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<document id="9851563EF17037F97CACAFF8A998C55D" ID-CLB-Dataset="20299" ID-DOI="10.5281/zenodo.5314350" ID-GBIF-Dataset="8839e56a-8836-4002-8a03-6fd4ee00cb99" ID-ISSN="0374-1036" ID-Zenodo-Dep="5314350" ID-ZooBank="44343D04-2985-45F4-BA26-4F5C3B481BDAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630121442887" checkinUser="marcus" docAuthor="Malenovský, Igor &amp; Burckhardt, Daniel" docDate="2014" docId="03C787CEFFB98E31FE12CC1B958D9602" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.23-61.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Diaphorina caliginosa Malenovský &amp; Burckhardt 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFFEFFB6FFB18E3FFFC7C81A945F9531" masterDocTitle="Jumping plant-lice of Socotra Island (Hemiptera: Psylloidea)" masterLastPageNumber="61" masterPageNumber="23" pageNumber="31" updateTime="1699165484273" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0BE37369E606688037D5ACA88863AD87" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5FEDA7610740C4E8C352C228BE14507F">
<mods:title id="640A42254B66BC2D62C06910CE214BB9">Jumping plant-lice of Socotra Island (Hemiptera: Psylloidea)</mods:title>
</mods:titleInfo>
<mods:name id="CBA3388C5970C3ED1BA194E65101BCA9" type="personal">
<mods:role id="B5FA674B87A3D38FCFCEE6D517D83FD0">
<mods:roleTerm id="3F59E4F79FD3FE03F04874EC23FAE106">Author</mods:roleTerm>
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<mods:namePart id="22DA635AA3A520B0CA3D1319DE9A7B40">Malenovský, Igor</mods:namePart>
<mods:affiliation id="644D9AF9D5F708D4235C95EBDC557B47">Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlářská 2, CZ- 611 37 Brno, Czech Republic &amp; Department of Entomology, Moravian Museum, Hviezdoslavova 29 a, CZ- 627 00 Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="AEF1680925A096D8A3C9B777837342FA" type="email">malenovsky@sci.muni.cz &amp; imalenovsky@mzm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="32F43445952415553D5F3EE81F1ABCF7" type="personal">
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</mods:role>
<mods:namePart id="B08359D064358A5F41F67E5B8A4A589C">Burckhardt, Daniel</mods:namePart>
<mods:affiliation id="682182186E222FB1348E5FC25F392A1A">Naturhistorisches Museum, Augustinergasse 2, CH- 4001 Basel, Switzerland</mods:affiliation>
<mods:nameIdentifier id="805D3013A20C32AA8C4470330BF94EE1" type="email">daniel.burckhardt@bs.ch</mods:nameIdentifier>
</mods:name>
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<mods:relatedItem id="A231B0E8F26F660F7B63D6B19A24C20A" type="host">
<mods:titleInfo id="14912DA29EBBCCE50F1BC48D0D1D5215">
<mods:title id="8014948978B262E4AC5383468A76F71E">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="F8B68D95FF287790D94B2FBBBBEDC89B">
<mods:date id="493EFB307BBB9575D99B05990FA16E3A">2014</mods:date>
<mods:detail id="6B7D8E2E60A791BD8473F69B25FB6A8D" type="pubDate">
<mods:number id="FECF562E5D4B331DDC9D62AA0073595F">2014-12-30</mods:number>
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<mods:classification id="642140481F4FD56AD98C3BCBD446B120">journal article</mods:classification>
<mods:identifier id="1BA68D0B2BFF39912023C4F4430DFF60" type="CLB-Dataset">20299</mods:identifier>
<mods:identifier id="440A1F08A2AACB04B6FFA95EE7AE0E1A" type="DOI">10.5281/zenodo.5314350</mods:identifier>
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<treatment id="03C787CEFFB98E31FE12CC1B958D9602" ID-DOI="http://doi.org/10.5281/zenodo.6344467" ID-GBIF-Taxon="193896470" ID-Zenodo-Dep="6344467" LSID="urn:lsid:plazi:treatment:03C787CEFFB98E31FE12CC1B958D9602" httpUri="http://treatment.plazi.org/id/03C787CEFFB98E31FE12CC1B958D9602" lastPageId="14" lastPageNumber="37" pageId="8" pageNumber="31">
<subSubSection id="C3746553FFB98E37FE12CC1B97779128" box="[469,808,1025,1049]" pageId="8" pageNumber="31" type="nomenclature">
<paragraph id="8BD136D8FFB98E37FE12CC1B97779128" blockId="8.[469,808,1025,1049]" box="[469,808,1025,1049]" pageId="8" pageNumber="31">
<heading id="D09981B4FFB98E37FE12CC1B97779128" bold="true" box="[469,808,1025,1049]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="31" reason="3">
<emphasis id="B91AEACAFFB98E37FE12CC1B97779128" bold="true" box="[469,808,1025,1049]" pageId="8" pageNumber="31">
<taxonomicName id="4C6E4D5BFFB98E37FE12CC1B96929128" ID-CoL="6CRM5" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[469,717,1025,1049]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFB98E37FE12CC1B96929128" bold="true" box="[469,717,1025,1049]" italics="true" pageId="8" pageNumber="31">Diaphorina caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFB98E37FD14CC1897779128" box="[723,808,1026,1049]" pageId="8" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFB98E37FDEFCC3A96899102" box="[552,726,1056,1075]" pageId="8" pageNumber="31" type="description">
<paragraph id="8BD136D8FFB98E37FDEFCC3A96899102" blockId="8.[552,726,1056,1075]" box="[552,726,1056,1075]" pageId="8" pageNumber="31">
(
<figureCitation id="13552A5DFFB98E37FDE9CC3A96CE9102" box="[558,657,1056,1075]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="8" pageNumber="31">Figs 1822</figureCitation>
,
<figureCitation id="13552A5DFFB98E37FD5ECC3A96909102" box="[665,719,1056,1075]" captionStart-0="Figs 2733" captionStart-1="Figs 3438" captionStartId-0="12.[151,188,1259,1278]" captionStartId-1="13.[113,150,1237,1256]" captionTargetBox-0="[184,1090,217,1226]" captionTargetBox-1="[120,1043,220,1210]" captionTargetId-0="figure-11@12.[167,1110,214,1247]" captionTargetId-1="figure-12@13.[113,1088,182,1217]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." captionText-1="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi-0="http://doi.org/10.5281/zenodo.5314364" figureDoi-1="http://doi.org/10.5281/zenodo.5314366" httpUri-0="https://zenodo.org/record/5314364/files/figure.png" httpUri-1="https://zenodo.org/record/5314366/files/figure.png" pageId="8" pageNumber="31">2738</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFB98E37FF50CC50961C91D3" pageId="8" pageNumber="31" type="materials_examined">
<paragraph id="8BD136D8FFB98E37FF50CC5094B79149" blockId="8.[151,1127,1098,1250]" pageId="8" pageNumber="31">
<emphasis id="B91AEACAFFB98E37FF50CC509552916C" bold="true" box="[151,269,1098,1117]" pageId="8" pageNumber="31">
<typeStatus id="54D5887AFFB98E37FF50CC50949C916C" box="[151,195,1098,1117]" pageId="8" pageNumber="31">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3797648FFB98E37FED7CC509512916C" box="[272,333,1098,1117]" name="Yemen" pageId="8" pageNumber="31">Yemen</collectingCountry>
, central
<collectingCountry id="F3797648FFB98E37FE56CC50958C916C" box="[401,467,1098,1117]" name="Yemen" pageId="8" pageNumber="31">Socotra</collectingCountry>
, Hagher mountains, Mt.Skand environs,
<geoCoordinate id="EE5A501FFFB98E37FCE0CC5097D4916C" box="[807,907,1098,1117]" degrees="12" direction="north" minutes="34" orientation="latitude" pageId="8" pageNumber="31" precision="15" seconds="36" value="12.576667">12°3436″N</geoCoordinate>
<geoCoordinate id="EE5A501FFFB98E37FC48CC5097AC916C" box="[911,1011,1098,1117]" degrees="54" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="31" precision="15" seconds="30" value="54.025">54°0130″E</geoCoordinate>
,
<quantity id="4C969B3DFFB98E37FC3ECC509067916C" box="[1017,1080,1098,1117]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="31" unit="m" value="1450.0">1450 m</quantity>
