164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document id="6ABABAFCD43AC8B307B7C8E6FA6C1D7C" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C787854614FFDA4251400532A9F832" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Biomphalodonta Herbert 2020, gen. nov." docType="treatment" docUuid="CE2A2B8F-D281-4E81-BF8D-5A705868D71A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="25" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="25" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="5973995E295EE5EF1D2BC42C2D8E5687" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FCAF13B897F73DC6A798DF8070DCC1E3">
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<mods:title id="577430192DE282332D8FDBC1D00BBFB8">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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</mods:titleInfo>
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<mods:name id="77C3696D5E7F2F331C568FD92D49EAF2" type="personal">
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<mods:role id="1CB867ADAEBA8D9188FE76D892041F2C">
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<mods:roleTerm id="337D306FFD531EA19A923BD30C316657">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8EF12FBADFD3F20024E18E896D35FF53">Herbert, David G.</mods:namePart>
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<mods:nameIdentifier id="1F23E1196FD2A8B9B09437E416AED4EA" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
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<mods:affiliation id="B6EA1154EF89337E4A9B59E480A66B18">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
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<mods:nameIdentifier id="1F4AA956DFBCABC7FA1602954C017E70" type="email">phasianella@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="D168B30F7C890204A942BE0B832D5012">text</mods:typeOfResource>
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<mods:titleInfo id="8C52B4CE822472DE84E64E611FA91A60">
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<mods:title id="712F2D2752828F36A719FF0BF39F7BA7">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="2D3D5C52401482699074E5B10FEF3B29">
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<mods:date id="605431F162CDA696F89F9B10CBE5D966">2020</mods:date>
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<mods:detail id="9E3ACB96E2B7BF343F2B21610B658AB8" type="pubDate">
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<mods:number id="07B48BABDD18036AFFD5B2D44D7796BA">2020-04-17</mods:number>
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</mods:detail>
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<mods:detail id="9E05B9E5F0C43F859AA247A4DE5EB748" type="volume">
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<mods:number id="D4EB4864A4365BEE1B7DCDEBA0920979">629</mods:number>
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</mods:detail>
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<mods:start id="4DA9F489E67FCB8212416FDAA318E7F1">1</mods:start>
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<mods:end id="AC99CD678DDC1C43BA1B92FD358A4720">55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="F7174041A3039695A1112536CD8ED1AD">journal article</mods:classification>
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<mods:identifier id="84492870B97C52F50B62F6822ABA5E72" type="CLB-Dataset">22877</mods:identifier>
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<mods:identifier id="0E00071B1DADE45E938C53458241C858" type="DOI">10.5852/ejt.2020.629</mods:identifier>
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<mods:identifier id="243583C29B37057EC00E8AF7F4FC80E2" type="GBIF-Dataset">3722aa65-b935-4294-ac20-0d1179da9015</mods:identifier>
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<mods:identifier id="ECDDD2F8E75EAF8913857322BBA3E9C1" type="Zenodo-Dep">3762280</mods:identifier>
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<mods:identifier id="476ED09DC78D05318A1FF72275B7A857" type="ZooBank">ECEBD539-6E3E-45BE-A0CB-264DF3270CC0</mods:identifier>
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</mods:mods>
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<treatment id="03C787854614FFDA4251400532A9F832" ID-DOI="http://doi.org/10.5281/zenodo.3804715" ID-GBIF-Taxon="163732375" ID-Zenodo-Dep="3804715" LSID="urn:lsid:zoobank.org:act:CE2A2B8F-D281-4E81-BF8D-5A705868D71A" httpUri="http://treatment.plazi.org/id/03C787854614FFDA4251400532A9F832" lastPageNumber="25" pageId="24" pageNumber="25">
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<subSubSection id="C37465184614FFDA4251400535F3FCBC" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BD136934614FFDA4251400532D9FCE7" blockId="24.[362,1227,803,869]" box="[595,993,803,830]" pageId="24" pageNumber="25">
