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<mods:title id="7BE9106C425F12750959ED21FBDD8F1E">A new species of Aegus Macleay, 1819 (Coleoptera: Lucanidae: Lucaninae: Aegini) from Thailand with checklist of the genus species from the country</mods:title>
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<taxonomicName id="4C3D1585990AFFDEFF2BFA6BFD8FF9A3" ID-CoL="64WWB" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[151,512,1524,1551]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="chelifer">
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<emphasis id="B949B214990AFFDEFF2BFA6BFEC9F9A3" bold="true" box="[151,326,1524,1551]" italics="true" pageId="9" pageNumber="350">Aegus chelifer</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFEF2FA6BFD8FF9A3" author="MacLeay, W. S." box="[334,512,1525,1551]" pageId="9" pageNumber="350" refId="ref9658" refString="MacLeay, W. S. (1819) Horae entomologicae: or essays on the annulose animals. Vol. 1. Part 1. S. Bagster, London, 160 pp. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">Macleay, 1819</bibRefCitation>
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(
<figureCitation id="13067283990AFFDEFF23F987FE84F998" box="[159,267,1561,1588]" captionStart="FIGURES 1–14" captionStartId="2.[151,264,1822,1846]" captionTargetBox="[216,1371,185,1794]" captionTargetId="figure-22@2.[212,1375,181,1798]" captionTargetPageId="2" captionText="FIGURES 1–14. Aegus spp. male habitus, dorsal view. 1–5, Aegus chelifer Macleay, 1819; 1, Nakhon Ratchasima Province, Thailand; 2, 4, Chiang Rai Province, Thailand; 3, Nakhon Si Thammarat Province, Thailand; 5, Chanthaburi Province, Thailand; 6–7, 9–10, Paratype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species, Ratchaburi Province, Thailand; 8, Holotype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species, Ratchaburi Province, Thailand; 11, Aegus rondoni Bomans, 1971, Sekong Province, Laos.; 12–14, Aegus acuminatus acuminatus (Fabricius, 1801), Nakhon Si Thammarat Province, Thailand." figureDoi="http://doi.org/10.5281/zenodo.11234120" httpUri="https://zenodo.org/record/11234120/files/figure.png" pageId="9" pageNumber="350">Figs. 1–5</figureCitation>
, 
<figureCitation id="13067283990AFFDEFEABF987FEBAF99F" box="[279,309,1561,1587]" captionStart="FIGURES 15–18" captionStartId="3.[151,264,715,739]" captionTargetBox="[171,1430,186,691]" captionTargetId="figure-17@3.[151,1435,181,691]" captionTargetPageId="3" captionText="FIGURES 15–18. Head and pronotum of Aegus spp. large sized male in dorsal view. 15, paratype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species; 16, Aegus rondoni Bomans, 1971; 17, Aegus acuminatus (Fabricius, 1801); 18, Aegus chelifer Macleay, 1819." figureDoi="http://doi.org/10.5281/zenodo.11234124" httpUri="https://zenodo.org/record/11234124/files/figure.png" pageId="9" pageNumber="350">18</figureCitation>
, 
<figureCitation id="13067283990AFFDEFEFDF987FEEFF99F" box="[321,352,1561,1587]" captionStart="FIGURES 19–22" captionStartId="4.[151,264,1945,1969]" captionTargetBox="[166,1430,252,1913]" captionTargetId="figure-54@4.[158,1430,237,1922]" captionTargetPageId="4" captionText="FIGURES 19–22. Head of Aegus spp. large sized male in lateral view. 19, paratype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species; 20, Aegus rondoni Bomans, 1971; 21, Aegus acuminatus (Fabricius, 1801); 22, Aegus chelifer Macleay, 1819." figureDoi="http://doi.org/10.5281/zenodo.11234128" httpUri="https://zenodo.org/record/11234128/files/figure.png" pageId="9" pageNumber="350">22</figureCitation>
, 
