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<document id="284A023B71DF9F3FC6B2630ECBD0EB91" ID-CLB-Dataset="5796" ID-DOI="10.2478/aemnp-2018-0033" ID-GBIF-Dataset="2593a8b2-8a13-4fe0-b5b8-58ea4834527a" ID-ISSN="1804-6487" ID-Zenodo-Dep="4504916" ID-ZooBank="2E9BB0E5-6412-422C-872A-60C2F4BF1D61" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612474442305" checkinUser="felipe" docAuthor="Lackner, Tomáš &amp; Seres, Gábor" docDate="2018" docId="03ED7B67FFE9470AFF49FC84CD85F7F9" docLanguage="en" docName="ActaEntMusNatPra.58.2.419-439.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2)" docSource="http://dx.doi.org/10.2478/aemnp-2018-0033" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Hypocaccus (Nessus) rubripes" docType="treatment" docVersion="8" lastPageNumber="432" masterDocId="FFD4031FFFE04707FFFCFFB9CF64FFEC" masterDocTitle="Revision of the subgenus Nessus of the genus Hypocaccus from Central Europe, with description of a new species (Coleoptera: Histeridae)" masterLastPageNumber="439" masterPageNumber="419" pageNumber="428" updateTime="1698927440595" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="8DF9E33B7FC46D75149F088DBBCE1EB4">Revision of the subgenus Nessus of the genus Hypocaccus from Central Europe, with description of a new species (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="AD58E2A8A2B4F046BE4738DF45DC2304">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="FE4BC090E53C3F25E9C55A8625CD6149">Zoologische Staatssammlung München, Münchhausenstrasse 21, DE- 81247, Germany; e-mail: tomaslackner @ me. com</mods:affiliation>
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<mods:namePart id="7E0E03C5B8EE15341B1A96FE3A0D12B9">Seres, Gábor</mods:namePart>
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<treatment id="03ED7B67FFE9470AFF49FC84CD85F7F9" ID-DOI="http://doi.org/10.5281/zenodo.5062337" ID-GBIF-Taxon="183237533" ID-Zenodo-Dep="5062337" LSID="urn:lsid:plazi:treatment:03ED7B67FFE9470AFF49FC84CD85F7F9" httpUri="http://treatment.plazi.org/id/03ED7B67FFE9470AFF49FC84CD85F7F9" lastPageId="13" lastPageNumber="432" pageId="9" pageNumber="428">
<subSubSection id="C35E99FAFFE9470EFF49FC84CDA0FCB9" box="[181,708,829,853]" pageId="9" pageNumber="428" type="nomenclature">
<paragraph id="8BFBCA71FFE9470EFF49FC84CDA0FCB9" blockId="9.[181,708,829,853]" box="[181,708,829,853]" pageId="9" pageNumber="428">
<heading id="D0B37D1DFFE9470EFF49FC84CDA0FCB9" box="[181,708,829,853]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="428" reason="3">
<taxonomicName id="4C44B1F2FFE9470EFF49FC84CDA0FCB9" ID-CoL="7VHVX" authority="(Erichson, 1834)" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[181,708,829,853]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFE9470EFF49FC84CDA0FCB9" bold="true" box="[181,708,829,853]" pageId="9" pageNumber="428">
<emphasis id="B9301663FFE9470EFF49FC84CE5EFCB8" bold="true" box="[181,314,829,852]" italics="true" pageId="9" pageNumber="428">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFE9470EFEB6FC84CEF3FCB8" bold="true" box="[330,407,829,852]" italics="true" pageId="9" pageNumber="428">Nessus</emphasis>
)
<emphasis id="B9301663FFE9470EFE5BFC84CD67FCB9" bold="true" box="[423,515,829,853]" italics="true" pageId="9" pageNumber="428">rubripes</emphasis>
(
<bibRefCitation id="EFD5B780FFE9470EFDEDFC87CDD9FCB9" author="ERICHSON W. F." box="[529,701,830,853]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFE9470EFE8DFCE5CD7AFC83" box="[369,542,860,879]" pageId="9" pageNumber="428" type="description">
<paragraph id="8BFBCA71FFE9470EFE8DFCE5CD7AFC83" blockId="9.[151,761,860,1313]" box="[369,542,860,879]" pageId="9" pageNumber="428">
(
<figureCitation id="137FD6F4FFE9470EFE8BFCE5CEBDFC83" box="[375,473,860,879]" captionStart-0="Figs 2528" captionStart-1="Figs 2930" captionStart-2="Figs 3132" captionStartId-0="10.[151,188,2030,2049]" captionStartId-1="11.[152,189,972,991]" captionStartId-2="11.[151,188,1619,1638]" captionTargetBox-0="[213,1403,494,1971]" captionTargetBox-1="[168,1435,194,945]" captionTargetBox-2="[172,742,1091,1585]" captionTargetId-0="figure-13@10.[151,1436,456,2005]" captionTargetId-1="figure-480@11.[148,1439,186,959]" captionTargetId-2="figure-518@11.[151,760,1066,1603]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionText-0="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." captionText-1="Figs 2930. Hypocaccus (Nessus) rubripes (Erichson, 1834). 29 Saprinus rubripes var. clermonti Auzat, 1920, holotype, habitus, dorsal view. 30 varieties of Hypocaccus (Nessus) rubripes as summed up by Jean Thérond." captionText-2="Figs 3132. Protibiae, dorsal view. 31 Hypocaccus (Nessus) rubripes (Erichson, 1834); 32 H. (N.) curtus (Rosenhauer, 1847) (both re-drawn from VIENNA 1980)." figureDoi-0="http://doi.org/10.5281/zenodo.4504934" figureDoi-1="http://doi.org/10.5281/zenodo.4504936" figureDoi-2="http://doi.org/10.5281/zenodo.4504938" httpUri-0="https://zenodo.org/record/4504934/files/figure.png" httpUri-1="https://zenodo.org/record/4504936/files/figure.png" httpUri-2="https://zenodo.org/record/4504938/files/figure.png" pageId="9" pageNumber="428">Figs 2531</figureCitation>
,
<figureCitation id="137FD6F4FFE9470EFE1EFCE5CD7DFC83" box="[482,537,860,879]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="9" pageNumber="428">3339</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFE9470EFF6BFCC7CE92FACD" pageId="9" pageNumber="428" type="reference_group">
<paragraph id="8BFBCA71FFE9470EFF6BFCC7CDC7FC7D" blockId="9.[151,761,860,1313]" box="[151,675,894,913]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFCC7CDC7FC7D" box="[151,675,894,913]" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFCC7CE86FC7D" author="ERICHSON W. F." box="[151,482,894,913]" page="193" pageId="9" pageNumber="428" year="1834">
<taxonomicName id="4C44B1F2FFE9470EFF6BFCC7CE86FC7D" ID-CoL="9N3BM" authority="Erichson, 1834: 193" authorityName="Erichson" authorityPageNumber="193" authorityYear="1834" box="[151,482,894,913]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B9301663FFE9470EFF6BFCC7CE54FC7D" box="[151,304,894,913]" italics="true" pageId="9" pageNumber="428">Saprinus rubripes</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFECAFCC7CE86FC7D" author="ERICHSON W. F." box="[310,482,894,913]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834: 193</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFC20CEFFFC0D" blockId="9.[151,761,860,1313]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFC20CEFFFC0D" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFC20CD2FFC40" author="MAZUR S." box="[151,587,921,940]" page="209" pageId="9" pageNumber="428" year="2011">
<taxonomicName id="4C44B1F2FFE9470EFF6BFC20CD46FC40" ID-CoL="7VHVX" authority=": MAZUR (2011)" authorityName=": MAZUR" authorityYear="2011" box="[151,546,921,940]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFE9470EFF6BFC20CF9BFC40" box="[151,255,921,940]" italics="true" pageId="9" pageNumber="428">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFE9470EFEF0FC20CE23FC40" box="[268,327,921,940]" italics="true" pageId="9" pageNumber="428">Nessus</emphasis>
)
<emphasis id="B9301663FFE9470EFEAFFC20CEFFFC40" box="[339,411,921,940]" italics="true" pageId="9" pageNumber="428">rubripes</emphasis>
:
<bibRefCitation id="EFD5B780FFE9470EFE5BFC20CD46FC40" author="MAZUR S." box="[423,546,921,940]" pageId="9" pageNumber="428" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">MAZUR (2011)</bibRefCitation>
</taxonomicName>
: 209
</treatmentCitation>
(new combination);
<treatmentCitation id="0AE5EC60FFE9470EFF49FC0ACEFCFC2A" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[181,408,947,966]" page="118" pageId="9" pageNumber="428" year="2015">
<bibRefCitation id="EFD5B780FFE9470EFF49FC0ACE14FC2A" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[181,368,947,966]" pageId="9" pageNumber="428" refId="ref21168" refString="LACKNER T., MAZUR S. &amp; NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. &amp; LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900." type="book" year="2015">LACKNER et al. (2015)</bibRefCitation>
: 118
</treatmentCitation>
(catalogue). For complete references see
<treatmentCitation id="0AE5EC60FFE9470EFF49FC77CE0EFC0D" author="LACKNER T." box="[181,362,974,993]" page="134" pageId="9" pageNumber="428" year="2010">
<bibRefCitation id="EFD5B780FFE9470EFF49FC77CE25FC0D" author="LACKNER T." box="[181,321,974,993]" pageId="9" pageNumber="428" pagination="1 - 254" refId="ref21136" refString="LACKNER T. 2010: Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 50 (Supplementum): 1 - 254." type="journal article" year="2010">LACKNER (2010)</bibRefCitation>
: 134
</treatmentCitation>
135.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFC50CEBDFBFA" blockId="9.[151,761,860,1313]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFC50CEBDFBFA" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFC50CE90FC10" author="ERICHSON W. F." box="[151,500,1001,1020]" page="191" pageId="9" pageNumber="428" year="1834">
<taxonomicName id="4C44B1F2FFE9470EFF6BFC50CE90FC10" ID-CoL="9N38Z" authority="Erichson, 1834: 191" authorityName="Erichson" authorityPageNumber="191" authorityYear="1834" box="[151,500,1001,1020]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">
<emphasis id="B9301663FFE9470EFF6BFC50CE5AFC10" box="[151,318,1001,1020]" italics="true" pageId="9" pageNumber="428">Saprinus granarius</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFEB9FC50CE90FC10" author="ERICHSON W. F." box="[325,500,1001,1020]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834: 191</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Fauvel in
<treatmentCitation id="0AE5EC60FFE9470EFF37FBBACE01FBFA" author="GOZIS M." box="[203,357,1027,1046]" page="202" pageId="9" pageNumber="428" year="1886">
<bibRefCitation id="EFD5B780FFE9470EFF37FBBACE59FBFA" author="GOZIS M." box="[203,317,1027,1046]" pageId="9" pageNumber="428" pagination="152 - 213" refId="ref20776" refString="GOZIS M. 1886: Les Histeridae Gallo-Rhenans. Tableaux traduits et abreges de l'allemand de Joh. Schmidt. Avec catalogue supplementaire par Albert Fauvel. Revue d'Entomologie 5: 152 - 213." type="journal article" year="1886">GOZIS (1886)</bibRefCitation>
: 202
</treatmentCitation>
(synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFBA7CEE9FBA0" blockId="9.[151,761,860,1313]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFBA7CEE9FBA0" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFBA7CE8EFBDD" author="MARSEUL S. A." box="[151,490,1054,1073]" page="691" pageId="9" pageNumber="428" year="1855">
<taxonomicName id="4C44B1F2FFE9470EFF6BFBA7CE8EFBDD" ID-CoL="9N38F" authority="Marseul, 1855: 691" authorityName="Marseul" authorityPageNumber="691" authorityYear="1855" box="[151,490,1054,1073]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="arenarius">
<emphasis id="B9301663FFE9470EFF6BFBA7CE5FFBDD" box="[151,315,1054,1073]" italics="true" pageId="9" pageNumber="428">Saprinus arenarius</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFEBDFBA7CE8EFBDD" author="MARSEUL S. A." box="[321,490,1054,1073]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855: 691</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<treatmentCitation id="0AE5EC60FFE9470EFD4DFBA7CE7EFBA0" author="SCHMIDT J." page="313" pageId="9" pageNumber="428" year="1885">
<bibRefCitation id="EFD5B780FFE9470EFD4DFBA7CF96FBA0" author="SCHMIDT J." pageId="9" pageNumber="428" pagination="279 - 330" refId="ref21874" refString="SCHMIDT J. 1885: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitschrift 29: 279 - 330." type="journal article" year="1885">SCHMIDT (1885)</bibRefCitation>
: 313
</treatmentCitation>
(synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFBEACEE9FB6D" blockId="9.[151,761,860,1313]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFBEACEE9FB6D" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFBEACEBDFB8A" author="MARSEUL S. A." box="[151,473,1107,1126]" page="688" pageId="9" pageNumber="428" year="1855">
<taxonomicName id="4C44B1F2FFE9470EFF6BFBEACEBDFB8A" ID-CoL="9N38R" authority="Marseul, 1855: 688" authorityName="Marseul" authorityPageNumber="688" authorityYear="1855" box="[151,473,1107,1126]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFE9470EFF6BFBEACE4BFB8A" box="[151,303,1107,1126]" italics="true" pageId="9" pageNumber="428">Saprinus corsicus</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFEC8FBEACEBDFB8A" author="MARSEUL S. A." box="[308,473,1107,1126]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855: 688</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<treatmentCitation id="0AE5EC60FFE9470EFD60FBEACE7EFB6D" author="BICKHARDT H." page="104" pageId="9" pageNumber="428" year="1910">
<bibRefCitation id="EFD5B780FFE9470EFD60FBEACF95FB6D" author="BICKHARDT H." pageId="9" pageNumber="428" refId="ref20474" refString="BICKHARDT H. 1910: Histeridae. In: SCHENKLING S. (ed.): Coleopterorum Catalogus. Pars 24. W. Junk, Berlin, 137 pp." type="book" year="1910">BICKHARDT (1910)</bibRefCitation>
: 104
</treatmentCitation>
(synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFB30CD94FB70" blockId="9.[151,761,860,1313]" box="[151,752,1161,1180]" pageId="9" pageNumber="428">
<treatmentCitationGroup id="AB54ED5FFFE9470EFF6BFB30CD94FB70" box="[151,752,1161,1180]" pageId="9" pageNumber="428">
<treatmentCitation id="0AE5EC60FFE9470EFF6BFB30CD4BFB70" author="AUZAT V." box="[151,559,1161,1180]" page="4" pageId="9" pageNumber="428" year="1920">
<taxonomicName id="4C44B1F2FFE9470EFF6BFB30CD4BFB70" authority="Auzat, 1920: 4" authorityName="Auzat" authorityPageNumber="4" authorityYear="1920" box="[151,559,1161,1180]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="variety" species="rubripes" variety="clermonti">
<emphasis id="B9301663FFE9470EFF6BFB30CE54FB70" box="[151,304,1161,1180]" italics="true" pageId="9" pageNumber="428">Saprinus rubripes</emphasis>
var.