a.s.l. (
<figureCitation id="13552A5DFFB98E37FF59CC7F94829149" box="[158,221,1125,1144]" captionStart="Figs 2326" captionStartId="11.[113,150,1165,1184]" captionTargetBox="[113,1088,219,1139]" captionTargetId="figure-103@11.[113,1091,219,1139]" captionTargetPageId="11" captionText="Figs 2326. Habitats of Diaphorina spp. in Socotra. 23 montane evergreen woodland in Skand area, Hagher Mts., with a Carissa spinarum tree in foreground (centre): type locality of D. caliginosa sp. nov. 24 detail of a Carissa spinarum twig, host plant of D. caliginosa sp. nov. 25 montane shrubland with open grassland patches in wadi Madar, Hagher Mts.: type locality of D. hagherensis sp. nov. 26 open dry woodland on limestone plateau near Firmihin with a Lycium sokotranum shrub in foreground:habitat and host plant of D. elegans Burckhardt &amp; Mifsud, 1998. Photographs by I. Malenovský (23, 24) and P. Kment (25, 26)." figureDoi="http://doi.org/10.5281/zenodo.5314360" httpUri="https://zenodo.org/record/5314360/files/figure.png" pageId="8" pageNumber="31">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BD136D8FFB98E37FF50CC65961C91D3" blockId="8.[151,1127,1098,1250]" pageId="8" pageNumber="31">
<emphasis id="B91AEACAFFB98E37FF50CC65954491A3" bold="true" box="[151,283,1151,1170]" pageId="8" pageNumber="31">Type material.</emphasis>
<materialsCitation id="3B063C85FFB98E37FED8CC65949B91F9" ID-GBIF-Occurrence="3699124301" collectedFrom="on Carissa spinarum" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectionCode="MMBC" collectorName="I. Malenovsky" country="Yemen" elevation="1450" latitude="12.576667" location="Hagher Mts." longLatPrecision="129" longitude="54.025" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="SOCOTRA Island" typeStatus="holotype">
<typeStatus id="54D5887AFFB98E37FED8CC65952C91A1" box="[287,371,1151,1170]" pageId="8" pageNumber="31" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D68FD51FFB98E37FEBACC6095D291A2" box="[381,397,1146,1171]" pageId="8" pageNumber="31" type="male"></specimenCount>
(
<collectionCode id="ED7FAE1DFFB98E37FE50CC65958291A3" box="[407,477,1151,1170]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="8" pageNumber="31">MMBC</collectionCode>
, dry-mounted),
<collectingCountry id="F3797648FFB98E37FDA0CC6596E691A3" box="[615,697,1151,1170]" name="Yemen" pageId="8" pageNumber="31">YEMEN</collectingCountry>
,
<collectingRegion id="49AAF83AFFB98E37FD07CC65970491A3" box="[704,859,1151,1170]" pageId="8" pageNumber="31">SOCOTRA Island</collectingRegion>
/
<location id="8EB16003FFB98E37FCADCC65978891A3" LSID="urn:lsid:plazi:treatment:03C787CEFFB98E31FE12CC1B958D9602:8EB16003FFB98E37FCADCC65978891A3" box="[874,983,1151,1170]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" name="Hagher Mts." pageId="8" pageNumber="31" stateProvince="SOCOTRA Island">Hagher Mts.</location>
,
<location id="8EB16003FFB98E37FC19CC65900291A3" LSID="urn:lsid:plazi:treatment:03C787CEFFB98E31FE12CC1B958D9602:8EB16003FFB98E37FC19CC65900291A3" box="[990,1117,1151,1170]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" name="Scand Mt. env." pageId="8" pageNumber="31" stateProvince="SOCOTRA Island">Scand Mt. env.</location>
/
<geoCoordinate id="EE5A501FFFB98E37FF50CC8094B0919C" box="[151,239,1178,1197]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="8" pageNumber="31" precision="92" value="12.576667">12°34.6N</geoCoordinate>
,
<geoCoordinate id="EE5A501FFFB98E37FF30CC809513919C" box="[247,332,1178,1197]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="8" pageNumber="31" precision="92" value="54.025">54°01.5E</geoCoordinate>
,
<quantity id="4C969B3DFFB98E37FE94CC8095CC919C" box="[339,403,1178,1197]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="31" unit="m" value="1450.0">
<elevation id="0043D1EBFFB98E37FE94CC8095CC919C" box="[339,403,1178,1197]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="31" unit="m" value="1450.0">1450 m</elevation>
</quantity>
/
<location id="8EB16003FFB98E37FE67CC8096C9919C" LSID="urn:lsid:plazi:treatment:03C787CEFFB98E31FE12CC1B958D9602:8EB16003FFB98E37FE67CC8096C9919C" box="[416,662,1178,1197]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" name="montane evergreen woodland" pageId="8" pageNumber="31" stateProvince="SOCOTRA Island">montane evergreen woodland</location>
/
<date id="FFD01018FFB98E37FD64CC809740919C" box="[675,799,1178,1197]" pageId="8" pageNumber="31" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">
<collectingDate id="EF94E9F0FFB98E37FD64CC809740919C" box="[675,799,1178,1197]" pageId="8" pageNumber="31" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">16.-18.vi.2012</collectingDate>
</date>
//
<collectedFrom id="38727525FFB98E37FCF5CC8097BE919C" box="[818,993,1178,1197]" pageId="8" pageNumber="31">
on
<taxonomicName id="4C6E4D5BFFB98E37FC8CCC8097BE919C" box="[843,993,1178,1197]" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="species" species="spinarum">
<emphasis id="B91AEACAFFB98E37FC8CCC8097BE919C" box="[843,993,1178,1197]" italics="true" pageId="8" pageNumber="31">Carissa spinarum</emphasis>
</taxonomicName>
</collectedFrom>
/
<collectorName id="269B530EFFB98E37FC29CC809038919C" box="[1006,1127,1178,1197]" pageId="8" pageNumber="31">I. Malenovský</collectorName>
leg..
</materialsCitation>
<materialsCitation id="3B063C85FFB98E37FF0FCCAF961C91D3" ID-GBIF-Occurrence="3699124318" collectedFrom="on Carissa spinarum" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectionCode="MMBC, NHMB, NMPC" collectorName="I. Malenovsky" country="Yemen" elevation="1450" latitude="12.576667" location="Hagher Mts." longLatPrecision="129" longitude="54.025" pageId="8" pageNumber="31" specimenCount="21" specimenCount-female="9" specimenCount-male="12" stateProvince="SOCOTRA Island" typeStatus="paratype">
<typeStatus id="54D5887AFFB98E37FF0FCCAF957F91F7" box="[200,288,1205,1224]" pageId="8" pageNumber="31" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D68FD51FFB98E37FEEDCCAF953C91F9" box="[298,355,1199,1224]" pageId="8" pageNumber="31" type="male">12 ♂♂</specimenCount>
<specimenCount id="9D68FD51FFB98E37FEAFCCAF95C991F6" box="[360,406,1198,1224]" pageId="8" pageNumber="31" type="female">9 ♀♀</specimenCount>
, 1 skin of fifth instar immature, same data as the
<typeStatus id="54D5887AFFB98E37FCF0CCAF97DD91F9" box="[823,898,1205,1224]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7FAE1DFFB98E37FC4ACCAF978C91F9" box="[909,979,1205,1224]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="8" pageNumber="31">MMBC</collectionCode>
,
<collectionCode id="ED7FAE1DFFB98E37FC1DCCAF904391F9" box="[986,1052,1205,1224]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="31">NHMB</collectionCode>
,
<collectionCode id="ED7FAE1DFFB98E37FBE2CCAF903B91F9" box="[1061,1124,1205,1224]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="31">NMPC</collectionCode>
, dry- and slide-mounted and preserved in alcohol).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFB98E34FF50CCE1907290E6" lastPageId="11" lastPageNumber="34" pageId="8" pageNumber="31" type="description">
<paragraph id="8BD136D8FFB98E36FF50CCE197469462" blockId="8.[151,1127,1274,1586]" lastBlockId="9.[113,1090,220,1619]" lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31">
<emphasis id="B91AEACAFFB98E37FF50CCE195369023" bold="true" box="[151,361,1274,1298]" pageId="8" pageNumber="31">
Description.
