treatments-xml/data/03/A1/EF/03A1EF5ECB3FD82DFC8298B45DEA0113.xml

<document id="4CC478A835CC1D2C3E011CE31E1103BF" ID-DOI="10.1590/1806-9665-RBENT-2023-0088" ID-ISSN="1806-9665" ID-Zenodo-Dep="10665223" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708000482662" checkinUser="felipe" docAuthor="Matson, Tanner Alex" docDate="2024" docId="03A1EF5ECB3FD82DFC8298B45DEA0113" docLanguage="en" docName="RevBrasEntomol.68.1.e20230088.pdf" docOrigin="Revista Brasileira de Entomologia (e 20230088) 68 (1)" docSource="http://dx.doi.org/10.1590/1806-9665-rbent-2023-0088" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Cyphoedma ashleyorum Matson 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FF989726CB3ED828FFA39D355C28035F" masterDocTitle="Cyphoedma n. gen. now described after a century of use, with the addition of a new species from Central America (Geometridae, Ennominae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1711378174356" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="622CAC9827939B02F6132EC48F599775">Cyphoedma n. gen. now described after a century of use, with the addition of a new species from Central America (Geometridae, Ennominae)</mods:title>
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<mods:namePart id="E70B58F4B15D022EC9C44B2BD44D33F8">Matson, Tanner Alex</mods:namePart>
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<treatment id="03A1EF5ECB3FD82DFC8298B45DEA0113" LSID="urn:lsid:plazi:treatment:03A1EF5ECB3FD82DFC8298B45DEA0113" httpUri="http://treatment.plazi.org/id/03A1EF5ECB3FD82DFC8298B45DEA0113" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="C3120DC3CB3FD829FC8298B459E90693" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BB75E48CB3FD829FC8298B4588506CA" blockId="1.[801,1197,1409,1429]" box="[801,1197,1409,1429]" pageId="1" pageNumber="2">
<heading id="D0FFE924CB3FD829FC8298B4588506CA" bold="true" box="[801,1197,1409,1429]" fontSize="8" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="B97C825ACB3FD829FC8298B4588506CA" bold="true" box="[801,1197,1409,1429]" pageId="1" pageNumber="2">
<taxonomicName id="4C0825CBCB3FD829FC8298B4584606CA" authority="Matson" authorityName="Matson" authorityYear="2024" box="[801,1134,1409,1429]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">Cyphoedma ashleyorum Matson</taxonomicName>
, 
<taxonomicNameLabel id="A24F3F21CB3FD829FBD498B7588506CA" box="[1143,1197,1410,1429]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB75E48CB3FD829FCE4988C59E90693" blockId="1.[800,1475,1465,1985]" box="[839,1473,1465,1484]" pageId="1" pageNumber="2">
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urn:lsid:zoobank.org:act: 
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<subSubSection id="C3120DC3CB3FD829FCE498E2586306B5" box="[839,1099,1495,1514]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BB75E48CB3FD829FCE498E2586306B5" blockId="1.[800,1475,1465,1985]" box="[839,1099,1495,1514]" pageId="1" pageNumber="2">
(
<figureCitation id="133342CDCB3FD829FCEC98E25F8006B5" box="[847,936,1495,1514]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figures 2</figureCitation>
a-2d, 3a-b, 4a, 5)
</paragraph>
</subSubSection>
<subSubSection id="C3120DC3CB3FD82CFCE498C159B20502" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BB75E48CB3FD829FCE498C15984044F" blockId="1.[800,1475,1465,1985]" pageId="1" pageNumber="2">
<emphasis id="B97C825ACB3FD829FCE498C15F850558" bold="true" box="[839,941,1524,1543]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="4C0825CBCB3FD829FC1298C158BE0558" authorityName="Matson" authorityYear="2024" box="[945,1174,1524,1543]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ashleyorum">Cyphoedma ashleyorum</taxonomicName>
is the predominant 
<taxonomicName id="4C0825CBCB3FD829FAEC98C159E90558" authorityName="Matson" authorityYear="2024" box="[1359,1473,1524,1543]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
in Central America and is not known to occur in South America. In males, the ground color is typically darker than congeners, and the darkened area of the hindwing underside does not extend as broadly down the outer margin to the tornus as seen in 
<taxonomicName id="4C0825CBCB3FD829FAAA9B5F59BA0522" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[1289,1426,1642,1661]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="mirafloresa">C. mirafloresa</taxonomicName>
(see arrows, 
<figureCitation id="133342CDCB3FD829FCD39BBD5FF205C4" box="[880,986,1672,1691]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figs. 2b, 2f</figureCitation>
). Males of both 
<taxonomicName id="4C0825CBCB3FD829FBDE9BBD592205C4" authorityName="Matson" authorityYear="2024" box="[1149,1290,1672,1691]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ashleyorum">C. ashleyorum</taxonomicName>
and 
<taxonomicName id="4C0825CBCB3FD829FA989BBD59EB05C4" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[1339,1475,1672,1691]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="mirafloresa">C. mirafloresa</taxonomicName>
also have more starkly contrasted hindwing undersides than seen in 
<taxonomicName id="4C0825CBCB3FD829FA0C9B905F8A058A" authorityName="Walker" authorityYear="1860" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata</taxonomicName>
(
<figureCitation id="133342CDCB3FD829FC0D9BF75805058A" box="[942,1069,1730,1749]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figs. 2b, 2f, 2j</figureCitation>
). Genitalia appear to be too similar among congeners to adequately diagnose, but COI barcode data readily supports the recognition of this new taxon (see Molecular characterization).
</paragraph>
<paragraph id="8BB75E48CB3FD82CFCE49A2E5DDB0091" blockId="1.[800,1475,1465,1985]" lastBlockId="4.[112,788,153,974]" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
<emphasis id="B97C825ACB3FD829FCE49A2E5FD40472" bold="true" box="[839,1020,1818,1838]" pageId="1" pageNumber="2">Male description (</emphasis>
<figureCitation id="133342CDCB3FD829FC5F9A2E581F0471" box="[1020,1079,1819,1838]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
a-b, 3a-b). Nearly redundant with generic description.Forewing length 
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. Head: Antenna filiform, brown scales above. Vertex dark brown with lighter brown scales near scape. Frons mostly dark brown, but with lighter brown scales medially; ventral margin white. Labial palpus porrect, slightly longer than diameter of eye, cream to white on inner surface, and variably brown and white on outer surface. Proboscis well-developed. Chaetosemata well separated. Thorax: Patagium mostly gray. Tegula brown. Mesothorax brown above, white below. Foreleg with large epiphysis; tibial spur formula 0–2–4; hindtibia swollen, bearing plume of long androconia folded into longitudinal groove on inner surface (when not deployed). Legs often whiter on internal surface and variably banded or speckled with yellow and dark brown on external surface; hind leg generally less speckled. Forewing: Predominantly brown with slightly falcate apex speckled with white and yellow. Costal and subapical areas given toward mustard-brown; costa sometimes with frosty highlights. Curved transverse silvery-white antemedial and postmedial lines. Dark brown medial transverse line through frosted central part of wing. Small black discal spot.Irregular, central, light brown patch distal to postmedial line.Subterminal area sometimes with jagged silvery-white line, especially toward apex.Outer margin with silvery-white highlight.Underside pale brown to white in proximal half; transitioning to dark brown in distal half. Fringe brown. Hindwing: Concolorous with forewing, but more frosted in basal half; bearing arcing, medial, transverse silvery-white line that joins with antemedial line of forewing. Small black discal spot present. Subterminal area sometimes with small, lighter brown flecks. Outer margin with silvery-white highlight. Underside pale brown to white in proximal two-thirds; transitioning to dark brown in distal third around apex. Fringe brown. Abdomen: Dark brown above, pale white to cream below. Third sternite with comb of setae. Male Genitalia: (
<figureCitation id="133342CDCB3AD82CFD0C9F2E5ECA0171" box="[687,738,539,558]" captionStart="Figure 3" captionStartId="2.[113,165,1710,1727]" captionTargetBox="[262,1353,1132,1673]" captionTargetId="figure-239@2.[249,1370,1125,1696]" captionTargetPageId="2" captionText="Figure 3 Cyphoedma male genitalia.(a)C.ashleyorum n.sp.,TAM-2023-288(USNM 154251),USNMENT01771290,paratype,USNM,genital capsule;(b) same,phallus;(c)C.mirafloresa (Dognin),TAM-2023-279 (USNM154250),USNMENT01771323,genital capsule;(d) same,phallus;(e)C.transvolutata (Walker),TAM-2023-277 (USNM 154248),USNMENT01771293, genital capsule; (f) same, phallus.Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10665231" httpUri="https://zenodo.org/record/10665231/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
a-b): Uncus short and tapered toward apex. Juxta plate-like and not strongly sclerotized. Vesica with two separate fields of small dentate cornuti on adjacent diverticula. 