<figureCitation id="13067283990AFFDEFED7F987FE3BF99F" box="[363,436,1561,1587]" captionStart="FIGURES 23–30" captionStartId="5.[151,264,1576,1600]" captionTargetBox="[152,1435,181,1553]" captionTargetId="figure-17@5.[151,1435,181,1553]" captionTargetPageId="5" captionText="FIGURES 23–30. Male genitalia of Aegus species in dorsal (23, 25, 27, 29) and ventral (24, 26, 28, 30) views. 23–24, holotype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species, Ratchaburi Province, Thailand; 25–26, Aegus rondoni Bomans, 1971, Sekong Province, Laos; 27–28, Aegus acuminatus (Fabricius, 1801), Nakhon Si Thammarat Province, Thailand; 29–30, Aegus chelifer Macleay, 1819, Chiang Mai Province, Thailand." figureDoi="http://doi.org/10.5281/zenodo.11234132" httpUri="https://zenodo.org/record/11234132/files/figure.png" pageId="9" pageNumber="350">29–30</figureCitation>
, 
<figureCitation id="13067283990AFFDEFE03F987FE52F99F" box="[447,477,1561,1587]" captionStart="FIGURES 31–34" captionStartId="8.[151,264,783,807]" captionTargetBox="[156,1431,185,755]" captionTargetId="figure-22@8.[151,1435,181,759]" captionTargetPageId="8" captionText="FIGURES 31–34. Female habitus of Aegus species in dorsal view. 31, paratype of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species, Ratchaburi Province, Thailand; 32, Aegus rondoni Bomans, 1971, Quang Binh, Vietnam; 33, Aegus acuminatus (Fabricius, 1801), Nakhon Si Thammarat Province, Thailand; 34, Aegus chelifer Macleay, 1819, Chanthaburi Province, Thailand." figureDoi="http://doi.org/10.5281/zenodo.11234134" httpUri="https://zenodo.org/record/11234134/files/figure.png" pageId="9" pageNumber="350">34</figureCitation>
, 
<figureCitation id="13067283990AFFDEFE55F987FD88F99F" box="[489,519,1561,1587]" captionStart="FIGURES 35–39" captionStartId="8.[151,264,1668,1692]" captionTargetBox="[151,1435,961,1644]" captionTargetId="figure-75@8.[151,1435,939,1644]" captionTargetPageId="8" captionText="FIGURES 35–39. Female genitalia of Aegus species. 35–36, paratypes of Aegus occidentalis Pathomwattananurak &amp; Jiaranaisakul, new species, Ratchaburi Province, Thailand; 37, Aegus rondoni Bomans, 1971, Quang Binh, Vietnam; 38, Aegus acuminatus (Fabricius, 1801), Nakhon Si Thammarat Province, Thailand; 39, Aegus chelifer Macleay, 1819, Nakhon Si Thammarat Province, Thailand." figureDoi="http://doi.org/10.5281/zenodo.11234136" httpUri="https://zenodo.org/record/11234136/files/figure.png" pageId="9" pageNumber="350">39</figureCitation>
, 
<figureCitation id="13067283990AFFDEFDAFF987FDD4F998" box="[531,603,1561,1588]" captionStart-0="FIGURES 45–47" captionStart-1="FIGURES 48–50" captionStartId-0="11.[151,264,943,967]" captionStartId-1="11.[151,264,1877,1901]" captionTargetBox-0="[154,1435,184,916]" captionTargetBox-1="[168,1435,1027,1853]" captionTargetId-0="figure-17@11.[151,1435,181,919]" captionTargetId-1="figure-39@11.[151,1435,1027,1853]" captionTargetPageId-0="11" captionTargetPageId-1="11" captionText-0="FIGURES 45–47. Syntype of Aegus chelifer Macleay, 1819 (image ©BMNH/Keita Matsumoto). 45, dorsal habitus; 46, ventral habitus; 47, labels." captionText-1="FIGURES 48–50. Holotype of Aegus chelifer crassodontus Bomans, 1992 (image ©BMNH/Keita Matsumoto). 48, dorsal habitus; 49, ventral habitus; 50, labels." figureDoi-0="http://doi.org/10.5281/zenodo.11234142" figureDoi-1="http://doi.org/10.5281/zenodo.11234146" httpUri-0="https://zenodo.org/record/11234142/files/figure.png" httpUri-1="https://zenodo.org/record/11234146/files/figure.png" pageId="9" pageNumber="350">45–50</figureCitation>
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<taxonomicName id="4C3D1585990AFFDEFF2BF9C0FD88F9DA" ID-CoL="64WWB" authority="Macleay, 1819: 113" authorityName="Macleay" authorityPageNumber="113" authorityYear="1819" box="[151,519,1629,1654]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B214990AFFDEFF2BF9C0FEA3F9D9" box="[151,300,1629,1653]" italics="true" pageId="9" pageNumber="350">Aegus chelifer</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFE8AF9C3FD88F9DA" author="MacLeay, W. S." box="[310,519,1629,1654]" pageId="9" pageNumber="350" refId="ref9658" refString="MacLeay, W. S. (1819) Horae entomologicae: or essays on the annulose animals. Vol. 1. Part 1. S. Bagster, London, 160 pp. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">Macleay, 1819: 113</bibRefCitation>
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locality: Australasia? 