<emphasis id="B91AEACAFFB98E37FEE1CCE095369023" box="[294,361,1274,1298]" italics="true" pageId="8" pageNumber="31">Adult.</emphasis>
</emphasis>
Coloration (
<figureCitation id="13552A5DFFB98E37FE29CCE1963A9023" box="[494,613,1275,1298]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="8" pageNumber="31">Figs 18, 19</figureCitation>
). Body partly covered with grey waxy secretions. Head including genal processes and thorax dark brown, pronotum medially and mesoscutellum slightly lighter brown, mesoscutum dorsally with blackish brown longitudinal stripes, lateral sclerites of thorax partly red. Eyes dark brown, ocelli red. Antenna dirty yellow, segments 1 and 2 brown, segment 8 fumous, segments 9 and 10 entirely dark brown to black. Legs with tarsi dirty yellow, apical tarsal segments slightly infuscate, tibiae largely dirty yellow, dark brown basally, femora and coxae dark brown. Forewing membrane opaque, off-white, covered from base to apex with many sharply contrasting dark brown irregular patches, fusing into a narrow continuous dark brown band at outer-posterior wing margin, extending from apical tip of r 1 to apex of cu 2 cells and leaving small light semioval patches medially in cells r 2, m 1, m 2 and cu 1 at wing margin (
<figureCitation id="13552A5DFFB88E36FE54C8C695BD95C2" box="[403,482,220,243]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="9" pageNumber="32">Fig. 20</figureCitation>
); veins light brown, short basal portions of M+Cu, M, R 1, R S and Cu 1a dark brown, wing margin with several short dark brown sections at apices of R 1 and Cu 1b and in apical wing quarter. Hind wing hyaline, C+Sc vein entirely and apical veins basally dark brown. Abdomen including terminalia dark brown.
</paragraph>
<paragraph id="8BD136D8FFB88E36FF57C94695629082" blockId="9.[113,1090,220,1619]" pageId="9" pageNumber="32">
<emphasis id="B91AEACAFFB88E36FF57C94694AD9442" box="[144,242,348,371]" italics="true" pageId="9" pageNumber="32">Structure</emphasis>
. Integument with coarse microsculpture, matt, covered with short greyish setae. Head slightly inclined from longitudinal body axis. Vertex flat with one distinct fovea on each side of median suture; lateral ocelli lying in plane of vertex, frontal ocellus clearly visible in dorsal view. Genal processes approximately as long as vertex along midline, with straight inner margins contiguous basally and strongly diverging in apical two thirds, lateral margins convex, and apex subacute (
<figureCitation id="13552A5DFFB88E36FE62C9E695A89722" box="[421,503,508,531]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="9" pageNumber="32">Fig. 22</figureCitation>
). Eyes subglobular. Antenna (
<figureCitation id="13552A5DFFB88E36FCFAC9E697D29722" box="[829,909,508,531]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 32</figureCitation>
) relatively long, about 0.9 as long as head width, with 10 segments; segments 37 nearly cylindrical, segment 8 strongly widening to apex; segment 3 longest, segments 46 each subequal in length, segments 7 and 8 each shorter than more proximal flagellar segments but each slightly longer than segment 9; single circular rhinarium subapically on segments 4, 6, 8 and 9; rhinaria on segments 4 and 6 each associated with simple seta; segment 10 with terminal setae greatly differing in length: one short (ca.
<quantity id="4C969B3DFFB88E36FE14CAA6966B97E2" box="[467,564,700,723]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="32" unit="mm" value="0.02">0.02 mm</quantity>
), stout, and truncate, the other long (ca.
<quantity id="4C969B3DFFB88E36FC1DCAA6906697E2" box="[986,1081,700,723]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="32" unit="mm" value="0.06">0.06 mm</quantity>
) and slender (
<figureCitation id="13552A5DFFB88E36FF31CAC6951B97C2" box="[246,324,732,755]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 33</figureCitation>
). Clypeus nearly flat, basally with few short inconspicuous setae; rostrum short, apical segment with two long setae. Metacoxa with relatively long, pointed meracanthus. Metatibia elongate, rugged basally but lacking genual spine, slightly widening to apex, with a crown of 7 or 8 dark sclerotised spurs apically. Metabasitarsus laterally bearing two dark sclerotised apical spurs. Forewing (
<figureCitation id="13552A5DFFB88E36FE35CB46961D9642" box="[498,578,860,883]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="9" pageNumber="32">Fig. 20</figureCitation>
) elongate, relatively narrow, broadest in apical quarter, broadly and almost symmetrically rounded apically, with apex lying in cell r 2 close to R S vein apex; R S vein sinuate in apical half; membrane densely covered in large, star-shaped surface spinules which are irregularly arranged ca. 7 (611) μm apart in middle of cells (
<figureCitation id="13552A5DFFB88E36FBD2CBA694D396C2" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="9" pageNumber="32">Fig. 21</figureCitation>
) and becoming smaller, simply radular and more densely arranged near veins and along apical wing margin; fore wing veins associated with many short setae inserted into membrane close to veins (
<figureCitation id="13552A5DFFB88E36FED4CC06953C9102" box="[275,355,1052,1075]" captionStart="Figs 1822" captionStartId="10.[151,187,1367,1386]" captionTargetBox="[151,1126,219,1349]" captionTargetId="figure-11@10.[151,1129,219,1352]" captionTargetPageId="10" captionText="Figs 1822.Diaphorina caliginosa sp. nov., adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m1 at M-fork; 22 head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="9" pageNumber="32">Fig. 21</figureCitation>
). Hind wing costal margin with 2 + 8 ungrouped setae basally and 1 seta apically. Male subgenital plate with slightly sinuate dorsal margin and few sparse short setae postero-ventrally (
<figureCitation id="13552A5DFFB88E36FEA8CC4695E29142" box="[367,445,1116,1139]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 27</figureCitation>
). Male proctiger relatively broad, with large, apically broadly rounded posterior lobes (
<figureCitation id="13552A5DFFB88E36FEB8CC66958F91A2" box="[383,464,1148,1171]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 27</figureCitation>
). Paramere, in lateral view, nearly parallel-sided, straight; apical part slightly asymmetrical with anterior margin more strongly narrowing to apex than posterior margin; apex narrowly rounded, bearing a small tooth on inner side; inner face covered with numerous long unsclerotised setae (
<figureCitation id="13552A5DFFB88E36FD46CCC6968D91C2" box="[641,722,1244,1267]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 28</figureCitation>
). Distal segment of aedeagus with a simple, obovate apical dilatation, ductus ejaculatorius short and sinuate (
<figureCitation id="13552A5DFFB88E36FC48CCE697809022" box="[911,991,1276,1299]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 29</figureCitation>
). Female terminalia with proctiger, in lateral view, with dorsal margin slightly concave, apex narrowly rounded; circumanal pore ring elliptic with two contiguous rows of pores; subgenital plate, in lateral view, with distinct ventral hump medially, in apical half straight and narrowing to pointed apex (
<figureCitation id="13552A5DFFB88E36FEC0CD66950990A2" box="[263,342,1404,1427]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 30</figureCitation>
); dorsal and ventral valvulae slightly curved ventrally, smooth, lacking any teeth (
<figureCitation id="13552A5DFFB88E36FF26CD86956F9082" box="[225,304,1436,1459]" captionStart="Figs 2733" captionStartId="12.[151,188,1259,1278]" captionTargetBox="[184,1090,217,1226]" captionTargetId="figure-11@12.[167,1110,214,1247]" captionTargetPageId="12" captionText="Figs 2733. Diaphorina caliginosa sp. nov., adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10." figureDoi="http://doi.org/10.5281/zenodo.5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="9" pageNumber="32">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BD136D8FFB88E36FF57CDA697219362" blockId="9.[113,1090,220,1619]" pageId="9" pageNumber="32">
<emphasis id="B91AEACAFFB88E36FF57CDA6957490E2" box="[144,299,1468,1491]" italics="true" pageId="9" pageNumber="32">Measurements</emphasis>
(in mm). Males (n = 2): HW 0.59, AL 0.520.57, WL 2.212.25, WW 0.850.89, TL 0.57, MPL 0.310.33, PL 0.240.25, AEL 0.170.18. Females (n = 2): HW 0.600.62, AL 0.510.54, WL 2.372.40, WW 0.940.95, TL 0.58, FPL 0.620.63, SL 0.470.48. Ratios: AL/HW 0.850.97, WL/HW 3.754.00, WL/WW 2.472.60, TL/HW 0.940.97, MPL/HW 0.530.56, FPL/HW 1.021.03, FPL/SL 1.291.34.