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<heading id="D09981FF4614FFDA4251400532D9FCE7" box="[595,993,803,830]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
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Genus
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<taxonomicName id="4C6E4D104614FFDA42A740023257FCE7" ID-CoL="7NZGW" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[677,879,803,830]" class="Gastropoda" family="Charopidae" genus="Biomphalodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B91AEA814614FFDA42A740023257FCE7" bold="true" box="[677,879,803,830]" italics="true" pageId="24" pageNumber="25">Biomphalodonta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA4374400532D9FCE7" box="[886,993,804,830]" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BD136934614FFDA4168406B35F3FCBC" blockId="24.[362,1227,803,869]" box="[362,1227,842,869]" pageId="24" pageNumber="25">
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<uri id="FFFF3A914614FFDA4168406B35F3FCBC" box="[362,1227,842,869]" pageId="24" pageNumber="25">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC80C464614FFDA4289406B35F3FCBC" box="[651,1227,842,869]" pageId="24" pageNumber="25">CE2A2B8F-D281-4E81-BF8D-5A705868D71A</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37465184614FFDA40BF40AE33FFFC02" pageId="24" pageNumber="25" type="type_taxon">
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<paragraph id="8BD136934614FFDA40BF40AE3060FC70" blockId="24.[189,344,911,937]" box="[189,344,911,937]" pageId="24" pageNumber="25">
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<heading id="D09981FF4614FFDA40BF40AE3060FC70" bold="true" box="[189,344,911,937]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B91AEA814614FFDA40BF40AE3060FC70" bold="true" box="[189,344,911,937]" pageId="24" pageNumber="25">
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<typeStatus id="54D588314614FFDA40BF40AE31C2FC70" box="[189,250,911,937]" pageId="24" pageNumber="25">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934614FFDA40BF40E033FFFC02" blockId="24.[189,711,961,987]" box="[189,711,961,987]" pageId="24" pageNumber="25">
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<taxonomicName id="4C6E4D104614FFDA40BF40E03337FC02" ID-CoL="7QBY7" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[189,527,961,987]" class="Gastropoda" family="Charopidae" genus="Biomphalodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="forticostata" status="gen. et sp. nov.">
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<emphasis id="B91AEA814614FFDA40BF40E03337FC02" box="[189,527,961,987]" italics="true" pageId="24" pageNumber="25">Biomphalodonta forticostata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA421440E033FFFC02" box="[534,711,961,987]" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465184614FFDA40BF472430DFFAE1" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8BD136934614FFDA40BF4724300EFBC6" blockId="24.[189,310,1029,1055]" box="[189,310,1029,1055]" pageId="24" pageNumber="25">
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<heading id="D09981FF4614FFDA40BF4724300EFBC6" bold="true" box="[189,310,1029,1055]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B91AEA814614FFDA40BF4724300EFBC6" bold="true" box="[189,310,1029,1055]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934614FFDA40BF471630DFFAE1" blockId="24.[189,1399,1079,1337]" pageId="24" pageNumber="25">
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Shell relatively large (max. diameter
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<quantity id="4C969B764614FFDA427D471933D8FB8B" box="[639,736,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="24" pageNumber="25" unit="mm" value="2.7">2.7 mm</quantity>
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), biconcave with deeply sunken spire; umbilicus wide; whorls deep and tightly coiled, strongly rounded apically and basally, less so at periphery; last adult whorl not descendant. Protoconch with fine, close-set, regularly spaced axial riblets; riblets crisp and simple. Teleoconch with relatively coarse, widely spaced, lamellate axial riblets; intervals between riblets with numerous, fine, intermediary axial threads and even finer spiral threads. Aperture narrowly lunate; parietal dentition absent; palatal region with 1–2 axially aligned rows of denticles set back up to ½ whorl behind outer lip.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465184614FFDA40BF4642355BFA0C" pageId="24" pageNumber="25" type="etymology">
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<paragraph id="8BD136934614FFDA40BF4642307CFAA4" blockId="24.[189,324,1379,1405]" box="[189,324,1379,1405]" pageId="24" pageNumber="25">
