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<document id="2D089E72B1BD0F126999E0522DD8D7C0" ID-DOI="10.5281/zenodo.13154240" ID-ISSN="0253-116X" ID-Zenodo-Dep="13154240" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722537210208" checkinUser="felipe" docAuthor="Riedel, Matthias &amp; Japoshvili, George" docDate="2021" docId="039CD078FFFBFFF5E5A2FE202C834B63" docLanguage="en" docName="LinzerbiolBeitr.2021.53.1.241-288.pdf" docOrigin="Linzer biologische Beiträge 53 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Diadegma (Neoangitia) caucasicum Riedel &amp; Japoshvili, 2021, nov.sp." docType="treatment" docVersion="1" lastPageNumber="278" masterDocId="FFA5A800FFDFFFD0E510FFEB2E374971" masterDocTitle="New records of Ichneumonidae from Northwestern Sakartvelo (Georgia), with additions to the fauna of Lagodekhi reserve" masterLastPageNumber="288" masterPageNumber="241" pageNumber="277" updateTime="1722539510771" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5DEF7DDE418949C865B808B0C1F1939F">New records of Ichneumonidae from Northwestern Sakartvelo (Georgia), with additions to the fauna of Lagodekhi reserve</mods:title>
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<mods:namePart id="9F2A8AE73281121B6E4DF11CBB6ACCA4">Riedel, Matthias</mods:namePart>
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<mods:namePart id="CC03D2D9B1C85D52ACA2D52CB761A944">Japoshvili, George</mods:namePart>
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<mods:title id="8F58742AB99A2B79DF19463A9C433ADE">Linzer biologische Beiträge</mods:title>
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<mods:date id="AB1EC7BA244C6D60D64F1030AAEB5237">2021</mods:date>
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<mods:identifier id="443A214AE7EB858F9F8370B24C52247F" type="ISSN">0253-116X</mods:identifier>
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<treatment id="039CD078FFFBFFF5E5A2FE202C834B63" LSID="urn:lsid:plazi:treatment:039CD078FFFBFFF5E5A2FE202C834B63" httpUri="http://treatment.plazi.org/id/039CD078FFFBFFF5E5A2FE202C834B63" lastPageId="37" lastPageNumber="278" pageId="36" pageNumber="277">
<subSubSection id="C32F32E5FFFBFFF4E5A2FE202C8A4B6E" pageId="36" pageNumber="277" type="nomenclature">
<paragraph id="8B8A616EFFFBFFF4E5A2FE202CD94890" blockId="36.[178,750,459,481]" box="[178,750,459,481]" pageId="36" pageNumber="277">
<heading id="D0C2D602FFFBFFF4E5A2FE202CD94890" box="[178,750,459,481]" fontSize="8" level="2" pageId="36" pageNumber="277" reason="2">
<emphasis id="B941BD7CFFFBFFF4E5A2FE202CD94890" bold="true" box="[178,750,459,481]" pageId="36" pageNumber="277">
<taxonomicName id="4C351AEDFFFBFFF4E5A2FE202C1D4891" authorityName="Riedel &amp; Japoshvili" authorityYear="2021" box="[178,554,459,481]" class="Insecta" family="Ichneumonidae" genus="Diadegma" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="277" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov." subGenus="Neoangitia">
<emphasis id="B941BD7CFFFBFFF4E5A2FE202F2A4890" bold="true" box="[178,285,459,481]" italics="true" pageId="36" pageNumber="277">Diadegma</emphasis>
(
<emphasis id="B941BD7CFFFBFFF4E43BFE202FA94891" bold="true" box="[299,414,459,480]" italics="true" pageId="36" pageNumber="277">Neoangitia</emphasis>
)
<emphasis id="B941BD7CFFFBFFF4E4BEFE202C1D4891" bold="true" box="[430,554,459,480]" italics="true" pageId="36" pageNumber="277">caucasicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2720007FFFBFFF4E720FE202C4B4891" box="[560,636,459,480]" pageId="36" pageNumber="277" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="130E7DEBFFFBFFF4E799FE202CD14890" box="[649,742,459,481]" captionStart-0="Figs 7-8" captionStart-1="Figs 9-13" captionStartId-0="45.[178,218,1514,1536]" captionStartId-1="46.[178,218,1339,1361]" captionTargetBox-0="[181,999,898,1503]" captionTargetBox-1="[179,1086,204,1306]" captionTargetId-0="figure-41@45.