<emphasis id="B9301663FFE9470EFEA7FB30CEC9FB70" box="[347,429,1161,1180]" italics="true" pageId="9" pageNumber="428">clermonti</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFE4DFB30CD4BFB70" author="AUZAT V." box="[433,559,1161,1180]" pageId="9" pageNumber="428" pagination="1 - 4" refId="ref20400" refString="AUZAT V. 1920: Quelques observations sur la sculpture superficielle des Histerides; varietes nouvelles, synonymies. L'Echange, Revue Linneenne 36 (399, Hors texte): 1 - 4." type="book chapter" year="1920">Auzat, 1920: 4</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF6BFB1ACE92FACD" blockId="9.[151,761,860,1313]" pageId="9" pageNumber="428">
<emphasis id="B9301663FFE9470EFF6BFB1ACFA3FB5A" bold="true" box="[151,199,1187,1206]" pageId="9" pageNumber="428">Note.</emphasis>
Complete synonymies and literature references of this species are given in
<bibRefCitation id="EFD5B780FFE9470EFF21FB07CED1FB3D" author="LACKNER T." box="[221,437,1214,1233]" pageId="9" pageNumber="428" pagination="1 - 254" refId="ref21136" refString="LACKNER T. 2010: Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 50 (Supplementum): 1 - 254." type="journal article" year="2010">LACKNER (2010:134135)</bibRefCitation>
and the reader is referred to them there. For the sake of completeness, and the full list of synonymies examined, however, above we list the references of all original descriptions and the works, in which they were synonymised.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFE9470BFF6BFA81CEC5F86C" lastPageId="12" lastPageNumber="431" pageId="9" pageNumber="428" type="materials_examined">
<paragraph id="8BFBCA71FFE9470EFF6BFA81CEA1F8C7" blockId="9.[151,761,1336,2075]" pageId="9" pageNumber="428">
<emphasis id="B9301663FFE9470EFF6BFA81CE13FAA7" bold="true" box="[151,375,1336,1355]" pageId="9" pageNumber="428">Type material examined.</emphasis>
<taxonomicName id="4C44B1F2FFE9470EFE87FA81CDFFFAA7" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[379,667,1336,1355]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B9301663FFE9470EFE87FA81CD70FAA7" box="[379,532,1336,1355]" italics="true" pageId="9" pageNumber="428">Saprinus rubripes</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFDE4FA81CDFFFAA7" author="ERICHSON W. F." box="[536,667,1336,1355]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54FF74D3FFE9470EFD5FFA81CD95FAA5" box="[675,753,1336,1355]" pageId="9" pageNumber="428" type="syntype">SYNTYPES</typeStatus>
: 1 ♁ (
<figureCitation id="137FD6F4FFE9470EFF3DFAEBCE64FA89" box="[193,256,1362,1381]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="9" pageNumber="428">Fig. 25</figureCitation>
), originally pinned with a pin-hole in its right elytron, glued onto a rectangular mounting card, genitalia examined and photographed by the senior author, unfortunately subsequently lost, 49207 [printed] // Hist. -Coll. (
<taxonomicName id="4C44B1F2FFE9470EFEEDFA1BCE15FA59" box="[273,369,1442,1461]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="428" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 49207 /
<taxonomicName id="4C44B1F2FFE9470EFE16FA1BCDFAFA59" authority="Er." authorityName="Erichson" authorityYear="1834" box="[490,670,1442,1461]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">Saprinus rubripes Er.</taxonomicName>
x / Sardin. -Lusitan / Zool.
<collectingRegion id="49800493FFE9470EFEDDFA04CE2EFA3C" box="[289,330,1469,1488]" country="Turkey" name="Mus" pageId="9" pageNumber="428">Mus</collectingRegion>
.
<collectingRegion id="49800493FFE9470EFEAEFA04CEE3FA3C" box="[338,391,1469,1488]" country="Germany" name="Berlin" pageId="9" pageNumber="428">Berlin</collectingRegion>
[black-framed, printed label] //
<typeStatus id="54FF74D3FFE9470EFD64FA04CD9CFA3C" box="[664,760,1469,1488]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
/
<taxonomicName id="4C44B1F2FFE9470EFF5DFA61CE52FA07" authorityName="Erichson" authorityYear="1834" box="[161,310,1496,1515]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">Saprinus rubripes</taxonomicName>
/
<bibRefCitation id="EFD5B780FFE9470EFEB8FA61CEA0FA07" author="ERICHSON W. F." box="[324,452,1496,1515]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
/ labelled by MFNB 2016 [red label, printed] // rubripes / Er. / Lusit. Hoffm [black-framed, written label] (
<collectionCode id="ED5552B4FFE9470EFF61F9B4CFBAF9CC" box="[157,222,1549,1568]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="428">MFNB</collectionCode>
);
<specimenCount id="9D4201F8FFE9470EFF10F9B4CE6DF9CD" box="[236,265,1546,1569]" pageId="9" pageNumber="428" type="female">1♀</specimenCount>
, glued onto a rectangular mounting card, originally pinned with a pin-hole in its right elytron, genitalia extracted and glued onto the same mounting card as the specimen, 22 [written] // 49207 [written] // Hist. -Coll. (
<taxonomicName id="4C44B1F2FFE9470EFEFFF9E4CE07F99C" box="[259,355,1629,1648]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="428" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 49207 /
<taxonomicName id="4C44B1F2FFE9470EFE1FF9E4CDFDF99C" authority="Er." authorityName="Erichson" authorityYear="1834" box="[483,665,1629,1648]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">Saprinus rubripes Er.</taxonomicName>
x / Sardin. -Lusitan / Zool.
<collectingRegion id="49800493FFE9470EFEDDF9C1CE2EF967" box="[289,330,1656,1675]" country="Turkey" name="Mus" pageId="9" pageNumber="428">Mus</collectingRegion>
.
<collectingRegion id="49800493FFE9470EFEAEF9C1CEE3F967" box="[338,391,1656,1675]" country="Germany" name="Berlin" pageId="9" pageNumber="428">Berlin</collectingRegion>
[black-framed, printed label] //
<typeStatus id="54FF74D3FFE9470EFD64F9C1CD9CF967" box="[664,760,1656,1675]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
/
<taxonomicName id="4C44B1F2FFE9470EFF5DF92BCE52F949" authorityName="Erichson" authorityYear="1834" box="[161,310,1682,1701]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">Saprinus rubripes</taxonomicName>
/
<bibRefCitation id="EFD5B780FFE9470EFEB8F92BCEA0F949" author="ERICHSON W. F." box="[324,452,1682,1701]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
/ labelled by MFNB 2016 [red label, printed] (
<collectionCode id="ED5552B4FFE9470EFF15F914CE4EF92C" box="[233,298,1709,1728]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="428">MFNB</collectionCode>
);
<specimenCount id="9D4201F8FFE9470EFECBF914CE30F92D" box="[311,340,1706,1729]" pageId="9" pageNumber="428" type="female">1♀</specimenCount>
, left protibia missing, left mesotarsus missing, left metatibia missing, glued onto a rectangular mounting card with the labels identical to those of the preceding
<typeStatus id="54FF74D3FFE9470EFE3BF95BCD6DF919" box="[455,521,1762,1781]" pageId="9" pageNumber="428" type="syntype">syntype</typeStatus>
(
<collectionCode id="ED5552B4FFE9470EFDEAF95BCD33F919" box="[534,599,1762,1781]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="428">MFNB</collectionCode>
). According to the MFNB staff, the
<typeStatus id="54FF74D3FFE9470EFED6F944CE1CF8FC" box="[298,376,1789,1808]" pageId="9" pageNumber="428" type="lectotype">lectotype</typeStatus>
designation of this species was not allowed, hence no
<typeStatus id="54FF74D3FFE9470EFF15F8A1CE53F8C7" box="[233,311,1816,1835]" pageId="9" pageNumber="428" type="lectotype">lectotype</typeStatus>
was designated.
</paragraph>
<paragraph id="8BFBCA71FFE9470EFF49F88BCB0EFEB0" blockId="9.[151,761,1336,2075]" lastBlockId="9.[827,1438,196,2081]" pageId="9" pageNumber="428">
<taxonomicName id="4C44B1F2FFE9470EFF49F88BCE8DF8A9" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[181,489,1842,1861]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">
<emphasis id="B9301663FFE9470EFF49F88BCE38F8A9" box="[181,348,1842,1861]" italics="true" pageId="9" pageNumber="428">Saprinus granarius</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFE98F88BCE8DF8A9" author="ERICHSON W. F." box="[356,489,1842,1861]" pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54FF74D3FFE9470EFE0FF88BCD5EF8A8" box="[499,570,1842,1861]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
: 1 ♁ (
<figureCitation id="137FD6F4FFE9470EFD8AF88BCDD3F8A9" box="[630,695,1842,1861]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="9" pageNumber="428">Fig. 26</figureCitation>
), originally pinned with a pin-hole in its right elytron, glued onto a rectangular mounting card, left antennal flagellum, left protarsus and right metatibia missing, genitalia extracted, disarticulated and glued to the same mounting card as the specimen, 49201 [printed] // Hist. -Coll. (
<taxonomicName id="4C44B1F2FFE9470EFD7AF824CD82F85C" box="[646,742,1949,1968]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="428" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 49201 /
<taxonomicName id="4C44B1F2FFE9470EFF01F801CED8F827" authority="Er." authorityName="Erichson" authorityYear="1834" box="[253,444,1976,1995]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">Saprinus granarius Er.</taxonomicName>
x /
<collectingCountry id="F3538AE1FFE9470EFE27F801CD78F827" box="[475,540,1976,1995]" name="Austria" pageId="9" pageNumber="428">Austria</collectingCountry>
, Dahl / Zool. Mus. Berlin [black-framed, printed label] //
<typeStatus id="54FF74D3FFE9470EFE6AF86BCE91F809" box="[406,501,2002,2021]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
/
<taxonomicName id="4C44B1F2FFE9470EFDFEF86BCDFFF809" authorityName="Erichson" authorityYear="1834" box="[514,667,2002,2021]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">Saprinus granarius</taxonomicName>
/
<bibRefCitation id="EFD5B780FFE9470EFD54F86BCFA5F7EC" author="ERICHSON W. F." pageId="9" pageNumber="428" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
/ labelled by MFNB 2016 [red label, printed] //
<taxonomicName id="4C44B1F2FFE9470EFDB1F854CDDDF7EC" authority="Er." authorityName="Erichson" authorityYear="1834" box="[589,697,2029,2048]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">granarius Er.</taxonomicName>
/ Austr. Dahl [black-framed, written label] (
<collectionCode id="ED5552B4FFE9470EFE38F7B1CD61F7F7" box="[452,517,2056,2075]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="428">MFNB</collectionCode>
).Another♁, without
<typeStatus id="54FF74D3FFE9470EFD4AF7B1CD9CF7F7" box="[694,760,2056,2075]" pageId="9" pageNumber="428" type="syntype">syntype</typeStatus>
status, identified as
<taxonomicName id="4C44B1F2FFE9470EFC0DFF7DCB24FF3B" authorityName="Erichson" authorityYear="1834" box="[1009,1088,196,215]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
granarius and labelled as Carthagena [pink label, written] //
<taxonomicName id="4C44B1F2FFE9470EFC02FF67CB2DFF1D" authorityName="Erichson" authorityYear="1834" box="[1022,1097,222,241]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ granarius / Er. [written] // Hist. -Coll. (
<taxonomicName id="4C44B1F2FFE9470EFCBDFF40CCC5FEE0" box="[833,929,249,268]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="428" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 49201 /
<taxonomicName id="4C44B1F2FFE9470EFBE4FF40CBB0FEE0" authority="Er." authorityName="Erichson" authorityYear="1834" box="[1048,1236,249,268]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="granarius">Saprinus granarius Er.</taxonomicName>
x /
<collectingCountry id="F3538AE1FFE9470EFB13FF40CA54FEE0" box="[1263,1328,249,268]" name="Austria" pageId="9" pageNumber="428">Austria</collectingCountry>
, Dahl / Zool. Mus. Berlin [black-framed, printed label] (
<collectionCode id="ED5552B4FFE9470EFB4AFEADCB93FECB" box="[1206,1271,276,295]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="428">MFNB</collectionCode>
). According to the MFNB staff, the
<typeStatus id="54FF74D3FFE9470EFC33FE97CB79FEAD" box="[975,1053,302,321]" pageId="9" pageNumber="428" type="lectotype">lectotype</typeStatus>
designation of this species was not allowed, hence no
<typeStatus id="54FF74D3FFE9470EFC71FEF0CCBFFEB0" box="[909,987,329,348]" pageId="9" pageNumber="428" type="lectotype">lectotype</typeStatus>
was designated.
</paragraph>
<paragraph id="8BFBCA71FFE9470EFCA6FEDDCC9FFD70" blockId="9.[827,1438,196,2081]" pageId="9" pageNumber="428">
<taxonomicName id="4C44B1F2FFE9470EFCA6FEDDCB1AFE9B" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[858,1150,356,375]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="arenarius">
<emphasis id="B9301663FFE9470EFCA6FEDDCC98FE9B" box="[858,1020,356,375]" italics="true" pageId="9" pageNumber="428">Saprinus arenarius</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFBFDFEDDCB1AFE9B" author="MARSEUL S. A." box="[1025,1150,356,375]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54FF74D3FFE9470EFB7AFEDDCBBBFE99" box="[1158,1247,356,375]" pageId="9" pageNumber="428" type="lectotype">LECTOTYPE</typeStatus>
(present designation): ♁ (
<figureCitation id="137FD6F4FFE9470EFCABFEC7CCF3FE7D" box="[855,919,382,401]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="9" pageNumber="428">Fig. 27</figureCitation>
), originally pinned, with a pin-hole in its right elytron, glued onto a rectangular mounting card, right metatarsomere missing, genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, 132 /
<taxonomicName id="4C44B1F2FFE9470EFC32FE77CB7DFE0D" authorityName="Erichson" authorityYear="1834" box="[974,1049,462,481]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ arenarius / Dej / Aust. / Dej. [round, written label] //
<taxonomicName id="4C44B1F2FFE9470EFC78FE50CCABFE10" authorityName="Erichson" authorityYear="1834" box="[900,975,489,508]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
132 / arenarius / Dej. / Aust. [yellow, written label] // TYPE [red-printed label] // MUSEUM PARIS / Coll. / DE MARSEUL 1890 [printed] //
<taxonomicName id="4C44B1F2FFE9470EFC2CFDA7CB10FDDD" authorityName="Marseul" authorityYear="1855" box="[976,1140,542,561]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="arenarius">
<emphasis id="B9301663FFE9470EFC2CFDA7CB10FDDD" box="[976,1140,542,561]" italics="true" pageId="9" pageNumber="428">Saprinus arenarius</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFD5B780FFE9470EFB7BFDA7CA60FDDD" author="MARSEUL S. A." box="[1159,1284,542,561]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
/
<typeStatus id="54FF74D3FFE9470EFAEAFDA7CAF4FDDD" box="[1302,1424,542,561]" pageId="9" pageNumber="428" type="lectotype">LECTOTYPE</typeStatus>
/ Des. T. Lackner 2017 [red label, printed] (
<collectionCode id="ED5552B4FFE9470EFB48FD80CB9EFDA0" box="[1204,1274,569,588]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
). This species was described based on unknown number of specimens and existence of other material cannot be excluded therefore we designate the
<typeStatus id="54FF74D3FFE9470EFAE7FDD7CA0DFD6D" box="[1307,1385,622,641]" pageId="9" pageNumber="428" type="lectotype">lectotype</typeStatus>
to fix its taxonomic identity.
</paragraph>
<paragraph id="8BFBCA71FFE9470EFCA6FD1DCBDEFC9D" blockId="9.[827,1438,196,2081]" pageId="9" pageNumber="428">
<taxonomicName id="4C44B1F2FFE9470EFCA6FD1DCB13FD5B" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[858,1143,676,695]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFE9470EFCA6FD1DCC97FD5B" box="[858,1011,676,695]" italics="true" pageId="9" pageNumber="428">Saprinus corsicus</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFC04FD1DCB13FD5B" author="MARSEUL S. A." box="[1016,1143,676,695]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54FF74D3FFE9470EFB83FD1DCBA2FD59" box="[1151,1222,676,695]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
?: ♁ (
<figureCitation id="137FD6F4FFE9470EFB0BFD1DCA53FD5B" box="[1271,1335,676,695]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="9" pageNumber="428">Fig. 28</figureCitation>
), glued to a rectangular mounting card, genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, [small, round golden label, which could be an indication that this was indeed Marseuls type specimen] //
<collectingCountry id="F3538AE1FFE9470EFC4CFCB7CC86FCCD" box="[944,994,782,801]" name="France" pageId="9" pageNumber="428">Corse</collectingCountry>
[printed] // type / Marseul [written] //
<taxonomicName id="4C44B1F2FFE9470EFABFFCB7CAEAFCCD" authorityName="Erichson" authorityYear="1834" box="[1347,1422,782,801]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ granarius [written] //
<taxonomicName id="4C44B1F2FFE9470EFC0EFC90CB59FCD0" authorityName="Erichson" authorityYear="1834" box="[1010,1085,809,828]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ rubripes /
<taxonomicName id="4C44B1F2FFE9470EFB59FC90CA58FCD0" authority=", Mars." authorityName="Mars." box="[1189,1340,809,828]" pageId="9" pageNumber="428" rank="variety" variety="corsicus">v. corsicus, Mars.</taxonomicName>
[written] //
<taxonomicName id="4C44B1F2FFE9470EFCC0FCFDCCB7FCBB" authorityName="Marseul" authorityYear="1855" box="[828,979,836,855]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFE9470EFCC0FCFDCCB7FCBB" box="[828,979,836,855]" italics="true" pageId="9" pageNumber="428">Saprinus corsicus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFD5B780FFE9470EFC1EFCFDCB38FCBB" author="MARSEUL S. A." box="[994,1116,836,855]" pageId="9" pageNumber="428" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
/
<typeStatus id="54FF74D3FFE9470EFB97FCFDCBAFFCBB" box="[1131,1227,836,855]" pageId="9" pageNumber="428" type="syntype">SYNTYPE</typeStatus>
? / Des. T. Lackner 2017 [red label, printed] (
<collectionCode id="ED5552B4FFE9470EFC10FCE7CB55FC9D" box="[1004,1073,862,881]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond).