<emphasis id="B97C825ACB3AD82CFEF99F415E3401D8" bold="true" box="[346,540,628,647]" pageId="4" pageNumber="5">Female description</emphasis>
(
<figureCitation id="133342CDCB3AD82CFD8A9F415E4D01D8" box="[553,613,628,647]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
c-d, 4a). Forewing length 
<quantity id="4CF0F3ADCB3AD82CFF179FA75D3501FA" box="[180,285,658,677]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="4" pageNumber="5" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23–26 mm</quantity>
.Head: As in male.Thorax: As in male; though lacking hind tibial androconia. Forewing: As in male, but often with lighter brown ground color and less pronounced central light brown patch distad to postmedial line above. Underside brown to tan in proximal half; transitioning to darker brown in distal half.Hindwing: As in male, but underside brown to tan in proximal half and transitioning to darker brown in distal half (less contrasted than male).Abdomen: As in male, but lacking third sternite comb of setae. Female Genitalia: (
<figureCitation id="133342CDCB3AD82CFDDB9E575E9C002A" box="[632,692,866,885]" captionStart="Figure 4" captionStartId="3.[83,137,1049,1066]" captionTargetBox="[220,1328,160,1008]" captionTargetId="figure-47@3.[207,1350,151,1032]" captionTargetPageId="3" captionText="Figure 4 Cyphoedma female genitalia. (a) C. ashleyorum n. sp., TAM-2023-291 (USNM 154253), USNMENT01771290, paratype; (b) C. mirafloresa (Dognin), TAM-2023-290 (USNM 154252), USNMENT01771322; (c) C. transvolutata (Walker), TAM-2023-278 (USNM 154249), USNMENT01771320. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10665233" httpUri="https://zenodo.org/record/10665233/files/figure.png" pageId="4" pageNumber="5">Fig. 4a</figureCitation>
): Papillae anales rounded; posterior apophysis 2.5x longer than anterior apophysis. Ductus bursae more heavily sclerotized at junction with corpus bursae. Corpus bursae pyriform; signum absent.
</paragraph>
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<paragraph id="8BB75E48CB3FD829FFF79AAE5E52049E" blockId="1.[83,756,1947,1985]" pageId="1" pageNumber="2">
<emphasis id="B97C825ACB3FD829FFF79AAE5CB204F3" bold="true" box="[84,154,1947,1964]" pageId="1" pageNumber="2">Figure 1</emphasis>
Natural resting posture of 
<taxonomicName id="4C0825CBCB3FD829FEDF9AAE5DF404F3" authorityName="Matson" authorityYear="2024" box="[380,476,1947,1964]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
. Image of female 
<taxonomicName id="4C0825CBCB3FD829FDD09AAE5CB1049E" authority="(Walker)" baseAuthorityName="Walker" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata (Walker)</taxonomicName>
taken in Santa 
<taxonomicName id="4C0825CBCB3FD829FEB19A855D7E049E" box="[274,342,1968,1985]" class="Insecta" family="Geometridae" genus="Catarina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Catarina</taxonomicName>
, Brazil.Photo credit: Douglas Meyer.
</paragraph>
</caption>
<caption id="DF770EC0CB3CD82AFFD39EFF5F860746" ID-DOI="http://doi.org/10.5281/zenodo.10665229" ID-Zenodo-Dep="10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="2" pageNumber="3" startId="2.[112,165,970,987]" targetBox="[135,1480,154,948]" targetPageId="2" targetType="figure">
<paragraph id="8BB75E48CB3CD82AFFD39EFF5F860746" blockId="2.[112,1505,970,1049]" pageId="2" pageNumber="3">
<emphasis id="B97C825ACB3CD82AFFD39EFF5C9D0084" bold="true" box="[112,181,970,987]" pageId="2" pageNumber="3">Figure 2</emphasis>
Adult habitus of 
<taxonomicName id="4C0825CBCB3CD82AFEE09EFF5D890084" authorityName="Matson" authorityYear="2024" box="[323,417,970,987]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
. Arrows pointing to diagnostic wing differences. (a) 
<taxonomicName id="4C0825CBCB3CD82AFCEA9EFF5F930084" authority="Matson, 2024" authorityName="Matson" authorityYear="2024" box="[841,955,970,987]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">C. ashleyorum</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3CD82AFC1C9EFF5FC30084" box="[959,1003,970,987]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
, male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) 
<taxonomicName id="4C0825CBCB3CD82AFF2F9EEA5CD500AF" authority="Matson, 2024" authorityName="Matson" authorityYear="2024" box="[140,253,991,1008]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">C. ashleyorum</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3CD82AFEA19EEA5D0600AF" box="[258,302,991,1008]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
, female paratype, USNM01771288, dorsal; (d) same, ventral; (e) 
<taxonomicName id="4C0825CBCB3CD82AFC909EEA5FC500AF" authority="(Dognin)" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[819,1005,991,1008]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mirafloresa">C. mirafloresa (Dognin)</taxonomicName>
, male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) 
<taxonomicName id="4C0825CBCB3CD82AFF339EC65D7E075B" authority="(Dognin)" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[144,342,1011,1028]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mirafloresa">C. mirafloresa (Dognin)</taxonomicName>
, female, USNMENT01771294, dorsal; (h) same, ventral; (i) 
<taxonomicName id="4C0825CBCB3CD82AFCE89EC65808075B" authority="(Walker)" baseAuthorityName="Walker" box="[843,1056,1011,1028]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata (Walker)</taxonomicName>
, male, USNMENT01771293, dorsal; (j) same, ventral; (k) 
<taxonomicName id="4C0825CBCB3CD82AFF2D993D5D760746" authority="(Walker)" baseAuthorityName="Walker" box="[142,350,1032,1049]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata (Walker)</taxonomicName>
, female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm.