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<heading id="D09981FF4614FFDA40BF4642307CFAA4" bold="true" box="[189,324,1379,1405]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B91AEA814614FFDA40BF4642307CFAA4" bold="true" box="[189,324,1379,1405]" pageId="24" pageNumber="25">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934614FFDA40BF46B4355BFA0C" blockId="24.[189,1398,1428,1494]" pageId="24" pageNumber="25">
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From the Latin
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<emphasis id="B91AEA814614FFDA417B46B430A2FA76" box="[377,410,1429,1455]" italics="true" pageId="24" pageNumber="25">bi-</emphasis>
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: two, twice and the Greek
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<emphasis id="B91AEA814614FFDA42E246B4326AFA76" box="[736,850,1429,1455]" italics="true" pageId="24" pageNumber="25">omphalos</emphasis>
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(
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<emphasis id="B91AEA814614FFDA436746B532F1FA76" box="[869,969,1428,1455]" italics="true" pageId="24" pageNumber="25">Ομφαλός</emphasis>
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): navel, and
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<emphasis id="B91AEA814614FFDA446546B43592FA76" box="[1127,1194,1429,1455]" italics="true" pageId="24" pageNumber="25">donta</emphasis>
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: a contraction of
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<taxonomicName id="4C6E4D104614FFDA40BF469A300BFA0C" authorityName="Melvill & Ponsonby" authorityYear="1908" box="[189,307,1467,1493]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
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<emphasis id="B91AEA814614FFDA40BF469A300BFA0C" box="[189,307,1467,1493]" italics="true" pageId="24" pageNumber="25">Afrodonta</emphasis>
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</taxonomicName>
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; with reference to the biconcave shell morphology. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465184614FFDA40BF452132A9F832" pageId="24" pageNumber="25" type="discussion">
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<paragraph id="8BD136934614FFDA40BF45213017F9C3" blockId="24.[189,303,1536,1562]" box="[189,303,1536,1562]" pageId="24" pageNumber="25">
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<heading id="D09981FF4614FFDA40BF45213017F9C3" bold="true" box="[189,303,1536,1562]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B91AEA814614FFDA40BF45213017F9C3" bold="true" box="[189,303,1536,1562]" pageId="24" pageNumber="25">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934614FFDA40BF451335D8F93F" blockId="24.[189,1399,1586,1766]" pageId="24" pageNumber="25">
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<taxonomicName id="4C6E4D104614FFDA40BF451330B9F995" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[189,385,1586,1612]" class="Gastropoda" family="Charopidae" genus="Biomphalodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B91AEA814614FFDA40BF451330B9F995" box="[189,385,1586,1612]" italics="true" pageId="24" pageNumber="25">Biomphalodonta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA4188451330CFF995" box="[394,503,1586,1612]" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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is proposed for a single highly characteristic, range-restricted species. The axially costate protoconch is similar to that found in
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<taxonomicName id="4C6E4D104614FFDA433F457932E3F9AB" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[829,987,1624,1650]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B91AEA814614FFDA433F457932E3F9AB" box="[829,987,1624,1650]" italics="true" pageId="24" pageNumber="25">Costulodonta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA43E74578356CF9AA" box="[997,1108,1625,1651]" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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, but the deep, relatively large, biconcave shell and coarse teleoconch sculpture set it apart from members of that genus. The shape of the shell and the iterodont palatal dentition resemble those of
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<taxonomicName id="4C6E4D104614FFDA43FC45843589F966" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1022,1201,1701,1727]" class="Gastropoda" family="Charopidae" genus="Amatholedonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B91AEA814614FFDA43FC45843589F966" box="[1022,1201,1701,1727]" italics="true" pageId="24" pageNumber="25">Amatholedonta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA44BB4587341EF919" box="[1209,1318,1702,1728]" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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, but in that genus the protoconch is smooth and globose, and the teleoconch sculpture much finer.
