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<document id="3A2EBDF97FA7A182885B956765B0D150" ID-CLB-Dataset="21809" ID-DOI="10.5852/ejt.2020.652" ID-GBIF-Dataset="64fd24bb-ee1d-44d7-ac6a-25e8d5f96777" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860236" ID-ZooBank="DC65C142-53F1-4416-A916-8F78C27DCF93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590592466678" checkinUser="valdenar" docAuthor="Garbin, Rafaella C., Bandyopadhyay, Saswati &amp; Joyce, Walter G." docDate="2020" docId="03C7CF0BC0381D7BFDABAF68FE299122" docLanguage="en" docName="ejt-652_garbin_bandyopadhyay_joyce.pdf.imf" docOrigin="European Journal of Taxonomy 652" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Batagur Gray 1856" docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="FFFEB773C0341D74FFD1A908FFA79736" masterDocTitle="A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan" masterLastPageNumber="67" masterPageNumber="1" pageNumber="13" updateTime="1698841626966" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A626C117A9EF06B9B798E1250DF8338F">A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan</mods:title>
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<mods:namePart id="E604EE0F60089A26F3B6EA9D55D3128F">Garbin, Rafaella C.</mods:namePart>
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<mods:affiliation id="0B3A4965EBC339A39964C67297C1CF41">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. &amp; urn: lsid: zoobank. org: author: A 8 FCEF 2 B- 7524 - 46 C 3 - 808 C-D 9 CBFBC 08017 &amp; Corresponding author: garbin. rafaella @ gmail. com</mods:affiliation>
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<mods:namePart id="42AE52411E6B93E54571B88C7AE99A8F">Bandyopadhyay, Saswati</mods:namePart>
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<mods:affiliation id="A0F844C9D8FE34B61792E5EC2AFC1510">Geological Studies Unit, Indian Statistical Institute, Kolkata; Barrackpore Trunk Road 233, Kolkata, India. &amp; Email: saswati @ isical. ac. in &amp; urn: lsid: zoobank. org: author: 02005 D 35 - 6 DA 9 - 4 A 7 E-B 838 - 16 F 4 FEF 108 DA</mods:affiliation>
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<mods:namePart id="4A0974A448851DE1C1802D77B2F80943">Joyce, Walter G.</mods:namePart>
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<treatment id="03C7CF0BC0381D7BFDABAF68FE299122" ID-DOI="http://doi.org/10.5281/zenodo.3861111" ID-GBIF-Taxon="164362568" ID-Zenodo-Dep="3861111" LSID="urn:lsid:plazi:treatment:03C7CF0BC0381D7BFDABAF68FE299122" httpUri="http://treatment.plazi.org/id/03C7CF0BC0381D7BFDABAF68FE299122" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<subSubSection id="C3742D96C0381D78FDABAF68FC1D914D" box="[634,954,1632,1659]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BD17E1DC0381D78FDABAF68FC1D914D" blockId="12.[634,954,1632,1694]" box="[634,954,1632,1659]" pageId="12" pageNumber="13">
<heading id="D099C971C0381D78FDABAF68FC1D914D" box="[634,954,1632,1659]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
Genus
<taxonomicName id="4C6E059EC0381D78FD1DAF69FC1D914D" authority="Gray, 1856" authorityName="Gray" authorityYear="1856" box="[716,954,1632,1659]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC0381D78FD1DAF69FC88914D" bold="true" box="[716,815,1633,1659]" italics="true" pageId="12" pageNumber="13">Batagur</emphasis>
<bibRefCitation id="EFFF03ECC0381D78FCE6AF68FC1D914D" author="Gray J. E." box="[823,954,1632,1659]" pageId="12" pageNumber="13" refId="ref23879" refString="Gray J. E. 1856 (&quot; 1855 &quot;). Catalogue of Shield Reptiles in the Collection of the British Museum, Part I - Testudinata (Tortoises). Order of the Trustees, London. https: // doi. org / 10.5962 / bhl. title. 5491" type="book" year="1856">Gray, 1856</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0381D78FD08AF8CFCFD91A8" box="[729,858,1668,1694]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BD17E1DC0381D78FD08AF8CFCFD91A8" blockId="12.[634,954,1632,1694]" box="[729,858,1668,1694]" pageId="12" pageNumber="13">
<figureCitation id="13556298C0381D78FD08AF8CFCFD91A8" box="[729,858,1668,1694]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1930,1956]" captionStartId-1="14.[189,232,1964,1990]" captionTargetBox-0="[197,1389,347,1872]" captionTargetBox-1="[195,1391,369,1908]" captionTargetId-0="figure@13.[189,1398,335,1883]" captionTargetId-1="figure@14.[189,1398,365,1916]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 10. BMNH 39834, holotype of Batagur cautleyi Lydekker 1885, here identified as Batagur sp. A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." captionText-1="Fig. 11. IM E.176, previously undescribed, here identified as Batagur sp. A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.3860260" figureDoi-1="http://doi.org/10.5281/zenodo.3860262" httpUri-0="https://zenodo.org/record/3860260/files/figure.png" httpUri-1="https://zenodo.org/record/3860262/files/figure.png" pageId="12" pageNumber="13">Figs 1011</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0381D78FF6CAFCEFDAA9039" pageId="12" pageNumber="13" type="type_taxon">
<paragraph id="8BD17E1DC0381D78FF6CAFCEFEFF91D6" blockId="12.[189,344,1734,1760]" box="[189,344,1734,1760]" pageId="12" pageNumber="13">
<heading id="D099C971C0381D78FF6CAFCEFEFF91D6" bold="true" box="[189,344,1734,1760]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91AA20FC0381D78FF6CAFCEFEFF91D6" bold="true" box="[189,344,1734,1760]" pageId="12" pageNumber="13">
<typeStatus id="54D5C0BFC0381D78FF6CAFCEFF5D91D6" box="[189,250,1734,1760]" pageId="12" pageNumber="13">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0381D78FF6CAFFEFDAA9039" blockId="12.[189,525,1781,1807]" box="[189,525,1781,1807]" pageId="12" pageNumber="13">
<taxonomicName id="4C6E059EC0381D78FF6CAFFEFDAD9039" authority="(Gray, 1830)" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[189,522,1781,1807]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="baska">
<emphasis id="B91AA20FC0381D78FF6CAFFEFECF9039" box="[189,360,1781,1807]" italics="true" pageId="12" pageNumber="13">Batagur baska</emphasis>
(
<bibRefCitation id="EFFF03ECC0381D78FEA9AFFDFDA79039" author="Gray J. E." box="[376,512,1781,1807]" pageId="12" pageNumber="13" refId="ref23595" refString="Gray J. E. 1830. A synopsis of the species of class Reptilia. In: Griffith E. &amp; Pidgeon E. (eds) The Class Reptilia Arranged by the Baron Cuvier, with Specific Descriptions. In: Griffith E. (ed.) The Animal Kingdom Arranged in Conformity with its Organization by the Baron Cuvier, with Additional Descriptions of all the Species Hitherto Named, and of Many not Before noticed. Vol. 9. Reptilia. Whittaker, Treacher and Co, London." type="book" year="1830">Gray, 1830</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0381D78FF6CAE3EFAD590DD" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BD17E1DC0381D78FF6CAE3EFCD09067" blockId="12.[189,887,1846,1873]" box="[189,887,1846,1873]" pageId="12" pageNumber="13">
<heading id="D099C971C0381D78FF6CAE3EFCD09067" bold="true" box="[189,887,1846,1873]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91AA20FC0381D78FF6CAE3EFCD09067" bold="true" box="[189,887,1846,1873]" pageId="12" pageNumber="13">Differential osteological diagnosis using shell characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0381D78FF6CAE6EFAD590DD" blockId="12.[189,1399,1894,2027]" pageId="12" pageNumber="13">
A representative of
<taxonomicName id="4C6E059EC0381D78FE73AE6FFDA590B6" authorityName="Gray" authorityYear="1856" box="[418,514,1895,1920]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC0381D78FE73AE6FFDA590B6" box="[418,514,1895,1920]" italics="true" pageId="12" pageNumber="13">Batagur</emphasis>
</taxonomicName>
can be diagnosed by the presence of a large carapace size (median carapace length of more than
<quantity id="4C96D3F8C0381D78FE67AE81FDA69092" box="[438,513,1929,1956]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="cm" value="40.0">40 cm</quantity>
in adults), a well-developed bridge, well-developed axillary and inguinal buttresses, neural scutes with anterior short sides, a long third neural bone, a short, anteriorly truncated gular scute, an entoplastron that is not intersected by the humeropectoral sulcus and a short anal notch.