</paragraph>
</caption>
<caption id="DF770EC0CB3CD82AFFD29B9B5E2E05B6" ID-DOI="http://doi.org/10.5281/zenodo.10665231" ID-Zenodo-Dep="10665231" httpUri="https://zenodo.org/record/10665231/files/figure.png" pageId="2" pageNumber="3" startId="2.[113,165,1710,1727]" targetBox="[262,1353,1132,1673]" targetPageId="2" targetType="figure">
<paragraph id="8BB75E48CB3CD82AFFD29B9B5E2E05B6" blockId="2.[112,1506,1710,1769]" pageId="2" pageNumber="3">
<emphasis id="B97C825ACB3CD82AFFD29B9B5C9A05E0" bold="true" box="[113,178,1710,1727]" pageId="2" pageNumber="3">Figure 3</emphasis>
<taxonomicName id="4C0825CBCB3CD82AFF169B9B5D3A05E0" authorityName="Matson" authorityYear="2024" box="[181,274,1710,1727]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
male genitalia.(a) 
<taxonomicName id="4C0825CBCB3CD82AFE3D9B9B5E2205E0" authority="Matson, 2024" authorityName="Matson" authorityYear="2024" box="[414,522,1710,1727]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">C.ashleyorum</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3CD82AFDAC9B9B5E1F05E0" box="[527,567,1710,1727]" pageId="2" pageNumber="3" rank="species">n.sp.</taxonomicNameLabel>
,TAM-2023-288(USNM 154251),USNMENT01771290,paratype,USNM,genital capsule;(b) same,phallus;(c) 
<taxonomicName id="4C0825CBCB3CD82AFAD59B9B5C90058B" authority="(Dognin)" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mirafloresa">C.mirafloresa (Dognin)</taxonomicName>
,TAM-2023-279 (USNM154250),USNMENT01771323,genital capsule;(d) same,phallus;(e) 
<taxonomicName id="4C0825CBCB3CD82AFCDF9BF65816058B" authority="(Walker)" baseAuthorityName="Walker" box="[892,1086,1731,1748]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="transvolutata">C.transvolutata (Walker)</taxonomicName>
,TAM-2023-277 (USNM 154248),USNMENT01771293, genital capsule; (f) same, phallus.Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="DF770EC0CB3DD82BFFF0992C58920760" ID-DOI="http://doi.org/10.5281/zenodo.10665233" ID-Zenodo-Dep="10665233" httpUri="https://zenodo.org/record/10665233/files/figure.png" pageId="3" pageNumber="4" startId="3.[83,137,1049,1066]" targetBox="[220,1328,160,1008]" targetPageId="3" targetType="figure">
<paragraph id="8BB75E48CB3DD82BFFF0992C58920760" blockId="3.[83,1474,1049,1087]" pageId="3" pageNumber="4">
<emphasis id="B97C825ACB3DD82BFFF0992C5CB00775" bold="true" box="[83,152,1049,1066]" pageId="3" pageNumber="4">Figure 4</emphasis>
<taxonomicName id="4C0825CBCB3DD82BFF3D992C5CD70775" authorityName="Matson" authorityYear="2024" box="[158,255,1049,1066]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
female genitalia. (a) 
<taxonomicName id="4C0825CBCB3DD82BFE0E992C5E080775" authority="Matson, 2024" authorityName="Matson" authorityYear="2024" box="[429,544,1049,1066]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">C. ashleyorum</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3DD82BFD86992C5E7A0775" box="[549,594,1049,1066]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
, TAM-2023-291 (USNM 154253), USNMENT01771290, paratype; (b) 
<taxonomicName id="4C0825CBCB3DD82BFB22992C59160775" authority="(Dognin)" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[1153,1342,1049,1066]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mirafloresa">C. mirafloresa (Dognin)</taxonomicName>
, TAM-2023-290 (USNM 154252), USNMENT01771322; (c) 
<taxonomicName id="4C0825CBCB3DD82BFE00991B5E470760" authority="(Walker)" baseAuthorityName="Walker" box="[419,623,1070,1087]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata (Walker)</taxonomicName>
, TAM-2023-278 (USNM 154249), USNMENT01771320. Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="DF770EC0CB3DD82BFFF09AAE5EAD049E" ID-DOI="http://doi.org/10.5281/zenodo.10665235" ID-Zenodo-Dep="10665235" httpUri="https://zenodo.org/record/10665235/files/figure.png" pageId="3" pageNumber="4" startId="3.[83,137,1947,1964]" targetBox="[296,1261,1167,1930]" targetPageId="3" targetType="figure">
<paragraph id="8BB75E48CB3DD82BFFF09AAE5EAD049E" blockId="3.[83,1474,1947,1985]" pageId="3" pageNumber="4">
<emphasis id="B97C825ACB3DD82BFFF09AAE5CB004F3" bold="true" box="[83,152,1947,1964]" pageId="3" pageNumber="4">Figure 5</emphasis>
Geographic distribution of 
<taxonomicName id="4C0825CBCB3DD82BFED59AAE5DF104F3" authorityName="Matson" authorityYear="2024" box="[374,473,1947,1964]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
: 
<taxonomicName id="4C0825CBCB3DD82BFE429AAE5E7A04F3" authority="Matson, 2024" authorityName="Matson" authorityYear="2024" box="[481,594,1947,1964]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ashleyorum" status="sp. nov.">C. ashleyorum</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3DD82BFDF59AAE5EAA04F3" box="[598,642,1947,1964]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
, green circles; 
<taxonomicName id="4C0825CBCB3DD82BFD5A9AAE5F4004F3" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[761,872,1947,1964]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mirafloresa" status="stat. nov.">C. mirafloresa</taxonomicName>
<taxonomicNameLabel id="A24F3F21CB3DD82BFCCF9AAE5F9804F3" box="[876,944,1947,1964]" pageId="3" pageNumber="4">rev. stat.</taxonomicNameLabel>
(Dognin), magenta circles; and 
<taxonomicName id="4C0825CBCB3DD82BFB0C9AAE595F04F3" authority="(Walker)" baseAuthorityName="Walker" box="[1199,1399,1947,1964]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="transvolutata">C. transvolutata (Walker)</taxonomicName>
, turquois circles. Single points may represent&gt;1 individuals. Scale bar = 500 km.
</paragraph>
</caption>
<paragraph id="8BB75E48CB3AD82CFFD2993F5CD00743" blockId="4.[113,248,1033,1053]" box="[113,248,1033,1053]" pageId="4" pageNumber="5">Type material</paragraph>
<paragraph id="8BB75E48CB3AD82CFFD2990B5CE3070E" blockId="4.[112,787,1086,1253]" box="[113,203,1086,1105]" pageId="4" pageNumber="5">
<typeStatus id="54B3E0EACB3AD82CFFD2990B5CE3070E" box="[113,203,1086,1105]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</paragraph>
<paragraph id="8BB75E48CB3AD82CFF3499695DAD07BA" blockId="4.[112,787,1086,1253]" pageId="4" pageNumber="5">
<materialsCitation id="3B605415CB3AD82CFF3499695DAD07BA" collectingDate="2016-09-08" collectingMethod="light trap" collectionCode="USNM" collectorName="H. Cambronero &amp; R. Franco" country="Costa Rica" elevation="295" location="Area Administrativa" municipality="Sector Santa Rosa" pageId="4" pageNumber="5" specimenCode="16-SRNP-105545, USNMENT01771276" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<collectingCountry id="F31F1ED8CB3AD82CFF3499695D2C0730" box="[151,260,1116,1135]" name="Costa Rica" pageId="4" pageNumber="5">COSTA RICA</collectingCountry>
• 
<specimenCount id="9D0E95C1CB3AD82CFEB299695D08072E" box="[273,288,1116,1137]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación Guanacaste], 
<collectingRegion id="49CC90AACB3AD82CFD0099695F270730" box="[675,783,1116,1135]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, 
<collectingMunicipality id="6BD3C432CB3AD82CFFD3994C5D0C07D2" box="[112,292,1145,1165]" pageId="4" pageNumber="5">Sector Santa Rosa</collectingMunicipality>
, 
<location id="8ED70893CB3AD82CFE93994C5DD107D3" LSID="urn:lsid:plazi:treatment:03A1EF5ECB3FD82DFC8298B45DEA0113:8ED70893CB3AD82CFE93994C5DD107D3" box="[304,505,1145,1164]" country="Costa Rica" municipality="Sector Santa Rosa" name="Area Administrativa" pageId="4" pageNumber="5" stateProvince="Guanacaste">Area Administrativa</location>
, (
<geoCoordinate id="EE3C388FCB3AD82CFDA8994C5E4A07D3" box="[523,610,1145,1164]" pageId="4" pageNumber="5">10.8376°</geoCoordinate>
, 
<geoCoordinate id="EE3C388FCB3AD82CFDCD994C5EFA07D3" box="[622,722,1145,1164]" pageId="4" pageNumber="5">-85.6187°</geoCoordinate>
); elev. 