<treatmentCitation id="0A9C4817990AFFDEFC8EF9C0FB8BF9DA" author="Van Roon, G." box="[818,1028,1630,1654]" page="41" pageId="9" pageNumber="350" year="1910">
<bibRefCitation id="EFAC13F7990AFFDEFC8EF9C0FB8BF9DA" author="Van Roon, G." box="[818,1028,1630,1654]" pageId="9" pageNumber="350" refId="ref10367" refString="Van Roon, G. (1910) Coleopterorum Catalogus, Vol. 8: Lucanidae. W. Junk, Berlin, 70 pp." type="book" year="1910">Van Roon 1910: 41</bibRefCitation>
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<treatmentCitation id="0A9C4817990AFFDEFBADF9C0FB46F9DA" author="Arrow, G. J." box="[1041,1225,1630,1654]" page="141" pageId="9" pageNumber="350" year="1943">
<bibRefCitation id="EFAC13F7990AFFDEFBADF9C0FB46F9DA" author="Arrow, G. J." box="[1041,1225,1630,1654]" pageId="9" pageNumber="350" pagination="133 - 143" refId="ref8374" refString="Arrow, G. J. (1943) On the genera and nomenclature of the lucanid Coleoptera, and descriptions of a few new species. Proceedings of the Royal Entomological Society of London, Series B, 12 (9 - 10), 133 - 143. https: // doi. org / 10.1111 / j. 1365 - 3113.1943. tb 00761. x" type="journal article" year="1943">Arrow 1943: 141</bibRefCitation>
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<treatmentCitation id="0A9C4817990AFFDEFB6BF9C3FA17F9DA" author="Benesh, B." box="[1239,1432,1629,1654]" page="100" pageId="9" pageNumber="350" year="1960">
<bibRefCitation id="EFAC13F7990AFFDEFB6BF9C3FA17F9DA" author="Benesh, B." box="[1239,1432,1629,1654]" pageId="9" pageNumber="350" pagination="1 - 178" refId="ref8502" refString="Benesh, B. (1960) Lucanidae. Coleopterorum Catalogus, Supplementa, 2 nd Edition, 8, 1 - 178." type="journal article" year="1960">Benesh 1960: 100</bibRefCitation>
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<bibRefCitation id="EFAC13F7990AFFDEFF7BF9E3FE06F938" author="Bomans, H. E." box="[199,393,1660,1684]" pageId="9" pageNumber="350" pagination="317 - 323" refId="ref8618" refString="Bomans, H. E. (1967) Contribution a l'etude des Coleopteres Lucanides. Lucanidae du Laos, 1 ere note. Bulletin et annales de la Societe royale d'Entomologie de Belgique, 103 (7 - 8), 317 - 323." type="journal article" year="1967">Bomans 1967: 321</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFE2FF9E3FDDBF938" author="Bomans, H. E." box="[403,596,1660,1684]" pageId="9" pageNumber="350" pagination="223 - 234" refId="ref8664" refString="Bomans, H. E. (1970) Contribution a l'etude des Coleopteres Lucanides. Lucanidae du Laos - 3 e note. Bulletin et annales de la Societe royale d'Entomologie de Belgique, 106, 223 - 234." type="journal article" year="1970">Bomans 1970: 231</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFDE2F9E3FC90F938" author="Bomans, H. E." box="[606,799,1660,1684]" pageId="9" pageNumber="350" pagination="293 - 305" refId="ref8792" refString="Bomans, H. E. (1991) Considerations sur le Genre Aegus Macleay et description d'especes nouvelles (Coleoptera Lucanidae) 2 eme note. (76 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 91 (3), 293 - 305." type="journal article" year="1991">Bomans 1991: 144</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFC95F9E2FBEEF938" author="Mizunuma, T. &amp; Nagai, T." box="[809,1121,1660,1684]" pageId="9" pageNumber="350" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai 1994: 281</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFBD7F9E2FA96F938" author="Krajcik, M." box="[1131,1305,1660,1684]" pageId="9" pageNumber="350" refId="ref9509" refString="Krajcik, M. (2003) Lucanidae of the World. Catalogue - Part II. Encyclopaedia of the Lucanidae (Coleoptera: Lucanidae). Privately published, Plzen, 197 pp." type="book" year="2003">Krajcik 2003: 12</bibRefCitation>
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<bibRefCitation id="EFAC13F7990AFFDEFA9FF9E2FA13F938" author="Pinratana, A. &amp; Maes, J. M." box="[1315,1436,1660,1684]" pageId="9" pageNumber="350" refId="ref10072" refString="Pinratana, A. &amp; Maes, J. M. (2003) Lucanidae of Thailand. Sunprinting, Bangkok, 248 pp." type="book" year="2003">Pinratana &amp;</bibRefCitation>
Maes 2003: 235; 