[181,1011,898,1503]" captionTargetId-1="figure-39@46.[179,1086,204,1328]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="Figs 7-8: Diadegma caucasicum nov.sp.: (7) apical tergites from lateral, (8) hind tibia and metatarsus from lateral." captionText-1="Figs 9-13: Diadegma caucasicum nov.sp.: (9) face from frontal, (10) head from dorsal, (11) mesopleuron from lateral, (12) propodeum from dorsal, (13) apical tergites from dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.13154248" figureDoi-1="http://doi.org/10.5281/zenodo.13154250" httpUri-0="https://zenodo.org/record/13154248/files/figure.png" httpUri-1="https://zenodo.org/record/13154250/files/figure.png" pageId="36" pageNumber="277">figs 7-13</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A616EFFFBFFF4E5A2FE192C8A4B6E" blockId="36.[178,1085,497,623]" pageId="36" pageNumber="277">
H o l o t y p u s: (
<specimenCount id="9D33AAE7FFFBFFF4E468FE1A2FB14B77" box="[376,390,497,518]" count="1" pageId="36" pageNumber="277" type="female">♀</specimenCount>
) &quot;
<collectingCountry id="F32221FEFFFBFFF4E48DFE1A2FDF4B76" box="[413,488,497,519]" name="Georgia" pageId="36" pageNumber="277">Georgia</collectingCountry>
, Tsageri, Dogurashi,
<quantity id="4CCDCC8BFFFBFFF4E7BDFE1A2CDE4B76" box="[685,745,497,519]" metricMagnitude="2" metricUnit="m" metricValue="9.97" pageId="36" pageNumber="277" unit="m" value="997.0">997 m</quantity>
alt.,
<geoCoordinate id="EE0107A9FFFBFFF4E60BFE1A2DA54B76" box="[795,914,497,519]" degrees="42.670582" direction="north" orientation="latitude" pageId="36" pageNumber="277" precision="1" value="42.67058">42.670582°N</geoCoordinate>
,
<geoCoordinate id="EE0107A9FFFBFFF4E68EFE1A2A234B77" box="[926,1044,497,518]" degrees="42.783149" direction="east" orientation="longitude" pageId="36" pageNumber="277" precision="1" value="42.78315">42.783149°E</geoCoordinate>
,
<date id="FF8B47AEFFFBFFF4E10FFE1A2F7B4B6E" pageId="36" pageNumber="277" value="2020-07-19" valueMax="2020-07-25" valueMin="2020-07-19">19- 25.VII.2020</date>
, Malaise trap, leg. G. Japoshvili&quot; (
<collectingRegion id="49F1AF8CFFFBFFF4E766FDE22C844B6E" box="[630,691,521,543]" country="Georgia" name="Tbilisi" pageId="36" pageNumber="277">Tbilisi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32F32E5FFFBFFF5E5A2FDC12C834B63" lastPageId="37" lastPageNumber="278" pageId="36" pageNumber="277" type="description">
<paragraph id="8B8A616EFFFBFFF4E5A2FDC12A344B1E" blockId="36.[178,1085,497,623]" pageId="36" pageNumber="277">
<materialsCitation id="3B5D6B33FFFBFFF4E5A2FDC12C9A4B27" collectingDate="2020-07-19" collectingDateMax="2020-07-25" collectingDateMin="2020-07-19" collectorName="G. Japoshvili" country="Georgia" elevation="997" latitude="42.67058" location="Tsageri" longLatPrecision="1" longitude="42.78315" municipality="Dogurashi" pageId="36" pageNumber="277" specimenCount="2" specimenCount-female="2">
P a r a t y p e s: (
<specimenCount id="9D33AAE7FFFBFFF4E461FDC22F4A4B4F" box="[369,381,553,574]" count="1" pageId="36" pageNumber="277" type="female">♀</specimenCount>
) &quot;
<collectingCountry id="F32221FEFFFBFFF4E488FDC22FE84B4E" box="[408,479,553,575]" name="Georgia" pageId="36" pageNumber="277">Georgia</collectingCountry>
,
<location id="8EEA37B5FFFBFFF4E4E0FDC22C184B4E" LSID="urn:lsid:plazi:treatment:039CD078FFFBFFF5E5A2FE202C834B63:8EEA37B5FFFBFFF4E4E0FDC22C184B4E" box="[496,559,553,575]" country="Georgia" latitude="42.67058" longLatPrecision="1" longitude="42.78315" municipality="Dogurashi" name="Tsageri" pageId="36" pageNumber="277">Tsageri</location>
,
<collectingMunicipality id="6BEEFB14FFFBFFF4E750FDC22CAE4B4E" box="[576,665,553,575]" pageId="36" pageNumber="277">Dogurashi</collectingMunicipality>
,
<quantity id="4CCDCC8BFFFBFFF4E7BAFDC22CD54B4E" box="[682,738,553,575]" metricMagnitude="2" metricUnit="m" metricValue="9.97" pageId="36" pageNumber="277" unit="m" value="997.0">