<quantity id="4CF0F3ADCB3AD82CFFD299A25C9007F5" box="[113,184,1175,1194]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="4" pageNumber="5" unit="m" value="295.0">
<elevation id="0025B97BCB3AD82CFFD299A25C9007F5" box="[113,184,1175,1194]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="4" pageNumber="5" unit="m" value="295.0">295 m</elevation>
</quantity>
; 
<date id="FFB67888CB3AD82CFF6799A25D6307F5" box="[196,331,1175,1194]" pageId="4" pageNumber="5" value="2016-09-08">
<collectingDate id="EFF28160CB3AD82CFF6799A25D6307F5" box="[196,331,1175,1194]" pageId="4" pageNumber="5" value="2016-09-08">08 Sep. 2016</collectingDate>
</date>
; 
<collectorName id="26FD3B9ECB3AD82CFEF499A25DD907F5" box="[343,497,1175,1194]" pageId="4" pageNumber="5">H. Cambronero</collectorName>
&amp; 
<collectorName id="26FD3B9ECB3AD82CFDB299A25E4707F5" box="[529,623,1175,1194]" pageId="4" pageNumber="5">R. Franco</collectorName>
leg.; 
<collectingMethod id="5249265FCB3AD82CFD0899A25F2607F5" box="[683,782,1175,1194]" pageId="4" pageNumber="5">light trap</collectingMethod>
; Voucher Code: 
<specimenCode id="DBAEF633CB3AD82CFEAE99815D9F0798" box="[269,439,1204,1223]" pageId="4" pageNumber="5">16-SRNP-105545</specimenCode>
; BOLD Process ID: 
<accessionNumber id="945BC3ABCB3AD82CFDD899815F270798" box="[635,783,1204,1223]" pageId="4" pageNumber="5">BLPAA1558-17</accessionNumber>
; 
<specimenCode id="DBAEF633CB3AD82CFFD299E75D1007BA" box="[113,312,1234,1253]" pageId="4" pageNumber="5">USNMENT01771276</specimenCode>
; 
<collectionCode id="ED19C68DCB3AD82CFEE399E75DA907BA" box="[320,385,1234,1253]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB75E48CB3AD82CFFD298175D01066B" blockId="4.[112,788,1313,1985]" box="[113,297,1313,1335]" pageId="4" pageNumber="5">
<typeStatus id="54B3E0EACB3AD82CFFD298175CF8066A" box="[113,208,1314,1333]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D0E95C1CB3AD82CFF7D98145CD20668" box="[222,250,1313,1335]" count="9" pageId="4" pageNumber="5" type="male">9♂</specimenCount>
, 
<specimenCount id="9D0E95C1CB3AD82CFEA698145D090668" box="[261,289,1313,1335]" count="9" pageId="4" pageNumber="5" type="female">9♀</specimenCount>
)
</paragraph>
<paragraph id="8BB75E48CB3AD82CFF34980A59B20502" blockId="4.[112,788,1313,1985]" lastBlockId="4.[830,1506,153,1985]" pageId="4" pageNumber="5">
<collectingCountry id="F31F1ED8CB3AD82CFF34980A5D30060D" box="[151,280,1343,1362]" name="Costa Rica" pageId="4" pageNumber="5">COSTA RICA</collectingCountry>
• 
<specimenCount id="9D0E95C1CB3AD82CFE90980A5D6C060B" box="[307,324,1343,1364]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFD2A980A5F20060D" box="[649,776,1343,1362]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFFD298685CC0062F" box="[113,232,1373,1392]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pitilla, Estacion Pitilla, (10.9893°, -85.4258°); elev. 
<quantity id="4CF0F3ADCB3AD82CFF04984F5CC506D1" box="[167,237,1402,1422]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="4" pageNumber="5" unit="m" value="675.0">675 m</quantity>
; 
<date id="FFB67888CB3AD82CFF5A984F5DAC06D2" box="[249,388,1402,1422]" pageId="4" pageNumber="5" value="2021-03-12">12 Mar. 2021</date>
; 
<collectionCode id="ED19C68DCB3AD82CFE33984E5D8806D1" box="[400,416,1403,1422]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
. Franco &amp; H. Cambronero leg.; light trap; Voucher Code: 21-SRNP-101527; USNMENT01771274; 
<collectionCode id="ED19C68DCB3AD82CFD6D98AD5F2706F4" box="[718,783,1432,1451]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFFD298835CA90694" box="[113,129,1462,1483]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFE3F98835E380696" box="[412,528,1462,1481]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFDBC98835EB90696" box="[543,657,1462,1481]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Santa Maria, Crater Bosque Sendero Adentro, (10.8035°, -85.3273°); elev. 
<quantity id="4CF0F3ADCB3AD82CFFD298C45C96055B" box="[113,190,1521,1540]" metricMagnitude="3" metricUnit="m" metricValue="1.594" pageId="4" pageNumber="5" unit="m" value="1594.0">1594 m</quantity>
; 
<date id="FFB67888CB3AD82CFF6B98C45D14055B" box="[200,316,1521,1540]" pageId="4" pageNumber="5" value="2017-07-21">21 Jul. 2017</date>
; S. Rios leg.; light trap; Voucher Code: 17-SRNP-104721; 
<collectionCode id="ED19C68DCB3AD82CFF629B3A5CDE057D" box="[193,246,1551,1570]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFEC59B3B5D99057E" box="[358,433,1550,1569]" pageId="4" pageNumber="5">BLPAA</collectionCode>
10532-17; USNMENT01771275; 
<collectionCode id="ED19C68DCB3AD82CFD729B3A5F38057D" box="[721,784,1551,1570]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFFD39B185CA8051D" box="[112,128,1581,1602]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFE3B9B195E240560" box="[408,524,1580,1599]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFDBA9B195EA30560" box="[537,651,1580,1599]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pitilla, Estacion Pitilla, (10.989°, -85.426°); elev. 
<quantity id="4CF0F3ADCB3AD82CFDB09B7F5E700502" box="[531,600,1610,1629]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="4" pageNumber="5" unit="m" value="675.0">675 m</quantity>
; 
<date id="FFB67888CB3AD82CFDC79B7F5ED80502" box="[612,752,1610,1629]" pageId="4" pageNumber="5" value="2006-03-03">03 Mar. 2006</date>
; H. Cambronero &amp; F.Quesada leg.; light trap; Voucher Code: 06-SRNP-102944; 
<collectionCode id="ED19C68DCB3AD82CFFD39BB05C8B05C7" box="[112,163,1669,1688]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFEAE9BB05D7D05C7" box="[269,341,1669,1688]" pageId="4" pageNumber="5">BLPAD</collectionCode>
125-06; GenBank:JN806613; USNMENT01771277; 
<collectionCode id="ED19C68DCB3AD82CFFD39B965C9805E9" box="[112,176,1699,1718]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFF1B9B965CE005E7" box="[184,200,1699,1720]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFE7F9B965E7805E9" box="[476,592,1699,1718]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFDFF9B965EE605E9" box="[604,718,1699,1718]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Del Oro, Serrano, (11°, -85.456°); elev. 
