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<document id="4B869A1B5D7682934C21A760A34B2BB2" ID-CLB-Dataset="3668" ID-DOI="10.1206/0003-0082(2007)475[1:CSAPOT]2.0.CO;2" ID-GBIF-Dataset="3202c158-e1c4-4cbf-835f-3c1435527f6c" ID-ISSN="0003-0082" ID-Zenodo-Dep="4735338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1620068818117" checkinUser="felipe" docAuthor="ENGEL, MICHAEL S. &amp; GRIMALDI, DAVID A." docDate="2007" docId="03C787F17A38A333FF47FE9CA931CD66" docLanguage="en" docName="AmMusNovit.2007.3568.1-16.pdf" docOrigin="American Museum Novitates 3568 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&amp;doi=10.1206%2F0003-0082(2007)475%5B1%3ACSAPOT%5D2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Boreobythus ENGEL &amp; GRIMALDI 2007, new genus" docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="FFFEFF897A3CA337FFB9FFD3AA55CC40" masterDocTitle="Cretaceous Scolebythidae and Phylogeny of the Family (Hymenoptera: Chrysidoidea)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="5" updateTime="1698938188972" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="19B4EFFAFE2DA07FF4097C94DDE09E46">Cretaceous Scolebythidae and Phylogeny of the Family (Hymenoptera: Chrysidoidea)</mods:title>
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<mods:namePart id="EB45322D3EA1D133C8B1C5CA4E5F6182">ENGEL, MICHAEL S.</mods:namePart>
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<mods:namePart id="4244A84CF031309501FAF4BB8BC536DB">GRIMALDI, DAVID A.</mods:namePart>
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<mods:title id="CB4E9ACACC0A34C6A5C3E0BB54BC3AF8">American Museum Novitates</mods:title>
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<treatment id="03C787F17A38A333FF47FE9CA931CD66" ID-DOI="http://doi.org/10.5281/zenodo.5655261" ID-GBIF-Taxon="190116383" ID-Zenodo-Dep="5655261" LSID="urn:lsid:plazi:treatment:03C787F17A38A333FF47FE9CA931CD66" httpUri="http://treatment.plazi.org/id/03C787F17A38A333FF47FE9CA931CD66" lastPageNumber="5" pageId="4" pageNumber="5">
<subSubSection id="C374656C7A38A333FF47FE9CA85ACD27" box="[254,527,335,359]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BD136E77A38A333FF47FE9CA85ACD27" blockId="4.[254,527,335,359]" box="[254,527,335,359]" pageId="4" pageNumber="5">
<heading id="D099818B7A38A333FF47FE9CA85ACD27" box="[254,527,335,359]" centered="true" fontSize="36" level="2" pageId="4" pageNumber="5" reason="8">
<taxonomicName id="4C6E4D647A38A333FF47FE9CABDCCD27" authority="ENGEL &amp; GRIMALDI, 2007" authorityName="ENGEL &amp; GRIMALDI" authorityYear="2007" box="[254,393,335,359]" class="Insecta" family="Scolebythidae" genus="Boreobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91AEAF57A38A333FF47FE9CABDCCD27" bold="true" box="[254,393,335,359]" italics="true" pageId="4" pageNumber="5">Boreobythus</emphasis>
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,
<taxonomicNameLabel id="A229578E7A38A333FE21FE9CA85ACD27" box="[408,527,335,359]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
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<subSubSection id="C374656C7A38A333FF23FE5BAA80CDFE" pageId="4" pageNumber="5" type="materials_examined">
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<typeStatus id="54D588457A38A333FF23FE5BAA87CDDE" box="[154,210,392,416]" pageId="4" pageNumber="5">TYPE</typeStatus>
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:
<emphasis id="B91AEAF57A38A333FEECFE5BA805CDE0" box="[341,592,392,416]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="4C6E4D647A38A333FEECFE5BA819CDE0" authority="ENGEL &amp; GRIMALDI, 2007" authorityName="ENGEL &amp; GRIMALDI" authorityYear="2007" box="[341,588,392,416]" class="Insecta" family="Scolebythidae" genus="Boreobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="turonius" status="sp. nov.">Boreobythus turonius</taxonomicName>
,
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<taxonomicNameLabel id="A229578E7A38A333FDD9FE5BAA84CDFE" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
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.
