treatments-xml/data/03/C7/F6/03C7F607FFFFFFC0A7AB3455EBB566F9.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5562918" ID-GBIF-Dataset="9de32fb5-ae5c-4552-afdb-9cfdcad56ce9" ID-GBIF-Taxon="188614993" ID-Zenodo-Dep="5562918" approvalRequired="2" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" checkinTime="1633713770872" checkinUser="carolina" docAuthor="Deschodt, Christian M. &amp; Davis, Adrian L. V." docDate="2017" docId="03C7F607FFFFFFC0A7AB3455EBB566F9" docLanguage="en" docName="AnnDNMNH.7.109-114.pdf.imf" docOrigin="Annals of the Ditsong National Museum of Natural History 7 (7)" docTitle="Byrrhidium OvaLe Harold 1869" docType="key" docVersion="6" lastPageId="4" lastPageNumber="113" masterDocId="FFFE8E7FFFFCFFC4A72A3246EF2D645E" masterDocTitle="Transfer of three species of Namakwanus Scholtz &amp; Howden to Versicorpus Deschodt, Davis &amp; Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="114" masterPageNumber="109" pageId="3" pageNumber="112" updateTime="1634154866724" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Transfer of three species of Namakwanus Scholtz &amp; Howden to Versicorpus Deschodt, Davis &amp; Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Deschodt, Christian M.</mods:namePart>
<mods:affiliation>Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa</mods:affiliation>
<mods:nameIdentifier type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Davis, Adrian L. V.</mods:namePart>
<mods:affiliation>Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa</mods:affiliation>
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<mods:date>2017</mods:date>
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<mods:number>2017-05-01</mods:number>
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<mods:number>7</mods:number>
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<paragraph blockId="3.[129,644,1555,1630]" pageId="3" pageNumber="112">
<heading level="2" pageId="3" pageNumber="112">
<emphasis bold="true" pageId="3" pageNumber="112">
NEW KEY TO THE GENERA AND SPECIES IN THE
<taxonomicName authorityName="OvaLe Harold" authorityYear="1869" box="[290,443,1581,1603]" class="Insecta" family="Scarabaeidae" genus="Byrrhidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="112" phylum="Arthropoda" rank="genus">BYRRHIDIUM</taxonomicName>
GROUP
</emphasis>
</heading>
(adapted from
<bibRefCitation author="DESCHODT, C. M. &amp; DAVIS, A. L. V. &amp; SCHOLTZ, C. H." box="[347,576,1608,1630]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="112" pagination="490 - 500" part="4147" refId="ref3603" refString="DESCHODT, C. M., DAVIS, A. L. V. and SCHOLTZ, C. H., 2016. A new monotypic genus, a species synonymy and nomenclatural corrections in the arid-adapted Canthonini (Scarabaeidae, Scarabaeinae) from the Succulent Karoo Biome of south-western Africa. Zootaxa 4147 (4): 490 - 500." title="A new monotypic genus, a species synonymy and nomenclatural corrections in the arid-adapted Canthonini (Scarabaeidae, Scarabaeinae) from the Succulent Karoo Biome of south-western Africa" type="journal article" year="2016">Deschodt et al., 2016</bibRefCitation>
)
</paragraph>
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<subSubSection lastPageId="4" lastPageNumber="113" pageId="3" pageNumber="112" type="key">
<key lastPageId="4" lastPageNumber="113" pageId="3" pageNumber="112">
<keyStep pageId="3" pageNumber="112">
<paragraph blockId="3.[126,644,1662,1716]" box="[126,644,1662,1684]" pageId="3" pageNumber="112">
<keyLead box="[126,644,1662,1684]" pageId="3" pageNumber="112">1 Foretibiawiththreeexternal denticles..... 2</keyLead>
</paragraph>
<paragraph blockId="3.[126,644,1662,1716]" box="[126,644,1694,1716]" pageId="3" pageNumber="112">
<keyLead box="[126,644,1694,1716]" pageId="3" pageNumber="112">— Foretibiawithtwoexternaldenticles ...... 8</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="112">
<paragraph blockId="3.[679,1197,1289,1426]" pageId="3" pageNumber="112">
<keyLead pageId="3" pageNumber="112">
2 Habitus relatively large, longer than
<quantity box="[1100,1173,1290,1312]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="112" unit="mm" value="10.0">10 mm</quantity>
and elongate
<taxonomicName authority="Deschodt, Davis andScholtz ........................." authorityName="Deschodt, Davis and Scholtz." class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[874,994,1317,1339]" italics="true" pageId="3" pageNumber="112">Versicorpus</emphasis>
Deschodt, Davis dScholtz
</taxonomicName>
......................... 3
</keyLead>
</paragraph>
<paragraph blockId="3.[679,1197,1289,1426]" pageId="3" pageNumber="112">
<keyLead pageId="3" pageNumber="112">
— Habitus smaller than
<quantity box="[948,1020,1377,1399]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="112" unit="mm" value="10.0">10 mm</quantity>
long and convex ................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="112">