</paragraph>
</subSubSection>
<caption id="DF112E95C0391D79FF6CAE82FE4090DD" ID-DOI="http://doi.org/10.5281/zenodo.3860260" ID-Zenodo-Dep="3860260" httpUri="https://zenodo.org/record/3860260/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1930,1956]" targetBox="[197,1389,347,1872]" targetPageId="13">
<paragraph id="8BD17E1DC0391D79FF6CAE82FE4090DD" blockId="13.[189,1398,1930,2027]" pageId="13" pageNumber="14">
<emphasis id="B91AA20FC0391D79FF6CAE82FEBA9092" bold="true" box="[189,285,1930,1956]" pageId="13" pageNumber="14">Fig. 10.</emphasis>
BMNH 39834, holotype of
<taxonomicName id="4C6E059EC0391D79FDA8AE83FBA69093" authority="Lydekker 1885" authorityName="Lydekker" authorityYear="1885" box="[633,1025,1930,1957]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="cautleyi">
<emphasis id="B91AA20FC0391D79FDA8AE83FCE69092" box="[633,833,1930,1956]" italics="true" pageId="13" pageNumber="14">Batagur cautleyi</emphasis>
Lydekker 1885
</taxonomicName>
, here identified as
<taxonomicName id="4C6E059EC0391D79FB3BAE83FAED9092" authorityName="Gray" authorityYear="1856" box="[1258,1354,1931,1956]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC0391D79FB3BAE83FAED9092" box="[1258,1354,1931,1956]" italics="true" pageId="13" pageNumber="14">Batagur</emphasis>
</taxonomicName>
sp.
<emphasis id="B91AA20FC0391D79FF6CAEA5FF7590F1" bold="true" box="[189,210,1965,1991]" pageId="13" pageNumber="14">A</emphasis>
. Photograph of carapace.
<emphasis id="B91AA20FC0391D79FDC3AEA6FD8190FE" bold="true" box="[530,550,1966,1992]" pageId="13" pageNumber="14">B</emphasis>
. Illustration of carapace.
<emphasis id="B91AA20FC0391D79FC8EAEA5FCD390F1" bold="true" box="[863,884,1965,1991]" pageId="13" pageNumber="14">C</emphasis>
. Photograph of plastron.
<emphasis id="B91AA20FC0391D79FB7AAEA6FB6790FE" bold="true" box="[1195,1216,1966,1992]" pageId="13" pageNumber="14">D</emphasis>
. Illustration of plastron. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="DF112E95C03A1D7AFF6CAEA4FA9590DC" ID-DOI="http://doi.org/10.5281/zenodo.3860262" ID-Zenodo-Dep="3860262" httpUri="https://zenodo.org/record/3860262/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1964,1990]" targetBox="[195,1391,369,1908]" targetPageId="14">
<paragraph id="8BD17E1DC03A1D7AFF6CAEA4FA9590DC" blockId="14.[189,1399,1964,2026]" pageId="14" pageNumber="15">
<emphasis id="B91AA20FC03A1D7AFF6CAEA4FEBD90F0" bold="true" box="[189,282,1964,1990]" pageId="14" pageNumber="15">Fig. 11.</emphasis>
IM E.176, previously undescribed, here identified as
<taxonomicName id="4C6E059EC03A1D7AFC4CAEA5FC5A90F0" authorityName="Gray" authorityYear="1856" box="[925,1021,1965,1990]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC03A1D7AFC4CAEA5FC5A90F0" box="[925,1021,1965,1990]" italics="true" pageId="14" pageNumber="15">Batagur</emphasis>
</taxonomicName>
sp.