<elevation id="0018865DFFFBFFF4E7BAFDC22CD54B4E" box="[682,738,553,575]" metricMagnitude="2" metricUnit="m" metricValue="9.97" pageId="36" pageNumber="277" unit="m" value="997.0">997 m</elevation>
</quantity>
alt.,
<geoCoordinate id="EE0107A9FFFBFFF4E609FDC22DA64B4E" box="[793,913,553,575]" degrees="42.670582" direction="north" orientation="latitude" pageId="36" pageNumber="277" precision="1" value="42.67058">42.670582°N</geoCoordinate>
,
<geoCoordinate id="EE0107A9FFFBFFF4E68EFDC22A244B4F" box="[926,1043,553,574]" degrees="42.783149" direction="east" orientation="longitude" pageId="36" pageNumber="277" precision="1" value="42.78315">42.783149°E</geoCoordinate>
,
<date id="FF8B47AEFFFBFFF4E10FFDC22F7B4B26" pageId="36" pageNumber="277" value="2020-07-19" valueMax="2020-07-25" valueMin="2020-07-19">
<collectingDate id="EFCFBE46FFFBFFF4E10FFDC22F7B4B26" pageId="36" pageNumber="277" value="2020-07-19" valueMax="2020-07-25" valueMin="2020-07-19">19- 25.VII.2020</collectingDate>
</date>
,
<collectingMethod id="52741979FFFBFFF4E448FDAA2FFD4B26" box="[344,458,577,599]" pageId="36" pageNumber="277">Malaise trap</collectingMethod>
, leg.
<collectorName id="26C004B8FFFBFFF4E4EFFDAA2C4B4B26" box="[511,636,577,599]" pageId="36" pageNumber="277">G. Japoshvili</collectorName>
&quot;; (
<specimenCount id="9D33AAE7FFFBFFF4E786FDAA2C934B27" box="[662,676,577,598]" count="1" pageId="36" pageNumber="277" type="female">♀</specimenCount>
)
</materialsCitation>
<materialsCitation id="3B5D6B33FFFBFFF4E7A6FDAA2DC84B1E" collectingDate="2020-07-19" collectingDateMax="2020-07-25" collectingDateMin="2020-07-19" collectorName="G. Japoshvili &amp; Riedel" country="Georgia" elevation="1489" latitude="42.69615" location="Tsageri" longLatPrecision="1" longitude="42.812965" municipality="Dogurashi" pageId="36" pageNumber="277" specimenCount="1">
&quot;
<collectingCountry id="F32221FEFFFBFFF4E7ACFDAA2D304B26" box="[700,775,577,599]" name="Georgia" pageId="36" pageNumber="277">Georgia</collectingCountry>
,
<location id="8EEA37B5FFFBFFF4E604FDAA2D6F4B26" LSID="urn:lsid:plazi:treatment:039CD078FFFBFFF5E5A2FE202C834B63:8EEA37B5FFFBFFF4E604FDAA2D6F4B26" box="[788,856,577,599]" country="Georgia" latitude="42.69615" longLatPrecision="1" longitude="42.812965" municipality="Dogurashi" name="Tsageri" pageId="36" pageNumber="277">Tsageri</location>
,
<collectingMunicipality id="6BEEFB14FFFBFFF4E674FDAA2DF64B26" box="[868,961,577,599]" pageId="36" pageNumber="277">Dogurashi</collectingMunicipality>
,
<quantity id="4CCDCC8BFFFBFFF4E6DDFDAA2A234B26" box="[973,1044,577,599]" metricMagnitude="3" metricUnit="m" metricValue="1.489" pageId="36" pageNumber="277" unit="m" value="1489.0">
<elevation id="0018865DFFFBFFF4E6DDFDAA2A234B26" box="[973,1044,577,599]" metricMagnitude="3" metricUnit="m" metricValue="1.489" pageId="36" pageNumber="277" unit="m" value="1489.0">1489 m</elevation>
</quantity>
alt.,
<geoCoordinate id="EE0107A9FFFBFFF4E5CFFDB22F604B1E" box="[223,343,601,623]" degrees="42.696151" direction="north" orientation="latitude" pageId="36" pageNumber="277" precision="1" value="42.69615">42.696151°N</geoCoordinate>
,
<geoCoordinate id="EE0107A9FFFBFFF4E44FFDB22FE24B1E" box="[351,469,601,623]" degrees="42.812965" direction="east" orientation="longitude" pageId="36" pageNumber="277" precision="1" value="42.812965">42.812965°E</geoCoordinate>
,
<date id="FF8B47AEFFFBFFF4E4CCFDB22C514B1E" box="[476,614,601,623]" pageId="36" pageNumber="277" value="2020-07-19" valueMax="2020-07-25" valueMin="2020-07-19">
<collectingDate id="EFCFBE46FFFBFFF4E4CCFDB22C514B1E" box="[476,614,601,623]" pageId="36" pageNumber="277" value="2020-07-19" valueMax="2020-07-25" valueMin="2020-07-19">19-25.VII.2020</collectingDate>
</date>
,
<collectingMethod id="52741979FFFBFFF4E77EFDB22CEA4B1E" box="[622,733,601,623]" pageId="36" pageNumber="277">Malaise trap</collectingMethod>
, leg.