<paragraph blockId="3.[679,1197,1437,1628]" pageId="3" pageNumber="112">
<keyLead pageId="3" pageNumber="112">
3 Mesometasternal suture curving forward; punctures on prosternum large; Erongo Mountain ....
<taxonomicName authority="Deschodt" authorityName="Deschodt, Davis &amp; Scholtz" authorityYear="2011" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="erongoense">
<emphasis box="[882,899,1491,1513]" italics="true" pageId="3" pageNumber="112">V.</emphasis>
<emphasis box="[901,1024,1491,1513]" italics="true" pageId="3" pageNumber="112">erongoense</emphasis>
Deschodt, Davis &amp; Scholtz
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[679,1197,1437,1628]" pageId="3" pageNumber="112">
<keyLead pageId="3" pageNumber="112">
— Mesometasternal suture straight; punctures on prosternum small; Tsondab Mountain ........................
<taxonomicName box="[1042,1197,1606,1628]" pageId="3" pageNumber="112">
<emphasis box="[1042,1114,1606,1628]" italics="true" pageId="3" pageNumber="112">V.streyi</emphasis>
(Frolov)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="4" lastPageNumber="113" pageId="3" pageNumber="112">
<paragraph blockId="3.[679,1197,1640,1716]" pageId="3" pageNumber="112">
<keyLead pageId="3" pageNumber="112">
4 Clypeal horns as long as, or longer than, distance between them .......
<taxonomicName authority="Scholtz &amp; Howden" authorityName="Scholtz &amp; Howden" authorityYear="1987" box="[824,1152,1694,1716]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[824,963,1694,1716]" italics="true" pageId="3" pageNumber="112">Namakwanus</emphasis>
Scholtz &amp; Howden
</taxonomicName>
… 5
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,201,277]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Clypeal horns shorter than distance between them ..
<taxonomicName authority="Deschodt &amp; Davis, 2017" authorityName="Deschodt &amp; Davis" authorityYear="2017" box="[204,599,255,277]" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[204,324,255,277]" italics="true" pageId="4" pageNumber="113">Namaphilus</emphasis>
<taxonomicNameLabel box="[328,419,255,277]" pageId="4" pageNumber="113" rank="genus">gen. nov.</taxonomicNameLabel>
Deschodt &amp; Davis
</taxonomicName>
… 6
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<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,645,288,584]" pageId="4" pageNumber="113">
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5 Space between clypeal horns about one and a half times more at base than the width of a single horn at apex; clypeal horns not upturned and slightly pointing inward at the apex; Windhoek area ................
<taxonomicName authority="Scholtz &amp; Howden" authorityName="Scholtz &amp; Howden" authorityYear="1987" box="[381,643,422,444]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="irishi">
<emphasis box="[381,405,422,444]" italics="true" pageId="4" pageNumber="113">N.</emphasis>
<emphasis box="[408,454,422,444]" italics="true" pageId="4" pageNumber="113">iriSsi</emphasis>
Scholtz &amp; Howden
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[126,645,288,584]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Spacebetween clypealhornsabouttwotimes greater at base than the width of a single horn at apex; clypeal horns more upturned and slightly pointing inward at the apex; Gobabebarea ........
<taxonomicName box="[433,641,562,584]" pageId="4" pageNumber="113">
<emphasis box="[433,537,562,584]" italics="true" pageId="4" pageNumber="113">N. scholtzi</emphasis>
spec. nov
</taxonomicName>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,645,595,678]" pageId="4" pageNumber="113">
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6 Area between clypeal teeth somewhat arcuate ..........