<quantity id="4CF0F3ADCB3AD82CFE789BF55E33058B" box="[475,539,1728,1748]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="4" pageNumber="5" unit="m" value="585.0">585 m</quantity>
; 
<date id="FFB67888CB3AD82CFD809BF45E8B058B" box="[547,675,1729,1748]" pageId="4" pageNumber="5" value="2007-11-08">08 Nov. 2007</date>
; F.Quesada &amp; S. Rios leg.; light trap; Voucher Code:07-SRNP-109827; 
<collectionCode id="ED19C68DCB3AD82CFD339BEB5EED05AE" box="[656,709,1758,1777]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFF369BC95CF50450" box="[149,221,1788,1807]" pageId="4" pageNumber="5">BLPCF</collectionCode>
522-08; GenBank: JQ562061; USNMENT01771278; 
<collectionCode id="ED19C68DCB3AD82CFD6D9BC95F270450" box="[718,783,1788,1807]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFFD29A2F5CA90470" box="[113,129,1818,1839]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFE099A2F5E360472" box="[426,542,1818,1837]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFD8C9A2F5E890472" box="[559,673,1818,1837]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Del Oro, Bosque Aguirre, (11.004°, -85.441°); elev. 
<quantity id="4CF0F3ADCB3AD82CFD929A025E580414" box="[561,624,1847,1867]" metricMagnitude="2" metricUnit="m" metricValue="5.71" pageId="4" pageNumber="5" unit="m" value="571.0">571 m</quantity>
; 
<date id="FFB67888CB3AD82CFDDA9A025ED30415" box="[633,763,1847,1867]" pageId="4" pageNumber="5" value="2009-11-18">18 Nov. 2009</date>
; F. Quesada &amp; S. Rios leg.; light trap; Voucher Code: 09-SRNP-110910; 
<collectionCode id="ED19C68DCB3AD82CFD7E9A605F3A0437" box="[733,786,1877,1896]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFF789A475D0404DA" box="[219,300,1906,1925]" pageId="4" pageNumber="5">BLPDM</collectionCode>
1533-10; GenBank: 
<collectionCode id="ED19C68DCB3AD82CFE799A465DD704D9" box="[474,511,1907,1926]" country="0" name="Hastings Museum" pageId="4" pageNumber="5">HM</collectionCode>
410374; USNMENT01771279; 
<collectionCode id="ED19C68DCB3AD82CFFD39AA55C9804FC" box="[112,176,1936,1955]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFF1B9AA45CE004F9" box="[184,200,1937,1958]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFE7F9AA55E7804FC" box="[476,592,1936,1955]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFDFF9AA55EE604FC" box="[604,718,1936,1955]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Del Oro, Bosque Aguirre, (11.004°, -85.441°); elev. 
<quantity id="4CF0F3ADCB3AD82CFDD59A9B5E91049E" box="[630,697,1966,1985]" metricMagnitude="2" metricUnit="m" metricValue="5.71" pageId="4" pageNumber="5" unit="m" value="571.0">571 m</quantity>
; 
<date id="FFB67888CB3AD82CFD659A9B5F5D03F2" box="[710,885,154,1985]" pageId="4" pageNumber="5" value="2009-11-15">15 Nov. 2009</date>
; H. Cambronero &amp; F. Quesada leg.; light trap; Voucher Code: 09-SRNP-110137; Genitalia: TAM-2023-302 (
<collectionCode id="ED19C68DCB3AD82CFAA39D8259690395" box="[1280,1345,183,202]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
154255); 
<collectionCode id="ED19C68DCB3AD82CFA099D8259C80395" box="[1450,1504,183,202]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFC0E9DE05FD503B7" box="[941,1021,213,232]" pageId="4" pageNumber="5">BLPDM</collectionCode>
760-10; GenBank: 
<collectionCode id="ED19C68DCB3AD82CFB079DE058E203B7" box="[1188,1226,213,232]" country="0" name="Hastings Museum" pageId="4" pageNumber="5">HM</collectionCode>
401743; USNMENT01771280; 
<collectionCode id="ED19C68DCB3AD82CFC9D9DC65F560259" box="[830,894,243,262]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC259DC65FBE0257" box="[902,918,243,264]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB099DC659360259" box="[1194,1310,243,262]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFA889DC659B50259" box="[1323,1437,243,262]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pitilla, Estacion Pitilla, (10.9893°, -85.4258°); elev. 
<quantity id="4CF0F3ADCB3AD82CFAEB9C2559A5027B" box="[1352,1421,272,292]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="4" pageNumber="5" unit="m" value="675.0">675 m</quantity>
; 
<date id="FFB67888CB3AD82CFA399C255F5A021E" pageId="4" pageNumber="5" value="2018-01-16">16 Jan. 2018</date>
; H. Cambronero &amp; 
<collectionCode id="ED19C68DCB3AD82CFB8D9C1B5816021E" box="[1070,1086,302,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
. Franco leg.; light trap; Voucher Code: 18- SRNP-100292; 
<collectionCode id="ED19C68DCB3AD82CFC729C79582F0200" box="[977,1031,332,351]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFBDA9C7958EE0200" box="[1145,1222,332,351]" pageId="4" pageNumber="5">BLPDV</collectionCode>
6328-18; USNMENT01771281; 
<collectionCode id="ED19C68DCB3AD82CFC9D9C5F5F560222" box="[830,894,362,381]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC259C5F5FBE0220" box="[902,918,362,383]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB099C5F59360222" box="[1194,1310,362,381]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFA889C5F59B50222" box="[1323,1437,362,381]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Santa Maria, Crater Bosque Sendero Adentro, (10.8035°, -85.3273°); elev. 
<quantity id="4CF0F3ADCB3AD82CFCD69C905FED02E7" box="[885,965,421,440]" metricMagnitude="3" metricUnit="m" metricValue="1.594" pageId="4" pageNumber="5" unit="m" value="1594.0">1594 m</quantity>
; 
<date id="FFB67888CB3AD82CFC719C90587002E7" box="[978,1112,421,440]" pageId="4" pageNumber="5" value="2018-05-17">17 May 2018</date>
; S. Rios &amp; H. Ramirrez leg.; light trap; Voucher Code: 18-SRNP-103045; Genitalia: TAM-2023-301 (
<collectionCode id="ED19C68DCB3AD82CFA3C9CF659C80289" box="[1439,1504,451,470]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
154254); 
<collectionCode id="ED19C68DCB3AD82CFC3A9CD55FE602AC" box="[921,974,480,499]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFB9D9CD558A002AC" box="[1086,1160,480,499]" pageId="4" pageNumber="5">BLPDV</collectionCode>
9746-18; USNMENT01771282; 
<collectionCode id="ED19C68DCB3AD82CFA3C9CD559F602AC" box="[1439,1502,480,499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC9D9CCB5F74014C" box="[830,860,510,531]" count="2" pageId="4" pageNumber="5" type="male">2♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFBCF9CCB58F6014E" box="[1132,1246,510,529]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFB489CCB5973014E" box="[1259,1371,510,529]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Cacao, Toma de Agua, (10.9296°, -85.4651°); elev. 