<emphasis id="B91AA20FC03A1D7AFBE0AEA4FBE190F0" bold="true" box="[1073,1094,1964,1990]" pageId="14" pageNumber="15">A</emphasis>
. Photograph of carapace.
<emphasis id="B91AA20FC03A1D7AFF6CAED8FF7690DC" bold="true" box="[189,209,2000,2026]" pageId="14" pageNumber="15">B</emphasis>
. Illustration of carapace.
<emphasis id="B91AA20FC03A1D7AFE2DAEC7FDB690DF" bold="true" box="[508,529,1999,2025]" pageId="14" pageNumber="15">C</emphasis>
. Photograph of plastron.
<emphasis id="B91AA20FC03A1D7AFCEDAED8FCF690DC" bold="true" box="[828,849,2000,2026]" pageId="14" pageNumber="15">D</emphasis>
. Illustration of plastron. Scale bar: 3 cm.
</paragraph>
</caption>
<subSubSection id="C3742D96C03B1D7BFF6CA803FCA8968C" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BD17E1DC03B1D7BFF6CA803FE0D9613" blockId="15.[189,426,267,293]" box="[189,426,267,293]" pageId="15" pageNumber="16">
<heading id="D099C971C03B1D7BFF6CA803FE0D9613" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B91AA20FC03B1D7BFF6CA803FE0D9613" bold="true" box="[189,426,267,293]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC03B1D7BFF6CA832FECA964E" blockId="15.[189,1398,314,376]" pageId="15" pageNumber="16">
<materialsCitation id="3B067440C03B1D7BFF6CA832FECE964E" ID-GBIF-Occurrence="2626092320" collectionCode="BMNH" country="India" location="Siwalik Hills" pageId="15" pageNumber="16" specimenCode="BMNH 39834" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F3793E8DC03B1D7BFF6CA832FEB79662" box="[189,272,314,340]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
<specimenCount id="9D68B594C03B1D7BFEFBA832FE149662" box="[298,435,314,340]" pageId="15" pageNumber="16" type="generic">1 specimen</specimenCount>
, holotype of
<taxonomicName id="4C6E059EC03B1D7BFD81A833FC7D9663" authority="Lydekker, 1885" authorityName="Lydekker" authorityYear="1885" box="[592,986,314,341]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cautleyi">
<emphasis id="B91AA20FC03B1D7BFD81A833FCB19662" box="[592,790,314,340]" italics="true" pageId="15" pageNumber="16">Batagur cautleyi</emphasis>
Lydekker, 1885
</taxonomicName>
;
<location id="8EB128C6C03B1D7BFC39A832FB209662" LSID="urn:lsid:plazi:treatment:03C7CF0BC0381D7BFDABAF68FE299122:8EB128C6C03B1D7BFC39A832FB209662" box="[1000,1159,314,340]" country="India" name="Siwalik Hills" pageId="15" pageNumber="16">Siwalik Hills</location>
; MiocenePliocene;
<specimenCode id="DBC8D666C03B1D7BFF6CA856FECE964E" box="[189,361,350,376]" collectionCode="BMNH" pageId="15" pageNumber="16">BMNH 39834</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD17E1DC03B1D7BFF6CA8A8FCA8968C" blockId="15.[189,783,416,442]" box="[189,783,416,442]" pageId="15" pageNumber="16">
<materialsCitation id="3B067440C03B1D7BFF6CA8A8FCAC968C" ID-GBIF-Occurrence="2626092321" box="[189,779,416,442]" collectionCode="IM" pageId="15" pageNumber="16" specimenCode="IM E.176" specimenCount="1">
COUNTRY UNKNOWN •
<specimenCount id="9D68B594C03B1D7BFDD2A8A8FD2B968C" box="[515,652,416,442]" pageId="15" pageNumber="16" type="generic">1 specimen</specimenCount>
;
<specimenCode id="DBC8D666C03B1D7BFD46A8A8FCAC968C" box="[663,779,416,442]" pageId="15" pageNumber="16">IM E.176</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C03B1D7BFF6CA8E9FE3C9222" pageId="15" pageNumber="16" type="description">