<taxonomicName authority="Deschodt &amp; Davis, 2017" authorityName="Deschodt &amp; Davis" authorityYear="2017" box="[393,645,622,644]" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="ameibensis" status="spec.nov.">
<emphasis box="[393,537,622,644]" italics="true" pageId="4" pageNumber="113">N. ameibensis</emphasis>
spec. nov.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[126,645,595,678]" box="[126,644,656,678]" pageId="4" pageNumber="113">
<keyLead box="[126,644,656,678]" pageId="4" pageNumber="113">— Areabetweenclypealteethstraight ...... 7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,689,878]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
7 Tips of parameres with two notches in frontal view; Naukluft Mountains to the south of the Tsondab River canyon ....
<taxonomicName authorityName="Deschodt &amp; Davis" authorityYear="2017" baseAuthorityName="Deschodt, Davis &amp; Scholtz" baseAuthorityYear="2011" box="[229,643,769,791]" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="endroedyi">
<emphasis box="[229,358,769,791]" italics="true" pageId="4" pageNumber="113">N. endroedyi</emphasis>
(Deschodt,Davis &amp; Scholtz)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,689,878]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Tips of parameres without notches in frontal view; HardapDam ....
<taxonomicName authorityName="Deschodt &amp; Davis" authorityYear="2017" baseAuthorityName="Deschodt &amp; Scholtz" baseAuthorityYear="2007" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="davisi">
<emphasis box="[427,512,829,851]" italics="true" pageId="4" pageNumber="113">N. davisi</emphasis>
(Deschodt &amp; Scholtz)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,889,998]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
8 Clypeus clearly bidentate with clypealhorns long ......
<taxonomicName authority="Frolov &amp; Scholtz" authorityName="deSChOdti Frolov &amp; Scholtz" authorityYear="2003" box="[308,599,916,938]" class="Insecta" family="Scarabaeidae" genus="Dicranocara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[308,432,916,938]" italics="true" pageId="4" pageNumber="113">Dicranocara</emphasis>
Frolov &amp; Scholtz
</taxonomicName>
… 9
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,889,998]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">— Clypeus quadridentate, with short medial clypealhorns ....................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,1010,1118]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">9 Aedeagus without lateral horns on parameres ............................. 10</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,1010,1118]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">— Aedeagus with lateral horns on parameres .................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,1130,1265]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
10 Apex of parameresbulky; northern Richtersveld......
<taxonomicName box="[297,644,1156,1178]" pageId="4" pageNumber="113">
<emphasis box="[297,440,1156,1178]" italics="true" pageId="4" pageNumber="113">D. inexpectata</emphasis>
Deschodt &amp; Scholtz
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,1130,1265]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Apex of parameres slender; southern Richtersveld, Boom River .............
<taxonomicName box="[349,644,1243,1265]" pageId="4" pageNumber="113">
<emphasis box="[349,477,1243,1265]" italics="true" pageId="4" pageNumber="113">D. deschodti</emphasis>
Frolov &amp; Scholtz
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,1277,1466]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
11 Side horns on aedeagus extending somewhat sideways, tip curved downwards; Richtersveld National Park in South Africa ..........
<taxonomicName box="[312,644,1357,1379]" pageId="4" pageNumber="113">
<emphasis box="[312,440,1357,1379]" italics="true" pageId="4" pageNumber="113">D. tatasensis</emphasis>
Deschodt &amp; Scholtz
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,1277,1466]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Side horns on aedeagus slightly forward, tip not curved downwards; Fish River Canyon .........
<taxonomicName box="[375,643,1444,1466]" pageId="4" pageNumber="113">
<emphasis box="[375,560,1444,1466]" italics="true" pageId="4" pageNumber="113">D. vandersmisseni</emphasis>
Moretto
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,644,1477,1666]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
12 Clypealhorns short, mesometasternal suture not well defined; south of Orange River .............
<taxonomicName authority="Harold" authorityName="OvaLe Harold" authorityYear="1869" box="[405,586,1531,1553]" class="Insecta" family="Scarabaeidae" genus="Byrrhidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[405,514,1531,1553]" italics="true" pageId="4" pageNumber="113">Byrrhidium</emphasis>
Harold
</taxonomicName>
… 13
</keyLead>
</paragraph>
<paragraph blockId="4.[126,644,1477,1666]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Clypealhorns of medium length, mesometasternal suture well defined; north of Orange River ......
<taxonomicName authorityName="Deschodt, Davis &amp; Scholtz" authorityYear="2016" box="[253,644,1644,1666]" class="Insecta" family="Scarabaeidae" genus="Drogo" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="species" species="stalsi">
<emphasis box="[253,372,1644,1666]" italics="true" pageId="4" pageNumber="113">Drogo stalsi</emphasis>
Deschodt, Davis &amp; Scholtz
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="113">
<paragraph blockId="4.[126,620,1677,1699]" lastBlockId="4.[679,1196,195,356]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
13 Aedeagus with side protrusions, fore tibia with outer teeth parallel; Richtersveld in South Africa ...........
<taxonomicName box="[876,1196,248,270]" pageId="4" pageNumber="113">
<emphasis box="[876,1007,248,270]" italics="true" pageId="4" pageNumber="113">B. convexum</emphasis>
Scholtz &amp; Howden
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[679,1196,195,356]" pageId="4" pageNumber="113">
<keyLead pageId="4" pageNumber="113">
— Aedeagus plain with no side protrusions, fore tibia with outer teeth diverging; Namaqualand..................