<bibRefCitation id="EFAC13F7990AFFDEFEC4F905FDADF91F" author="Fujita, H." box="[376,546,1691,1715]" pageId="9" pageNumber="350" refId="ref9102" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010: 318</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFD91F905FCA0F91F" author="Huang, H. &amp; Chen, C. C." box="[557,815,1691,1715]" pageId="9" pageNumber="350" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen 2017:249</bibRefCitation>
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</paragraph>
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<taxonomicName id="4C3D1585990AFFDEFF2BF924FE64F97E" authority="Boileau, 1899: 321" authorityName="Boileaui" authorityPageNumber="321" authorityYear="1899" box="[151,491,1721,1746]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B949B214990AFFDEFF2BF924FEAFF97D" box="[151,288,1721,1745]" italics="true" pageId="9" pageNumber="350">Aegus nitidus</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFE9BF927FE64F97E" author="Boileau, H." box="[295,491,1721,1746]" pageId="9" pageNumber="350" pagination="319 - 322" refId="ref8526" refString="Boileau, H. (1899) Descriptions sommaires d'Aegus nouveaux. Bulletin de la Societe entomologique de France, 48, 319 - 322. https: // doi. org / 10.3406 / bsef. 1899.22441" type="journal article" year="1899">Boileau, 1899: 321</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="5486D0A4990AFFDEFE48F924FDA9F97E" box="[500,550,1722,1746]" pageId="9" pageNumber="350">Type</typeStatus>
locality: Borneo. 
<bibRefCitation id="EFAC13F7990AFFDEFD62F927FC16F97D" author="Boileau, H." box="[734,921,1721,1746]" pageId="9" pageNumber="350" pagination="213 - 272" refId="ref8564" refString="Boileau, H. (1913) Note sur Lucanides conserves dans les collections de l'Universite d'Oxford et du British Museum. Transactions of the entomological Society London, 1913, 213 - 272. https: // doi. org / 10.1111 / j. 1365 - 2311.1913. tb 01776. x" type="journal article" year="1913">Boileau 1913: 256</bibRefCitation>
(as synonym).
</paragraph>
<paragraph id="8B826E06990AFFDEFF2BF946FA13F95C" blockId="9.[151,1437,1629,1930]" box="[151,1436,1752,1777]" pageId="9" pageNumber="350">
<taxonomicName id="4C3D1585990AFFDEFF2BF946FD97F95C" authority="Jakovlev, 1900: 633" authorityName="Jakovlev" authorityPageNumber="633" authorityYear="1900" box="[151,536,1752,1776]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="specularis">
<emphasis id="B949B214990AFFDEFF2BF946FECCF95C" box="[151,323,1752,1776]" italics="true" pageId="9" pageNumber="350">Aegus specularis</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFEF6F946FD97F95C" author="Jakovlev, B. E." box="[330,536,1752,1776]" pageId="9" pageNumber="350" pagination="631 - 642" refId="ref9443" refString="Jakovlev, B. E. (1900) Description de quelques nouvelles especes de la famille des Lucanides. Horae Societatis Entomologicae Rossicae, 34, 631 - 642." type="journal article" year="1900">Jakovlev, 1900: 633</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="5486D0A4990AFFDEFD9DF947FDDCF95D" box="[545,595,1753,1777]" pageId="9" pageNumber="350">Type</typeStatus>
locality: Cochinchine: 
<collectingRegion id="49F9A0E4990AFFDEFCFDF946FC04F95C" box="[833,907,1752,1776]" country="Vietnam" name="Ho Chi Minh City" pageId="9" pageNumber="350">Saigon</collectingRegion>
. 
<bibRefCitation id="EFAC13F7990AFFDEFC2EF946FBCBF95C" author="Arrow, G. J." box="[914,1092,1752,1776]" pageId="9" pageNumber="350" pagination="133 - 143" refId="ref8374" refString="Arrow, G. J. (1943) On the genera and nomenclature of the lucanid Coleoptera, and descriptions of a few new species. Proceedings of the Royal Entomological Society of London, Series B, 12 (9 - 10), 133 - 143. https: // doi. org / 10.1111 / j. 1365 - 3113.1943. tb 00761. x" type="journal article" year="1943">Arrow 1943: 141</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFBF2F946FA8AF95C" author="Benesh, B." box="[1102,1285,1752,1776]" pageId="9" pageNumber="350" pagination="1 - 178" refId="ref8502" refString="Benesh, B. (1960) Lucanidae. Coleopterorum Catalogus, Supplementa, 2 nd Edition, 8, 1 - 178." type="journal article" year="1960">Benesh 1960: 100</bibRefCitation>
(as synonym).