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<subSubSection id="C374656C7A38A333FF23FE17AB42C979" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BD136E77A38A333FF23FE17AB42C979" blockId="4.[128,654,392,1754]" pageId="4" pageNumber="5">
DIAGNOSIS: Head elongate (
<figureCitation id="13552A627A38A333FE53FE17A87DCD9C" box="[490,552,452,476]" captionStart="Figs" captionStartId="3.[112,159,1457,1479]" captionTargetBox="[207,1068,210,1422]" captionTargetId="figure-55@3.[207,1068,210,1423]" captionTargetPageId="3" captionText="Figs. 13. Photomicrographs of Cretaceous Scolebythidae. 1. Female holotype of Boreobythus turonius, new genus and species (AMNH). 2. Female holotype of Zapenesia libanica, new genus and species (Azar 146- A). 3. Male holotype of Uliobythus terpsichore, new genus and species (Azar 157-A)." figureDoi="http://doi.org/10.5281/zenodo.4735340" httpUri="https://zenodo.org/record/4735340/files/figure.png" pageId="4" pageNumber="5">fig. 2</figureCitation>
); frontal prominence absent; clypeal apex weakly produced; malar space apparently more than onehalf basal mandibular width; face lateral to antennal torulus flat; compound eyes relatively small, length only slightly more than distance from upper tangent of compound eyes to preoccipital ridge; inner margins of compound eyes very minutely converging below; ocelli almost entirely obscured by fine layer of microscopic bubbles but apparently near upper tangent of compound eyes; occipital carina absent; pronotal collar present, dorsal surface of pronotum nearly 1.5 times as long as mesoscutum; propleura well developed, anteriorly forming a short neck; notauli absent; prosternum large, exposed, broadly triangular posteriorly (
<figureCitation id="13552A627A38A333FE3FFC6EAB91CF95" box="[390,452,957,981]" captionStart="Figs" captionStartId="3.[112,159,1457,1479]" captionTargetBox="[207,1068,210,1422]" captionTargetId="figure-55@3.[207,1068,210,1423]" captionTargetPageId="3" captionText="Figs. 13. Photomicrographs of Cretaceous Scolebythidae. 1. Female holotype of Boreobythus turonius, new genus and species (AMNH). 2. Female holotype of Zapenesia libanica, new genus and species (Azar 146- A). 3. Male holotype of Uliobythus terpsichore, new genus and species (Azar 157-A)." figureDoi="http://doi.org/10.5281/zenodo.4735340" httpUri="https://zenodo.org/record/4735340/files/figure.png" pageId="4" pageNumber="5">fig. 2</figureCitation>
); parapsidal lines absent; forewing with open marginal cell; Rs long, much longer than pterostigma, terminating near anterior wing margin (
<figureCitation id="13552A627A38A333FDFBFBC5A8D5C86E" box="[578,640,1046,1070]" captionStart="Figs" captionStartId="3.[112,159,1457,1479]" captionTargetBox="[207,1068,210,1422]" captionTargetId="figure-55@3.[207,1068,210,1423]" captionTargetPageId="3" captionText="Figs. 13. Photomicrographs of Cretaceous Scolebythidae. 1. Female holotype of Boreobythus turonius, new genus and species (AMNH). 2. Female holotype of Zapenesia libanica, new genus and species (Azar 146- A). 3. Male holotype of Uliobythus terpsichore, new genus and species (Azar 157-A)." figureDoi="http://doi.org/10.5281/zenodo.4735340" httpUri="https://zenodo.org/record/4735340/files/figure.png" pageId="4" pageNumber="5">fig. 2</figureCitation>
), tubular over entire length; R
<subScript id="17EA34A27A38A333FE7EFBEDAB84C80E" attach="left" box="[455,465,1086,1102]" fontSize="7" pageId="4" pageNumber="5">1</subScript>
present on wing margin; pterostigma relatively small, margin within marginal cell convex; 1m-cu present, tubular; Rs + M tubular; Cu distad separation from M nebulous (i.e., abscissae basad and distad 1m-cu nebulous); first submarginal cell short; metafemur not flattened; tibial spur formula indeterminate from fossil; pretarsal claws simple.
</paragraph>
</subSubSection>
<subSubSection id="C374656C7A38A333FF23FAECA870CA49" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BD136E77A38A333FF23FAECA870CA49" blockId="4.[128,654,392,1754]" pageId="4" pageNumber="5">
ETYMOLOGY: The genus-group name is a combination of
<emphasis id="B91AEAF57A38A333FED2FA8EABE6C935" box="[363,435,1373,1397]" italics="true" pageId="4" pageNumber="5">boreas</emphasis>
(Greek, meaning north wind) and
<emphasis id="B91AEAF57A38A333FECCFAA8ABE8C9D3" box="[373,445,1403,1427]" italics="true" pageId="4" pageNumber="5">bythus</emphasis>
(root of the
<typeStatus id="54D588457A38A333FDE4FAA8A8D8C9D3" box="[605,653,1403,1427]" pageId="4" pageNumber="5">type</typeStatus>
genus for the family,
<taxonomicName id="4C6E4D647A38A333FEC4FA4AA855C9F1" authorityName="Evans" authorityYear="1963" box="[381,512,1433,1457]" class="Insecta" family="Scolebythidae" genus="Scolebythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF57A38A333FEC4FA4AA855C9F1" box="[381,512,1433,1457]" italics="true" pageId="4" pageNumber="5">Scolebythus</emphasis>
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). The name refers to the fact that this fossil comes from the furthest north paleolatitude among Cretaceous scolebythids. The name is masculine.
</paragraph>
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<subSubSection id="C374656C7A38A333FF23F9C3A931CD66" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BD136E77A38A333FF23F9C3A931CD66" blockId="4.[128,654,392,1754]" lastBlockId="4.[701,1228,211,294]" pageId="4" pageNumber="5">
COMMENTS:
<taxonomicName id="4C6E4D647A38A333FE8CF9C3AB95CA68" authorityName="ENGEL &amp; GRIMALDI" authorityYear="2007" box="[309,448,1552,1576]" class="Insecta" family="Scolebythidae" genus="Boreobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF57A38A333FE8CF9C3AB95CA68" box="[309,448,1552,1576]" italics="true" pageId="4" pageNumber="5">Boreobythus</emphasis>
</taxonomicName>
has, perhaps, the most plesiomorphic habitus of any scolebythid, overall resembling a primitive bethylid, particularly in the shape of the head. However, despite the superficial shape of the head, the greatly enlarged and exposed prosternum, the laterally articulated protrochanters, and the enlarged propleura that form a short neck attest to its placement in
<taxonomicName id="4C6E4D647A38A333FD04FEDDA90ACD66" authorityName="Evans" authorityYear="1963" box="[701,863,270,294]" class="Insecta" family="Scolebythidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Scolebythidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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