<quantity id="4CF0F3ADCB3AD82CFB499F29591D0170" box="[1258,1333,540,559]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="5" unit="m" value="1160.0">1160 m</quantity>
; 
<date id="FFB67888CB3AD82CFAE29F2959EC0170" box="[1345,1476,540,559]" pageId="4" pageNumber="5" value="2010-08-10">10 Aug. 2010</date>
; S. Rios &amp; F.Quesada leg.; light trap; Voucher Codes: 10-SRNP-112668, 10- SRNP-112669; 
<collectionCode id="ED19C68DCB3AD82CFC6B9F625FD40135" box="[968,1020,599,618]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFBCB9F62589E0135" box="[1128,1206,599,618]" pageId="4" pageNumber="5">BLPDW</collectionCode>
776-11, 
<collectionCode id="ED19C68DCB3AD82CFB579F62596D0135" box="[1268,1349,599,618]" pageId="4" pageNumber="5">BLPDW</collectionCode>
777-11; GenBank: JN267525, JN267526; USNMENT01771283, USNMENT01771284; 
<collectionCode id="ED19C68DCB3AD82CFA3E9F4059F401D7" box="[1437,1500,629,648]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC9D9FA65F6601F7" box="[830,846,659,680]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFBC59FA658F201F9" box="[1126,1242,659,678]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFB4B9FA6597201F9" box="[1256,1370,659,678]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pailas, Manta Rio Blanco, (10.7746°, -85.35°); elev. 
<quantity id="4CF0F3ADCB3AD82CFB579F85591F019B" box="[1268,1335,688,708]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="5" unit="m" value="790.0">790 m</quantity>
; 
<date id="FFB67888CB3AD82CFAE09F8559EC019C" box="[1347,1476,688,707]" pageId="4" pageNumber="5" value="2010-10-05">05 Oct. 2010</date>
; S. Rios &amp; 
<collectionCode id="ED19C68DCB3AD82CFC239FFB5FB801BE" box="[896,912,718,737]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
. Franco leg.; light trap; Voucher Code: 10-SRNP-114210; 
<collectionCode id="ED19C68DCB3AD82CFA099FFB59C801BE" box="[1450,1504,718,737]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFC0F9FD95FD201A0" box="[940,1018,748,767]" pageId="4" pageNumber="5">BLPDX</collectionCode>
1083-11; GenBank:JN267567; USNMENT01771285; 
<collectionCode id="ED19C68DCB3AD82CFC9D9E3F5F560042" box="[830,894,778,797]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC259E3F5FBE0040" box="[902,918,778,799]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB099E3F59360042" box="[1194,1310,778,797]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFA889E3F59B50042" box="[1323,1437,778,797]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pailas, Palmeras, (10.8107°, -85.347°); elev. 
<quantity id="4CF0F3ADCB3AD82CFB489E1259110064" box="[1259,1337,807,827]" metricMagnitude="3" metricUnit="m" metricValue="1.368" pageId="4" pageNumber="5" unit="m" value="1368.0">1368 m</quantity>
; 
<date id="FFB67888CB3AD82CFAE69E1259EC0065" box="[1349,1476,807,826]" pageId="4" pageNumber="5" value="2010-10-07">07 Oct. 2010</date>
; S. Rios &amp; 
<collectionCode id="ED19C68DCB3AD82CFC229E705FB80007" box="[897,912,837,856]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
. Franco leg.; light trap; Voucher Code: 10-SRNP-114609; 
<collectionCode id="ED19C68DCB3AD82CFA0F9E7059CA0007" box="[1452,1506,837,856]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFC129E575FD5002A" box="[945,1021,866,885]" pageId="4" pageNumber="5">BLPDY</collectionCode>
057-11; GenBank: JN267652; USNMENT01771286; 
<collectionCode id="ED19C68DCB3AD82CFCE39EB55FA800CC" box="[832,896,896,915]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC259EB45FBE00C9" box="[902,918,897,918]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB099EB5593600CC" box="[1194,1310,896,915]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFA889EB559B500CC" box="[1323,1437,896,915]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pailas, Manta Rio Blanco, (10.7746°, -85.35°); elev. 
<quantity id="4CF0F3ADCB3AD82CFAB89EAB597300EE" box="[1307,1371,926,945]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="5" unit="m" value="790.0">790 m</quantity>
; 
<date id="FFB67888CB3AD82CFAC09EAB59F400EE" box="[1379,1500,926,945]" pageId="4" pageNumber="5" value="2010-10-08">08 Oct. 2010</date>
; S. Rios &amp; 
<collectionCode id="ED19C68DCB3AD82CFC339E895FB70090" box="[912,927,956,975]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
.Franco leg.; light trap; Voucher Code:10-SRNP-114696; 
<collectionCode id="ED19C68DCB3AD82CFA089E8959C80090" box="[1451,1504,956,975]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFC129EEF5FD600B2" box="[945,1022,986,1005]" pageId="4" pageNumber="5">BLPDY</collectionCode>
144-11; GenBank: JN267659; USNMENT01771287; 
<collectionCode id="ED19C68DCB3AD82CFCE29EC25FAC0755" box="[833,900,1015,1034]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC2D9ECD5FB60752" box="[910,926,1016,1037]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB7B9EC2597A0755" box="[1240,1362,1015,1034]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFAC79EC259F40755" box="[1380,1500,1015,1034]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, Sector Pailas Dos, PDL#5, (10.7627°, -85.334°); elev. 
<quantity id="4CF0F3ADCB3AD82CFAEB992059A30777" box="[1352,1419,1045,1064]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="4" pageNumber="5" unit="m" value="825.0">825 m</quantity>
; 
<date id="FFB67888CB3AD82CFA3499205F5B0719" pageId="4" pageNumber="5" value="2013-11-01">01 Nov. 2013</date>
; S. Rios &amp; H. Cambronero leg.; light trap; Voucher Code: 13- SRNP-103007; 
<collectionCode id="ED19C68DCB3AD82CFC7199655820073C" box="[978,1032,1104,1123]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFBDD996558EE073C" box="[1150,1222,1104,1123]" pageId="4" pageNumber="5">BLPEE</collectionCode>
3571-14; USNMENT01771288; 
<collectionCode id="ED19C68DCB3AD82CFC9C995B5FA807DE" box="[831,896,1134,1153]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC28995B5FB307DC" box="[907,923,1134,1155]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFB61995B591E07DE" box="[1218,1334,1134,1153]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFAE4995B59BF07DE" box="[1351,1431,1134,1153]" country="Costa Rica" name="Alajuela" pageId="4" pageNumber="5">Alajuela</collectingRegion>
, Sector San Cristobal, Sendero Vivero, (10.867°, -85.387°); elev. 
<quantity id="4CF0F3ADCB3AD82CFAF899B959B307C0" box="[1371,1435,1164,1183]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="4" pageNumber="5" unit="m" value="730.0">730 m</quantity>
; Gloria Sihezar leg.; prepupa collected; prepupa: 
<date id="FFB67888CB3AD82CFB56999F595707E2" box="[1269,1407,1194,1213]" pageId="4" pageNumber="5" value="1998-11-17">17 Nov. 1998</date>
; eclosed: 
<date id="FFB67888CB3AD82CFC9C99F25F970785" box="[831,959,1223,1243]" pageId="4" pageNumber="5" value="1998-12-11">11 Dec. 1998</date>
; food plant: 
<taxonomicName id="4C0825CBCB3AD82CFB9899F258BF0785" box="[1083,1175,1223,1242]" class="Magnoliopsida" family="Annonaceae" genus="Guatteria" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">Guatteria</taxonomicName>
verrucosa 
<collectionCode id="ED19C68DCB3AD82CFAA199FD593A0784" box="[1282,1298,1224,1243]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
.E.Fr.; food plant det.: Gloria Sihezar; Voucher Code: 98-SRNP-15091; 
<collectionCode id="ED19C68DCB3AD82CFA8D99D0594E07A7" box="[1326,1382,1253,1272]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFC9C98375FB2064A" box="[831,922,1282,1301]" pageId="4" pageNumber="5">MHAGA</collectionCode>
850-06; GenBank: GU154099; USNMENT01771289; 
<collectionCode id="ED19C68DCB3AD82CFA3F983659F50649" box="[1436,1501,1283,1302]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC9C98145F660669" box="[831,846,1313,1334]" count="1" pageId="4" pageNumber="5" type="male">♂</specimenCount>
; ACG [Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFBF5981558ED066C" box="[1110,1221,1312,1331]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
], 
<collectingRegion id="49CC90AACB3AD82CFB7298155936066C" box="[1233,1310,1312,1331]" country="Costa Rica" name="Alajuela" pageId="4" pageNumber="5">Alajuela</collectingRegion>
, Sector San Cristobal, Rio Blanco Abajo, (10.9°, -85.373°); elev. 