</paragraph>
<paragraph id="8B826E06990AFFDEFF2BF969FB41F8A3" blockId="9.[151,1437,1629,1930]" box="[151,1230,1783,1807]" pageId="9" pageNumber="350">
<taxonomicName id="4C3D1585990AFFDEFF2BF969FD1DF8A3" authority="Kriesche, 1920: 101" authorityName="Kriesche" authorityPageNumber="101" authorityYear="1920" box="[151,658,1783,1807]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B214990AFFDEFF2BF969FE33F8A3" box="[151,444,1783,1807]" italics="true" pageId="9" pageNumber="350">Aegus specularis tonkinensis</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFE7EF969FD1DF8A3" author="Kriesche, R." box="[450,658,1783,1807]" pageId="9" pageNumber="350" pagination="92 - 107" refId="ref9544" refString="Kriesche, R. (1920) Zur Kenntnis der Lucaniden. Archiv fur Naturgeschichte, 86 A (8), 92 - 107." type="journal article" year="1920">Kriesche, 1920: 101</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="5486D0A4990AFFDEFD27F969FD42F8A3" box="[667,717,1783,1807]" pageId="9" pageNumber="350">Type</typeStatus>
locality: 
<collectingCountry id="F32A2E96990AFFDEFC91F969FCF7F8A3" box="[813,888,1783,1807]" name="Vietnam" pageId="9" pageNumber="350">Tonkin</collectingCountry>
. 
<bibRefCitation id="EFAC13F7990AFFDEFC3DF969FBB7F8A3" author="Benesh, B." box="[897,1080,1783,1807]" pageId="9" pageNumber="350" pagination="1 - 178" refId="ref8502" refString="Benesh, B. (1960) Lucanidae. Coleopterorum Catalogus, Supplementa, 2 nd Edition, 8, 1 - 178." type="journal article" year="1960">Benesh 1960: 101</bibRefCitation>
(as synonym).
</paragraph>
<paragraph id="8B826E06990AFFDEFF2BF888FF7BF8E0" blockId="9.[151,1437,1629,1930]" pageId="9" pageNumber="350">
<taxonomicName id="4C3D1585990AFFDEFF2BF888FEFCF881" authorityName="Boileau" authorityYear="1899" box="[151,371,1813,1837]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="nitidus">
<emphasis id="B949B214990AFFDEFF2BF888FEFCF881" box="[151,371,1813,1837]" italics="true" pageId="9" pageNumber="350">Aegus chelifer nitidus</emphasis>
</taxonomicName>
: 
<bibRefCitation id="EFAC13F7990AFFDEFEC3F88BFD3CF882" author="Mizunuma, T. &amp; Nagai, T." box="[383,691,1813,1838]" pageId="9" pageNumber="350" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai 1994: 281</bibRefCitation>
(as subspecies); 
<bibRefCitation id="EFAC13F7990AFFDEFCE1F88BFC6CF882" author="Krajcik, M." box="[861,995,1813,1838]" pageId="9" pageNumber="350" refId="ref9474" refString="Krajcik, M. (2001) Lucanidae of the World. Catalogue - Part I. Encyclopaedia of the Lucanidae (Coleoptera: Lucanidae). Privately published, Most, 197 pp." type="book" year="2001">Krajcik 2001</bibRefCitation>
, 55; 
<bibRefCitation id="EFAC13F7990AFFDEFBAFF88BFB33F882" author="Fujita, H." box="[1043,1212,1813,1838]" pageId="9" pageNumber="350" refId="ref9102" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010: 319</bibRefCitation>
; 
<bibRefCitation id="EFAC13F7990AFFDEFB79F888FA12F882" author="Huang, H. &amp; Chen, C. C." box="[1221,1437,1813,1838]" pageId="9" pageNumber="350" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen 2017:</bibRefCitation>
252.