<paragraph id="8BD17E1DC03B1D7BFF6CA8E9FEE996CD" blockId="15.[189,334,481,507]" box="[189,334,481,507]" pageId="15" pageNumber="16">
<heading id="D099C971C03B1D7BFF6CA8E9FEE996CD" bold="true" box="[189,334,481,507]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B91AA20FC03B1D7BFF6CA8E9FEE996CD" bold="true" box="[189,334,481,507]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC03B1D7BFF6CAB19FAD094A7" blockId="15.[189,1399,529,913]" pageId="15" pageNumber="16">
BMNH 39834 (
<figureCitation id="13556298C03B1D7BFEA3AB19FE6F951D" box="[370,456,529,555]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[197,1389,347,1872]" captionTargetId="figure@13.[189,1398,335,1883]" captionTargetPageId="13" captionText="Fig. 10. BMNH 39834, holotype of Batagur cautleyi Lydekker 1885, here identified as Batagur sp. A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860260" httpUri="https://zenodo.org/record/3860260/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
),
<typeStatus id="54D5C0BFC03B1D7BFE0DAB19FDE4951D" box="[476,579,529,555]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6E059EC03B1D7BFDB7AB1AFC8E951D" authorityName="Lydekker" authorityYear="1885" box="[614,809,529,555]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cautleyi">
<emphasis id="B91AA20FC03B1D7BFDB7AB1AFC8E951D" box="[614,809,529,555]" italics="true" pageId="15" pageNumber="16">Batagur cautleyi</emphasis>
</taxonomicName>
This an almost complete shell from the Miocene/ Pliocene of the Siwalik Hills, likely of
<collectingCountry id="F3793E8DC03B1D7BFD59AB3CFD609578" box="[648,711,564,590]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
, showing the majority of sulci and of the sutures along the center of the carapace. It was originally figured by
<bibRefCitation id="EFFF03ECC03B1D7BFCFCAB50FC5B9545" author="Lydekker R." box="[813,1020,600,627]" pageId="15" pageNumber="16" pagination="155 - 208" refId="ref24414" refString="Lydekker R. 1885 a. Indian Tertiary and post-Tertiary Vertebrata, Part III - Siwalik and Narbada Chelonia. Memoirs of the Geological Survey of India, Palaeontologia Indica 10 (3): 155 - 208." type="journal article" year="1885">Lydekker (1885a</bibRefCitation>
: pl. 24.1), but our observations reveal many additional sutures. There are no signs of growth annuli. The specimen likely is an adult female due to its large size (carapace length greater than
<quantity id="4C96D3F8C03B1D7BFCA1AB97FC18958C" box="[880,959,671,698]" metricMagnitude="-1" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="16" unit="cm" value="54.0">54 cm</quantity>
). A small protuberance on neural IV suggests the previous presence of a median keel. A cervical scute is present. The first vertebral scute is longer than wide and shows a lateral constriction. Neurals IIIV have anteriorly short sides, but neural V has a short left posterior side, which is probably abnormal. The sulcus between the second pleural and the third vertebral is almost straight. The anterior plastral margin is straight and lacks an inflection on the contact of the gularohumeral sulcus. The pectoroabdominal sulcus contacts the sulcus between fifth and sixth marginal scutes. The anal notch is not preserved in this specimen. No plastral sutures can be seen.