<quantity id="4CF0F3ADCB3AD82CFB77980B5930060E" box="[1236,1304,1342,1361]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="500.0">500 m</quantity>
; 
<taxonomicName id="4C0825CBCB3AD82CFA87980B5986060E" authority="Cano" authorityName="Cano" box="[1316,1454,1342,1361]" class="Insecta" family="Scarabaeidae" genus="Carolina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Carolina Cano</taxonomicName>
leg.; found as pupa; pupa: 
<date id="FFB67888CB3AD82CFBAF986958AA0630" box="[1036,1154,1372,1391]" pageId="4" pageNumber="5" value="2005-01-18">18 Jan.2005</date>
; eclosed: 
<date id="FFB67888CB3AD82CFB7E986959710630" box="[1245,1369,1372,1391]" pageId="4" pageNumber="5" value="2005-02-03">03 Feb. 2005</date>
; 
<date id="FFB67888CB3AD82CFAC2986959F40630" box="[1377,1500,1372,1391]" pageId="4" pageNumber="5" value="2005-02-03">03 Feb.2005</date>
; presumed food plant: 
<taxonomicName id="4C0825CBCB3AD82CFBBD984C585206D3" box="[1054,1146,1401,1420]" class="Magnoliopsida" family="Annonaceae" genus="Guatteria" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">Guatteria</taxonomicName>
verrucosa 
<collectionCode id="ED19C68DCB3AD82CFB4B984F58D006D2" box="[1256,1272,1402,1421]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
.E.Fr.; host det.: Yesenia Mendoza; Voucher Code:05-SRNP-155; Genitalia:TAM-2023-288 (
<collectionCode id="ED19C68DCB3AD82CFA0098A259C806F5" box="[1443,1504,1431,1450]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
154251); 
<collectionCode id="ED19C68DCB3AD82CFC3C98805FFE0697" box="[927,982,1461,1480]" pageId="4" pageNumber="5">BOLD</collectionCode>
Process ID: 
<collectionCode id="ED19C68DCB3AD82CFBF4988058910697" box="[1111,1209,1461,1480]" pageId="4" pageNumber="5">MHMXN</collectionCode>
675-07; GenBank: GU160999; USNMENT01771290; 
<collectionCode id="ED19C68DCB3AD82CFBAD98E6586606B9" box="[1038,1102,1491,1510]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFBF798E6584C06B7" box="[1108,1124,1491,1512]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; Juan Vinas; May; Schaus &amp; Barnes leg.; Genitalia slide: TAM-2023-291 (
<collectionCode id="ED19C68DCB3AD82CFBD798C5589C055C" box="[1140,1204,1520,1539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
154253); USNMENT01771291; 
<collectionCode id="ED19C68DCB3AD82CFC9C9B3B5FA8057E" box="[831,896,1550,1569]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC289B3B5FB3057C" box="[907,923,1550,1571]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; Sitio; May; WmSchaus leg.; USNMENT01771292; 
<collectionCode id="ED19C68DCB3AD82CFA3E9B3B59F6057E" box="[1437,1502,1550,1569]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D0E95C1CB3AD82CFC9C9B195F67051E" box="[831,847,1580,1601]" count="1" pageId="4" pageNumber="5" type="female">♀</specimenCount>
; 
<collectingRegion id="49CC90AACB3AD82CFCFF9B195FCA0560" box="[860,994,1580,1599]" country="Costa Rica" name="Alajuela" pageId="4" pageNumber="5">Alajuela Prov.</collectingRegion>
, 
<quantity id="4CF0F3ADCB3AD82CFC4A9B19583E0560" box="[1001,1046,1580,1599]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="km" value="8.0">8km</quantity>
N. Vara Blanca; elev. 
<quantity id="4CF0F3ADCB3AD82CFB419B1959070560" box="[1250,1327,1580,1599]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="m" value="1400.0">1400 m</quantity>
; 
<date id="FFB67888CB3AD82CFA9B9B1959920560" box="[1336,1466,1580,1599]" pageId="4" pageNumber="5" value="1992-03-25">25 Mar. 1992</date>
; 
<collectionCode id="ED19C68DCB3AD82CFA609B1959C80560" box="[1475,1504,1580,1599]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="4" pageNumber="5">UV</collectionCode>
&amp; 
<collectionCode id="ED19C68DCB3AD82CFCF09B7F5F5D0502" box="[851,885,1610,1629]" country="0" name="University of Montana Museum" pageId="4" pageNumber="5">MV</collectionCode>
lights; J. McCarty &amp; J. Powell leg.; 
<collectionCode id="ED19C68DCB3AD82CFB619B7C592B0503" box="[1218,1283,1609,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="5" type="University or college">EMEC</collectionCode>
1748466; 
<collectionCode id="ED19C68DCB3AD82CFAF89B7F59BE0502" box="[1371,1430,1610,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="5" type="University or college">EMEC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3120DC3CB3AD82CFCC69B52594705C7" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BB75E48CB3AD82CFCC69B52594705C7" blockId="4.[830,1506,153,1985]" pageId="4" pageNumber="5">
<emphasis id="B97C825ACB3AD82CFCC69B525FC20525" bold="true" box="[869,1002,1639,1658]" pageId="4" pageNumber="5">Distribution.</emphasis>
<taxonomicName id="4C0825CBCB3AD82CFC519B5258CC0525" authorityName="Matson" authorityYear="2024" box="[1010,1252,1639,1658]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ashleyorum">Cyphoedma ashleyorum</taxonomicName>
is known from the midelevation montane rain forests of 
<collectingCountry id="F31F1ED8CB3AD82CFB2B9BB058C405C7" box="[1160,1260,1669,1688]" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
and 
<collectingCountry id="F31F1ED8CB3AD82CFAB89BB0594305C7" box="[1307,1387,1669,1688]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3120DC3CB3AD82CFCC69B9758E604D9" pageId="4" pageNumber="5" type="biology_ecology">
<paragraph id="8BB75E48CB3AD82CFCC69B9758E604D9" blockId="4.[830,1506,153,1985]" pageId="4" pageNumber="5">
<emphasis id="B97C825ACB3AD82CFCC69B975F9F05EA" bold="true" box="[869,951,1698,1717]" pageId="4" pageNumber="5">Biology.</emphasis>
<taxonomicName id="4C0825CBCB3AD82CFC1F9B97588305EA" authorityName="Matson" authorityYear="2024" box="[956,1195,1698,1717]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ashleyorum">Cyphoedma ashleyorum</taxonomicName>
flies throughout the year. It has been reared from the Costa Rican endemic 
<taxonomicName id="4C0825CBCB3AD82CFB7A9BF5591D058C" box="[1241,1333,1728,1747]" class="Magnoliopsida" family="Annonaceae" genus="Guatteria" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">Guatteria</taxonomicName>
verrucosa 
<collectionCode id="ED19C68DCB3AD82CFA3D9BF45986058B" box="[1438,1454,1729,1748]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
. E. Fr. (
<taxonomicName id="4C0825CBCB3AD82CFCE79BEB5F9F05AE" box="[836,951,1758,1777]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
) (