<collectorName id="26C004B8FFFBFFF4E61AFDB22D484B1E" box="[778,895,601,623]" pageId="36" pageNumber="277">G. Japoshvili</collectorName>
&quot; (coll.
<collectorName id="26C004B8FFFBFFF4E6ACFDB22DCE4B1E" box="[956,1017,601,623]" pageId="36" pageNumber="277">Riedel</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8A616EFFFBFFF4E5A1FD942A194A38" blockId="36.[177,1085,638,841]" pageId="36" pageNumber="277">
D e s c r i p t i o n:
<specimenCount id="9D33AAE7FFFBFFF4E487FD942F9B4BE5" box="[407,428,639,660]" count="1" pageId="36" pageNumber="277" type="female">♀</specimenCount>
: Body length
<quantity id="4CCDCC8BFFFBFFF4E72FFD952C8E4BE5" box="[575,697,638,660]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="36" pageNumber="277" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5-5.5 mm</quantity>
. Flagellum with 22-24 flagellomeres, stout; preapical flagellomere 1.2× longer than wide. Head granulate, dull, covered with fine whitish hairs. Temple moderately and roundly narrowed behind eyes, dorsally 0.65×as long as eye. Distance between lateral ocellus and eye 0.8-0.9× and between lateral ocelli 1.1-1.2× ocellar diameter. Sides of face slightly convergent ventrally, minimally 0.47× as wide as head. Malar space 0.6× as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base, both carinae low.
</paragraph>
<paragraph id="8B8A616EFFFBFFF4E5A2FCB22FE44D0F" blockId="36.[178,1086,857,1150]" pageId="36" pageNumber="277">Side of pronotum finely rugose dorsally, with fine longitudinal striation ventrally. Mesoscutum granulate, dull. Mesopleural disc granulate, without distinct punctation, dull, dorsally with few fine rugae; speculum with microsculpture but strongly shining. Propodeum granulate. Area basalis triangular, about as long as wide, with long apical stalk. Area superomedia slightly wider than long and confluent with area petiolaris, caudal sides slightly divergent. Anterior transverse carina (costula) absent. Area petiolaris moderately impressed medially, granulate, without rugae. Hind femur 4.4-4.6× as long as wide. Claws as long as their pulvilli; hind claw with two or three strong basal teeth. Areolet ± oblique, stalked frontally; vein 2m-cu distad its middle; vein 1cu-a postfurcal by 2-3× its width.</paragraph>
<paragraph id="8B8A616EFFFBFFF4E5A2FB642FD44CE4" blockId="36.[178,1086,1163,1429]" pageId="36" pageNumber="277">
Metasoma rather slender. 1
<superScript id="7C40CC26FFFBFFF4E4C5FB602FD74DE8" attach="left" box="[469,480,1163,1177]" fontSize="6" pageId="36" pageNumber="277">st</superScript>
tergite 2.2-2.4× longer than wide, with strong glymma. Postpetiolus 0.90× as long as wide, with rounded sides. 2
<superScript id="7C40CC26FFFBFFF4E7E6FB422D314DC6" attach="left" box="[758,774,1193,1207]" fontSize="6" pageId="36" pageNumber="277">nd</superScript>
tergite 1.25-1.40× longer than wide. 3
<superScript id="7C40CC26FFFBFFF4E410FB2C2F3A4DA4" attach="left" box="[256,269,1223,1237]" fontSize="6" pageId="36" pageNumber="277">rd</superScript>
tergite 0.85-0.90× as long as wide. Basal tergites granulate and dull. Apical tergites from the 4
<superScript id="7C40CC26FFFBFFF4E46BFB0E2FB04D82" attach="left" box="[379,391,1253,1267]" fontSize="6" pageId="36" pageNumber="277">th</superScript>
on with fine microsculpture dorsally, smooth and with fine and scattered setiferous punctures laterally and apically. 6
<superScript id="7C40CC26FFFBFFF4E7C5FAE82CD64C60" attach="left" box="[725,737,1283,1297]" fontSize="6" pageId="36" pageNumber="277">th</superScript>
tergite distinctly excavate medio-dorsally; 7
<superScript id="7C40CC26FFFBFFF4E436FACA2F054C5E" attach="left" box="[294,306,1313,1327]" fontSize="6" pageId="36" pageNumber="277">th</superScript>
tergite with a deep, rectangular to trapezoid incision medio-dorsally. Ovipositor sheath 1.15-1.25× longer than hind tibia. Ovipositor continuously and distinctly curved upwards, not thickened medially; ventral valve smooth, tip of dorsal valve with triangular incision.