</paragraph>
<paragraph id="8BFBCA71FFE9470EFCA6FCC0CAF1FB8D" blockId="9.[827,1438,196,2081]" pageId="9" pageNumber="428">
<taxonomicName id="4C44B1F2FFE9470EFCA6FCC0CB8BFC60" authority="Auzat, 1920" authorityName="Auzat" authorityYear="1920" box="[858,1263,889,908]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="variety" species="rubripes" variety="clermonti">
<emphasis id="B9301663FFE9470EFCA6FCC0CC93FC60" box="[858,1015,889,908]" italics="true" pageId="9" pageNumber="428">Saprinus rubripes</emphasis>
var.
<emphasis id="B9301663FFE9470EFBDBFCC0CB1DFC60" box="[1063,1145,889,908]" italics="true" pageId="9" pageNumber="428">clermonti</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFB7CFCC0CB8BFC60" author="AUZAT V." box="[1152,1263,889,908]" pageId="9" pageNumber="428" pagination="1 - 4" refId="ref20400" refString="AUZAT V. 1920: Quelques observations sur la sculpture superficielle des Histerides; varietes nouvelles, synonymies. L'Echange, Revue Linneenne 36 (399, Hors texte): 1 - 4." type="book chapter" year="1920">Auzat, 1920</bibRefCitation>
</taxonomicName>
:
<typeStatus id="54FF74D3FFE9470EFB05FCC0CA29FC66" box="[1273,1357,889,908]" pageId="9" pageNumber="428" type="holotype">HOLOTYPE</typeStatus>
: ♁ (
<figureCitation id="137FD6F4FFE9470EFA85FCC0CC30FC4B" captionStart="Figs 2930" captionStartId="11.[152,189,972,991]" captionTargetBox="[168,1435,194,945]" captionTargetId="figure-480@11.[148,1439,186,959]" captionTargetPageId="11" captionText="Figs 2930. Hypocaccus (Nessus) rubripes (Erichson, 1834). 29 Saprinus rubripes var. clermonti Auzat, 1920, holotype, habitus, dorsal view. 30 varieties of Hypocaccus (Nessus) rubripes as summed up by Jean Thérond." figureDoi="http://doi.org/10.5281/zenodo.4504936" httpUri="https://zenodo.org/record/4504936/files/figure.png" pageId="9" pageNumber="428">Fig. 29</figureCitation>
), right antennal funicle, two left metatarsomeres missing, glued onto a rectangular mounting card, genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, Arcachon / Cap Ferret [written] // var./ Clermonti / Type / Dr.Auzat det. 1920 [printed-written] // Coll./ Dr.Auzat [light-green label, written] // TYPE [red label, printed] //
<taxonomicName id="4C44B1F2FFE9470EFCB0FBA0CC81FBC0" authorityName="Erichson" authorityYear="1834" box="[844,997,1049,1068]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B9301663FFE9470EFCB0FBA0CC81FBC0" box="[844,997,1049,1068]" italics="true" pageId="9" pageNumber="428">Saprinus rubripes</emphasis>
</taxonomicName>
var. /
<taxonomicName id="4C44B1F2FFE9470EFBE5FBA0CBB2FBC0" authority="Auzat, 1920" authorityName="Auzat" authorityYear="1920" box="[1049,1238,1049,1068]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="clermonti">
<emphasis id="B9301663FFE9470EFBE5FBA0CB0FFBC0" box="[1049,1131,1049,1068]" italics="true" pageId="9" pageNumber="428">clermonti</emphasis>
<bibRefCitation id="EFD5B780FFE9470EFB92FBA0CBB2FBC0" author="AUZAT V." box="[1134,1238,1049,1068]" pageId="9" pageNumber="428" pagination="1 - 4" refId="ref20400" refString="AUZAT V. 1920: Quelques observations sur la sculpture superficielle des Histerides; varietes nouvelles, synonymies. L'Echange, Revue Linneenne 36 (399, Hors texte): 1 - 4." type="book chapter" year="1920">Auzat, 1920</bibRefCitation>
</taxonomicName>
/
<typeStatus id="54FF74D3FFE9470EFB1AFBA0CA33FBC0" box="[1254,1367,1049,1068]" pageId="9" pageNumber="428" type="holotype">HOLOTYPE</typeStatus>
/ Des. t. Lackner 2017 [red label, printed] (
<collectionCode id="ED5552B4FFE9470EFB90FB8DCBD6FBAB" box="[1132,1202,1076,1095]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
). This taxon was described based on a single specimen, which is therefore
<typeStatus id="54FF74D3FFE9470EFB32FBF7CA7DFB8D" box="[1230,1305,1102,1121]" pageId="9" pageNumber="428" type="holotype">holotype</typeStatus>
by monotypy.
</paragraph>
<paragraph id="8BFBCA71FFE9470CFCC0FBD0CBF8FADF" blockId="9.[827,1438,196,2081]" lastBlockId="11.[828,1438,1066,1331]" lastPageId="11" lastPageNumber="430" pageId="9" pageNumber="428">
<emphasis id="B9301663FFE9470EFCC0FBD0CA0FFB90" bold="true" box="[828,1387,1129,1148]" pageId="9" pageNumber="428">
Additional material examined.
<collectingCountry id="F3538AE1FFE9470EFB9DFBD0CBADFB90" box="[1121,1225,1129,1148]" name="Algeria" pageId="9" pageNumber="428">ALGERIA</collectingCountry>
:
<collectingRegion id="49800493FFE9470EFB29FBD0CA7EFB96" box="[1237,1306,1129,1148]" country="Algeria" name="Bechar" pageId="9" pageNumber="428">BÉCHAR</collectingRegion>
REGION:
</emphasis>
Bèni Abbès, Sahara,
<date id="FFFAECB1FFE9470EFC38FB3DCB7EFB7B" box="[964,1050,1156,1175]" pageId="9" pageNumber="428" value="1980-10-20">20.x.1980</date>
, 1 ♁, A. Olexa lgt. (
<collectionCode id="ED5552B4FFE9470EFB2DFB3DCA69FB7B" box="[1233,1293,1156,1175]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFAE1FB3DCAF8FB7B" bold="true" box="[1309,1436,1156,1175]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFAE1FB3DCA29FB79" box="[1309,1357,1156,1175]" name="Algeria" pageId="9" pageNumber="428">ORAN</collectingCountry>
REGION:
</emphasis>
Oran,no further data,1♁ (
<collectionCode id="ED5552B4FFE9470EFBE9FB27CB3FFB5D" box="[1045,1115,1182,1201]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="9" pageNumber="428">MMBC</collectionCode>
).
<emphasis id="B9301663FFE9470EFB9AFB27CAF8FB5D" bold="true" box="[1126,1436,1182,1201]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFB9AFB27CB96FB5D" box="[1126,1266,1182,1201]" name="Azerbaijan" pageId="9" pageNumber="428">AZERBAIJAN</collectingCountry>
:
<collectingRegion id="49800493FFE9470EFB05FB27CA36FB5C" box="[1273,1362,1182,1201]" country="Azerbaijan" name="Lankaran" pageId="9" pageNumber="428">LANKARAN</collectingRegion>
REGION:
</emphasis>
Mamusta env.,
<date id="FFFAECB1FFE9470EFC46FB00CB6AFB20" box="[954,1038,1209,1228]" pageId="9" pageNumber="428" value="2001-05-12">12.v.2001</date>
,
<specimenCount id="9D4201F8FFE9470EFBE9FB00CB36FB20" box="[1045,1106,1209,1228]" pageId="9" pageNumber="428" type="generic">5 spec.</specimenCount>
, T. Lackner lgt. (
<collectionCode id="ED5552B4FFE9470EFB27FB00CA72FB20" box="[1243,1302,1209,1228]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFAD8FB00CB65FB0B" bold="true" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFAD8FB00CAFDFB20" box="[1316,1433,1209,1228]" name="Bulgaria" pageId="9" pageNumber="428">BULGARIA</collectingCountry>
:
<collectingRegion id="49800493FFE9470EFCC0FB6DCCD1FB09" box="[828,949,1236,1255]" country="Bulgaria" name="Blagoevgrad" pageId="9" pageNumber="428">BLAGOEVGRAD</collectingRegion>
REGION:
</emphasis>
Struma River valley, Sandanski,
<date id="FFFAECB1FFE9470EFAE7FB6DCAFDFB0B" box="[1307,1433,1236,1255]" pageId="9" pageNumber="428" value="1987-04-21" valueMax="1987-04-22" valueMin="1987-04-21">21.22.iv.1987</date>
,
<specimenCount id="9D4201F8FFE9470EFCC0FB57CC1EFAED" box="[828,890,1262,1281]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, J. Mertlik lgt. (
<collectionCode id="ED5552B4FFE9470EFBF8FB57CB5BFAED" box="[1028,1087,1262,1281]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFBB2FB57CBB9FAED" bold="true" box="[1102,1245,1262,1281]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFBB2FB57CBB2FAEC" box="[1102,1238,1262,1281]" country="Bulgaria" name="Burgas" pageId="9" pageNumber="428">BURGAS REGION</collectingRegion>
:
</emphasis>
Burgas, on the beach,
<date id="FFFAECB1FFE9470EFCC0FAB0CCFCFAF0" box="[828,920,1289,1308]" pageId="9" pageNumber="428" value="1981-08-05">5.viii.1981</date>
,
<specimenCount id="9D4201F8FFE9470EFC5EFAB0CCA7FAF0" box="[930,963,1285,1308]" pageId="9" pageNumber="428" type="female">1 ♀</specimenCount>
, collector unknown (
<collectionCode id="ED5552B4FFE9470EFB86FAB0CBA0FAF0" box="[1146,1220,1289,1308]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="428" type="Museum">NHMW</collectionCode>
); Nessebar,
<date id="FFFAECB1FFE9470EFAD7FAB0CAE3FAF0" box="[1323,1415,1289,1308]" pageId="9" pageNumber="428" value="1994-08-04">4.viii.1994</date>
,
<specimenCount id="9D4201F8FFE9470EFA6EFAB0CC0DFADB" pageId="9" pageNumber="428" type="generic">2 spec.</specimenCount>
,
<date id="FFFAECB1FFE9470EFC8FFA9DCCBDFADB" box="[883,985,1316,1335]" pageId="9" pageNumber="428" value="1996-08-22">22.viii.1996</date>
,
<specimenCount id="9D4201F8FFE9470EFC18FA9DCB40FADB" box="[996,1060,1316,1335]" pageId="9" pageNumber="428" type="generic">4 spec.</specimenCount>
, T. Lackner lgt. (
<collectionCode id="ED5552B4FFE9470EFB46FA9DCB91FADB" box="[1210,1269,1316,1335]" pageId="9" pageNumber="428">CTLA</collectionCode>
); Arkutino,
<date id="FFFAECB1FFE9470EFAA1FA9DCC19FABD" pageId="9" pageNumber="428" value="1988-09-14" valueMax="1988-09-16" valueMin="1988-09-14">14.16. ix.1988</date>
,
<specimenCount id="9D4201F8FFE9470EFC79FA87CCA5FABD" box="[901,961,1342,1361]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, J. Růžička lgt. (
<collectionCode id="ED5552B4FFE9470EFBB4FA87CBE7FABD" box="[1096,1155,1342,1361]" pageId="9" pageNumber="428">CTLA</collectionCode>
); Sozopol env.,
<date id="FFFAECB1FFE9470EFAFAFA87CA31FABD" box="[1286,1365,1342,1361]" pageId="9" pageNumber="428" value="2014-04-01">1.iv.2014</date>
,
<specimenCount id="9D4201F8FFE9470EFAA2FA87CAFEFABD" box="[1374,1434,1342,1361]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, P.Kylies lgt. (
<collectionCode id="ED5552B4FFE9470EFC52FAE0CC8DFA80" box="[942,1001,1369,1388]" pageId="9" pageNumber="428">CTLA</collectionCode>
); Primorsko,
<date id="FFFAECB1FFE9470EFBABFAE0CBF9FA80" box="[1111,1181,1369,1388]" pageId="9" pageNumber="428" value="1980-07">vii.1980</date>
,
<specimenCount id="9D4201F8FFE9470EFB59FAE0CBA6FA80" box="[1189,1218,1365,1388]" pageId="9" pageNumber="428" type="female">1♀</specimenCount>
, J. Pokorný lgt.(
<collectionCode id="ED5552B4FFE9470EFAB1FAE0CAF7FA80" box="[1357,1427,1369,1388]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="9" pageNumber="428">MMBC</collectionCode>
).
<emphasis id="B9301663FFE9470EFCC0FACDCCA0FA6B" bold="true" box="[828,964,1396,1415]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFCC0FACDCCD9FA69" box="[828,957,1396,1415]" country="Bulgaria" name="Varna" pageId="9" pageNumber="428">VARNA REGION</collectingRegion>
:
</emphasis>
Škorpilovici,
<date id="FFFAECB1FFE9470EFBBCFACDCBE3FA6B" box="[1088,1159,1396,1415]" pageId="9" pageNumber="428" value="1983-07">vii.1983</date>
,
<specimenCount id="9D4201F8FFE9470EFB6EFACDCBD7FA6B" box="[1170,1203,1392,1415]" pageId="9" pageNumber="428" type="female">1 ♀</specimenCount>
, J. Pokorný lgt. (
<collectionCode id="ED5552B4FFE9470EFAB1FACDCAF7FA6B" box="[1357,1427,1396,1415]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="9" pageNumber="428">MMBC</collectionCode>
).
<emphasis id="B9301663FFE9470EFCC0FA37CCFFFA4D" bold="true" box="[828,923,1422,1441]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFCC0FA37CCF3FA4D" box="[828,919,1422,1441]" name="France" pageId="9" pageNumber="428">FRANCE</collectingCountry>
:
</emphasis>
Gallia, no further data,
<specimenCount id="9D4201F8FFE9470EFB95FA37CBE8FA4E" box="[1129,1164,1419,1442]" pageId="9" pageNumber="428" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5552B4FFE9470EFB6BFA37CBB9FA4D" box="[1175,1245,1422,1441]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="9" pageNumber="428">MMBC</collectionCode>
).
<emphasis id="B9301663FFE9470EFB10FA37CAF8FA4D" bold="true" box="[1260,1436,1422,1441]" pageId="9" pageNumber="428">
BOUCHES-
<subScript id="17C0C834FFE9470EFAC3FA2ACA3EFA4D" attach="none" box="[1343,1370,1422,1441]" fontSize="6" pageId="9" pageNumber="428">DU-</subScript>
RHÔNE:
</emphasis>
St.Maries, Camargue,
<date id="FFFAECB1FFE9470EFC04FA10CB26FA50" box="[1016,1090,1449,1468]" pageId="9" pageNumber="428" value="1928-10-09">9.x.1928</date>
,
<specimenCount id="9D4201F8FFE9470EFBB6FA10CB03FA50" box="[1098,1127,1445,1468]" pageId="9" pageNumber="428" type="female">1♀</specimenCount>
, L.Puel lgt. (
<collectionCode id="ED5552B4FFE9470EFB29FA10CA7DFA50" box="[1237,1305,1449,1468]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond); Grau du Roi,
<date id="FFFAECB1FFE9470EFC54FA7DCC99FA3B" box="[936,1021,1476,1495]" pageId="9" pageNumber="428" value="1939-02-10">10.ii.1939</date>
, 1♁,
<specimenCount id="9D4201F8FFE9470EFBD6FA7DCB22FA3B" box="[1066,1094,1472,1495]" pageId="9" pageNumber="428" type="female">1♀</specimenCount>
,
<date id="FFFAECB1FFE9470EFBACFA7DCBCEFA3B" box="[1104,1194,1476,1495]" pageId="9" pageNumber="428" value="1933-06-19">19.vi.1933</date>
, 1♁, J.Thérond lgt.(
<collectionCode id="ED5552B4FFE9470EFAA9FA7DCAFDFA3B" box="[1365,1433,1476,1495]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond).
<emphasis id="B9301663FFE9470EFC40FA67CC9EFA1D" bold="true" box="[956,1018,1502,1521]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFC40FA67CC97FA1C" box="[956,1011,1502,1521]" name="France" pageId="9" pageNumber="428">CORSE</collectingCountry>
:
</emphasis>
Oletta,
<date id="FFFAECB1FFE9470EFBC7FA67CBC6FA1D" box="[1083,1186,1502,1521]" pageId="9" pageNumber="428" value="1981-08-13">13.viii.1981</date>
,
<specimenCount id="9D4201F8FFE9470EFB56FA67CBACFA1E" box="[1194,1224,1499,1522]" pageId="9" pageNumber="428" type="female">1 ♀</specimenCount>
, Wewalka lgt. (
<collectionCode id="ED5552B4FFE9470EFAB5FA67CAF7FA1D" box="[1353,1427,1502,1521]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="428" type="Museum">NHMW</collectionCode>
); Bonifacio, Révélière,
<specimenCount id="9D4201F8FFE9470EFC04FA40CB7DF9E0" box="[1016,1049,1525,1548]" pageId="9" pageNumber="428" type="female">1 ♀</specimenCount>
, Col. A. Grouvelle (
<collectionCode id="ED5552B4FFE9470EFB32FA40CA76F9E0" box="[1230,1298,1529,1548]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
, coll. Thérond); Corse, 1 ♁,
<specimenCount id="9D4201F8FFE9470EFC54F9ADCCBDF9CB" box="[936,985,1552,1575]" pageId="9" pageNumber="428" type="female">2 ♀♀</specimenCount>
, coll. Croissandeau (
<collectionCode id="ED5552B4FFE9470EFB6CF9ADCBB0F9CB" box="[1168,1236,1556,1575]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond); Corse, 1 ♁, coll. Bonnaire (
<collectionCode id="ED5552B4FFE9470EFC10F997CB54F9AD" box="[1004,1072,1582,1601]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond).
<emphasis id="B9301663FFE9470EFB42F997CA75F9AD" bold="true" box="[1214,1297,1582,1601]" pageId="9" pageNumber="428">HÉRAULT:</emphasis>
Sète, no further data, 1 ♁ (
<collectionCode id="ED5552B4FFE9470EFC65F9F0CCB9F9B0" box="[921,989,1609,1628]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="428">MNHN</collectionCode>
; coll. Thérond).
<emphasis id="B9301663FFE9470EFB90F9F0CA76F9B0" bold="true" box="[1132,1298,1609,1628]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFB90F9F0CBACF9B0" box="[1132,1224,1609,1628]" name="Greece" pageId="9" pageNumber="428">GREECE</collectingCountry>
: CORFU:
</emphasis>
Acharawi west,
<date id="FFFAECB1FFE9470EFCC0F9DDCCF3F99B" box="[828,919,1636,1655]" pageId="9" pageNumber="428" value="2017-06-24">24.vi.2017</date>
,
<specimenCount id="9D4201F8FFE9470EFC5CF9DDCCBAF99B" box="[928,990,1636,1655]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, O. Majzlan lgt. (
<collectionCode id="ED5552B4FFE9470EFB8EF9DDCBC9F99B" box="[1138,1197,1636,1655]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFB40F9DDCA71F99B" bold="true" box="[1212,1301,1636,1655]" pageId="9" pageNumber="428">KAVAVALA:</emphasis>
Thassos Island, SW Potos env., saline,
<date id="FFFAECB1FFE9470EFBF5F9C7CB08F97D" box="[1033,1132,1662,1681]" pageId="9" pageNumber="428" value="2004-07-10">10.vii.2004</date>
,
<specimenCount id="9D4201F8FFE9470EFB8BF9C7CBD3F97D" box="[1143,1207,1662,1681]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, P. Bulirsch lgt., (
<collectionCode id="ED5552B4FFE9470EFAABF9C7CAF7F97D" box="[1367,1427,1662,1681]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFCC0F920CCF0F940" bold="true" box="[828,916,1689,1708]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFCC0F920CCE9F946" box="[828,909,1689,1708]" country="Greece" name="Thessalia" pageId="9" pageNumber="428">THESSALY</collectingRegion>
:
</emphasis>
Leptokaria,
<date id="FFFAECB1FFE9470EFBFCF920CBC1F940" box="[1024,1189,1689,1708]" pageId="9" pageNumber="428" value="1993-07-30" valueMax="1993-08-15" valueMin="1993-07-30">30.vii.15.viii.1993</date>
, 2 ♁♁, J. Háva lgt. (
<collectionCode id="ED5552B4FFE9470EFAA4F920CAF0F940" box="[1368,1428,1689,1708]" pageId="9" pageNumber="428">CTLA</collectionCode>
).
<emphasis id="B9301663FFE9470EFCC0F90DCB00F92B" bold="true" box="[828,1124,1716,1735]" pageId="9" pageNumber="428">
<collectingCountry id="F3538AE1FFE9470EFCC0F90DCCCDF92B" box="[828,937,1716,1735]" name="Hungary" pageId="9" pageNumber="428">HUNGARY</collectingCountry>
: BÁCS- KISKUN MEGYE:
</emphasis>
Kalocsa, no further data, 1 ♁, Speiser (
<collectionCode id="ED5552B4FFE9470EFCBEF977CCE6F90D" box="[834,898,1742,1761]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="428">NMPC</collectionCode>
);Bócsa,in sand-hills,
<date id="FFFAECB1FFE9470EFBC4F977CBF7F90D" box="[1080,1171,1742,1761]" pageId="9" pageNumber="428" value="1956-06-17">17.vi.1956</date>
,
<specimenCount id="9D4201F8FFE9470EFB66F977CBB2F90D" box="[1178,1238,1742,1761]" pageId="9" pageNumber="428" type="generic">3 spec.</specimenCount>
, Kaszab &amp; Székessy lgt. (
<collectionCode id="ED5552B4FFE9470EFCBEF950CCE3F910" box="[834,903,1769,1788]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Kéleshalom,
<date id="FFFAECB1FFE9470EFBF8F950CB21F910" box="[1028,1093,1769,1788]" pageId="9" pageNumber="428" value="1955-06">vi.1955</date>
,
<specimenCount id="9D4201F8FFE9470EFBB2F950CBE8F910" box="[1102,1164,1769,1788]" pageId="9" pageNumber="428" type="generic">7 spec.</specimenCount>
, Dr. Lenci lgt. (
<collectionCode id="ED5552B4FFE9470EFAECF950CA31F910" box="[1296,1365,1769,1788]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Bócsa,
<date id="FFFAECB1FFE9470EFCC0F8BDCCF3F8FB" box="[828,919,1796,1815]" pageId="9" pageNumber="428" value="1956-06-17">17.vi.1956</date>
,
<specimenCount id="9D4201F8FFE9470EFC5CF8BDCC8CF8FB" box="[928,1000,1796,1815]" pageId="9" pageNumber="428" type="generic">10 spec.</specimenCount>
, Dr. Lenci lgt. (
<collectionCode id="ED5552B4FFE9470EFB90F8BDCBD5F8FB" box="[1132,1201,1796,1815]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Kiskunsági National Park, Fülöpháza sand-hills,
<date id="FFFAECB1FFE9470EFC0EF8A7CB2AF8DD" box="[1010,1102,1822,1841]" pageId="9" pageNumber="428" value="1978-06-22">22.vi.1978</date>
,
<specimenCount id="9D4201F8FFE9470EFBAAF8A7CBF5F8DD" box="[1110,1169,1822,1841]" pageId="9" pageNumber="428" type="generic">9 spec.</specimenCount>
(pitfall trap baited with cheese), Ádám &amp; Hámori lgt. (
<collectionCode id="ED5552B4FFE9470EFC0CF880CB51F8A0" box="[1008,1077,1849,1868]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Kalocsa,
<specimenCount id="9D4201F8FFE9470EFB73F880CBAFF8A0" box="[1167,1227,1849,1868]" pageId="9" pageNumber="428" type="generic">4 spec.</specimenCount>
, Speiser lgt., coll. Speiser (
<collectionCode id="ED5552B4FFE9470EFCBDF8EDCCE1F88B" box="[833,901,1876,1895]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
);Soltvadkert,
<quantity id="4CBC6794FFE9470EFC05F8EDCB49F88B" box="[1017,1069,1876,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="428" unit="m" value="100.0">100 m</quantity>
,sandy pasture,
<date id="FFFAECB1FFE9470EFB57F8EDCA62F88B" box="[1195,1286,1876,1895]" pageId="9" pageNumber="428" value="1975-03-31">31.iii.1975</date>
,
<specimenCount id="9D4201F8FFE9470EFAF0F8EDCA21F88B" box="[1292,1349,1876,1895]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
(from cow dung), O. Merkl lgt. (
<collectionCode id="ED5552B4FFE9470EFC17F8D7CB54F86D" box="[1003,1072,1902,1921]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
).
<emphasis id="B9301663FFE9470EFBC0F8D7CBB3F86D" bold="true" box="[1084,1239,1902,1921]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFBC0F8D7CBB4F86C" box="[1084,1232,1902,1921]" country="Hungary" name="Csongrad" pageId="9" pageNumber="428">CSONGRÁD MEGYE</collectingRegion>
:
</emphasis>
Nagyszéksós,
<date id="FFFAECB1FFE9470EFAAEF8D7CAFDF86D" box="[1362,1433,1902,1921]" pageId="9" pageNumber="428" value="1922-07">vii.1922</date>
,
<specimenCount id="9D4201F8FFE9470EFCC0F830CC13F870" box="[828,887,1929,1948]" pageId="9" pageNumber="428" type="generic">2 spec.</specimenCount>
, Szabó-Patay (
<collectionCode id="ED5552B4FFE9470EFC11F830CB56F870" box="[1005,1074,1929,1948]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
).
<emphasis id="B9301663FFE9470EFBC1F830CBD5F870" bold="true" box="[1085,1201,1929,1948]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFBC1F830CBCEF876" box="[1085,1194,1929,1948]" country="Hungary" name="Fejer" pageId="9" pageNumber="428">FEJÉR MEGYE</collectingRegion>
:
</emphasis>
Budapest env., Martonvásár, no date,
<specimenCount id="9D4201F8FFE9470EFC82F81DCCDCF85B" box="[894,952,1956,1975]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, H.Diener coll.(
<collectionCode id="ED5552B4FFE9470EFBC1F81DCBE5F85B" box="[1085,1153,1956,1975]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
).
<emphasis id="B9301663FFE9470EFB70F81DCB91F85B" bold="true" box="[1164,1269,1956,1975]" pageId="9" pageNumber="428">
<collectingRegion id="49800493FFE9470EFB70F81DCB89F859" box="[1164,1261,1956,1975]" country="Hungary" name="Pest" pageId="9" pageNumber="428">PEST MEGYE</collectingRegion>
:
</emphasis>
Budapest, no further data, 1 ♁, Gammel lgt. (
<collectionCode id="ED5552B4FFE9470EFBF9F807CB2FF83D" box="[1029,1099,1982,2001]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="9" pageNumber="428">MMBC</collectionCode>
); Budapest,
<date id="FFFAECB1FFE9470EFB4DF807CA77F83D" box="[1201,1299,1982,2001]" pageId="9" pageNumber="428" value="1934-12-12">12.xii.1934</date>
,
<specimenCount id="9D4201F8FFE9470EFAE6F807CA33F83D" box="[1306,1367,1982,2001]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
, Kaszab lgt. (
<collectionCode id="ED5552B4FFE9470EFC9DF860CCC2F800" box="[865,934,2009,2028]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Budapest, no further data,
<specimenCount id="9D4201F8FFE9470EFB71F860CBACF800" box="[1165,1224,2009,2028]" pageId="9" pageNumber="428" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED5552B4FFE9470EFB2CF860CA71F800" box="[1232,1301,2009,2028]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Budapest env., Újpest-Alag, no date,
<specimenCount id="9D4201F8FFE9470EFC0AF84DCB50F7EB" box="[1014,1076,2036,2055]" pageId="9" pageNumber="428" type="generic">2 spec.</specimenCount>
, coll. H. Diener (
<collectionCode id="ED5552B4FFE9470EFB3BF84DCA68F7EB" box="[1223,1292,2036,2055]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Budapest, Pest, no further data,
<specimenCount id="9D4201F8FFE9470EFC3CF7B7CC98F7CD" box="[960,1020,2062,2081]" pageId="9" pageNumber="428" type="generic">3 spec.</specimenCount>
, Gimmel lgt. (
<collectionCode id="ED5552B4FFE9470EFB8EF7B7CBD3F7CD" box="[1138,1207,2062,2081]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="428">HNHM</collectionCode>
); Budapest, no further data,
<specimenCount id="9D4201F8FFEA470DFF6BFF07CFB3FF3D" box="[151,215,190,209]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
, Kuthy lgt. (
<collectionCode id="ED5552B4FFEA470DFEB9FF07CEEEFF3D" box="[325,394,190,209]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Pest m., Pusztavacs, Strázsa hill,
<quantity id="4CBC6794FFEA470DFD47FF07CD91FF3D" box="[699,757,190,209]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="429" unit="m" value="100.0">100 m</quantity>
, Festucetum vaginatae danubiale, from plant debris,
<date id="FFFAECB1FFEA470DFDB6FF60CDC0FF00" box="[586,676,217,236]" pageId="10" pageNumber="429" value="1990-06-29">29.vi.1990</date>
,
<specimenCount id="9D4201F8FFEA470DFD52FF60CD92FF00" box="[686,758,217,236]" pageId="10" pageNumber="429" type="generic">30 spec.</specimenCount>
, L. Ádám lgt. (
<collectionCode id="ED5552B4FFEA470DFEF2FF4ACE37FEEA" box="[270,339,243,262]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Táborfalva,
<date id="FFFAECB1FFEA470DFE34FF4ACD7CFEEA" box="[456,536,243,262]" pageId="10" pageNumber="429" value="2015-06-05">5.vi.2015</date>
,
<geoCoordinate id="EE70ACB6FFEA470DFDDDFF4ACD1EFEEA" box="[545,634,243,262]" degrees="47" direction="north" minutes="3" orientation="latitude" pageId="10" pageNumber="429" precision="15" seconds="39" value="47.060833">47º339″N</geoCoordinate>
,
<geoCoordinate id="EE70ACB6FFEA470DFD7FFF4ACD80FEEA" box="[643,740,243,262]" degrees="19" direction="east" minutes="27" orientation="longitude" pageId="10" pageNumber="429" precision="15" seconds="36" value="19.460001">19º2736″E</geoCoordinate>
,
<specimenCount id="9D4201F8FFEA470DFD11FF4ACFA7FECD" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
(car-netting on the driving course) (
<collectionCode id="ED5552B4FFEA470DFE01FEB7CD26FECD" box="[509,578,270,289]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Nagykőrös, Csókás forest,
<date id="FFFAECB1FFEA470DFF2EFE90CE21FED0" box="[210,325,297,316]" pageId="10" pageNumber="429" value="2010-04-04" valueMax="2010-05-09" valueMin="2010-04-04">4.iv9.v.2010</date>
,
<specimenCount id="9D4201F8FFEA470DFEB3FE90CEE8FED0" box="[335,396,297,316]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
(on sand in the forest),
<date id="FFFAECB1FFEA470DFDA4FE90CD87FED0" box="[600,739,297,316]" pageId="10" pageNumber="429" value="2010-08-10" valueMax="2010-08-27" valueMin="2010-08-10">10.27.viii.2010</date>
,
<specimenCount id="9D4201F8FFEA470DFD11FE90CFA7FEBA" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
(on sand in the forest), Tallósi lgt. (
<collectionCode id="ED5552B4FFEA470DFE1FFEFACD4CFEBA" box="[483,552,323,342]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
);Táborfalva,
<date id="FFFAECB1FFEA470DFD65FEFACD91FEBA" box="[665,757,323,342]" pageId="10" pageNumber="429" value="2012-06-30">30.vi.2012</date>
,
<specimenCount id="9D4201F8FFEA470DFF6BFEE7CFB6FE9D" box="[151,210,350,369]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
(car-netting on the driving course), O. Merkl lgt. (
<collectionCode id="ED5552B4FFEA470DFD8EFEE7CDD3FE9D" box="[626,695,350,369]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Budapest-Rákos, no date,
<specimenCount id="9D4201F8FFEA470DFEB6FEC0CEECFE60" box="[330,392,377,396]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
, H. Diener coll. (
<collectionCode id="ED5552B4FFEA470DFDE0FEC0CD05FE60" box="[540,609,377,396]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Óbuda,
<date id="FFFAECB1FFEA470DFD49FEC0CD91FE60" box="[693,757,377,396]" pageId="10" pageNumber="429" value="1903-04">iv.1903</date>
,
<specimenCount id="9D4201F8FFEA470DFF6BFE2ACFBCFE4A" box="[151,216,403,422]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
, coll. H. Diener (
<collectionCode id="ED5552B4FFEA470DFE8FFE2ACEDDFE4A" box="[371,441,403,422]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Budapest, no further data,
<specimenCount id="9D4201F8FFEA470DFD49FE2ACD91FE4A" box="[693,757,403,422]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
, coll. Pillich (
<collectionCode id="ED5552B4FFEA470DFC56FF07CC8BFF3D" box="[938,1007,190,209]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
).
<emphasis id="B9301663FFEA470DFC02FF07CBEEFF3D" bold="true" box="[1022,1162,190,209]" pageId="10" pageNumber="429">
<collectingRegion id="49800493FFEA470DFC02FF07CBE6FF23" box="[1022,1154,190,209]" country="Hungary" name="Somogy" pageId="10" pageNumber="429">SOMOGY MEGYE</collectingRegion>
:
</emphasis>
Zamárdi, Balatonszéplak shore,
<date id="FFFAECB1FFEA470DFCC0FF60CC85FF00" box="[828,993,217,236]" pageId="10" pageNumber="429" value="1951-07-17" valueMax="1951-08-10" valueMin="1951-07-17">17.vii.10.viii.1951</date>
,
<specimenCount id="9D4201F8FFEA470DFC10FF60CB4DFF00" box="[1004,1065,217,236]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
(on sandy meadow), Kaszab lgt. (
<collectionCode id="ED5552B4FFEA470DFAB2FF60CAF7FF00" box="[1358,1427,217,236]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Siófok, no further data,
<specimenCount id="9D4201F8FFEA470DFBF8FF4ACB26FEEA" box="[1028,1090,243,262]" pageId="10" pageNumber="429" type="generic">3 spec.</specimenCount>
, Lichtneckert lgt. (
<collectionCode id="ED5552B4FFEA470DFB23FF4ACA40FEEA" box="[1247,1316,243,262]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Balatonlelle, no further data,
<specimenCount id="9D4201F8FFEA470DFC3AFEB7CB60FECD" box="[966,1028,270,289]" pageId="10" pageNumber="429" type="generic">6 spec.</specimenCount>
, coll. Peregi (
<collectionCode id="ED5552B4FFEA470DFB86FEB7CBDBFECD" box="[1146,1215,270,289]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
); Öszöd, no date,
<specimenCount id="9D4201F8FFEA470DFAA7FEB7CAFEFECD" box="[1371,1434,270,289]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
,
<date id="FFFAECB1FFEA470DFCC0FE90CCECFED0" box="[828,904,297,316]" pageId="10" pageNumber="429" value="1903-08">viii.1903</date>
,
<specimenCount id="9D4201F8FFEA470DFC6EFE90CCB4FED0" box="[914,976,297,316]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
,
<date id="FFFAECB1FFEA470DFC24FE90CB7BFED0" box="[984,1055,297,316]" pageId="10" pageNumber="429" value="1905-07">vii.1905</date>
,
<specimenCount id="9D4201F8FFEA470DFBD4FE90CB03FED0" box="[1064,1127,297,316]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
,
<date id="FFFAECB1FFEA470DFB93FE90CBD1FED0" box="[1135,1205,297,316]" pageId="10" pageNumber="429" value="1906-07">vii.1906</date>
,
<specimenCount id="9D4201F8FFEA470DFB43FE90CB9AFED0" box="[1215,1278,297,316]" pageId="10" pageNumber="429" type="generic">2 spec.</specimenCount>
,
<date id="FFFAECB1FFEA470DFAFAFE90CA37FED0" box="[1286,1363,297,316]" pageId="10" pageNumber="429" value="1906-08">viii.1906</date>
,
<specimenCount id="9D4201F8FFEA470DFAA0FE90CAFEFED0" box="[1372,1434,297,316]" pageId="10" pageNumber="429" type="generic">2 spec.</specimenCount>
, Ehmnann lgt., coll. Dr. R. Streda (
<collectionCode id="ED5552B4FFEA470DFBA4FEFACBF9FEBA" box="[1112,1181,323,342]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="429">HNHM</collectionCode>
).
<emphasis id="B9301663FFEA470DFB56FEFACAE6FEBA" bold="true" box="[1194,1410,323,342]" pageId="10" pageNumber="429">
<collectingCountry id="F3538AE1FFEA470DFB56FEFACB8FFEBA" box="[1194,1259,323,342]" name="India" pageId="10" pageNumber="429">INDIA</collectingCountry>
: ANDRA PRADESH:
</emphasis>
<quantity id="4CBC6794FFEA470DFA7BFEFACC33FE9D" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="429" unit="km" value="35.0">35 km</quantity>
SE of Rajahmundry, Kottipale, Godavari River bank,
<date id="FFFAECB1FFEA470DFAECFEE7CAEEFE9D" box="[1296,1418,350,369]" pageId="10" pageNumber="429" value="1994-02-23" valueMax="1994-02-24" valueMin="1994-02-23">23.24.ii.1994</date>
,
<specimenCount id="9D4201F8FFEA470DFA6DFEE7CC0DFE60" pageId="10" pageNumber="429" type="generic">4 spec.</specimenCount>
, Z. Kejval lgt. (
<collectionCode id="ED5552B4FFEA470DFC1EFEC0CB79FE60" box="[994,1053,377,396]" pageId="10" pageNumber="429">CTLA</collectionCode>
).
<emphasis id="B9301663FFEA470DFBD5FEC0CB17FE60" bold="true" box="[1065,1139,377,396]" pageId="10" pageNumber="429">
<collectingRegion id="49800493FFEA470DFBD5FEC0CB09FE66" box="[1065,1133,377,396]" country="India" name="Kerala" pageId="10" pageNumber="429">KERALA</collectingRegion>
:
</emphasis>
<quantity id="4CBC6794FFEA470DFB8BFEC0CBCEFE60" box="[1143,1194,377,396]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="429" unit="km" value="10.0">10 km</quantity>
E of Punalur,bank of Kallada River,
<geoCoordinate id="EE70ACB6FFEA470DFC8AFE2ACCD6FE4A" box="[886,946,403,422]" degrees="8" direction="north" minutes="59" orientation="latitude" pageId="10" pageNumber="429" precision="925" value="8.983334">8º59N</geoCoordinate>
,
<geoCoordinate id="EE70ACB6FFEA470DFC41FE2ACB64FE4A" box="[957,1024,403,422]" degrees="77" direction="east" minutes="01" orientation="longitude" pageId="10" pageNumber="429" precision="925" value="77.01667">77º01E</geoCoordinate>
,
<date id="FFFAECB1FFEA470DFBF7FE2ACBE5FE4A" box="[1035,1153,403,422]" pageId="10" pageNumber="429" value="1994-01-20" valueMax="1994-01-21" valueMin="1994-01-20">20.21.i.1994</date>
,
<specimenCount id="9D4201F8FFEA470DFB70FE2ACBAFFE4A" box="[1164,1227,403,422]" pageId="10" pageNumber="429" type="generic">1 spec.</specimenCount>
, Z. Kejval lgt. (
<collectionCode id="ED5552B4FFEA470DFAABFE2ACAF7FE4A" box="[1367,1427,403,422]" pageId="10" pageNumber="429">CTLA</collectionCode>
); Shoranur, Ponnani River,
<geoCoordinate id="EE70ACB6FFEB470CFE8CF95DCED1F91B" box="[368,437,1764,1783]" degrees="10" direction="north" minutes="46" orientation="latitude" pageId="11" pageNumber="430" precision="925" value="10.766666">10º46N</geoCoordinate>
,
<geoCoordinate id="EE70ACB6FFEB470CFE42F95DCD66F91B" box="[446,514,1764,1783]" degrees="76" direction="east" minutes="16" orientation="longitude" pageId="11" pageNumber="430" precision="925" value="76.26667">76º16E</geoCoordinate>
,
<date id="FFFAECB1FFEB470CFDF6F95DCD3EF91B" box="[522,602,1764,1783]" pageId="11" pageNumber="430" value="1994-01-31">31.i.1994</date>
,
<specimenCount id="9D4201F8FFEB470CFD98F95DCDC5F91B" box="[612,673,1764,1783]" pageId="11" pageNumber="430" type="generic">3 spec.</specimenCount>
, Z. Kejval lgt.(
<collectionCode id="ED5552B4FFEB470CFF47F946CF92F8FE" box="[187,246,1791,1810]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFEFEF946CE27F8FE" bold="true" box="[258,323,1791,1810]" pageId="11" pageNumber="430">
<collectingRegion id="49800493FFEB470CFEFEF946CE58F8FC" box="[258,316,1791,1810]" name="ORISSA" pageId="11" pageNumber="430">ORISSA</collectingRegion>
:
</emphasis>
<quantity id="4CBC6794FFEB470CFEBBF946CE1EF8FE" box="[327,378,1791,1810]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="430" unit="km" value="30.0">30 km</quantity>
NE of Jaleswar, riverbank of the Balasor River,
<date id="FFFAECB1FFEB470CFF6BF8A0CF8AF8C0" box="[151,238,1817,1836]" pageId="11" pageNumber="430" value="1999-02-13">13.ii.1999</date>
, 1 ♁, Z. Kejval lgt. (
<collectionCode id="ED5552B4FFEB470CFE66F8A0CEB1F8C0" box="[410,469,1817,1836]" pageId="11" pageNumber="430">CTLA</collectionCode>
); Kalasandhapur, N of Berhampur, on a river bank,
<date id="FFFAECB1FFEB470CFEE0F88DCEF3F8AB" box="[284,407,1844,1863]" pageId="11" pageNumber="430" value="1994-02-20" valueMax="1994-02-21" valueMin="1994-02-20">20.21.ii.1994</date>
,
<specimenCount id="9D4201F8FFEB470CFE63F88DCEBFF8AB" box="[415,475,1844,1863]" pageId="11" pageNumber="430" type="generic">1 spec.</specimenCount>
, Z. Kejval lgt. (
<collectionCode id="ED5552B4FFEB470CFDA6F88DCDF1F8AB" box="[602,661,1844,1863]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFD5EF88DCD9CF8AB" bold="true" box="[674,760,1844,1863]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFD5EF88DCD97F8AB" box="[674,755,1844,1863]" name="Israel" pageId="11" pageNumber="430">ISRAEL</collectingCountry>
:
</emphasis>
Arvat Sedom,
<date id="FFFAECB1FFEB470CFEF0F8F6CE1AF88E" box="[268,382,1871,1890]" pageId="11" pageNumber="430" value="2014-04-08" valueMax="2014-04-29" valueMin="2014-04-08">8.29.iv.2014</date>
,
<specimenCount id="9D4201F8FFEB470CFE79F8F6CEA4F88E" box="[389,448,1871,1890]" pageId="11" pageNumber="430" type="generic">1 spec.</specimenCount>
,I. Renan lgt.,spring (
<collectionCode id="ED5552B4FFEB470CFD91F8F6CDCCF88E" box="[621,680,1871,1890]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFD48F8F6CF80F890" bold="true" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFD48F8F6CD90F88E" box="[692,756,1871,1890]" name="Italy" pageId="11" pageNumber="430">ITALY</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFF6BF8D0CFB8F897" box="[151,220,1897,1916]" country="Italy" name="GORIZIA" pageId="11" pageNumber="430">GORIZIA</collectingRegion>
:
</emphasis>
Grado,
<specimenCount id="9D4201F8FFEB470CFED4F8D0CE2CF891" box="[296,328,1894,1917]" pageId="11" pageNumber="430" type="female">1 ♀</specimenCount>
, J. Matcha lgt. (
<collectionCode id="ED5552B4FFEB470CFE2CF8D0CD74F890" box="[464,528,1897,1916]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="430">NMPC</collectionCode>
).
<emphasis id="B9301663FFEB470CFDE3F8D0CD15F890" bold="true" box="[543,625,1897,1916]" pageId="11" pageNumber="430">
<collectingRegion id="49800493FFEB470CFDE3F8D0CD0EF897" box="[543,618,1897,1916]" country="Italy" name="Sardegna" pageId="11" pageNumber="430">SARDINIA</collectingRegion>
:
</emphasis>
no further data,
<specimenCount id="9D4201F8FFEB470CFF6BF83DCFB0F87B" box="[151,212,1924,1943]" pageId="11" pageNumber="430" type="generic">1 spec.</specimenCount>
, Gené (
<collectionCode id="ED5552B4FFEB470CFEEDF83DCE35F87B" box="[273,337,1924,1943]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="11" pageNumber="430">MFNB</collectionCode>
).
<emphasis id="B9301663FFEB470CFEA2F83DCED4F87B" bold="true" box="[350,432,1924,1943]" pageId="11" pageNumber="430">
<collectingRegion id="49800493FFEB470CFEA2F83DCECDF879" box="[350,425,1924,1943]" country="Italy" name="Toscana" pageId="11" pageNumber="430">TUSCANY</collectingRegion>
:
</emphasis>
Pisa-Calambrone,
<date id="FFFAECB1FFEB470CFDB3F83DCDFAF87B" box="[591,670,1924,1943]" pageId="11" pageNumber="430" value="2014-04-09">9.iv.2014</date>
,
<specimenCount id="9D4201F8FFEB470CFD5AF83DCD86F87B" box="[678,738,1924,1943]" pageId="11" pageNumber="430" type="generic">2 spec.</specimenCount>
, P. Kylies (
<collectionCode id="ED5552B4FFEB470CFF25F826CE70F85E" box="[217,276,1951,1970]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFEDFF826CD13F85E" bold="true" box="[291,631,1951,1970]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFEDFF826CEDEF85E" box="[291,442,1951,1970]" name="Kazakhstan" pageId="11" pageNumber="430">KAZAKHSTAN</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFE3FF826CD4EF85C" box="[451,554,1951,1970]" country="Kazakhstan" name="AKTYUBINSK REGION" pageId="11" pageNumber="430">AKTYUBINSK</collectingRegion>
REGION:
</emphasis>
Khobda River,
<date id="FFFAECB1FFEB470CFF6BF800CF8DF820" box="[151,233,1977,1996]" pageId="11" pageNumber="430" value="2000-05-25">25.v.2000</date>
,
<specimenCount id="9D4201F8FFEB470CFF0CF800CE4EF820" box="[240,298,1977,1996]" pageId="11" pageNumber="430" type="generic">3 spec.</specimenCount>
,collector unknown (
<collectionCode id="ED5552B4FFEB470CFE2CF800CD6FF820" box="[464,523,1977,1996]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFDEAF800CD9CF820" bold="true" box="[534,760,1977,1996]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFDEAF800CDD0F820" box="[534,692,1977,1996]" name="Serbia" pageId="11" pageNumber="430">MONTENEGRO</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFD47F800CD95F827" box="[699,753,1977,1996]" country="Serbia" name="BUDVA" pageId="11" pageNumber="430">BUDVA</collectingRegion>
:
</emphasis>
Budva, no further data,1♁ (
<collectionCode id="ED5552B4FFEB470CFE87F86DCEDFF80B" box="[379,443,2004,2023]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="430">NMPC</collectionCode>
).
<emphasis id="B9301663FFEB470CFE3AF86DCD9CF80B" bold="true" box="[454,760,2004,2023]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFE3AF86DCD5FF80B" box="[454,571,2004,2023]" name="Morocco" pageId="11" pageNumber="430">MOROCCO</collectingCountry>
: FAS-
<collectingRegion id="49800493FFEB470CFD9BF86DCD94F809" box="[615,752,2004,2023]" country="Morocco" name="MEKNAS REGION" pageId="11" pageNumber="430">MEKNAS REGION</collectingRegion>
:
</emphasis>
MoyenAtlas,AguelmameAzegza Lake,lake shore,
<date id="FFFAECB1FFEB470CFDC5F856CDD1F7EE" box="[569,693,2031,2050]" pageId="11" pageNumber="430" value="1998-06-22" valueMax="1998-06-26" valueMin="1998-06-22">22.26.vi.1998</date>
,
<specimenCount id="9D4201F8FFEB470CFD47F856CD92F7EE" box="[699,758,2031,2050]" pageId="11" pageNumber="430" type="generic">1 spec.</specimenCount>
, T. Lackner lgt. (
<collectionCode id="ED5552B4FFEB470CFEE6F7B0CE31F7F0" box="[282,341,2057,2076]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFE9EF7B0CD69F7F0" bold="true" box="[354,525,2057,2076]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFE9EF7B0CEC7F7F0" box="[354,419,2057,2076]" name="Spain" pageId="11" pageNumber="430">SPAIN</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFE57F7B0CD63F7F7" box="[427,519,2057,2076]" country="Spain" name="Catalunya" pageId="11" pageNumber="430">CATALONIA</collectingRegion>
:
</emphasis>
Girona, Sant Pere Pescador,
<geoCoordinate id="EE70ACB6FFEB470CFCC0FB89CCCCFBAF" box="[828,936,1072,1091]" degrees="42" direction="north" minutes="10.628" orientation="latitude" pageId="11" pageNumber="430" precision="1" value="42.17713">42°10.628N</geoCoordinate>
,
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,
<date id="FFFAECB1FFEB470CFBEBFB89CB15FBAF" box="[1047,1137,1072,1091]" pageId="11" pageNumber="430" value="2010-04-24">24.iv.2010</date>
, 1 ♁, J. Krátký lgt. (
<collectionCode id="ED5552B4FFEB470CFAE4FB89CA37FBAF" box="[1304,1363,1072,1091]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFA9DFB89CB68FBB2" bold="true" pageId="11" pageNumber="430">TUNISIA: JENDOUBA REGION:</emphasis>
Chemtou env., Mejerda River,
<date id="FFFAECB1FFEB470CFAEBFBF2CAFDFBB2" box="[1303,1433,1099,1118]" pageId="11" pageNumber="430" value="1997-04-30" valueMax="1997-05-01" valueMin="1997-04-30">30.iv.1.v.1997</date>
,
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, J. Mertlik lgt. (
<collectionCode id="ED5552B4FFEB470CFBF6FBDFCB22FB95" box="[1034,1094,1126,1145]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFBA9FBDFCA29FB95" bold="true" box="[1109,1357,1126,1145]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFBA9FBDFCBD5FB95" box="[1109,1201,1126,1145]" name="Turkey" pageId="11" pageNumber="430">TURKEY</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFB40FBDFCB9AFB9B" box="[1212,1278,1126,1145]" country="Turkey" name="Samsun" pageId="11" pageNumber="430">SAMSUN</collectingRegion>
REGION:
</emphasis>
Samsun,
<quantity id="4CBC6794FFEB470CFCC0FB39CC15FB7F" box="[828,881,1152,1171]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="430" unit="km" value="17.0">17 km</quantity>
N of Çarsamba beach,
<date id="FFFAECB1FFEB470CFBC4FB39CBE8FB7F" box="[1080,1164,1152,1171]" pageId="11" pageNumber="430" value="1989-05-19">19.v.1989</date>
,
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, P. Kanaar lgt. (
<collectionCode id="ED5552B4FFEB470CFAA4FB39CAF0FB7F" box="[1368,1428,1152,1171]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
<emphasis id="B9301663FFEB470CFCC0FB22CB3AFB42" bold="true" box="[828,1118,1179,1198]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFCC0FB22CCC7FB42" box="[828,931,1179,1198]" name="Ukraine" pageId="11" pageNumber="430">UKRAINE</collectingCountry>
: CHERKASSY REGION:
</emphasis>
Dniper River,
<date id="FFFAECB1FFEB470CFB1FFB22CA0EFB42" box="[1251,1386,1179,1198]" pageId="11" pageNumber="430" value="2000-07-10" valueMax="2000-07-25" valueMin="2000-07-10">10.25.vii.2000</date>
,
<specimenCount id="9D4201F8FFEB470CFA89FB22CAF2FB43" box="[1397,1430,1176,1199]" pageId="11" pageNumber="430" type="female">1 ♀</specimenCount>
, Vasko lgt. (
<collectionCode id="ED5552B4FFEB470CFC63FB0FCC8DFB25" box="[927,1001,1206,1225]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430" type="Museum">NHMW</collectionCode>
).
<emphasis id="B9301663FFEB470CFC05FB0FCB26FB25" bold="true" box="[1017,1090,1206,1225]" pageId="11" pageNumber="430">
<collectingRegion id="49800493FFEB470CFC05FB0FCB5EFB2B" box="[1017,1082,1206,1225]" country="Ukraine" name="Crimea" pageId="11" pageNumber="430">CRIMEA</collectingRegion>
:
</emphasis>
Zurzut,
<date id="FFFAECB1FFEB470CFB72FB0FCB86FB25" box="[1166,1250,1206,1225]" pageId="11" pageNumber="430" value="1999-05-28">28.v.1999</date>
, 2 ♁♁, Putchkov lgt. (
<collectionCode id="ED5552B4FFEB470CFCBDFB69CCEEFB0F" box="[833,906,1232,1251]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430" type="Museum">NHMW</collectionCode>
).
<emphasis id="B9301663FFEB470CFC6AFB69CB4CFB0F" bold="true" box="[918,1064,1232,1251]" pageId="11" pageNumber="430">
<collectingRegion id="49800493FFEB470CFC6AFB69CB45FB0E" box="[918,1057,1232,1251]" country="Ukraine" name="Kherson" pageId="11" pageNumber="430">KHERSON REGION</collectingRegion>
:
</emphasis>
Golopristansky district, Bolshevik,
<date id="FFFAECB1FFEB470CFAB6FB69CAFDFB0F" box="[1354,1433,1232,1251]" pageId="11" pageNumber="430" value="2000-06-01">1.vi.2000</date>
,
<specimenCount id="9D4201F8FFEB470CFCC0FB52CC3DFB13" box="[828,857,1256,1279]" pageId="11" pageNumber="430" type="female">1♀</specimenCount>
, Putchkov lgt. (
<collectionCode id="ED5552B4FFEB470CFC24FB52CB46FB12" box="[984,1058,1259,1278]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430" type="Museum">NHMW</collectionCode>
).
<emphasis id="B9301663FFEB470CFBD2FB52CAF8FB12" bold="true" box="[1070,1436,1259,1278]" pageId="11" pageNumber="430">
<collectingCountry id="F3538AE1FFEB470CFBD2FB52CA57FB12" box="[1070,1331,1259,1278]" name="United Arab Emirates" pageId="11" pageNumber="430">UNITED ARAB EMIRATES</collectingCountry>
:
<collectingRegion id="49800493FFEB470CFAC6FB52CA39FB10" box="[1338,1373,1259,1278]" country="United Arab Emirates" name="Abu Zaby" pageId="11" pageNumber="430">ABU</collectingRegion>
DHABI:
</emphasis>
Abu Dhabi,Channel Street,Al Raha Beach,
<geoCoordinate id="EE70ACB6FFEB470CFB60FABFCA7CFAF5" box="[1180,1304,1286,1305]" degrees="24" direction="north" minutes="26" orientation="latitude" pageId="11" pageNumber="430" precision="1" seconds="07.62" value="24.43545">24°2607.62″N</geoCoordinate>
,
<geoCoordinate id="EE70ACB6FFEB470CFAE3FABFCAFDFAF5" box="[1311,1433,1286,1305]" degrees="54" direction="east" minutes="33" orientation="longitude" pageId="11" pageNumber="430" precision="1" seconds="42.26" value="54.561737">54°3342.26″E</geoCoordinate>
,
<date id="FFFAECB1FFEB470CFCC0FA99CCD8FADF" box="[828,956,1312,1331]" pageId="11" pageNumber="430" value="2015-03-22" valueMax="2015-03-31" valueMin="2015-03-22">22.31.iii.2015</date>
, 1 ♁, A. Pütz lgt. (
<collectionCode id="ED5552B4FFEB470CFBAAFA99CBF6FADF" box="[1110,1170,1312,1331]" pageId="11" pageNumber="430">CTLA</collectionCode>
).
</paragraph>
<caption id="DF3B9AF9FFEA470DFF6BF857CB53F7F0" ID-DOI="http://doi.org/10.5281/zenodo.4504934" ID-Zenodo-Dep="4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="10" pageNumber="429" startId="10.[151,188,2030,2049]" targetBox="[213,1403,494,1971]" targetPageId="10">
<paragraph id="8BFBCA71FFEA470DFF6BF857CB53F7F0" blockId="10.[151,1437,2030,2076]" pageId="10" pageNumber="429">
Figs 2528.
<taxonomicName id="4C44B1F2FFEA470DFEFEF857CD6EF7ED" authorityName=": MAZUR" authorityYear="2011" box="[258,522,2030,2049]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEA470DFEFEF857CE0EF7ED" box="[258,362,2030,2049]" italics="true" pageId="10" pageNumber="429">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEA470DFE84F857CED7F7ED" box="[376,435,2030,2049]" italics="true" pageId="10" pageNumber="429">Nessus</emphasis>
)
<emphasis id="B9301663FFEA470DFE3CF857CD6EF7ED" box="[448,522,2030,2049]" italics="true" pageId="10" pageNumber="429">rubripes</emphasis>
</taxonomicName>
<emphasis id="B9301663FFEA470DFDF6F857CD6AF7ED" box="[522,526,2030,2049]" italics="true" pageId="10" pageNumber="429">,</emphasis>
dorsal habitus. 25
<taxonomicName id="4C44B1F2FFEA470DFD3EF857CC83F7ED" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[706,999,2030,2049]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B9301663FFEA470DFD3EF857CC38F7ED" box="[706,860,2030,2049]" italics="true" pageId="10" pageNumber="429">Saprinus rubripes</emphasis>
<bibRefCitation id="EFD5B780FFEA470DFC9FF857CC83F7ED" author="ERICHSON W. F." box="[867,999,2030,2049]" pageId="10" pageNumber="429" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
</taxonomicName>
, syntype. 26
<taxonomicName id="4C44B1F2FFEA470DFB95F857CAFDF7ED" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[1129,1433,2030,2049]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="granarius">
<emphasis id="B9301663FFEA470DFB95F857CA6BF7ED" box="[1129,1295,2030,2049]" italics="true" pageId="10" pageNumber="429">Saprinus granarius</emphasis>
<bibRefCitation id="EFD5B780FFEA470DFAE9F857CAFDF7ED" author="ERICHSON W. F." box="[1301,1433,2030,2049]" pageId="10" pageNumber="429" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
</taxonomicName>
, syntype. 27
<taxonomicName id="4C44B1F2FFEA470DFEF3F7B0CD51F7F0" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[271,565,2057,2076]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="arenarius">
<emphasis id="B9301663FFEA470DFEF3F7B0CED6F7F0" box="[271,434,2057,2076]" italics="true" pageId="10" pageNumber="429">Saprinus arenarius</emphasis>
<bibRefCitation id="EFD5B780FFEA470DFE44F7B0CD51F7F0" author="MARSEUL S. A." box="[440,565,2057,2076]" pageId="10" pageNumber="429" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
, lectotype. 28
<taxonomicName id="4C44B1F2FFEA470DFD3DF7B0CCB8F7F0" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[705,988,2057,2076]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFEA470DFD3DF7B0CC3DF7F0" box="[705,857,2057,2076]" italics="true" pageId="10" pageNumber="429">Saprinus corsicus</emphasis>
<bibRefCitation id="EFD5B780FFEA470DFCA3F7B0CCB8F7F0" author="MARSEUL S. A." box="[863,988,2057,2076]" pageId="10" pageNumber="429" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
,?syntype.
</paragraph>
</caption>
<caption id="DF3B9AF9FFEB470CFF64FC75CC7EFC16" ID-DOI="http://doi.org/10.5281/zenodo.4504936" ID-Zenodo-Dep="4504936" httpUri="https://zenodo.org/record/4504936/files/figure.png" pageId="11" pageNumber="430" startId="11.[152,189,972,991]" targetBox="[168,1435,194,945]" targetPageId="11">
<paragraph id="8BFBCA71FFEB470CFF64FC75CC7EFC16" blockId="11.[151,1437,972,1018]" pageId="11" pageNumber="430">
Figs 2930.
<taxonomicName id="4C44B1F2FFEB470CFEFFFC75CDC6FC33" authority="(Erichson, 1834)" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[259,674,972,991]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEB470CFEFFFC75CE0FFC33" box="[259,363,972,991]" italics="true" pageId="11" pageNumber="430">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEB470CFE84FC75CED7FC33" box="[376,435,972,991]" italics="true" pageId="11" pageNumber="430">Nessus</emphasis>
)
<emphasis id="B9301663FFEB470CFE3DFC75CD6DFC33" box="[449,521,972,991]" italics="true" pageId="11" pageNumber="430">rubripes</emphasis>
(
<bibRefCitation id="EFD5B780FFEB470CFDE9FC75CDF8FC33" author="ERICHSON W. F." box="[533,668,972,991]" pageId="11" pageNumber="430" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
)
</taxonomicName>
. 29
<taxonomicName id="4C44B1F2FFEB470CFD2BFC75CB04FC33" authority="Auzat, 1920" authorityName="Auzat" authorityYear="1920" box="[727,1120,972,991]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="variety" species="rubripes" variety="clermonti">
<emphasis id="B9301663FFEB470CFD2BFC75CC15FC33" box="[727,881,972,991]" italics="true" pageId="11" pageNumber="430">Saprinus rubripes</emphasis>
var.
<emphasis id="B9301663FFEB470CFC61FC75CC8BFC33" box="[925,1007,972,991]" italics="true" pageId="11" pageNumber="430">clermonti</emphasis>
<bibRefCitation id="EFD5B780FFEB470CFC08FC75CB04FC33" author="AUZAT V." box="[1012,1120,972,991]" pageId="11" pageNumber="430" pagination="1 - 4" refId="ref20400" refString="AUZAT V. 1920: Quelques observations sur la sculpture superficielle des Histerides; varietes nouvelles, synonymies. L'Echange, Revue Linneenne 36 (399, Hors texte): 1 - 4." type="book chapter" year="1920">Auzat, 1920</bibRefCitation>
</taxonomicName>
, holotype, habitus, dorsal view. 30 varieties of
<taxonomicName id="4C44B1F2FFEB470CFF00FC5ECD64FC16" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[252,512,999,1018]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEB470CFF00FC5ECE00FC16" box="[252,356,999,1018]" italics="true" pageId="11" pageNumber="430">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEB470CFE8CFC5ECECFFC16" box="[368,427,999,1018]" italics="true" pageId="11" pageNumber="430">Nessus</emphasis>
)
<emphasis id="B9301663FFEB470CFE44FC5ECD64FC16" box="[440,512,999,1018]" italics="true" pageId="11" pageNumber="430">rubripes</emphasis>
</taxonomicName>
as summed up by Jean Thérond.