</paragraph>
<caption id="DF116650FFBB8E35FF50CD4D976690B4" ID-DOI="http://doi.org/10.5281/zenodo.5314358" ID-Zenodo-Dep="5314358" httpUri="https://zenodo.org/record/5314358/files/figure.png" pageId="10" pageNumber="33" startId="10.[151,187,1367,1386]" targetBox="[151,1126,219,1349]" targetPageId="10">
<paragraph id="8BD136D8FFBB8E35FF50CD4D976690B4" blockId="10.[151,1127,1367,1418]" pageId="10" pageNumber="33">
Figs 1822.
<taxonomicName id="4C6E4D5BFFBB8E35FF3ACD4D95E3905B" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[253,444,1367,1386]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBB8E35FF3ACD4D95E3905B" box="[253,444,1367,1386]" italics="true" pageId="10" pageNumber="33">Diaphorina caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBB8E35FE07CD4D965C905B" box="[448,515,1367,1386]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
, adult. 18 habitus, lateral view; 19 habitus, dorsal view; 20 forewing; 21 detail of forewing membrane in cell m
<subScript id="17EA349DFFBB8E35FDCDCD65964F90BB" attach="left" box="[522,528,1407,1418]" fontSize="5" pageId="10" pageNumber="33">1</subScript>
at M-fork; 22 head, dorsal view.
</paragraph>
</caption>
<paragraph id="8BD136D8FFBB8E34FF72CDA0953690A6" blockId="10.[151,1127,1466,1618]" lastBlockId="11.[113,1089,1376,1591]" lastPageId="11" lastPageNumber="34" pageId="10" pageNumber="33">
<emphasis id="B91AEACAFFBB8E35FF72CDA095F590E2" bold="true" box="[181,426,1466,1491]" italics="true" pageId="10" pageNumber="33">Fifth instar immature</emphasis>
(
<figureCitation id="13552A5DFFBB8E35FE7FCDA1965590E3" box="[440,522,1467,1490]" captionStart="Figs 3438" captionStartId="13.[113,150,1237,1256]" captionTargetBox="[120,1043,220,1210]" captionTargetId="figure-12@13.[113,1088,182,1217]" captionTargetPageId="13" captionText="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi="http://doi.org/10.5281/zenodo.5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="10" pageNumber="33">Fig. 34</figureCitation>
). Body brownish, broad, flat. Margins of head and wing pads lacking any visible setae, caudal plate margin with 120 densely arranged, relatively long, pointed lanceolate setae (
<figureCitation id="13552A5DFFBB8E35FE2ACDE196609323" box="[493,575,1531,1554]" captionStart="Figs 3438" captionStartId="13.[113,150,1237,1256]" captionTargetBox="[120,1043,220,1210]" captionTargetId="figure-12@13.[113,1088,182,1217]" captionTargetPageId="13" captionText="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi="http://doi.org/10.5281/zenodo.5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="10" pageNumber="33">Fig. 35</figureCitation>
). Antenna with 3 segments; apical segment posteriorly with three pointed lanceolate setae and four rhinaria (
<figureCitation id="13552A5DFFBB8E35FCE9CE0197DD9303" box="[814,898,1563,1586]" captionStart="Figs 3438" captionStartId="13.[113,150,1237,1256]" captionTargetBox="[120,1043,220,1210]" captionTargetId="figure-12@13.[113,1088,182,1217]" captionTargetPageId="13" captionText="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi="http://doi.org/10.5281/zenodo.5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="10" pageNumber="33">Fig. 36</figureCitation>
). Forewing pad with broadly rounded humeral lobe. Tarsal arolium broadly triangular (
<figureCitation id="13552A5DFFBB8E35FCAACE21979F9363" box="[877,960,1595,1618]" captionStart="Figs 3438" captionStartId="13.[113,150,1237,1256]" captionTargetBox="[120,1043,220,1210]" captionTargetId="figure-12@13.[113,1088,182,1217]" captionTargetPageId="13" captionText="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi="http://doi.org/10.5281/zenodo.5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="10" pageNumber="33">Fig. 37</figureCitation>
). Anus ventral, outer circumanal ring elongate with lateral sides slightly bent forwards, consisting of single row of pores (
<figureCitation id="13552A5DFFBA8E34FECDCD9A950290A6" box="[266,349,1408,1431]" captionStart="Figs 3438" captionStartId="13.[113,150,1237,1256]" captionTargetBox="[120,1043,220,1210]" captionTargetId="figure-12@13.[113,1088,182,1217]" captionTargetPageId="13" captionText="Figs 3438. Diaphorina caliginosa sp. nov., fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only)." figureDoi="http://doi.org/10.5281/zenodo.5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="11" pageNumber="34">Fig. 38</figureCitation>
).
</paragraph>
<caption id="DF116650FFBA8E34FFB6CC9796E39014" ID-DOI="http://doi.org/10.5281/zenodo.5314360" ID-Zenodo-Dep="5314360" httpUri="https://zenodo.org/record/5314360/files/figure.png" pageId="11" pageNumber="34" startId="11.[113,150,1165,1184]" targetBox="[113,1088,219,1139]" targetPageId="11">
<paragraph id="8BD136D8FFBA8E34FFB6CC9796E39014" blockId="11.[113,1089,1165,1317]" pageId="11" pageNumber="34">
Figs 2326. Habitats of
<taxonomicName id="4C6E4D5BFFBA8E34FEFBCC9795969191" box="[316,457,1165,1184]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91AEACAFFBA8E34FEFBCC9795FF9191" box="[316,416,1165,1184]" italics="true" pageId="11" pageNumber="34">Diaphorina</emphasis>
spp.