<bibRefCitation id="EF9923B9CB3AD82CFC699BEB587405AE" author="Maas, P. &amp; Westra, L. &amp; Guerrero, S. &amp; Lobao, A. &amp; Scharf, U. &amp; Zamora, N. A. &amp; Erkens, R." box="[970,1116,1758,1777]" pageId="4" pageNumber="5" pagination="1 - 219" refId="ref6535" refString="Maas, P., Westra, L., Guerrero, S., Lobao, A., Scharf, U., Zamora, N. A., Erkens, R., 2015. Confronting a morphological nightmare: revision of the Neotropical genus Guatteria (Annonaceae). Blumea 60 (1 - 2), 1 - 219. https: // doi. org / 10.3767 / 000651915 X 690341." type="journal article" year="2015">Maas et al., 2015</bibRefCitation>
). This food plant record was first published in the doctoral dissertation of Gunnar Brehm (
<bibRefCitation id="EF9923B9CB3AD82CFAAB9BC959AE0450" author="Brehm, G." box="[1288,1414,1788,1807]" pageId="4" pageNumber="5" refId="ref5918" refString="Brehm, G., 2002. Diversity of Geometrid Moths in a Montane Rainforest in Ecuador. Doctoral dissertation, University of Bayreuth." type="book" year="2002">Brehm, 2002</bibRefCitation>
), but the source of this record (Voucher: 98-SRNP-15091/USNMENT01771289) belongs to the caterpillar rearing efforts of Dan Janzen, Winnie Hallwachs, and parataxonomists in the Area de Conservación 
<collectingRegion id="49CC90AACB3AD82CFA8D9A6059880437" box="[1326,1440,1877,1896]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="5">Guanacaste</collectingRegion>
, 
<collectingCountry id="F31F1ED8CB3AD82CFA099A605F4204DA" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
(
<bibRefCitation id="EF9923B9CB3AD82CFCD59A4658A004D9" author="Janzen, D. H. &amp; Hallwachs, W." box="[886,1160,1906,1926]" pageId="4" pageNumber="5" refId="ref6299" refString="Janzen, D. H., Hallwachs, W., 2009. Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available in: http: // janzen. sas. upenn. edu / (accessed 13 June 2023)." type="url" year="2009">Janzen and Hallwachs, 2009</bibRefCitation>
, 
<bibRefCitation id="EF9923B9CB3AD82CFB319A4758E904DA" author="Janzen, D. H. &amp; Hallwachs, W." box="[1170,1217,1906,1925]" pageId="4" pageNumber="5" pagination="641 - 660" refId="ref6369" refString="Janzen, D. H., Hallwachs, W., 2016. DNA barcoding the Lepidoptera inventory of a large complex tropical conserved wildland, Area de Conservacion Guanacaste, northwestern Costa Rica. Genome 59 (9), 641 - 660. https: // doi. org / 10.1139 / gen- 2016 - 0005." type="journal article" year="2016">2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3120DC3CB3AD82DFCC69AA55EDD0201" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BB75E48CB3AD82DFCC69AA55D6B03B7" blockId="4.[830,1506,153,1985]" lastBlockId="5.[82,758,153,588]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B97C825ACB3AD82CFCC69AA55FFB04FC" bold="true" box="[869,979,1936,1955]" pageId="4" pageNumber="5">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="4C0825CBCB3AD82CFB2E9AA558D404FC" authorityName="Matson" authorityYear="2024" box="[1165,1276,1936,1955]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ashleyorum">ashleyorum</taxonomicName>
was chosen to recognize the invaluable contributions of Richard and Rita Ashley, who have generously supported the 
<collectingRegion id="49CC90AACB3BD82DFEF49DAF5DEF03F2" box="[343,455,154,173]" country="Costa Rica" name="Guanacaste" pageId="5" pageNumber="6">Guanacaste</collectingRegion>
Dry Forest Conservation Fund and made significant contributions to the conservation and study of Costa Rican biodiversity.
</paragraph>
<paragraph id="8BB75E48CB3BD82DFFDA9DC65EDD0201" blockId="5.[82,758,153,588]" pageId="5" pageNumber="6">
<emphasis id="B97C825ACB3BD82DFFDA9DC65DAF025A" bold="true" box="[121,391,242,262]" pageId="5" pageNumber="6">Molecular characterization.</emphasis>
<taxonomicName id="4C0825CBCB3BD82DFE299DC75E45025A" authorityName="Matson" authorityYear="2024" box="[394,621,242,261]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ashleyorum">Cyphoedma ashleyorum</taxonomicName>
is represented in 
<collectionCode id="ED19C68DCB3BD82DFFCB9C255CB5027C" box="[104,157,272,291]" pageId="5" pageNumber="6">BOLD</collectionCode>
by the 
<collectionCode id="ED19C68DCB3BD82DFF7C9C245D2E027B" box="[223,262,273,292]" pageId="5" pageNumber="6">BIN</collectionCode>
: 
<collectionCode id="ED19C68DCB3BD82DFEAF9C255D6E027C" box="[268,326,272,291]" pageId="5" pageNumber="6">BOLD</collectionCode>
:AAA0681 (n = 17, 
<collectingCountry id="F31F1ED8CB3BD82DFE4A9C255E63027C" box="[489,587,272,291]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
).The uncorrected pairwise distance to the nearest neighbor, 
<taxonomicName id="4C0825CBCB3BD82DFDA69C1B5EDB021E" authorityName="Matson" authorityYear="2024" baseAuthorityName="Dognin" baseAuthorityYear="1892" box="[517,755,302,321]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mirafloresa">Cyphoedma mirafloresa</taxonomicName>
(
<collectionCode id="ED19C68DCB3BD82DFFFA9C7E5CBA0201" box="[89,146,331,350]" pageId="5" pageNumber="6">BOLD</collectionCode>
: 
<collectionCode id="ED19C68DCB3BD82DFF379C7E5CE80201" box="[148,192,331,350]" pageId="5" pageNumber="6">AAB</collectionCode>
9560,n = 16, 
<collectingCountry id="F31F1ED8CB3BD82DFE8E9C7E5DAE0201" box="[301,390,331,350]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
, 
<collectingCountry id="F31F1ED8CB3BD82DFE2F9C7E5DFF0201" box="[396,471,331,350]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
, 
<collectingCountry id="F31F1ED8CB3BD82DFE7D9C795E210200" box="[478,521,332,351]" name="Peru" pageId="5" pageNumber="6">Peru</collectingCountry>
, 
<collectingCountry id="F31F1ED8CB3BD82DFDB39C7E5E590201" box="[528,625,331,350]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
),is about 5.1%.
</paragraph>
</subSubSection>
<subSubSection id="C3120DC3CB3BD82DFFDA9C5C5DEA0113" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BB75E48CB3BD82DFFDA9C5C5DEA0113" blockId="5.[82,758,153,588]" pageId="5" pageNumber="6">
<emphasis id="B97C825ACB3BD82DFFDA9C5C5CF10223" bold="true" box="[121,217,361,380]" pageId="5" pageNumber="6">Remarks.</emphasis>
A photograph was not taken of the caterpillar in the only known rearing of this species. However, collector Gloria Sihezar, wrote the following brief description of the caterpillar, “green with fine white dots, with three brown and white pathes on body in the form of rings, green head.” The reared male moth (Voucher: 98-SRNP-15091/USNMENT01771289) was barcoded and is in the 
<typeStatus id="54B3E0EACB3BD82DFDC19CCB5EA5014E" box="[610,653,510,529]" pageId="5" pageNumber="6">type</typeStatus>
series. It is smaller than the rest of the 
<typeStatus id="54B3E0EACB3BD82DFEC19F295DA60170" box="[354,398,540,559]" pageId="5" pageNumber="6">type</typeStatus>
series, as usual for inventory-reared specimens owing to suboptimal food.
</paragraph>
</subSubSection>
</treatment>
</document>