</paragraph>
</caption>
<paragraph id="8BFBCA71FFEB470BFCC7FAF5CEC5F86C" blockId="11.[827,1437,1356,2083]" lastBlockId="12.[151,760,1449,2080]" lastPageId="12" lastPageNumber="431" pageId="11" pageNumber="430">
<emphasis id="B9301663FFEB470CFCC7FAF5CB25FA8F" bold="true" box="[827,1089,1356,1379]" pageId="11" pageNumber="430">Diagnostic description.</emphasis>
This taxon was redescribed and figured in detail by
<bibRefCitation id="EFD5B780FFEB470CFBEDFAD5CBD8FA6F" author="LACKNER T." box="[1041,1212,1388,1411]" pageId="11" pageNumber="430" pagination="1 - 254" refId="ref21136" refString="LACKNER T. 2010: Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 50 (Supplementum): 1 - 254." type="journal article" year="2010">LACKNER (2010)</bibRefCitation>
and the reader is referred for the detailed redescription there. For the sake of better recognition we provide here only a short diagnostic description supplemented by a habitus image (
<figureCitation id="137FD6F4FFEB470CFAE2FA75CA0EFA0F" box="[1310,1386,1484,1507]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="11" pageNumber="430">Fig. 25</figureCitation>
) and genitalia drawings (
<figureCitation id="137FD6F4FFEB470CFBE8FA55CBF0F9EF" box="[1044,1172,1516,1539]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="11" pageNumber="430">Figs 3339</figureCitation>
). Clypeus with elevated anterior margin, somewhat margined laterally, rugulose- lacunose; frontal stria well impressed, straight, carinate; continued as a well-impressed carinate supraorbital stria; frontal disc normally with irregular longitudinal rugae intermingled with sparse microscopic punctation; eyes flat, inconspicuous from above. Pronotal disc laterally with coarse punctation, between it and pronotal margin present a smooth longitudinal band; medially punctation much finer and sparser. First dorsal elytral stria the longest, usually reaching approximately three-fourths of elytral length apically; second, third and fourth dorsal elytral striae about the same length, reaching approximately elytral half apically; fourth dorsal elytral stria basally connected with sutural elytral stria; sutural stria well impressed, in shallow punctures, shortened on its apical tenth. Elytral disc on apical half (except for elytral flanks) with coarse and dense punctation, punctures separated by about their own diameter, anteriorly reaching about half of elytral length; basal half with only fine microscopic punctation; extreme apex of elytra with an impunctate band. Protibia (
<figureCitation id="137FD6F4FFEC470BFF63F9B0CF94F9CC" box="[159,240,1545,1568]" captionStart="Figs 3132" captionStartId="11.[151,188,1619,1638]" captionTargetBox="[172,742,1091,1585]" captionTargetId="figure-518@11.[151,760,1066,1603]" captionTargetPageId="11" captionText="Figs 3132. Protibiae, dorsal view. 31 Hypocaccus (Nessus) rubripes (Erichson, 1834); 32 H. (N.) curtus (Rosenhauer, 1847) (both re-drawn from VIENNA 1980)." figureDoi="http://doi.org/10.5281/zenodo.4504938" httpUri="https://zenodo.org/record/4504938/files/figure.png" pageId="12" pageNumber="431">Fig. 31</figureCitation>
) flattened and somewhat dilated, outer margin with four low teeth topped with short denticle followed by three minuscule denticles. Male genitalia. Sternite VIII (
<figureCitation id="137FD6F4FFEC470BFF63F9D0CE7FF96C" box="[159,283,1641,1664]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Figs 3334</figureCitation>
) longitudinally separated medially, apically with tiny inflatable membrane (velum); fringed with single short seta; tergite VIII and sternite VIII not fused laterally (
<figureCitation id="137FD6F4FFEC470BFF63F970CF94F90C" box="[159,240,1737,1760]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Fig. 35</figureCitation>
). Morphology of tergite IX (
<figureCitation id="137FD6F4FFEC470BFDD9F970CDC4F90C" box="[549,672,1737,1760]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Figs 3839</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="137FD6F4FFEC470BFDCDF950CDE5F8EC" box="[561,641,1769,1792]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Fig. 37</figureCitation>
) expanded on both ends. Basal piece of aedeagus (
<figureCitation id="137FD6F4FFEC470BFDCFF8B0CDC8F8CC" box="[563,684,1801,1824]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Figs 3637</figureCitation>
) rather short, ratio of its length to length of parameres equals to 1: 3; parameres fused along their basal two-thirds; aedeagus curved ventrad (
<figureCitation id="137FD6F4FFEC470BFEB9F8D0CEF1F86C" box="[325,405,1897,1920]" captionStart="Figs 3339" captionStartId="12.[152,189,1301,1320]" captionTargetBox="[340,1249,191,1281]" captionTargetId="figure-368@12.[313,1274,168,1298]" captionTargetPageId="12" captionText="Figs 3339. Male genitalia of Hypocaccus (Nessus) rubripes (Erichson, 1834): 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3B9AF9FFEB470CFF6BF9EACE25F977" ID-DOI="http://doi.org/10.5281/zenodo.4504938" ID-Zenodo-Dep="4504938" httpUri="https://zenodo.org/record/4504938/files/figure.png" pageId="11" pageNumber="430" startId="11.[151,188,1619,1638]" targetBox="[172,742,1091,1585]" targetPageId="11">
<paragraph id="8BFBCA71FFEB470CFF6BF9EACE25F977" blockId="11.[151,760,1619,1691]" pageId="11" pageNumber="430">
Figs 3132. Protibiae, dorsal view. 31
<taxonomicName id="4C44B1F2FFEB470CFE0EF9EACE43F96C" authority="(Erichson, 1834)" baseAuthorityName="Erichson" baseAuthorityYear="1834" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEB470CFE0EF9EACD3EF98A" box="[498,602,1619,1638]" italics="true" pageId="11" pageNumber="430">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEB470CFD9BF9EACDC6F98A" box="[615,674,1619,1638]" italics="true" pageId="11" pageNumber="430">Nessus</emphasis>
)
<emphasis id="B9301663FFEB470CFD53F9EACD93F98A" box="[687,759,1619,1638]" italics="true" pageId="11" pageNumber="430">rubripes</emphasis>
(
<bibRefCitation id="EFD5B780FFEB470CFF61F9D4CE45F96C" author="ERICHSON W. F." box="[157,289,1645,1664]" pageId="11" pageNumber="430" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
)
</taxonomicName>
; 32
<taxonomicName id="4C44B1F2FFEB470CFEABF9D4CD11F96C" authority="(Rosenhauer, 1847)" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[343,629,1645,1664]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFEB470CFEABF9D4CE09F96C" box="[343,365,1645,1664]" italics="true" pageId="11" pageNumber="430">H.</emphasis>
(
<emphasis id="B9301663FFEB470CFE8BF9D4CEE8F96C" box="[375,396,1645,1664]" italics="true" pageId="11" pageNumber="430">N.</emphasis>
)
<emphasis id="B9301663FFEB470CFE6AF9D4CEAFF96C" box="[406,459,1645,1664]" italics="true" pageId="11" pageNumber="430">curtus</emphasis>
(
<bibRefCitation id="EFD5B780FFEB470CFE29F9D4CD14F96C" author="ROSENHAUER W. G." box="[469,624,1645,1664]" pageId="11" pageNumber="430" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
)
</taxonomicName>
(both re-drawn from
<bibRefCitation id="EFD5B780FFEB470CFF3AF931CE53F977" author="VIENNA P." box="[198,311,1672,1691]" pageId="11" pageNumber="430" refId="ref21978" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">VIENNA 1980</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF3B9AF9FFEC470BFF64FAACCDB2FAB1" ID-DOI="http://doi.org/10.5281/zenodo.4504940" ID-Zenodo-Dep="4504940" httpUri="https://zenodo.org/record/4504940/files/figure.png" pageId="12" pageNumber="431" startId="12.[152,189,1301,1320]" targetBox="[340,1249,191,1281]" targetPageId="12">
<paragraph id="8BFBCA71FFEC470BFF64FAACCDB2FAB1" blockId="12.[151,1437,1301,1373]" pageId="12" pageNumber="431">
Figs 3339. Male genitalia of
<taxonomicName id="4C44B1F2FFEC470BFE64FAACCC57FAC4" authority="(Erichson, 1834)" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[408,819,1301,1320]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="431" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEC470BFE64FAACCD64FAC4" box="[408,512,1301,1320]" italics="true" pageId="12" pageNumber="431">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEC470BFDF0FAACCD23FAC4" box="[524,583,1301,1320]" italics="true" pageId="12" pageNumber="431">Nessus</emphasis>
)
<emphasis id="B9301663FFEC470BFDA8FAACCDF8FAC4" box="[596,668,1301,1320]" italics="true" pageId="12" pageNumber="431">rubripes</emphasis>
(
<bibRefCitation id="EFD5B780FFEC470BFD5BFAACCC4AFAC4" author="ERICHSON W. F." box="[679,814,1301,1320]" pageId="12" pageNumber="431" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
)
</taxonomicName>
: 32 VIII sternite and tergite, ventral view; 33 ditto, dorsal view; 34 ditto, lateral view; 35 aedeagus, dorsal view; 36 ditto, lateral view; 37 IX &amp; X tergites, dorsal view + IX sternite (spiculum gastrale), ventral view; 38 IX &amp; X tergites + IX sternite (spiculum gastrale), lateral view.
</paragraph>
</caption>
<subSubSection id="C35E99FAFFEC470BFF6BF830CBDBFA0C" pageId="12" pageNumber="431" type="distribution">
<paragraph id="8BFBCA71FFEC470BFF6BF830CBDBFA0C" blockId="12.[151,760,1449,2080]" lastBlockId="12.[828,1437,1449,2080]" pageId="12" pageNumber="431">
<emphasis id="B9301663FFEC470BFF6BF830CE4DF84C" bold="true" box="[151,297,1929,1952]" pageId="12" pageNumber="431">Distribution.</emphasis>
<taxonomicName id="4C44B1F2FFEC470BFED2F830CD0BF84C" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[302,623,1929,1952]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="431" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEC470BFED2F830CED4F84C" box="[302,432,1929,1952]" italics="true" pageId="12" pageNumber="431">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEC470BFE41F830CD63F84C" box="[445,519,1929,1952]" italics="true" pageId="12" pageNumber="431">Nessus</emphasis>
)
<emphasis id="B9301663FFEC470BFDE9F830CD0BF84C" box="[533,623,1929,1952]" italics="true" pageId="12" pageNumber="431">rubripes</emphasis>
</taxonomicName>
, as presently understood, covers a large area from
<collectingCountry id="F3538AE1FFEC470BFDDFF810CD19F82C" box="[547,637,1961,1984]" name="Portugal" pageId="12" pageNumber="431">Portugal</collectingCountry>
in the west to the Russian Far East in the east. It is spread in the entire Mediterranean subregion,
<collectingCountry id="F3538AE1FFEC470BFE4FF850CD3BF7EC" box="[435,607,2025,2048]" name="Netherlands" pageId="12" pageNumber="431">the Netherlands</collectingCountry>
, Central, Eastern and Western Europe,
<collectingCountry id="F3538AE1FFEC470BFE4CF7B0CE99F7CC" box="[432,509,2057,2080]" name="Turkey" pageId="12" pageNumber="431">Turkey</collectingCountry>
, Georgia,
<collectingCountry id="F3538AE1FFEC470BFD90F7B0CDADF7CC" box="[620,713,2057,2080]" name="Armenia" pageId="12" pageNumber="431">Armenia</collectingCountry>
and
<collectingRegion id="49800493FFEC470BFCC0FA10CCD0FA2C" box="[828,948,1449,1472]" country="Iran" name="Azarbayjan-e Sharqi" pageId="12" pageNumber="431">Azerbaijan</collectingRegion>
,
<collectingCountry id="F3538AE1FFEC470BFC41FA10CC8FFA2C" box="[957,1003,1449,1472]" name="Iran" pageId="12" pageNumber="431">Iran</collectingCountry>
, Middle Asia,
<collectingCountry id="F3538AE1FFEC470BFB78FA10CB8AFA2C" box="[1156,1262,1449,1472]" name="Mongolia" pageId="12" pageNumber="431">Mongolia</collectingCountry>
,
<collectingCountry id="F3538AE1FFEC470BFB05FA10CA54FA2C" box="[1273,1328,1449,1472]" name="India" pageId="12" pageNumber="431">India</collectingCountry>
as well as entire tropical Africa (
<bibRefCitation id="EFD5B780FFEC470BFBD4FA70CBD7FA0C" author="MAZUR S." box="[1064,1203,1481,1504]" pageId="12" pageNumber="431" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">MAZUR 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEC470BFCC0FA50CA09F9AC" pageId="12" pageNumber="431" type="biology_ecology">
<paragraph id="8BFBCA71FFEC470BFCC0FA50CA09F9AC" blockId="12.[828,1437,1449,2080]" pageId="12" pageNumber="431">
<emphasis id="B9301663FFEC470BFCC0FA50CCF3F9EC" bold="true" box="[828,919,1513,1536]" pageId="12" pageNumber="431">Biology.</emphasis>
This species is most often found in sandy soils, often on riverbanks and seashores where it can be encountered on dung, carcass or under decaying vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEC470AFCC0F9F0CD85F7F9" lastPageId="13" lastPageNumber="432" pageId="12" pageNumber="431" type="discussion">
<paragraph id="8BFBCA71FFEC470BFCC0F9F0CAF9F8AC" blockId="12.[828,1437,1449,2080]" pageId="12" pageNumber="431">
<emphasis id="B9301663FFEC470BFCC0F9F0CCC8F98C" bold="true" box="[828,940,1609,1632]" pageId="12" pageNumber="431">Remarks.</emphasis>
As noted already by
<bibRefCitation id="EFD5B780FFEC470BFB59F9F0CA0FF98C" author="REICHARDT A." box="[1189,1387,1609,1632]" pageId="12" pageNumber="431" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
and
<bibRefCitation id="EFD5B780FFEC470BFCC0F9D0CC8FF96C" author="LACKNER T." box="[828,1003,1641,1664]" pageId="12" pageNumber="431" pagination="1 - 254" refId="ref21136" refString="LACKNER T. 2010: Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 50 (Supplementum): 1 - 254." type="journal article" year="2010">LACKNER (2010)</bibRefCitation>
this species exhibits a large degree of variation regarding its colouration of the cuticle and other external morphological characters, and might represent a complex of cryptic species. In this study, we try to sum up and depict (
<figureCitation id="137FD6F4FFEC470BFC22F950CB3DF8EC" box="[990,1113,1769,1792]" captionStart-0="Figs 2528" captionStart-1="Figs 2930" captionStartId-0="10.[151,188,2030,2049]" captionStartId-1="11.[152,189,972,991]" captionTargetBox-0="[213,1403,494,1971]" captionTargetBox-1="[168,1435,194,945]" captionTargetId-0="figure-13@10.[151,1436,456,2005]" captionTargetId-1="figure-480@11.[148,1439,186,959]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." captionText-1="Figs 2930. Hypocaccus (Nessus) rubripes (Erichson, 1834). 29 Saprinus rubripes var. clermonti Auzat, 1920, holotype, habitus, dorsal view. 30 varieties of Hypocaccus (Nessus) rubripes as summed up by Jean Thérond." figureDoi-0="http://doi.org/10.5281/zenodo.4504934" figureDoi-1="http://doi.org/10.5281/zenodo.4504936" httpUri-0="https://zenodo.org/record/4504934/files/figure.png" httpUri-1="https://zenodo.org/record/4504936/files/figure.png" pageId="12" pageNumber="431">Figs 2529</figureCitation>
) the most common variations (forms most of them originally described as species) of
<taxonomicName id="4C44B1F2FFEC470BFCC0F890CCBBF8AC" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[828,991,1833,1856]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="432" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFEC470BFCC0F890CC2BF8AC" box="[828,847,1833,1856]" italics="true" pageId="12" pageNumber="431">H</emphasis>
. (
<emphasis id="B9301663FFEC470BFC9EF890CC17F8AC" box="[866,883,1833,1856]" italics="true" pageId="12" pageNumber="431">N</emphasis>
.)
<emphasis id="B9301663FFEC470BFC7BF890CCBBF8AC" box="[903,991,1833,1856]" italics="true" pageId="12" pageNumber="431">rubripes</emphasis>
</taxonomicName>
as elaborated already by
<bibRefCitation id="EFD5B780FFEC470BFB22F890CAFDF8AC" author="REICHARDT A." box="[1246,1433,1833,1856]" pageId="12" pageNumber="431" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
.
</paragraph>
<paragraph id="8BFBCA71FFEC470BFCA6F8F0CCFBF82C" blockId="12.[828,1437,1449,2080]" pageId="12" pageNumber="431">
<taxonomicName id="4C44B1F2FFEC470BFCA6F8F0CA45F88C" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[858,1313,1865,1888]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="431" phylum="Arthropoda" rank="species" species="arenarius" subGenus="Nessus">
<emphasis id="B9301663FFEC470BFCA6F8F0CCB8F88C" box="[858,988,1865,1888]" italics="true" pageId="12" pageNumber="431">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEC470BFC11F8F0CB63F88C" box="[1005,1031,1865,1888]" italics="true" pageId="12" pageNumber="431">N.</emphasis>
)
<emphasis id="B9301663FFEC470BFBEBF8F0CB1AF88C" box="[1047,1150,1865,1888]" italics="true" pageId="12" pageNumber="431">arenarius</emphasis>
<bibRefCitation id="EFD5B780FFEC470BFB7AF8F0CA45F88C" author="MARSEUL S. A." box="[1158,1313,1865,1888]" pageId="12" pageNumber="431" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
is a darker form (
<figureCitation id="137FD6F4FFEC470BFC82F8D0CCA8F86C" box="[894,972,1897,1920]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="12" pageNumber="431">Fig. 27</figureCitation>
) without metallic hue, whose colour can be attributed to the specimens age and has no taxonomic meaning.