</paragraph>
<paragraph id="8B826E06990AFFDEFF2BF8CDFB28F8C7" blockId="9.[151,1437,1629,1930]" box="[151,1191,1875,1899]" pageId="9" pageNumber="350">
<taxonomicName id="4C3D1585990AFFDEFF2BF8CDFE2EF8C7" authorityName="Kriesche" authorityYear="1920" box="[151,417,1875,1899]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="tonkinensis">
<emphasis id="B949B214990AFFDEFF2BF8CDFE2EF8C7" box="[151,417,1875,1899]" italics="true" pageId="9" pageNumber="350">Aegus chelifer tonkinensis</emphasis>
</taxonomicName>
: 
<bibRefCitation id="EFAC13F7990AFFDEFE12F8CDFDFBF8C7" author="Bomans, H. E." box="[430,628,1875,1899]" pageId="9" pageNumber="350" pagination="179 - 196" refId="ref8841" refString="Bomans, H. E. (1992) Considerations sur le genre Aegus Macleay et description d'especes nouvelles 3 eme note. (77 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 92 (2), 179 - 196." type="journal article" year="1992">Bomans, 1992: 186</bibRefCitation>
(as variety); 
<bibRefCitation id="EFAC13F7990AFFDEFD47F8CDFB8FF8C7" author="Huang, H. &amp; Chen, C. C." box="[763,1024,1875,1899]" pageId="9" pageNumber="350" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen 2017: 249</bibRefCitation>
(as subspecies).
</paragraph>
<paragraph id="8B826E06990AFFDEFF2BF8ECFAAEF826" blockId="9.[151,1437,1629,1930]" box="[151,1313,1905,1930]" pageId="9" pageNumber="350">
<taxonomicName id="4C3D1585990AFFDEFF2BF8ECFD0BF825" authority="Bomans, 1992: 186" authorityName="Bomans" authorityPageNumber="186" authorityYear="1992" box="[151,644,1905,1930]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B214990AFFDEFF2BF8ECFE3AF825" box="[151,437,1905,1929]" italics="true" pageId="9" pageNumber="350">Aegus chelifer crassodontus</emphasis>
<bibRefCitation id="EFAC13F7990AFFDEFE07F8ECFD0BF825" author="Bomans, H. E." box="[443,644,1905,1930]" pageId="9" pageNumber="350" pagination="179 - 196" refId="ref8841" refString="Bomans, H. E. (1992) Considerations sur le genre Aegus Macleay et description d'especes nouvelles 3 eme note. (77 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 92 (2), 179 - 196." type="journal article" year="1992">Bomans, 1992: 186</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="5486D0A4990AFFDEFD31F8ECFD30F826" box="[653,703,1906,1930]" pageId="9" pageNumber="350">Type</typeStatus>
locality: 
<collectingCountry id="F32A2E96990AFFDEFCA2F8ECFCE3F826" box="[798,876,1906,1930]" name="Vietnam" pageId="9" pageNumber="350">Annam</collectingCountry>
. 
<bibRefCitation id="EFAC13F7990AFFDEFCC9F8ECFBF1F825" author="Huang, H. &amp; Chen, C. C." box="[885,1150,1905,1930]" pageId="9" pageNumber="350" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen 2017: 252</bibRefCitation>
; 
<emphasis id="B949B214990AFFDEFB34F8ECFAAEF826" bold="true" box="[1160,1313,1906,1930]" pageId="9" pageNumber="350">new synonym.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3273D8D9909FFDDFF2BFF09FC73FD05" pageId="10" pageNumber="351" type="materials_examined">
<paragraph id="8B826E069909FFDDFF2BFF09FD0EFE25" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
<emphasis id="B949B2149909FFDDFF2BFF09FE49FF1C" bold="true" box="[151,454,150,177]" pageId="10" pageNumber="351">Type material examined.</emphasis>
<materialsCitation id="3B55645B9909FFDDFE6CFF08FA16FF55" collectionCode="BMNH" collectorName="Keita Matsumoto" determinerName="H. Bomans" pageId="10" pageNumber="351" specimenCount="2" specimenCount-male="2" typeStatus="syntype">
<emphasis id="B949B2149909FFDDFE6CFF08FDBFFF1C" bold="true" box="[464,560,150,176]" pageId="10" pageNumber="351">
<typeStatus id="5486D0A49909FFDDFE6CFF08FDBFFF1C" box="[464,560,150,176]" pageId="10" pageNumber="351" type="syntype">Syntype</typeStatus>
</emphasis>
of 
<taxonomicName id="4C3D15859909FFDDFDE1FF09FC3CFF1D" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[605,947,151,177]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFDE1FF09FC8EFF1D" box="[605,769,151,177]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFCB7FF09FC3CFF1D" author="MacLeay, W. S." box="[779,947,151,177]" pageId="10" pageNumber="351" refId="ref9658" refString="MacLeay, W. S. (1819) Horae entomologicae: or essays on the annulose animals. Vol. 1. Part 1. S. Bagster, London, 160 pp. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
(
<specimenCount id="9D3BA58F9909FFDDFC7BFF09FC51FF1D" box="[967,990,151,177]" count="1" pageId="10" pageNumber="351" type="male">♂</specimenCount>
, 
<collectionCode id="ED2CF6C39909FFDDFC51FF09FBCBFF1D" box="[1005,1092,151,177]" pageId="10" pageNumber="351">BMNH</collectionCode>
, 
<figureCitation id="130672839909FFDDFBEFFF09FB51FF1D" box="[1107,1246,150,177]" captionStart="FIGURES 45–47" captionStartId="11.[151,264,943,967]" captionTargetBox="[154,1435,184,916]" captionTargetId="figure-17@11.[151,1435,181,919]" captionTargetPageId="11" captionText="FIGURES 45–47. Syntype of Aegus chelifer Macleay, 1819 (image ©BMNH/Keita Matsumoto). 45, dorsal habitus; 46, ventral habitus; 47, labels." figureDoi="http://doi.org/10.5281/zenodo.11234142" httpUri="https://zenodo.org/record/11234142/files/figure.png" pageId="10" pageNumber="351">Figs. 45–47</figureCitation>
), labeled: “Type // 1736 // 
<taxonomicName id="4C3D15859909FFDDFEB0FF25FD95FF79" authority="Macleay" authorityName="Macleay" authorityYear="1819" box="[268,538,187,213]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFEB0FF25FE3EFF79" box="[268,433,187,213]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
Macleay
</taxonomicName>
australasia. 1736 // 
<taxonomicName id="4C3D15859909FFDDFCBBFF25FCD2FF79" authorityName="Macleay" authorityYear="1819" box="[775,861,187,213]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFCBBFF25FCD2FF79" box="[775,861,187,213]" italics="true" pageId="10" pageNumber="351">chelifer</emphasis>
</taxonomicName>
. // 
<determinerName id="7E5E82C29909FFDDFC3BFF25FB86FF79" box="[903,1033,187,213]" pageId="10" pageNumber="351">H. Bomans</determinerName>
det., 1983 
<taxonomicName id="4C3D15859909FFDDFB33FF25FAF1FF79" authority="Macl." authorityName="Macleay" authorityYear="1819" box="[1167,1406,187,213]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFB33FF25FABCFF79" box="[1167,1331,187,213]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
Macl.
</taxonomicName>
<specimenCount id="9D3BA58F9909FFDDFA3BFF25FA13FF79" box="[1415,1436,187,213]" count="1" pageId="10" pageNumber="351" type="male">♂</specimenCount>
// BMNH(E) #610724 // NHMUK015534103”, examined through photographs provided by 
<collectorName id="26C80BD09909FFDDFB71FF41FA16FF55" box="[1229,1433,223,249]" pageId="10" pageNumber="351">Keita Matsumoto</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B55645B9909FFDDFF2BFE9DFDF2FE25" collectingDate="1920-05-25" collectionCode="BMNH" collectorName="H. E. Bomans" country="Vietnam" determinerName="Lacro J. P. Lacroix &amp; H. Bomans" location="ANNAM" pageId="10" pageNumber="351" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B949B2149909FFDDFF2BFE9DFE8BFEB1" bold="true" box="[151,260,259,285]" pageId="10" pageNumber="351">
<typeStatus id="5486D0A49909FFDDFF2BFE9DFE8BFEB1" box="[151,260,259,285]" pageId="10" pageNumber="351" type="holotype">Holotype</typeStatus>
</emphasis>
of 
<taxonomicName id="4C3D15859909FFDDFE94FE9DFC86FEB1" authority="Bomans, 1992" authorityName="Bomans" authorityYear="1992" box="[296,777,259,285]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B2149909FFDDFE94FE9DFDEDFEB1" box="[296,610,259,285]" italics="true" pageId="10" pageNumber="351">Aegus chelifer crassodontus</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFDD4FE9DFC86FEB1" author="Bomans, H. E." box="[616,777,259,285]" pageId="10" pageNumber="351" pagination="179 - 196" refId="ref8841" refString="Bomans, H. E. (1992) Considerations sur le genre Aegus Macleay et description d'especes nouvelles 3 eme note. (77 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 92 (2), 179 - 196." type="journal article" year="1992">Bomans, 1992</bibRefCitation>
