber="45">O. acutum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D4CFF94BE63AD12A740FACD6255" box="[1108,1285,1795,1820]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF94BE63AD12A740FACD6255" box="[1108,1285,1795,1820]" italics="true" pageId="44" pageNumber="45">O. quadricolor</emphasis>
</taxonomicName>
). Distal segment of genital ligula (
<figureCitation id="137F2A4AFF94BE63A8E3A769FE34620D" box="[421,508,1834,1860]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="44" pageNumber="45">Fig. 52</figureCitation>
) approximately as wide at apex as at base with two contiguous pairs of lateroapical lobes (unique) recurved and directed basally, small accessory latero-medial lobes and one (as in
<taxonomicName id="4C444D4CFF94BE63AC01A710FED462DD" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="prothoracicum">
<emphasis id="B930EADDFF94BE63AC01A710FED462DD" italics="true" pageId="44" pageNumber="45">O. prothoracicum</emphasis>
</taxonomicName>
) ental membranous faint transverse fold (
<figureCitation id="137F2A4AFF94BE63AA54A739FCA462DD" box="[786,876,1914,1940]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="44" pageNumber="45">Fig. 52</figureCitation>
c). Dorsal process of male cercus much shorter than ventral process, represented by a blunt setose tubercle (
<figureCitation id="137F2A4AFF94BE63AA1DA7E1FC7662F5" box="[859,958,1954,1980]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 89</figureCitation>
; 94; unique). Dorsum of S1 dark brown, posteriorly with a transverse oval greenish blue spot; of S2 orange becoming diffuse brown anteriorly; of S3– 5 pale orange-red; of S6 orange antero-laterally, black dorso-posteriorly; of S7 black with a narrow reddish lateral margin over the anterior three fourths; of S8–9 blue, laterally reddish, both colors separated through a black longitudinal stripe; of S10 black. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF95BE62A980A14CFD896753" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BFB36CFFF95BE62A980A14CFD896753" blockId="45.[151,1436,152,738]" pageId="45" pageNumber="46">
<emphasis id="B930EADDFF95BE62A980A14CFE8E6460" bold="true" box="[198,326,271,297]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
This species, known only from the mature
<typeStatus id="54FF886DFF95BE62AA0CA153FC786463" box="[842,944,272,298]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
male, is unique in that the dorsal process of the cercus is represented by a blunt setose tubercle (
<figureCitation id="137F2A4AFF95BE62AA59A17BFC4A641B" box="[799,898,312,338]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="45" pageNumber="46">Figs. 89</figureCitation>
; 94). In all other species the dorsal process is either long and digit-like (
<taxonomicName id="4C444D4CFF95BE62A889A123FD826430" box="[463,586,352,377]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B930EADDFF95BE62A889A123FD826430" box="[463,586,352,377]" italics="true" pageId="45" pageNumber="46">O. acutum</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62AB1DA123FD756433" box="[603,701,352,378]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="45" pageNumber="46">Figs. 85</figureCitation>
; 90;
<taxonomicName id="4C444D4CFF95BE62ABBFA123FC556430" box="[761,925,352,377]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF95BE62ABBFA123FC556430" box="[761,925,352,377]" italics="true" pageId="45" pageNumber="46">O. thelkterion</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62AAEBA123FBC76433" box="[941,1039,352,378]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="45" pageNumber="46">Figs. 86</figureCitation>
; 91) or acute if small (
<taxonomicName id="4C444D4CFF95BE62AC66A123FF3F64E8" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="prothoracicum">
<emphasis id="B930EADDFF95BE62AC66A123FF3F64E8" italics="true" pageId="45" pageNumber="46">O. prothoracicum</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62A84EA1CBFEA464EB" box="[264,364,392,418]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="45" pageNumber="46">Figs. 87</figureCitation>
; 92;
<taxonomicName id="4C444D4CFF95BE62A8EDA1CBFD9364E8" box="[427,603,392,417]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF95BE62A8EDA1CBFD9364E8" box="[427,603,392,417]" italics="true" pageId="45" pageNumber="46">O. quadricolor</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62AB2BA1CBFD1864EB" box="[621,720,392,418]" captionStart="FIGURES 85 – 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 – 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="45" pageNumber="46">Figs. 88</figureCitation>
a; 93). The distal segment of the genital ligula (