</taxonomicName>
in Socotra. 23 montane evergreen woodland in Skand area, Hagher Mts., with a
<taxonomicName id="4C6E4D5BFFBA8E34FF6CCCBD951B918B" box="[171,324,1191,1210]" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="34" phylum="Tracheophyta" rank="species" species="spinarum">
<emphasis id="B91AEACAFFBA8E34FF6CCCBD951B918B" box="[171,324,1191,1210]" italics="true" pageId="11" pageNumber="34">Carissa spinarum</emphasis>
</taxonomicName>
tree in foreground (centre): type locality of
<taxonomicName id="4C6E4D5BFFBA8E34FD7CCCBD9771918B" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[699,814,1191,1210]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBA8E34FD7CCCBD9771918B" box="[699,814,1191,1210]" italics="true" pageId="11" pageNumber="34">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBA8E34FCF4CCBD9729918B" box="[819,886,1191,1210]" pageId="11" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
24 detail of a
<taxonomicName id="4C6E4D5BFFBA8E34FC39CCBD949D91E4" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="34" phylum="Tracheophyta" rank="species" species="spinarum">
<emphasis id="B91AEACAFFBA8E34FC39CCBD949D91E4" italics="true" pageId="11" pageNumber="34">Carissa spinarum</emphasis>
</taxonomicName>
twig, host plant of
<taxonomicName id="4C6E4D5BFFBA8E34FEAACCD895BE91E4" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[365,481,1218,1237]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBA8E34FEAACCD895BE91E4" box="[365,481,1218,1237]" italics="true" pageId="11" pageNumber="34">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBA8E34FE20CCD8967491E4" box="[487,555,1218,1237]" pageId="11" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
25 montane shrubland with open grassland patches in wadi Madar, Hagher Mts.: type locality of
<taxonomicName id="4C6E4D5BFFBA8E34FE71CCC7966791C1" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[438,568,1245,1264]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="hagherensis" status="sp. nov.">
<emphasis id="B91AEACAFFBA8E34FE71CCC7966791C1" box="[438,568,1245,1264]" italics="true" pageId="11" pageNumber="34">D. hagherensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBA8E34FDF9CCC796DC91C1" box="[574,643,1245,1264]" pageId="11" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
26 open dry woodland on limestone plateau near Firmihin with a
<taxonomicName id="4C6E4D5BFFBA8E34FF3ECCED95FF903B" box="[249,416,1271,1290]" class="Magnoliopsida" family="Solanaceae" genus="Lycium" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="34" phylum="Tracheophyta" rank="species" species="sokotranum">
<emphasis id="B91AEACAFFBA8E34FF3ECCED95FF903B" box="[249,416,1271,1290]" italics="true" pageId="11" pageNumber="34">Lycium sokotranum</emphasis>
</taxonomicName>
shrub in foreground:habitat and host plant of
<taxonomicName id="4C6E4D5BFFBA8E34FCE2CCED94C19014" authority="Burckhardt &amp; Mifsud, 1998" authorityName="Burckhardt &amp; Mifsud" authorityYear="1998" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B91AEACAFFBA8E34FCE2CCED97DF903B" box="[805,896,1271,1290]" italics="true" pageId="11" pageNumber="34">D. elegans</emphasis>
<bibRefCitation id="EFFF4B29FFBA8E34FC43CCED94C19014" author="Burckhardt &amp; Mifsud" firstAuthor="Burckhardt" pageId="11" pageNumber="34" pagination="7 - 49" refId="ref15906" refString="BURCKHARDT D. &amp; MIFSUD D. 1998: Jumping plant-lice (Insecta: Hemiptera) of the Arabian Peninsula. Fauna of Arabia 17: 7 - 49." type="journal article" year="1998">Burckhardt &amp; Mifsud, 1998</bibRefCitation>
</taxonomicName>
. Photographs by I. Malenovský (23, 24) and P. Kment (25, 26).
</paragraph>
</caption>
<paragraph id="8BD136D8FFBA8E34FF57CDBA907290E6" blockId="11.[113,1089,1376,1591]" pageId="11" pageNumber="34">
<emphasis id="B91AEACAFFBA8E34FF57CDBA95749086" box="[144,299,1440,1463]" italics="true" pageId="11" pageNumber="34">Measurements</emphasis>
(n = 1; in mm). BL 1.67, BW 1.66, AL 0.35, FL 1.04, TL 0.34, CPL 0.50, CPW 0.82, CRW 0.18. Ratios: BL/BW 1.01, AL/FL 0.34, CL/CW 0.61, CRW/CPW 0.22.
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFBA8E31FFB6CDFA97E79462" lastPageId="14" lastPageNumber="37" pageId="11" pageNumber="34" type="diagnosis">
<paragraph id="8BD136D8FFBA8E31FFB6CDFA97E79462" blockId="11.[113,1089,1376,1591]" lastBlockId="14.[151,1127,220,819]" lastPageId="14" lastPageNumber="37" pageId="11" pageNumber="34">
<emphasis id="B91AEACAFFBA8E34FFB6CDFA953290C6" bold="true" box="[113,365,1504,1527]" pageId="11" pageNumber="34">Differential diagnosis.</emphasis>
<taxonomicName id="4C6E4D5BFFBA8E34FEB2CDFA95AE90C6" authorityName="Low" authorityYear="1880" box="[373,497,1504,1527]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEACAFFBA8E34FEB2CDFA95AE90C6" box="[373,497,1504,1527]" italics="true" pageId="11" pageNumber="34">Diaphorina</emphasis>
</taxonomicName>
is a large genus with some 80 species described from tropical and subtropical regions of the Old World (
<bibRefCitation id="EFFF4B29FFBA8E34FD59CE1A971A9326" author="LOGINOVA" box="[670,837,1536,1559]" firstAuthor="LOGINOVA" pageId="11" pageNumber="34" pagination="30 - 123" refId="ref17154" refString="LOGINOVA M. M. 1978: Novye vidy psillid (Homoptera, Psylloidea). [New species of psyllids (Homoptera, Psylloidea).] In: MEDVEDEV G. S. (ed.): Novye vidy zhivotnykh. [New animal species.] Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 61: 30 - 123 (in Russian)." type="journal article" year="1978">LOGINOVA 1978</bibRefCitation>
,
<bibRefCitation id="EFFF4B29FFBA8E34FC95CE1A97A89326" author="OUVRARD" box="[850,1015,1536,1559]" firstAuthor="OUVRARD" pageId="11" pageNumber="34" refId="ref17603" refString="OUVRARD D. 2014: Psyl'list - the world Psylloidea database. Available online: http: // www. hemiptera-databases. com / psyllist (accessed on 5 July 2014)." type="url" year="2014">OUVRARD 2014</bibRefCitation>
). Only one of them,
<taxonomicName id="4C6E4D5BFFBA8E34FF39CE3A96299306" authority="(Pettey, 1924)" baseAuthorityName="Pettey" baseAuthorityYear="1924" box="[254,630,1568,1591]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="carissae">
<emphasis id="B91AEACAFFBA8E34FF39CE3A95859306" box="[254,474,1568,1591]" italics="true" pageId="11" pageNumber="34">Diaphorina carissae</emphasis>
(
<bibRefCitation id="EFFF4B29FFBA8E34FE20CE3A96319306" author="Pettey" box="[487,622,1568,1591]" firstAuthor="Pettey" pageId="11" pageNumber="34" pagination="21 - 30" refId="ref17751" refString="PETTEY F. W. 1924: South African psyllids. Entomology Memoirs of the Department of Agriculture of the University of South Africa 2: 21 - 30." type="journal article" year="1924">Pettey, 1924</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F3797648FFBA8E34FD71CE3A971F9306" box="[694,832,1568,1591]" name="South Africa" pageId="11" pageNumber="34">South Africa</collectingCountry>
, has been reported from
<taxonomicName id="4C6E4D5BFFBD8E33FF50CD87954C9085" box="[151,275,1437,1460]" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="35" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91AEACAFFBD8E33FF50CD8794B59085" box="[151,234,1437,1460]" italics="true" pageId="12" pageNumber="35">Carissa</emphasis>
sp.