</paragraph>
<paragraph id="8BFBCA71FFEC470AFCA6F870CD79FEF9" blockId="12.[828,1437,1449,2080]" lastBlockId="13.[151,761,190,2069]" lastPageId="13" lastPageNumber="432" pageId="12" pageNumber="431">
<taxonomicName id="4C44B1F2FFEC470BFCA6F870CB9CF80C" authority="Auzat, 1920" authorityName="Auzat" authorityYear="1920" box="[858,1272,1993,2016]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="431" phylum="Arthropoda" rank="species" species="clermonti" subGenus="Nessus">
<emphasis id="B9301663FFEC470BFCA6F870CCB8F80C" box="[858,988,1993,2016]" italics="true" pageId="12" pageNumber="431">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEC470BFC15F870CB67F80C" box="[1001,1027,1993,2016]" italics="true" pageId="12" pageNumber="431">N.</emphasis>
)
<emphasis id="B9301663FFEC470BFBF3F870CB11F80C" box="[1039,1141,1993,2016]" italics="true" pageId="12" pageNumber="431">clermonti</emphasis>
<bibRefCitation id="EFD5B780FFEC470BFB85F870CB9CF80C" author="AUZAT V." box="[1145,1272,1993,2016]" pageId="12" pageNumber="431" pagination="1 - 4" refId="ref20400" refString="AUZAT V. 1920: Quelques observations sur la sculpture superficielle des Histerides; varietes nouvelles, synonymies. L'Echange, Revue Linneenne 36 (399, Hors texte): 1 - 4." type="book chapter" year="1920">Auzat, 1920</bibRefCitation>
</taxonomicName>
is another form (
<figureCitation id="137FD6F4FFEC470BFCB8F850CCF2F7EC" box="[836,918,2025,2048]" captionStart="Figs 2930" captionStartId="11.[152,189,972,991]" captionTargetBox="[168,1435,194,945]" captionTargetId="figure-480@11.[148,1439,186,959]" captionTargetPageId="11" captionText="Figs 2930. Hypocaccus (Nessus) rubripes (Erichson, 1834). 29 Saprinus rubripes var. clermonti Auzat, 1920, holotype, habitus, dorsal view. 30 varieties of Hypocaccus (Nessus) rubripes as summed up by Jean Thérond." figureDoi="http://doi.org/10.5281/zenodo.4504936" httpUri="https://zenodo.org/record/4504936/files/figure.png" pageId="12" pageNumber="431">Fig. 29</figureCitation>
), which represents specimens worn-out by age; its frontal disc is almost glabrous (missing the numerous elongate rugae of the typical form) and the fore tibiae are devoid of teeth and denticles (worn off by age). This form has likewise no taxonomic meaning.
</paragraph>
<paragraph id="8BFBCA71FFED470AFF49FEA7CDF5FDF9" blockId="13.[151,761,190,2069]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFF49FEA7CD09FED9" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[181,621,286,309]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="corsicus" subGenus="Nessus">
<emphasis id="B9301663FFED470AFF49FEA7CE53FED9" box="[181,311,286,309]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFED470AFEB4FEA7CE06FED9" box="[328,354,286,309]" italics="true" pageId="13" pageNumber="432">N.</emphasis>
)
<emphasis id="B9301663FFED470AFE8DFEA7CEAEFED9" box="[369,458,286,309]" italics="true" pageId="13" pageNumber="432">corsicus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFE2EFEA7CD09FED9" author="MARSEUL S. A." box="[466,621,286,309]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
occurring in
<collectingCountry id="F3538AE1FFED470AFF6BFE87CF8FFEB9" box="[151,235,318,341]" name="France" pageId="13" pageNumber="432">Corsica</collectingCountry>
, Sardinia and Sicily is a viable candidate for a subspecies (
<figureCitation id="137FD6F4FFED470AFEE5FEE7CE0CFE99" box="[281,360,350,373]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="13" pageNumber="432">Fig. 28</figureCitation>
) and in fact has been treated as such (even as a
<emphasis id="B9301663FFED470AFEFFFEC7CE00FE79" box="[259,356,382,405]" italics="true" pageId="13" pageNumber="432">bona fide</emphasis>
species by several authors (e.g. SAINT- CLAIRE- DEVILLE 1907)). In this form, the sutural elytral stria is missing, the elytral punctation is much finer and its aedeagus is somewhat shorter and less dilated. In
<collectingCountry id="F3538AE1FFED470AFD5CFE67CD90FE19" box="[672,756,478,501]" name="France" pageId="13" pageNumber="432">Corsica</collectingCountry>
, this form occurs together with the typical form.
</paragraph>
<paragraph id="8BFBCA71FFED470AFF49FDA7CEABFA39" blockId="13.[151,761,190,2069]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFF49FDA7CD03FDD9" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[181,615,542,565]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="granarius" subGenus="Nessus">
<emphasis id="B9301663FFED470AFF49FDA7CE57FDD9" box="[181,307,542,565]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFED470AFEBCFDA7CE3CFDD9" box="[320,344,542,565]" italics="true" pageId="13" pageNumber="432">N.</emphasis>
)
<emphasis id="B9301663FFED470AFE98FDA7CEADFDD9" box="[356,457,542,565]" italics="true" pageId="13" pageNumber="432">granarius</emphasis>
<bibRefCitation id="EFD5B780FFED470AFE31FDA7CD03FDD9" author="ERICHSON W. F." box="[461,615,542,565]" pageId="13" pageNumber="432" refId="ref20612" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F. (ed.): Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book" year="1834">Erichson, 1834</bibRefCitation>
</taxonomicName>
occurs chiefly in southern
<collectingCountry id="F3538AE1FFED470AFEEBFD87CE3BFDB9" box="[279,351,574,597]" name="Russia" pageId="13" pageNumber="432">Russia</collectingCountry>
, in the Caucasian republics (
<collectingCountry id="F3538AE1FFED470AFD68FD87CD90FDB9" box="[660,756,574,597]" name="Armenia" pageId="13" pageNumber="432">Armenia</collectingCountry>
,
<collectingCountry id="F3538AE1FFED470AFF6BFDE7CE6BFD99" box="[151,271,606,629]" name="Azerbaijan" pageId="13" pageNumber="432">Azerbaijan</collectingCountry>
,
<collectingCountry id="F3538AE1FFED470AFEE1FDE7CE10FD99" box="[285,372,606,629]" name="Georgia" pageId="13" pageNumber="432">Georgia</collectingCountry>
) and is characterized by anteriorly effaced sutural elytral stria, occasionally not connected with the fourth dorsal elytral stria (
<figureCitation id="137FD6F4FFED470AFE24FD27CD4DFD59" box="[472,553,670,693]" captionStart="Figs 2528" captionStartId="10.[151,188,2030,2049]" captionTargetBox="[213,1403,494,1971]" captionTargetId="figure-13@10.[151,1436,456,2005]" captionTargetPageId="10" captionText="Figs 2528. Hypocaccus (Nessus) rubripes, dorsal habitus. 25 Saprinus rubripes Erichson, 1834, syntype. 26 Saprinus granarius Erichson, 1834, syntype. 27 Saprinus arenarius Marseul, 1855, lectotype. 28 Saprinus corsicus Marseul, 1855,?syntype." figureDoi="http://doi.org/10.5281/zenodo.4504934" httpUri="https://zenodo.org/record/4504934/files/figure.png" pageId="13" pageNumber="432">Fig. 26</figureCitation>
).
<bibRefCitation id="EFD5B780FFED470AFDC1FD27CFA5FD39" author="REICHARDT A." pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932: 129</bibRefCitation>
, figs 16: A, B), when elaborating on different forms and variation of this species put the most emphasis on the different pronotal shapes between (geographically) different populations. The two extremes of the pronotum are represented by acute (
<bibRefCitation id="EFD5B780FFED470AFEBEFC87CD44FCB9" author="REICHARDT A." box="[322,544,830,853]" pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT 1932: 129</bibRefCitation>
, fig. 16:A)
<emphasis id="B9301663FFED470AFD6EFC87CDCBFCB9" box="[658,687,830,853]" italics="true" pageId="13" pageNumber="432">vs.</emphasis>
obtuse (
<bibRefCitation id="EFD5B780FFED470AFF62FCE7CE1DFC99" author="REICHARDT A." box="[158,377,862,885]" pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT 1932: 129</bibRefCitation>
, fig. 16:B) anterior pronotal angles. In case of acute anterior pronotal angles the marginal pronotal stria is well marked and easily discernible, whereas in case of obtuse anterior pronotal angles the marginal pronotal stria is rather difficult to distinguish.
<bibRefCitation id="EFD5B780FFED470AFDCAFC67CD9CFC19" author="REICHARDT A." box="[566,760,990,1013]" pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
noted that the form with acute anterior pronotal angles is much more frequent and almost all specimens from southern
<collectingCountry id="F3538AE1FFED470AFF6BFB87CFBBFBB9" box="[151,223,1086,1109]" name="Russia" pageId="13" pageNumber="432">Russia</collectingCountry>
, Crimea and Caucasian republics belong to it, as well as most specimens identified as the
<taxonomicName id="4C44B1F2FFED470AFDDCFBE7CDE3FB99" authorityName="Erichson" authorityYear="1834" box="[544,647,1118,1141]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="granarius">
<emphasis id="B9301663FFED470AFDDCFBE7CDE3FB99" box="[544,647,1118,1141]" italics="true" pageId="13" pageNumber="432">granarius</emphasis>
</taxonomicName>
form. On the other hand, most specimens from North Africa belong to the form with obtuse anterior pronotal angles. Although there is no clear-cut difference between these two forms, specimens of both extremes are very different. A thorough morphological as well as molecular study of the possible superspecies
<taxonomicName id="4C44B1F2FFED470AFECDFAA7CEBFFAD9" baseAuthorityName="Erichson" baseAuthorityYear="1834" box="[305,475,1310,1333]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="rubripes" subGenus="Nessus">
<emphasis id="B9301663FFED470AFECDFAA7CE2FFAD9" box="[305,331,1310,1333]" italics="true" pageId="13" pageNumber="432">H.</emphasis>
(
<emphasis id="B9301663FFED470AFEA6FAA7CE17FAD9" box="[346,371,1310,1333]" italics="true" pageId="13" pageNumber="432">N.</emphasis>
)
<emphasis id="B9301663FFED470AFE7EFAA7CEBFFAD9" box="[386,475,1310,1333]" italics="true" pageId="13" pageNumber="432">rubripes</emphasis>
</taxonomicName>
could possibly resolve this conundrum. In fact, the small handwritten label (
<figureCitation id="137FD6F4FFED470AFD5EFA87CD95FAB9" box="[674,753,1342,1365]" captionStart="Figs 2930" captionStartId="11.[152,189,972,991]" captionTargetBox="[168,1435,194,945]" captionTargetId="figure-480@11.[148,1439,186,959]" captionTargetPageId="11" captionText="Figs 2930. Hypocaccus (Nessus) rubripes (Erichson, 1834). 29 Saprinus rubripes var. clermonti Auzat, 1920, holotype, habitus, dorsal view. 30 varieties of Hypocaccus (Nessus) rubripes as summed up by Jean Thérond." figureDoi="http://doi.org/10.5281/zenodo.4504936" httpUri="https://zenodo.org/record/4504936/files/figure.png" pageId="13" pageNumber="432">Fig. 30</figureCitation>
) found in Théronds collection (housed in MNHN) summarizes the situation with different forms of this species up nicely. It is believed that Jean Thérond wrote up this label (Y. Gomy pers. comm. 2017).
</paragraph>
<paragraph id="8BFBCA71FFED470AFF49FA67CE57F899" blockId="13.[151,761,190,2069]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFF49FA67CD70FA19" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" box="[181,532,1502,1525]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFED470AFF49FA67CE17FA19" box="[181,371,1502,1525]" italics="true" pageId="13" pageNumber="432">Saprinus corsicus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFE84FA67CD70FA19" author="MARSEUL S. A." box="[376,532,1502,1525]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855</bibRefCitation>
</taxonomicName>
. The examined specimen does not bear the labels typical for Marseuls types, but it bears the tiny round golden label, which could indicate that it actually is the original Marseuls type specimen. The description of
<bibRefCitation id="EFD5B780FFED470AFE9CF9E7CD20F999" author="MARSEUL S. A." box="[352,580,1630,1653]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">MARSEUL (1855: 688)</bibRefCitation>
generally agrees with this specimen, because of the uncertainty of the exact status of the specimen we refrain from designating it as the
<typeStatus id="54FF74D3FFED470AFF3CF907CE41F939" box="[192,293,1726,1749]" pageId="13" pageNumber="432" type="lectotype">lectotype</typeStatus>
. In the general collection of the
<taxonomicName id="4C44B1F2FFED470AFD76F907CD9CF939" authorityName="Gyllenhaal" authorityYear="1808" box="[650,760,1726,1749]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
housed at MNHN (which contains Marseuls collection) the type specimen(s) of
<taxonomicName id="4C44B1F2FFED470AFE69F947CD36F8F9" authorityName="Marseul" authorityYear="1855" box="[405,594,1790,1813]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="corsicus">
<emphasis id="B9301663FFED470AFE69F947CD36F8F9" box="[405,594,1790,1813]" italics="true" pageId="13" pageNumber="432">Saprinus corsicus</emphasis>
</taxonomicName>
is missing. It is possible that this specimen was part of Auzats collection, which was later purchased by Thérond (N. Dégallier, pers. comm., 2017).
</paragraph>
<paragraph id="8BFBCA71FFED470AFF49F8C7CD85F7F9" blockId="13.[151,761,190,2069]" pageId="13" pageNumber="432">
After the publication of
<bibRefCitation id="EFD5B780FFED470AFE3FF8C7CD14F879" author="LACKNER T." box="[451,624,1918,1941]" pageId="13" pageNumber="432" pagination="1 - 254" refId="ref21136" refString="LACKNER T. 2010: Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 50 (Supplementum): 1 - 254." type="journal article" year="2010">LACKNER (2010)</bibRefCitation>
this species (together with the rest of the taxa included in the subgenus
<taxonomicName id="4C44B1F2FFED470AFF6BF807CEF3F839" authority="Reichardt, 1932" authorityName="Reichardt" authorityYear="1932" box="[151,407,1982,2005]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">
<emphasis id="B9301663FFED470AFF6BF807CF85F839" box="[151,225,1982,2005]" italics="true" pageId="13" pageNumber="432">Nessus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFF16F807CEF3F839" author="REICHARDT A." box="[234,407,1982,2005]" pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">Reichardt, 1932</bibRefCitation>
</taxonomicName>
) was transferred from the genus
<taxonomicName id="4C44B1F2FFED470AFF6BF867CEBAF819" authority="Bickhardt, 1914" authorityName="Bickhardt" authorityYear="1914" box="[151,478,2014,2037]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9301663FFED470AFF6BF867CE4AF819" box="[151,302,2014,2037]" italics="true" pageId="13" pageNumber="432">Hypocacculus</emphasis>
Bickhardt, 1914
</taxonomicName>
into genus
<taxonomicName id="4C44B1F2FFED470AFDABF867CE5AF7F9" authority="C. Thomson, 1867" authorityName="C. Thomson" authorityYear="1867" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9301663FFED470AFDABF867CDBDF819" box="[599,729,2014,2037]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
C.
<bibRefCitation id="EFD5B780FFED470AFF6BF847CE5AF7F9" author="THOMSON C. P." box="[151,318,2046,2069]" pageId="13" pageNumber="432" refId="ref21955" refString="THOMSON C. P. 1867: Skandinaviens Coleoptera, synoptiskt bearbetade, IX. Berlingska Bocktryckeriet, Lund, 407 pp." type="book" year="1867">Thomson, 1867</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFD5B780FFED470AFE9BF847CE9AF7F9" author="MAZUR S." box="[359,510,2046,2069]" pageId="13" pageNumber="432" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">MAZUR (2011)</bibRefCitation>
without explanation.
</paragraph>
</subSubSection>
</treatment>
</document>