<taxonomicName box="[1044,1196,334,356]" pageId="4" pageNumber="113">
<emphasis box="[1044,1123,334,356]" italics="true" pageId="4" pageNumber="113">B. ovale</emphasis>
Harold
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection pageId="4" pageNumber="113" type="discussion">
<paragraph blockId="4.[679,1201,389,1699]" pageId="4" pageNumber="113">
<heading level="2" pageId="4" pageNumber="113">
<emphasis bold="true" pageId="4" pageNumber="113">
NOTES ON THE CURRENTLY KNOWN SPECIES IN THE
<taxonomicName authorityName="OvaLe Harold" authorityYear="1869" box="[875,1028,415,437]" class="Insecta" family="Scarabaeidae" genus="Byrrhidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="genus">BYRRHIDIUM</taxonomicName>
GROUP
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="113" type="discussion">
<paragraph blockId="4.[679,1201,389,1699]" pageId="4" pageNumber="113">
The
<taxonomicName authorityName="OvaLe Harold" authorityYear="1869" box="[742,851,444,466]" class="Insecta" family="Scarabaeidae" genus="Byrrhidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[742,851,444,466]" italics="true" pageId="4" pageNumber="113">Byrrhidium</emphasis>
</taxonomicName>
group is confined to the arid west of Namibia and South Africa (
<figureCitation box="[1001,1068,470,492]" captionStart="Fig" captionStartId="3.[639,669,1187,1204]" captionTargetBox="[218,1104,185,1174]" captionTargetId="figure-305@3.[230,1104,185,1174]" captionText="Fig. 2 Map showing the collecting localities of all the known species of the Byrrhidium group.The black outline indicates the distribution of rock hyraxes (shapefile supplied by IUCN). See online version for map in colour." figureDoi="http://doi.org/10.5281/zenodo.5562916" httpUri="https://zenodo.org/record/5562916/files/figure.png" pageId="4" pageNumber="113">Fig. 2</figureCitation>
). It currently consists of six genera and 14 described species. These species (Deschodt
<emphasis box="[950,993,524,546]" italics="true" pageId="4" pageNumber="113">et al</emphasis>
., 2007, 2011, 2016) may mostly be associated with dung middens of rock hyraxes (
<taxonomicName authority="CapenSiS (Pallas, 1766" authorityName="Storr" authorityYear="1780" box="[822,1149,577,599]" class="Mammalia" family="Procaviidae" genus="Procavia" kingdom="Animalia" order="Hyracoidea" pageId="4" pageNumber="113" phylum="Chordata" rank="genus">
<emphasis box="[822,1008,577,599]" italics="true" pageId="4" pageNumber="113">Procavia capensis</emphasis>
(
<bibRefCitation author="PALLAS, P. S." bookContentInfo="224 pp." box="[1019,1149,577,599]" journalOrPublisher="A. P. van Cleef, La Haye" pageId="4" pageNumber="113" refId="ref4046" refString="PALLAS, P. S., 1766. Miscellanea Zoologica. A. P. van Cleef, La Haye, 224 pp." title="Miscellanea Zoologica" type="book" year="1766">Pallas, 1766</bibRefCitation>
</taxonomicName>
)). As there are gaps in the known range of this group relative to that of rock hyraxes (
<figureCitation box="[949,1009,631,653]" captionStart="Fig" captionStartId="3.[639,669,1187,1204]" captionTargetBox="[218,1104,185,1174]" captionTargetId="figure-305@3.[230,1104,185,1174]" captionText="Fig. 2 Map showing the collecting localities of all the known species of the Byrrhidium group.The black outline indicates the distribution of rock hyraxes (shapefile supplied by IUCN). See online version for map in colour." figureDoi="http://doi.org/10.5281/zenodo.5562916" httpUri="https://zenodo.org/record/5562916/files/figure.png" pageId="4" pageNumber="113">Fig. 2</figureCitation>
), new related dung beetle species and genera may await discovery.
</paragraph>
</subSubSection>
</treatment>
</document>