</taxonomicName>
, i.e. the same or related host plant as
<taxonomicName id="4C6E4D5BFFBD8E33FD7FCD8797159085" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[696,842,1437,1460]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBD8E33FD7FCD8797159085" box="[696,842,1437,1460]" italics="true" pageId="12" pageNumber="35">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBD8E33FC94CD8797F79085" box="[851,936,1437,1460]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
According to the published description,
<taxonomicName id="4C6E4D5BFFBD8E33FE4FCDA7965F90E5" baseAuthorityName="Pettey" baseAuthorityYear="1924" box="[392,512,1469,1492]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="carissae">
<emphasis id="B91AEACAFFBD8E33FE4FCDA7965F90E5" box="[392,512,1469,1492]" italics="true" pageId="12" pageNumber="35">D. carissae</emphasis>
</taxonomicName>
is generally smaller (HW =
<quantity id="4C969B3DFFBD8E33FCECCDA7972190E5" box="[811,894,1469,1492]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="35" unit="mm" value="0.4">0.4 mm</quantity>
, WL =
<quantity id="4C969B3DFFBD8E33FC0CCDA7907490E5" box="[971,1067,1469,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="35" unit="mm" value="1.65">1.65 mm</quantity>
, WW =
<quantity id="4C969B3DFFBD8E33FF6BCDC794A290C5" box="[172,253,1501,1524]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="35" unit="mm" value="0.7">0.7 mm</quantity>
for female), with lighter head and abdomen (pale yellowish brown to yellow), and shorter genal processes (only a little more than half as long as vertex) than
<taxonomicName id="4C6E4D5BFFBD8E33FC75CDE7901D9325" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[946,1090,1533,1556]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBD8E33FC75CDE7901D9325" box="[946,1090,1533,1556]" italics="true" pageId="12" pageNumber="35">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBD8E33FB8FCDE7949A9305" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
(
<bibRefCitation id="EFFF4B29FFBD8E33FF15CE0795049305" author="PETTEY" box="[210,347,1565,1588]" firstAuthor="PETTEY" pageId="12" pageNumber="35" pagination="21 - 30" refId="ref17751" refString="PETTEY F. W. 1924: South African psyllids. Entomology Memoirs of the Department of Agriculture of the University of South Africa 2: 21 - 30." type="journal article" year="1924">PETTEY 1924</bibRefCitation>
,
<bibRefCitation id="EFFF4B29FFBD8E33FEA1CE0795C09305" author="PETTEY" box="[358,415,1565,1588]" firstAuthor="PETTEY" pageId="12" pageNumber="35" pagination="125 - 142" refId="ref17781" refString="PETTEY F. W. 1925: New South African psyllids. South African Journal of Natural History 5: 125 - 142." type="journal article" year="1925">1925</bibRefCitation>
). Both species, however, share a similar forewing shape and maculation (the dark pattern is perhaps more expanded in
<taxonomicName id="4C6E4D5BFFBC8E32FD5DCD48974C9058" baseAuthorityName="Pettey" baseAuthorityYear="1924" box="[666,787,1362,1385]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="carissae">
<emphasis id="B91AEACAFFBC8E32FD5DCD48974C9058" box="[666,787,1362,1385]" italics="true" pageId="13" pageNumber="36">D. carissae</emphasis>
</taxonomicName>
) and may be closely related. A similar forewing shape and pattern is also present in
<taxonomicName id="4C6E4D5BFFBC8E32FD7DCD68979E90B8" authority="Capener, 1970" authorityName="Capener" authorityYear="1970" box="[698,961,1394,1417]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="petteyi">
<emphasis id="B91AEACAFFBC8E32FD7DCD68977E90B8" box="[698,801,1394,1417]" italics="true" pageId="13" pageNumber="36">D. petteyi</emphasis>
<bibRefCitation id="EFFF4B29FFBC8E32FCE0CD68979E90B8" author="Capener" box="[807,961,1394,1417]" firstAuthor="Capener" pageId="13" pageNumber="36" pagination="201 - 226" refId="ref16108" refString="CAPENER A. L. 1970: Southern African Psyllidae (Homoptera) - 2: Some new species of Diaphorina Low. Journal of the Entomological Society of Southern Africa 33: 201 - 226." type="journal article" year="1970">Capener, 1970</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3797648FFBC8E32FBC5CD6894EA9098" name="South Africa" pageId="13" pageNumber="36">South Africa</collectingCountry>
which is, however, slightly larger compared to
<taxonomicName id="4C6E4D5BFFBC8E32FD6DCD8897669098" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[682,825,1426,1449]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBC8E32FD6DCD8897669098" box="[682,825,1426,1449]" italics="true" pageId="13" pageNumber="36">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBC8E32FCF8CD8897CC9098" box="[831,915,1426,1449]" pageId="13" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, paler in general coloration, and its immatures have circumanal pore ring composed of four rows of pores and are associated with a different plant (
<taxonomicName id="4C6E4D5BFFBC8E32FE3CCDC8969490D8" box="[507,715,1490,1513]" class="Magnoliopsida" family="Polygalaceae" genus="Polygala" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="36" phylum="Tracheophyta" rank="species" species="myrtifolia">
<emphasis id="B91AEACAFFBC8E32FE3CCDC8969490D8" box="[507,715,1490,1513]" italics="true" pageId="13" pageNumber="36">Polygala myrtifolia</emphasis>
</taxonomicName>
L.,
<taxonomicName id="4C6E4D5BFFBC8E32FD31CDC897DB90D8" box="[758,900,1490,1513]" class="Magnoliopsida" family="Polygalaceae" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="36" phylum="Tracheophyta" rank="family">Polygalaceae</taxonomicName>
) (
<bibRefCitation id="EFFF4B29FFBC8E32FC5DCDC8906990D8" author="CAPENER" box="[922,1078,1490,1513]" firstAuthor="CAPENER" pageId="13" pageNumber="36" pagination="201 - 226" refId="ref16108" refString="CAPENER A. L. 1970: Southern African Psyllidae (Homoptera) - 2: Some new species of Diaphorina Low. Journal of the Entomological Society of Southern Africa 33: 201 - 226." type="journal article" year="1970">CAPENER 1970</bibRefCitation>
). The coarse surface spinulation of the forewing membrane of
<taxonomicName id="4C6E4D5BFFBC8E32FCC7CDE897CF9338" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[768,912,1522,1545]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBC8E32FCC7CDE897CF9338" box="[768,912,1522,1545]" italics="true" pageId="13" pageNumber="36">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBC8E32FC50CDE897B59338" box="[919,1002,1522,1545]" pageId="13" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C6E4D5BFFBC8E32FF75CE08959A9318" authority="Kuwayama, 1907" authorityName="Kuwayama" authorityYear="1907" box="[178,453,1554,1577]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="citri">
<emphasis id="B91AEACAFFBC8E32FF75CE08955F9318" box="[178,256,1554,1577]" italics="true" pageId="13" pageNumber="36">D. citri</emphasis>
Kuwayama, 1907
</taxonomicName>
and
<taxonomicName id="4C6E4D5BFFBC8E32FE3CCE08977D9318" authority="Mathur, 1975" authorityName="Mathur" authorityYear="1975" box="[507,802,1554,1577]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="communis">
<emphasis id="B91AEACAFFBC8E32FE3CCE0896D69318" box="[507,649,1554,1577]" italics="true" pageId="13" pageNumber="36">D. communis</emphasis>
<bibRefCitation id="EFFF4B29FFBC8E32FD56CE08977D9318" author="Mathur" box="[657,802,1554,1577]" firstAuthor="Mathur" pageId="13" pageNumber="36" refId="ref17381" refString="MATHUR R. N. 1975: Psyllidae of the Indian subcontinent. Indian Council of Agricultural Research, New Delhi, 429 pp." type="book" year="1975">Mathur, 1975</bibRefCitation>
</taxonomicName>
which are associated with
<taxonomicName id="4C6E4D5BFFBF8E31FF50C8C694A995C2" box="[151,246,220,243]" class="Magnoliopsida" family="Rutaceae" genus="Murraya" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEACAFFBF8E31FF50C8C694A995C2" box="[151,246,220,243]" italics="true" pageId="14" pageNumber="37">Murraya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D5BFFBF8E31FEEDC8C6964695C2" box="[298,537,220,243]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91AEACAFFBF8E31FEEDC8C6953295C2" box="[298,365,220,243]" italics="true" pageId="14" pageNumber="37">Citrus</emphasis>
spp. (Rutaceae)
</taxonomicName>
and differ, besides other characters, in a more angulate apical portion of anterior forewing margin (
<bibRefCitation id="EFFF4B29FFBF8E31FD9CC8E696B09422" author="MATHUR" box="[603,751,252,275]" firstAuthor="MATHUR" pageId="14" pageNumber="37" refId="ref17381" refString="MATHUR R. N. 1975: Psyllidae of the Indian subcontinent. Indian Council of Agricultural Research, New Delhi, 429 pp." type="book" year="1975">MATHUR 1975</bibRefCitation>
,
<bibRefCitation id="EFFF4B29FFBF8E31FD30C8E697C69422" author="LOGINOVA" box="[759,921,252,275]" firstAuthor="LOGINOVA" pageId="14" pageNumber="37" pagination="30 - 123" refId="ref17154" refString="LOGINOVA M. M. 1978: Novye vidy psillid (Homoptera, Psylloidea). [New species of psyllids (Homoptera, Psylloidea).] In: MEDVEDEV G. S. (ed.): Novye vidy zhivotnykh. [New animal species.] Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 61: 30 - 123 (in Russian)." type="journal article" year="1978">LOGINOVA 1978</bibRefCitation>
), which is regularly rounded in
<taxonomicName id="4C6E4D5BFFBF8E31FED4C90695FC9402" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[275,419,284,307]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBF8E31FED4C90695FC9402" box="[275,419,284,307]" italics="true" pageId="14" pageNumber="37">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBF8E31FE6CC90695A09402" box="[427,511,284,307]" pageId="14" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
From other three
<taxonomicName id="4C6E4D5BFFBF8E31FD04C906972C9402" box="[707,883,284,307]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91AEACAFFBF8E31FD04C90697609402" box="[707,831,284,307]" italics="true" pageId="14" pageNumber="37">Diaphorina</emphasis>
spp.