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</paragraph>
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<paragraph id="8BD136934614FFDA40BF443032A9F832" blockId="24.[189,1399,1808,2027]" pageId="24" pageNumber="25">
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A number of small, edentate charopid species currently referred to
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<taxonomicName id="4C6E4D104614FFDA44044431347EF8F3" authority="Pilsbry, 1893" authorityName="Pilsbry" authorityYear="1893" box="[1030,1350,1808,1834]" class="Bryopsida" family="Mniaceae" genus="Trachycystis" kingdom="Plantae" order="Bryales" pageId="24" pageNumber="25" phylum="Bryophyta" rank="genus">
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<emphasis id="B91AEA814614FFDA4404443135A1F8F3" box="[1030,1177,1808,1834]" italics="true" pageId="24" pageNumber="25">Trachycystis</emphasis>
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Pilsbry, 1893
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</taxonomicName>
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are conchologically similar to
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<taxonomicName id="4C6E4D104614FFDA41FF4419339FF888" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[509,679,1847,1873]" class="Gastropoda" family="Charopidae" genus="Biomphalodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="forticostata" status="gen. et sp. nov.">
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<emphasis id="B91AEA814614FFDA41FF4419339FF888" box="[509,679,1847,1873]" italics="true" pageId="24" pageNumber="25">B. forticostata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA42B344163252F888" box="[689,874,1847,1873]" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, except for their lack of apertural dentition. These include,
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<emphasis id="B91AEA814614FFDA4169447F30EEF8AE" box="[363,470,1885,1911]" italics="true" pageId="24" pageNumber="25">inter alia</emphasis>
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,
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<taxonomicName id="4C6E4D104614FFDA41E0447C357AF8AE" authority="(Melvill & Ponsonby, 1892)" baseAuthorityName="Melvill & Ponsonby" baseAuthorityYear="1892" box="[482,1090,1885,1911]" class="Bryopsida" family="Mniaceae" genus="Trachycystis" kingdom="Plantae" order="Bryales" pageId="24" pageNumber="25" phylum="Bryophyta" rank="species" species="bathycoele">
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<emphasis id="B91AEA814614FFDA41E0447C33C1F8AE" box="[482,761,1885,1911]" italics="true" pageId="24" pageNumber="25">Trachycystis bathycoele</emphasis>
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(Melvill & Ponsonby, 1892)
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</taxonomicName>
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and
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<taxonomicName id="4C6E4D104614FFDA4475447F31C3F847" authority="Connolly, 1932" authorityName="Connolly" authorityYear="1932" class="Bryopsida" family="Mniaceae" genus="Trachycystis" kingdom="Plantae" order="Bryales" pageId="24" pageNumber="25" phylum="Bryophyta" rank="species" species="bifoveata">
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<emphasis id="B91AEA814614FFDA4475447F35C6F8AE" box="[1143,1278,1885,1911]" italics="true" pageId="24" pageNumber="25">T. bifoveata</emphasis>
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Connolly, 1932
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</taxonomicName>
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. Further study is needed in order to establish whether these also belong to
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<taxonomicName id="4C6E4D104614FFDA447A44A53404F847" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1144,1340,1924,1950]" class="Gastropoda" family="Charopidae" genus="Biomphalodonta" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B91AEA814614FFDA447A44A53404F847" box="[1144,1340,1924,1950]" italics="true" pageId="24" pageNumber="25">Biomphalodonta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4614FFDA454744A531D7F81C" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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It may be that the presence/absence of apertural dentition is of limited phylogenetic significance and that the genus contains both dentate and edentate species.
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</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |