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<document id="7BBF618089EACA10DEA418FA6BE840B3" ID-DOI="10.5281/zenodo.5300375" ID-ISSN="0374-1036" ID-Zenodo-Dep="5300375" ID-ZooBank="631EB78E-94D0-4882-A91D-4532A5CCA923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630096395332" checkinUser="marcus" docAuthor="Nabozhenko, Maxim, Keskin, Bekir &amp; Ras, Institute of Arid Zones Ssc" docDate="2014" docId="03DA8330FFDBFFA01A8AFEC3FD4FF9E6" docLanguage="en" docName="ActaEntMusNatPra.2014.54.1.243-249.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Zophohelops montanatolicus Nabozhenko &amp; Keskin &amp; Ras 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="246" masterDocId="FFE3FB48FFD9FFA31B3BFF81FF9DFFD1" masterDocTitle="New data about nalassoid genera from south-eastern Anatolia with description of a new species of Zophohelops (Coleoptera: Tenebrionidae)" masterLastPageNumber="249" masterPageNumber="243" pageNumber="245" updateTime="1699143690612" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="69ABC08B54772E2103C29262E44AFB29">New data about nalassoid genera from south-eastern Anatolia with description of a new species of Zophohelops (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="26717F2D0574C5D37010BB37CB06E49A">Nabozhenko, Maxim</mods:namePart>
<mods:affiliation id="60E14C3EA18824DA79871BDA0CE9C705">Institute of Arid Zones SSC RAS, Southern Federal University, 41 Chekhov av., 105 Bolshaya Sadovaya str., Rostov-on-Don 344006, Russia; e-mail: nalassus @ mail. ru</mods:affiliation>
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<mods:namePart id="2956E9DD4021E3D08156C2A2E2E83B89">Keskin, Bekir</mods:namePart>
<mods:affiliation id="2B2614EDA25B26AA59986271D10EBC49">Zoology Department, Biology Section, Ege University, Bornova-Izmir 35100, Turkey; e-mail: bekir. keskin. phd @ gmail. com</mods:affiliation>
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<mods:namePart id="2CA9454DDFF74B560C3B23950795344D">Ras, Institute of Arid Zones Ssc</mods:namePart>
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<taxonomicName id="4C7349A5FFDBFFA11A8AFEC3FD6DFE8B" authority="Nabozhenko &amp; Keskin &amp; Ras, 2014" authorityName="Nabozhenko &amp; Keskin &amp; Ras" authorityYear="2014" box="[433,752,322,346]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="montanatolicus" status="sp. nov.">
<emphasis id="B907EE34FFDBFFA11A8AFEC3FD6DFE8B" bold="true" box="[433,752,322,346]" italics="true" pageId="2" pageNumber="245">Zophohelops montanatolicus</emphasis>
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<paragraph id="8BCC3226FFDBFFA11968FEE0FD36FEA5" blockId="2.[150,1127,353,468]" box="[595,683,353,372]" pageId="2" pageNumber="245">
(
<figureCitation id="13482EA3FFDBFFA11962FEE0FD38FEA5" box="[601,677,353,372]" captionStart-0="Figs 28" captionStart-1="Fig" captionStartId-0="3.[113,150,1265,1284]" captionStartId-1="4.[771,802,767,786]" captionTargetBox-0="[275,921,219,1251]" captionTargetBox-1="[151,751,218,946]" captionTargetId-0="figure-14@3.[267,945,216,1257]" captionTargetId-1="figure-223@4.[151,752,217,946]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs 28. Zophohelops montanatolicus sp. nov. 2 male; 3 female; 4 aedeagus, ventral view; 5 same, lateral view; 67 gastral spicula (6 ventral view, 7 lateral view); 8 sternite VIII of male." captionText-1="Fig. 9. Distribution of the genus Zophohelops and the subgenus Helopocerodes (Nalassus) in south-eastern Anatolia. Black circle N. faldermanni (Faldermann, 1837); white circle N. kaszabi Nabozhenko, 2001; square Z. montanatolicus sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.5300377" figureDoi-1="http://doi.org/10.5281/zenodo.5300379" httpUri-0="https://zenodo.org/record/5300377/files/figure.png" httpUri-1="https://zenodo.org/record/5300379/files/figure.png" pageId="2" pageNumber="245">Figs 29</figureCitation>
)
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<emphasis id="B907EE34FFDBFFA11BACFE0AFE86FE4F" bold="true" box="[151,283,395,414]" pageId="2" pageNumber="245">Type material.</emphasis>
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<typeStatus id="54C88C84FFDBFFA11A1BFE0AFEEEFE4D" box="[288,371,395,414]" pageId="2" pageNumber="245" type="holotype">HOLOTYPE</typeStatus>
: J(
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) and
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(
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: 4 JJ
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;
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: 1 J
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(dry specimens), 3 JJ
<specimenCount id="9D75F9AFFFDBFFA11F67FE0AFF2FFE68" pageId="2" pageNumber="245" type="female">4 ♀♀</specimenCount>
(in ethanol)):
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<collectingCountry id="F36472B6FFDBFFA11A10FE27FE1AFE68" box="[299,391,422,441]" name="Turkey" pageId="2" pageNumber="245">TURKEY</collectingCountry>
:
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PROVINCE:
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,
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<collectingDate id="EF89ED0EFFDBFFA119F1FE27FCBDFE68" box="[714,800,422,441]" pageId="2" pageNumber="245" value="2013-06-03">3.06.2013</collectingDate>
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,
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,
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(
<collectorName id="268657F0FFDBFFA11BA6FE40FF7EFE05" box="[157,227,449,468]" pageId="2" pageNumber="245">Leg. M.</collectorName>
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. &amp; S.
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.
<collectorName id="268657F0FFDBFFA11A03FE40FE22FE05" box="[312,447,449,468]" pageId="2" pageNumber="245">Nabozhenko, B</collectorName>
.
<collectorName id="268657F0FFDBFFA11AF3FE40FDBDFE05" box="[456,544,449,468]" pageId="2" pageNumber="245">Keskin, A</collectorName>
. Pektaú)
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.
</paragraph>
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<subSubSection id="C36961ADFFDBFFA11BACFE6DFDA6FA52" pageId="2" pageNumber="245" type="description">
<paragraph id="8BCC3226FFDBFFA11BACFE6DFD52FDD2" blockId="2.[151,1128,492,1603]" box="[151,719,492,515]" pageId="2" pageNumber="245">
<emphasis id="B907EE34FFDBFFA11BACFE6DFEBFFDD2" bold="true" box="[151,290,492,515]" pageId="2" pageNumber="245">Description.</emphasis>
Body length
<quantity id="4C8B9FC3FFDBFFA11A89FE6DFD90FDD2" box="[434,525,492,515]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="245" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
, width
<quantity id="4C8B9FC3FFDBFFA11960FE6DFD56FDD2" box="[603,715,492,515]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="2" pageNumber="245" unit="mm" value="3.15" valueMax="3.3" valueMin="3.0">33.3 mm</quantity>
.
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<paragraph id="8BCC3226FFDBFFA11B8EFD8AFCF8FCD2" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">
<emphasis id="B907EE34FFDBFFA11B8EFD8AFF69FDF2" bold="true" box="[181,244,523,547]" pageId="2" pageNumber="245">
<emphasis id="B907EE34FFDBFFA11B8EFD8AFF70FDF2" bold="true" box="[181,237,523,547]" italics="true" pageId="2" pageNumber="245">Male</emphasis>
.
</emphasis>
Body robust, dark brown, elytra almost black; pronotum and head shining, elytra dull. Head widest at level of eyes. Ratio of head width at level of eyes to distance between eyes 1.37. Eyes small, weakly convex, widely spaced. Anterior margin of clypeus straight, with weakly projected angles. Outer margin of head between genae and clypeus weakly sinuate. Head with moderately coarse and dense punctation (puncture diameters equal or little wider than distance between them). Surface of head with short recumbent hairs. Temple groove absent. Antennae short, antennomeres 1011 extending beyond base of pronotum when directed backwards, antennomeres 811 more widened than others.