</taxonomicName>
currently known from
<collectingCountry id="F3797648FFBF8E31FF50C92694B49462" box="[151,235,316,339]" name="Yemen" pageId="14" pageNumber="37">Socotra</collectingCountry>
,
<taxonomicName id="4C6E4D5BFFBF8E31FF31C92695D99462" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[246,390,316,339]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBF8E31FF31C92695D99462" box="[246,390,316,339]" italics="true" pageId="14" pageNumber="37">D. caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBF8E31FE4BC92695809462" box="[396,479,316,339]" pageId="14" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
differs in characters given in the key below.
</paragraph>
</subSubSection>
<caption id="DF116650FFBD8E33FF50CCF19563907F" ID-DOI="http://doi.org/10.5281/zenodo.5314364" ID-Zenodo-Dep="5314364" httpUri="https://zenodo.org/record/5314364/files/figure.png" pageId="12" pageNumber="35" startId="12.[151,188,1259,1278]" targetBox="[184,1090,217,1226]" targetPageId="12">
<paragraph id="8BD136D8FFBD8E33FF50CCF19563907F" blockId="12.[151,1127,1259,1358]" pageId="12" pageNumber="35">
Figs 2733.
<taxonomicName id="4C6E4D5BFFBD8E33FEC5CCF1959B91CF" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[258,452,1259,1278]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBD8E33FEC5CCF1959B91CF" box="[258,452,1259,1278]" italics="true" pageId="12" pageNumber="35">Diaphorina caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBD8E33FE0DCCF1965091CF" box="[458,527,1259,1278]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
, adult. 27 male terminalia, outer lateral view; 28 paramere, lateral view, inner face; 29 distal segment of aedeagus, lateral view; 30 female terminalia, lateral view, with detail of circumanal ring structure; 31 detail of dorsal and apex of ventral valvulae, lateral view; 32 antenna; 33 antennal segments 9 and 10.
</paragraph>
</caption>
<caption id="DF116650FFBC8E32FFB6CCCF901D902C" ID-DOI="http://doi.org/10.5281/zenodo.5314366" ID-Zenodo-Dep="5314366" httpUri="https://zenodo.org/record/5314366/files/figure.png" pageId="13" pageNumber="36" startId="13.[113,150,1237,1256]" targetBox="[120,1043,220,1210]" targetPageId="13">
<paragraph id="8BD136D8FFBC8E32FFB6CCCF901D902C" blockId="13.[113,1090,1237,1309]" pageId="13" pageNumber="36">
Figs 3438.
<taxonomicName id="4C6E4D5BFFBC8E32FF1FCCCF95C891D9" authority="Malenovský &amp; Burckhardt, 2014" authorityName="Malenovský &amp; Burckhardt" authorityYear="2014" box="[216,407,1237,1256]" class="Insecta" family="Liviidae" genus="Diaphorina" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
<emphasis id="B91AEACAFFBC8E32FF1FCCCF95C891D9" box="[216,407,1237,1256]" italics="true" pageId="13" pageNumber="36">Diaphorina caliginosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957B1FFBC8E32FE5CCCCF958191D9" box="[411,478,1237,1256]" pageId="13" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, fifth instar immature.34 body, dorsal view (left side only, with position and relative size of ventral circumanal pore ring indicated on right side of midline); 35 detail of lanceolate setae on caudal plate margin; 36 antenna; 37 detail of tibiotarsus apex; 38 detail of circumanal pore ring (right side only).
</paragraph>
</caption>
<subSubSection id="C3746553FFBF8E31FF50C94696C89482" pageId="14" pageNumber="37" type="etymology">
<paragraph id="8BD136D8FFBF8E31FF50C94696C89482" blockId="14.[151,1127,220,819]" pageId="14" pageNumber="37">
<emphasis id="B91AEACAFFBF8E31FF50C94695489442" bold="true" box="[151,279,348,371]" pageId="14" pageNumber="37">Etymology.</emphasis>
Derived from the Latin adjective
<emphasis id="B91AEACAFFBF8E31FDB4C94696B49442" box="[627,747,348,371]" italics="true" pageId="14" pageNumber="37">caliginosus</emphasis>
(= covered with mist, foggy, cloudy, dark, gloomy), referring to the weather conditions under which the
<typeStatus id="54D5887AFFBF8E31FCA7C96697D194A2" box="[864,910,380,403]" pageId="14" pageNumber="37">type</typeStatus>
series was collected and the generally dark coloration of the species.
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFBF8E31FF50C9A690389782" pageId="14" pageNumber="37" type="biology_ecology">
<paragraph id="8BD136D8FFBF8E31FF50C9A6976894E2" blockId="14.[151,1127,220,819]" box="[151,823,444,467]" pageId="14" pageNumber="37">
<emphasis id="B91AEACAFFBF8E31FF50C9A6954A94E2" bold="true" box="[151,277,444,467]" pageId="14" pageNumber="37">
<collectingRegion id="49AAF83AFFBF8E31FF50C9A6949394E2" box="[151,204,444,467]" country="Afghanistan" name="Khost" pageId="14" pageNumber="37">Host</collectingRegion>
plant.
</emphasis>
<taxonomicName id="4C6E4D5BFFBF8E31FEDBC9A6958594E2" box="[284,474,444,467]" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="species" species="spinarum">
<emphasis id="B91AEACAFFBF8E31FEDBC9A6958594E2" box="[284,474,444,467]" italics="true" pageId="14" pageNumber="37">Carissa spinarum</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C6E4D5BFFBF8E31FDC1C9A696C694E2" box="[518,665,444,467]" class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="family">Apocynaceae</taxonomicName>
) (
<figureCitation id="13552A5DFFBF8E31FD69C9A6977394E2" box="[686,812,444,467]" captionStart="Figs 2326" captionStartId="11.[113,150,1165,1184]" captionTargetBox="[113,1088,219,1139]" captionTargetId="figure-103@11.[113,1091,219,1139]" captionTargetPageId="11" captionText="Figs 2326. Habitats of Diaphorina spp. in Socotra. 23 montane evergreen woodland in Skand area, Hagher Mts., with a Carissa spinarum tree in foreground (centre): type locality of D. caliginosa sp. nov. 24 detail of a Carissa spinarum twig, host plant of D. caliginosa sp. nov. 25 montane shrubland with open grassland patches in wadi Madar, Hagher Mts.: type locality of D. hagherensis sp. nov. 26 open dry woodland on limestone plateau near Firmihin with a Lycium sokotranum shrub in foreground:habitat and host plant of D. elegans Burckhardt &amp; Mifsud, 1998. Photographs by I. Malenovský (23, 24) and P. Kment (25, 26)." figureDoi="http://doi.org/10.5281/zenodo.5314360" httpUri="https://zenodo.org/record/5314360/files/figure.png" pageId="14" pageNumber="37">Figs 2324</figureCitation>
).