<figureCitation id="137F2A4AFF95BE62AC57A1CBFAA164EB" box="[1297,1385,392,418]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="45" pageNumber="46">Fig. 52</figureCitation>
a) is uniquely armed with a pair of small contiguous latero-apical lobes (a single long latero-apical recurved lobe in
<taxonomicName id="4C444D4CFF95BE62A9D1A19BFEF364B8" box="[151,315,472,497]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF95BE62A9D1A19BFEF364B8" box="[151,315,472,497]" italics="true" pageId="45" pageNumber="46">O. thelkterion</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62A80AA19BFE6B64BB" box="[332,419,472,498]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="45" pageNumber="46">Fig. 49</figureCitation>
b;
<taxonomicName id="4C444D4CFF95BE62A886A19BFDB964B8" box="[448,625,472,497]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF95BE62A886A19BFDB964B8" box="[448,625,472,497]" italics="true" pageId="45" pageNumber="46">O. quadricolor</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62ABC7A19BFD1064BB" box="[641,728,472,498]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="45" pageNumber="46">Fig. 50</figureCitation>
b; and
<taxonomicName id="4C444D4CFF95BE62AA6EA19BFC6C64B8" box="[808,932,472,497]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B930EADDFF95BE62AA6EA19BFC6C64B8" box="[808,932,472,497]" italics="true" pageId="45" pageNumber="46">O. acutum</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62AAF2A19BFBC364BB" box="[948,1035,472,498]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="45" pageNumber="46">Fig. 51</figureCitation>
b; and a single small latero-apical lobe in
<taxonomicName id="4C444D4CFF95BE62A9A9A243FE086750" box="[239,448,512,537]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="prothoracicum">
<emphasis id="B930EADDFF95BE62A9A9A243FE086750" box="[239,448,512,537]" italics="true" pageId="45" pageNumber="46">O. prothoracicum</emphasis>
</taxonomicName>
,
<figureCitation id="137F2A4AFF95BE62A888A243FDEB6753" box="[462,547,512,538]" captionStart="FIGURES 49 – 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 – 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="45" pageNumber="46">Fig. 53</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF95BE62A980A264FC9A670B" box="[198,850,551,578]" pageId="45" pageNumber="46" type="biology_ecology">
<paragraph id="8BFB36CFFF95BE62A980A264FC9A670B" blockId="45.[151,1436,152,738]" box="[198,850,551,578]" pageId="45" pageNumber="46">
<emphasis id="B930EADDFF95BE62A980A264FEE26708" bold="true" box="[198,298,551,577]" pageId="45" pageNumber="46">Biology.</emphasis>
Unknown; see remarks under generic account.
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF95BE62A980A20CFBA667AB" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BFB36CFFF95BE62A980A20CFBA667AB" blockId="45.[151,1436,152,738]" pageId="45" pageNumber="46">
<emphasis id="B930EADDFF95BE62A980A20CFEAF6720" bold="true" box="[198,359,591,617]" pageId="45" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="F353765FFF95BE62A836A213FE2C6723" box="[368,484,592,618]" name="Colombia" pageId="45" pageNumber="46">Colombia</collectingCountry>
(Valle del Cauca Department) at
<quantity id="4CBC9B2AFF95BE62AA31A213FC1A6723" box="[887,978,592,618]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="45" pageNumber="46" unit="m" value="2300.0">2300 m</quantity>
above sea level (
<figureCitation id="137F2A4AFF95BE62ADE2A213FAC56723" box="[1188,1293,592,618]" captionStart="FIGURE 104" captionStartId="38.[151,255,1925,1949]" captionTargetBox="[170,1430,210,1889]" captionTargetId="figure@38.[151,1436,194,1902]" captionTargetPageId="38" captionText="FIGURE 104. Distribution of species of Oreiallagma in South America." httpUri="https://zenodo.org/record/182691/files/figure.png" pageId="45" pageNumber="46">Fig. 104</figureCitation>
). Following information about this locality is provided by
<bibRefCitation id="EFD54B3EFF95BE62AB8CA23BFC8767DB" author="Ris" box="[714,847,632,658]" pageId="45" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris (1918)</bibRefCitation>
: &quot;......Santa Margarita ('situated deep in the high mountain forest at
<quantity id="4CBC9B2AFF95BE62A835A2E3FE0367F3" box="[371,459,672,698]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="45" pageNumber="46" unit="m" value="2200.0">2200 m</quantity>
is the last house Santa Margarita, on upper bank of Rio Cali, the estate of a Colombian general'); out from here the gold mine Monte Socorro is reachable with effort.&quot;
</paragraph>
</subSubSection>
</treatment>
</document>
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