</paragraph>
<paragraph id="8B8A616EFFFBFFF4E5A2FA4E2A0A4F45" blockId="36.[178,1086,1445,1588]" pageId="36" pageNumber="277">
Color: black. Palps and tegula cream-yellow. Mandible reddish-yellow medially. Tergites entirely blackish. Coxae black; fore and mid trochanters yellowish, weakly infuscate basally, hind trochanter black. Trochantelli yellowish. Legs otherwise mainly reddish; hind tibia reddish-yellow, cream-yellow basally, with weak subbasal and distinct but narrow apical blackish bands. Hind metatarsus and 2
<superScript id="7C40CC26FFFBFFF4E682F9F12D954F59" attach="left" box="[914,930,1562,1576]" fontSize="6" pageId="36" pageNumber="277">nd</superScript>
tarsomeres ±
</paragraph>
<paragraph id="8B8A616EFFFAFFF5E5A2FF302C73487E" blockId="37.[178,1084,219,271]" pageId="37" pageNumber="278">yellowish basally, with infuscate tips; following hind tarsomeres blackish. Wings hyaline; pterostigma hyaline-yellowish.</paragraph>
<paragraph id="8B8A616EFFFAFFF5E5A2FEF62F064844" blockId="37.[178,305,285,309]" box="[178,305,285,309]" pageId="37" pageNumber="278">♁ unknown.</paragraph>
<paragraph id="8B8A616EFFFAFFF5E5A2FEAE2A2548E4" blockId="37.[178,1085,324,405]" pageId="37" pageNumber="278">
R e m a r k: This new species belongs to the subgenus
<taxonomicName id="4C351AEDFFFAFFF5E625FEAE2F254809" authority="HORSTMANN. Members" authorityName="HORSTMANN. Members" pageId="37" pageNumber="278" rank="subGenus" subGenus="Neoangitia">
<emphasis id="B941BD7CFFFAFFF5E625FEAE2D91482B" box="[821,934,325,346]" italics="true" pageId="37" pageNumber="278">Neoangitia</emphasis>
HORSTMANN. Members
</taxonomicName>
of this subgenus are characterized by the strongly shining apical tergites and the deep and rectangular or trapezoid form of the medio-dorsal excision of 7
<superScript id="7C40CC26FFFAFFF5E6A7FE902DF448F8" attach="left" box="[951,963,379,393]" fontSize="6" pageId="37" pageNumber="278">th</superScript>
tergite.
</paragraph>
<paragraph id="8B8A616EFFFAFFF5E5A2FE4E2C834B63" blockId="37.[178,1085,420,530]" pageId="37" pageNumber="278">
Due to its long ovipositor and weak dark coloration of hind tibia, the new species runs to
<taxonomicName id="4C351AEDFFFAFFF5E5A2FE292C5748A6" authority="(RATZEBURG)" baseAuthorityName="RATZEBURG" box="[178,608,450,471]" class="Insecta" family="Ichneumonidae" genus="Diadegma" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="278" phylum="Arthropoda" rank="species" species="coleophorarum">
<emphasis id="B941BD7CFFFAFFF5E5A2FE292FF348A6" box="[178,452,450,471]" italics="true" pageId="37" pageNumber="278">Diadegma coleophorarum</emphasis>
(RATZEBURG)
</taxonomicName>
, but differs by a slightly longer ovipositor sheath, apically open area superomedia, distinct medio-dorsal excavation of 6
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tergite, black fore and mid coxae and entirely black scape.
</paragraph>
</subSubSection>
</treatment>
</document>
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