</paragraph>
<paragraph id="8BCC3226FFDBFFA11B8EFC8DFB89FBD2" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">Pronotum transverse (1.3 times as wide as long), widest in middle, rectangular, 1.75 times as wide as head. Lateral margins and base of pronotum weakly rounded, anterior margin almost straight. Anterior angles of pronotum rounded, weakly obtuse. Disc moderately convex with narrowly flattened margins. Lateral and basal bead of pronotum thickened, anterior margin with narrow bead interrupted in middle. Punctation moderately coarse, not dense (puncture diameters 23 times lower than distance between them); punctures slightly elongated on sides of disc. Prothoracic hypomeron with very ¿ne longitudinal wrinkles, outer margin distinctly flattened. Prosternal process very weakly convex, with small tubercle in apical part.</paragraph>
<paragraph id="8BCC3226FFDBFFA11B8EFB8DFDD8FB12" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">Elytra distinctly convex, moderately broad (1.4 times as wide as long), widest before middle, 1.14 times as wide and 2.17× as long as pronotum, 2× as wide as head. Punctures in striae oval, not connected in grooves; elytral intervals flat, with ¿ne and sparse punctation. Intervals 1 and 9 connected at apex. Epipleural carina very wide, especially at base, completely visible dorsally, almost reaching elytral apex. Epipleura broad, depressed from base to half of length, almost reaching sutural angle.</paragraph>
<paragraph id="8BCC3226FFDBFFA11B8EFB4DFCC7FAD2" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">Legs short, tibiae straight, densely pubescent on inner side. Protarsi not widened, tarsomeres 2 and 3 transverse. All trochanters with dense brush of short hairs.</paragraph>
<paragraph id="8BCC3226FFDBFFA11B8EFA8DFC84FA92" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">Ventral side of body with short setation.Abdominal ventrites with ¿ne punctation, ventrite 1 with small hair brush in middle, ventrite 5 beaded at apex.</paragraph>
<paragraph id="8BCC3226FFDBFFA11B8EFACAFDA6FA52" blockId="2.[151,1128,492,1603]" pageId="2" pageNumber="245">
<emphasis id="B907EE34FFDBFFA11B8EFACAFE92FAB2" bold="true" box="[181,271,1355,1379]" italics="true" pageId="2" pageNumber="245">Female.</emphasis>
Body more robust, antennae shorter, pronotum widest behind middle, abdominal ventrite 1 without hair brush in middle.
</paragraph>
</subSubSection>
<subSubSection id="C36961ADFFDBFFA01BACFA0DFEB8FA66" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245" type="diagnosis">
<paragraph id="8BCC3226FFDBFFA01BACFA0DFEB8FA66" blockId="2.[151,1128,492,1603]" lastBlockId="3.[113,1089,1375,1591]" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245">
<emphasis id="B907EE34FFDBFFA11BACFA0DFE12FA72" bold="true" box="[151,399,1420,1443]" pageId="2" pageNumber="245">Differential diagnosis.</emphasis>
The new species is closely related to
<taxonomicName id="4C7349A5FFDBFFA11829FA0DFF51FA12" authority="Skopin, 1964" authorityName="Skopin" authorityYear="1964" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="protzenkoi">
<emphasis id="B907EE34FFDBFFA11829FA0DFB92FA72" box="[786,1039,1420,1443]" italics="true" pageId="2" pageNumber="245">Zophohelops protzenkoi</emphasis>
Skopin, 1964
</taxonomicName>
(Tien Shan: Chatkal Range) which also has widened and dorsally visible epipleural carina and hair brush on the ¿rst abdominal ventrite.
<taxonomicName id="4C7349A5FFDBFFA119B1FA4DFC5DFA32" authority="Nabozhenko &amp; Keskin &amp; Ras, 2014" authorityName="Nabozhenko &amp; Keskin &amp; Ras" authorityYear="2014" box="[650,960,1484,1507]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="montanatolicus" status="sp. nov.">
<emphasis id="B907EE34FFDBFFA119B1FA4DFC5DFA32" box="[650,960,1484,1507]" italics="true" pageId="2" pageNumber="245">Zophohelops montanatolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234534FFFDBFFA118FCFA4DFB86FA32" box="[967,1051,1484,1507]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C7349A5FFDBFFA11BE9FA6DFEC3F9D2" authorityName="Skopin" authorityYear="1964" box="[210,350,1516,1539]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="protzenkoi">
<emphasis id="B907EE34FFDBFFA11BE9FA6DFEC3F9D2" box="[210,350,1516,1539]" italics="true" pageId="2" pageNumber="245">Z. protzenkoi</emphasis>
</taxonomicName>
in robust body of male, pro- and mesotarsi of male not widened, shorter antennae and pronotum not sinuate at base. The new species also differs from
<taxonomicName id="4C7349A5FFDBFFA118E6F98DFE2BF992" authority="(Reitter, 1902)" baseAuthorityName="Reitter" baseAuthorityYear="1902" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="humeridens">
<emphasis id="B907EE34FFDBFFA118E6F98DFE8EF992" italics="true" pageId="2" pageNumber="245">Zophohelops humeridens</emphasis>
(
<bibRefCitation id="EFE24FD7FFDBFFA11A1AF9ADFE32F992" author="REITTER E." box="[289,431,1580,1603]" pageId="2" pageNumber="245" pagination="209 - 224" refId="ref3284" refString="REITTER E. 1902: Verschiedenes uber die Coleopteren der Tenebrioniden-Abtheilung Helopina. Deutsche Entomologische Zeitschrift 1901: 209 - 224." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
in dark brown body with dull elytra (
<taxonomicName id="4C7349A5FFDBFFA1187DF9ADFC43F992" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[838,990,1580,1603]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="humeridens">
<emphasis id="B907EE34FFDBFFA1187DF9ADFC43F992" box="[838,990,1580,1603]" italics="true" pageId="2" pageNumber="245">Z. humeridens</emphasis>
</taxonomicName>
has strongly shining, lacquered body), pubescent head, presence of hair spot on the ¿rst abdominal ventrite of male, absence of protruded humeral angles in male, and dorsally visible epipleural carina in female.