</paragraph>
<paragraph id="8BD136D8FFBF8E31FF50C9C690389782" blockId="14.[151,1127,220,819]" pageId="14" pageNumber="37">
<emphasis id="B91AEACAFFBF8E31FF50C9C694AC94C2" bold="true" box="[151,243,476,499]" pageId="14" pageNumber="37">Biology.</emphasis>
Adults and one skin of a fifth instar immature were found close to the central vein on lower surface of leaves of the host plant. Adults, often immerged in water drops from condensed fog, were attached to the leaf only with their fore body, with abdomen oriented almost vertically towards the ground. No deformations of the host plant could be observed.
<emphasis id="B91AEACAFFBF8E31FF50CA4695FB9742" bold="true" box="[151,420,604,627]" pageId="14" pageNumber="37">
Occurrence in
<collectingCountry id="F3797648FFBF8E31FE82CA4695C19742" box="[325,414,604,627]" name="Yemen" pageId="14" pageNumber="37">Socotra</collectingCountry>
.
</emphasis>
Found only in one small area in the highest part of the Hagher mountains, in a dense montane evergreen woodland belonging to the
<emphasis id="B91AEACAFFBF8E31FCB0CA6695F79782" italics="true" pageId="14" pageNumber="37">
<taxonomicName id="4C6E4D5BFFBF8E31FCB0CA66979C97A2" box="[887,963,636,659]" class="Magnoliopsida" family="Lamiaceae" genus="Leucas" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="genus">Leucas</taxonomicName>
hagghierensis- -
<taxonomicName id="4C6E4D5BFFBF8E31FF59CA8695F79782" box="[158,424,668,691]" class="Magnoliopsida" family="Pittosporaceae" genus="Pittosporum" kingdom="Plantae" order="Apiales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="species" species="viridiflorum">Pittosporum viridiflorum</taxonomicName>
</emphasis>
community (
<bibRefCitation id="EFFF4B29FFBF8E31FDF3CA86977F9782" author="KURSCHNER" box="[564,800,668,691]" etAl="et al." firstAuthor="KURSCHNER" pageId="14" pageNumber="37" pagination="1 - 176" refId="ref16771" refString="KURSCHNER H., HEIN P., KILIAN N. &amp; HUBAISHAN M. 2006: Diversity and zonation of forests and woodlands of the mountains of northern Socotra, Yemen. Pp. 11 - 55. In: KILIAN N. &amp; HUBAISHAN M. (eds): Biodiversity of Socotra: forests, woodlands and bryophytes. Englera 28: 1 - 176 + 16 pls." type="journal article" year="2006">KÜRSCHNER et al. 2006</bibRefCitation>
,
<bibRefCitation id="EFFF4B29FFBF8E31FCEECA86905B9782" author="BROWN &amp; MIES" box="[809,1028,668,691]" firstAuthor="BROWN" pageId="14" pageNumber="37" refId="ref15725" refString="BROWN G. &amp; MIES B. A. 2012: Vegetation Ecology of Socotra. Springer, Dordrecht - Heidelberg - New York - London, 379 pp." type="book" year="2012">BROWN &amp; MIES 2012</bibRefCitation>
;
<figureCitation id="13552A5DFFBF8E31FBCACA8690059782" box="[1037,1114,668,691]" captionStart="Figs 2326" captionStartId="11.[113,150,1165,1184]" captionTargetBox="[113,1088,219,1139]" captionTargetId="figure-103@11.[113,1091,219,1139]" captionTargetPageId="11" captionText="Figs 2326. Habitats of Diaphorina spp. in Socotra. 23 montane evergreen woodland in Skand area, Hagher Mts., with a Carissa spinarum tree in foreground (centre): type locality of D. caliginosa sp. nov. 24 detail of a Carissa spinarum twig, host plant of D. caliginosa sp. nov. 25 montane shrubland with open grassland patches in wadi Madar, Hagher Mts.: type locality of D. hagherensis sp. nov. 26 open dry woodland on limestone plateau near Firmihin with a Lycium sokotranum shrub in foreground:habitat and host plant of D. elegans Burckhardt &amp; Mifsud, 1998. Photographs by I. Malenovský (23, 24) and P. Kment (25, 26)." figureDoi="http://doi.org/10.5281/zenodo.5314360" httpUri="https://zenodo.org/record/5314360/files/figure.png" pageId="14" pageNumber="37">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3746553FFBF8E31FF50CAA6958D9602" pageId="14" pageNumber="37" type="distribution">
<paragraph id="8BD136D8FFBF8E31FF50CAA6958D9602" blockId="14.[151,1127,220,819]" pageId="14" pageNumber="37">
<emphasis id="B91AEACAFFBF8E31FF50CAA6957597E2" bold="true" box="[151,298,700,723]" pageId="14" pageNumber="37">Distribution.</emphasis>
So far only known from the
<collectingCountry id="F3797648FFBF8E31FD9ECAA696A997E2" box="[601,758,700,723]" name="Socotra Island" pageId="14" pageNumber="37">Socotra Island</collectingCountry>
. The host plant,
<taxonomicName id="4C6E4D5BFFBF8E31FC65CAA6903F97E2" box="[930,1120,700,723]" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="37" phylum="Tracheophyta" rank="species" species="spinarum">
<emphasis id="B91AEACAFFBF8E31FC65CAA6903F97E2" box="[930,1120,700,723]" italics="true" pageId="14" pageNumber="37">Carissa spinarum</emphasis>
</taxonomicName>
, in its currently accepted taxonomic definition, is very widely distributed throughout Africa,
<collectingCountry id="F3797648FFBF8E31FF50CAE695459622" box="[151,282,764,787]" name="Madagascar" pageId="14" pageNumber="37">Madagascar</collectingCountry>
,
<collectingCountry id="F3797648FFBF8E31FEE1CAE695C49622" box="[294,411,764,787]" name="Seychelles" pageId="14" pageNumber="37">Seychelles</collectingCountry>
, Mascarenes, Arabia,
<collectingCountry id="F3797648FFBF8E31FD4ECAE6969D9622" box="[649,706,764,787]" name="India" pageId="14" pageNumber="37">India</collectingCountry>
,
<collectingCountry id="F3797648FFBF8E31FD08CAE6974E9622" box="[719,785,764,787]" name="China" pageId="14" pageNumber="37">China</collectingCountry>
, South
<collectingCountry id="F3797648FFBF8E31FCA4CAE697939622" box="[867,972,764,787]" name="China" pageId="14" pageNumber="37">East Asia</collectingCountry>
,
<collectingCountry id="F3797648FFBF8E31FC11CAE690679622" box="[982,1080,764,787]" name="Australia" pageId="14" pageNumber="37">Australia</collectingCountry>
and New
<collectingCountry id="F3797648FFBF8E31FF17CB0695429602" box="[208,285,796,819]" name="Guinea-Bissau" pageId="14" pageNumber="37">Guinea</collectingCountry>
(
<bibRefCitation id="EFFF4B29FFBF8E31FEEBCB0695999602" author="HASSLER" box="[300,454,796,819]" firstAuthor="HASSLER" pageId="14" pageNumber="37" pagination="2000" refId="ref16498" refString="HASSLER M. 2014: World Plants: Synonymic checklists of the vascular plants of the World (version March 2014). In: ROSKOV Y., KUNZE T., ORRELL T., ABUCAY L., CULHAM A., BAILLY N., KIRK P., BOURGOIN T., DECOCK W., DE WEVER A. (eds): Species 2000 &amp; ITIS Catalogue of Life, 28 th June 2014. Digital resource at www. catalogueoflife. org / col. Species 2000: Naturalis, Leiden." type="book chapter" year="2014">HASSLER 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>