</paragraph>
<paragraph id="8BD17E1DC03B1D7BFF6CAAB1FE3C9222" blockId="15.[189,1399,952,1300]" pageId="15" pageNumber="16">
IM E.176 (
<figureCitation id="13556298C03B1D7BFE90AAB0FE3094E4" box="[321,407,952,978]" captionStart="Fig" captionStartId="14.[189,232,1964,1990]" captionTargetBox="[195,1391,369,1908]" captionTargetId="figure@14.[189,1398,365,1916]" captionTargetPageId="14" captionText="Fig. 11. IM E.176, previously undescribed, here identified as Batagur sp. A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860262" httpUri="https://zenodo.org/record/3860262/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
) This is a well-preserved specimen that lacks provenance data and appears to be unpublished. The majority of sutures and sulci are clearly visible. This is an adult specimen due to its large size (carapace length greater than
<quantity id="4C96D3F8C03B1D7BFD1BAAF7FCBD932C" box="[714,794,1023,1050]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="cm" value="35.0">35 cm</quantity>
), but its gender is unknown. A median carapacial keel is present on the posterior parts of the carapace and is strongly marked over neurals IV and VI. The carapace is rather domed at its center, with no signs of growth annuli. Vertebral scutes are subrectangular and have equally sized anterior and posterior margins. Neurals II to VI are anteriorly shortsided. Gular scutes appear to be wider than long and overlap part of the entoplastron. The humeropectoral sulcus is located posterior to the entoplastron. The pectoroabdominal sulcus does not intersect the hyohypoplastron suture and has two lateral notches, suggesting the former presence of parasagital plastral keels at the bridge.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C03B1D7BFF6CAC34FE299122" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BD17E1DC03B1D7BFF6CAC34FEE39260" blockId="15.[189,324,1340,1366]" box="[189,324,1340,1366]" pageId="15" pageNumber="16">
<heading id="D099C971C03B1D7BFF6CAC34FEE39260" bold="true" box="[189,324,1340,1366]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B91AA20FC03B1D7BFF6CAC34FEE39260" bold="true" box="[189,324,1340,1366]" pageId="15" pageNumber="16">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC03B1D7BFF6CAC63FE299122" blockId="15.[189,1399,1387,1556]" pageId="15" pageNumber="16">
We attributed BMNH 39834 and IM E.176, the
<typeStatus id="54D5C0BFC03B1D7BFD3FAC63FC8692B3" box="[750,801,1387,1413]" pageId="15" pageNumber="16">type</typeStatus>
series of
<taxonomicName id="4C6E059EC03B1D7BFC43AC64FBF192B3" authorityName="Lydekker" authorityYear="1885" box="[914,1110,1387,1413]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cautleyi">
<emphasis id="B91AA20FC03B1D7BFC43AC64FBF192B3" box="[914,1110,1387,1413]" italics="true" pageId="15" pageNumber="16">Batagur cautleyi</emphasis>
</taxonomicName>
, to
<taxonomicName id="4C6E059EC03B1D7BFB52AC64FB4492B3" authorityName="Gray" authorityYear="1856" box="[1155,1251,1388,1413]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC03B1D7BFB52AC64FB4492B3" box="[1155,1251,1388,1413]" italics="true" pageId="15" pageNumber="16">Batagur</emphasis>
</taxonomicName>
indet. based on their massive size, highly domed carapace, and presence of an axillary notch and rather straight anterior plastral margin. However, we cannot identify these specimens to species level even though they display unique character combinations. See Discussion (
<taxonomicName id="4C6E059EC03B1D7BFCB2ACDFFB8D92C6" authorityName="Lydekker" authorityYear="1885" box="[867,1066,1494,1520]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cautleyi">
<emphasis id="B91AA20FC03B1D7BFCB2ACDFFB8D92C6" box="[867,1066,1494,1520]" italics="true" pageId="15" pageNumber="16">Batagur cautleyi</emphasis>
</taxonomicName>
from the Siwalik Hills) for additional details.
</paragraph>
</subSubSection>
</treatment>
</document>