</paragraph>
</subSubSection>
<caption id="DF0C62AEFFDAFFA01B4AFB70FCFFFACE" ID-DOI="http://doi.org/10.5281/zenodo.5300377" ID-Zenodo-Dep="5300377" httpUri="https://zenodo.org/record/5300377/files/figure.png" pageId="3" pageNumber="246" startId="3.[113,150,1265,1284]" targetBox="[275,921,219,1251]" targetPageId="3">
<paragraph id="8BCC3226FFDAFFA01B4AFB70FCFFFACE" blockId="3.[113,1088,1265,1311]" pageId="3" pageNumber="246">
Figs 28.
<taxonomicName id="4C7349A5FFDAFFA01BFDFB70FE23FAD5" authority="Nabozhenko &amp; Keskin &amp; Ras, 2014" authorityName="Nabozhenko &amp; Keskin &amp; Ras" authorityYear="2014" box="[198,446,1265,1284]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="montanatolicus" status="sp. nov.">
<emphasis id="B907EE34FFDAFFA01BFDFB70FE23FAD5" box="[198,446,1265,1284]" italics="true" pageId="3" pageNumber="246">Zophohelops montanatolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234534FFFDAFFA01AF8FB70FD9BFAD5" box="[451,518,1265,1284]" pageId="3" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
2 male; 3 female; 4 aedeagus, ventral view; 5 same, lateral view; 67 gastral spicula (6 ventral view, 7 lateral view); 8 sternite VIII of male.
</paragraph>
</caption>
<subSubSection id="C36961ADFFDAFFA01B4AFA41FD4FF9E6" pageId="3" pageNumber="246" type="etymology">
<paragraph id="8BCC3226FFDAFFA01B4AFA41FF7CFA26" blockId="3.[113,1089,1375,1591]" pageId="3" pageNumber="246">
<emphasis id="B907EE34FFDAFFA01B4AFA41FF6FFA06" bold="true" box="[113,242,1472,1495]" pageId="3" pageNumber="246">Etymology.</emphasis>
The name
<taxonomicName id="4C7349A5FFDAFFA01A5DFA41FD97FA06" authorityName="Nabozhenko &amp; Keskin &amp; Ras" authorityYear="2014" box="[358,522,1472,1495]" class="Insecta" family="Tenebrionidae" genus="Zophohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="montanatolicus">
<emphasis id="B907EE34FFDAFFA01A5DFA41FD97FA06" box="[358,522,1472,1495]" italics="true" pageId="3" pageNumber="246">montanatolicus</emphasis>
</taxonomicName>
is formed from two words:
<emphasis id="B907EE34FFDAFFA0180AFA41FC04FA06" box="[817,921,1472,1495]" italics="true" pageId="3" pageNumber="246">montanus</emphasis>
and
<emphasis id="B907EE34FFDAFFA018F0FA41FBA7FA06" box="[971,1082,1472,1495]" italics="true" pageId="3" pageNumber="246">anatolicus</emphasis>
. Adjective.
</paragraph>
<paragraph id="8BCC3226FFDAFFA01B4AF981FD4FF9E6" blockId="3.[113,1089,1375,1591]" pageId="3" pageNumber="246">
<emphasis id="B907EE34FFDAFFA01B4AF981FF4DF9C6" bold="true" box="[113,208,1536,1559]" pageId="3" pageNumber="246">Habitat.</emphasis>
The species was found in high mountain alpine plateau under very small stones without lichens, in areas with very dense and short grass.
</paragraph>
</subSubSection>
</treatment>
</document>