</taxonomicName>
(
<specimenCount id="9D3BA58F9909FFDDFCA5FE9DFCBFFEB1" box="[793,816,259,285]" count="1" pageId="10" pageNumber="351" type="male">♂</specimenCount>
, 
<collectionCode id="ED2CF6C39909FFDDFC87FE9DFC1DFEB1" box="[827,914,259,285]" pageId="10" pageNumber="351">BMNH</collectionCode>
, 
<figureCitation id="130672839909FFDDFC20FE9DFBACFEB1" box="[924,1059,258,285]" captionStart="FIGURES 48–50" captionStartId="11.[151,264,1877,1901]" captionTargetBox="[168,1435,1027,1853]" captionTargetId="figure-39@11.[151,1435,1027,1853]" captionTargetPageId="11" captionText="FIGURES 48–50. Holotype of Aegus chelifer crassodontus Bomans, 1992 (image ©BMNH/Keita Matsumoto). 48, dorsal habitus; 49, ventral habitus; 50, labels." figureDoi="http://doi.org/10.5281/zenodo.11234146" httpUri="https://zenodo.org/record/11234146/files/figure.png" pageId="10" pageNumber="351">Figs. 48–50</figureCitation>
), labeled: “ 
<typeStatus id="5486D0A49909FFDDFB23FE9DFA82FEB1" box="[1183,1293,259,285]" pageId="10" pageNumber="351" type="holotype">Holotype</typeStatus>
. // 
<date id="FF8348C69909FFDDFA90FE9CFA13FEB1" box="[1324,1436,258,285]" pageId="10" pageNumber="351" value="1920-05-25">
<collectingDate id="EFC7B12E9909FFDDFA90FE9CFA13FEB1" box="[1324,1436,258,285]" pageId="10" pageNumber="351" value="1920-05-25">25.5.1920</collectingDate>
</date>
// 
<typeStatus id="5486D0A49909FFDDFF12FEB9FE99FEED" box="[174,278,295,321]" pageId="10" pageNumber="351" type="holotype">Holotype</typeStatus>
// 6136 // 
<taxonomicName id="4C3D15859909FFDDFE36FEB9FE41FEED" authorityName="Macleay" authorityYear="1819" box="[394,462,295,321]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B949B2149909FFDDFE36FEB9FE41FEED" box="[394,462,295,321]" italics="true" pageId="10" pageNumber="351">Aegus</emphasis>
</taxonomicName>
<specimenCount id="9D3BA58F9909FFDDFE6AFEB9FE64FEED" box="[470,491,295,321]" count="1" pageId="10" pageNumber="351" type="male">♂</specimenCount>
<emphasis id="B949B2149909FFDDFE4EFEB9FD0CFEED" box="[498,643,295,321]" italics="true" pageId="10" pageNumber="351">macrodontus</emphasis>
<collectingCountry id="F32A2E969909FFDDFD35FEB9FD79FEED" box="[649,758,295,321]" name="Vietnam" pageId="10" pageNumber="351">ANNAM</collectingCountry>
, 
<determinerName id="7E5E82C29909FFDDFCBDFEB9FC41FEED" box="[769,974,295,321]" pageId="10" pageNumber="351">Lacro J.P. Lacroix</determinerName>
det. 1977 // 
<determinerName id="7E5E82C29909FFDDFBE7FEB9FB55FEED" box="[1115,1242,295,321]" pageId="10" pageNumber="351">H. Bomans</determinerName>
det., 1992 
<taxonomicName id="4C3D15859909FFDDFAE4FEB9FE33FEC9" authority="Pathomwattananurak &amp; Jiaranaisakul, 2024" authorityName="Pathomwattananurak &amp; Jiaranaisakul" authorityYear="2024" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="variety" species="chelifer" status="sp. nov." variety="crassodontus">
<emphasis id="B949B2149909FFDDFAE4FEB9FF62FEC9" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
var. 
<emphasis id="B949B2149909FFDDFE94FED5FE33FEC9" box="[296,444,331,357]" italics="true" pageId="10" pageNumber="351">crassodontus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A0F6F9909FFDDFE79FED5FE75FEC9" box="[453,506,331,357]" pageId="10" pageNumber="351" rank="species">n.sp.</taxonomicNameLabel>
// 
<collectorName id="26C80BD09909FFDDFDA0FED5FD3BFEC9" box="[540,692,331,357]" pageId="10" pageNumber="351">H.E. Bomans</collectorName>
BMNH(E) 1999-248 // NHMUK015534102”, examined through photographs provided by Keita Matsumoto
</materialsCitation>
.
</paragraph>
<paragraph id="8B826E069909FFDDFF7BFE0CFC73FD05" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
<emphasis id="B949B2149909FFDDFF7BFE0CFDBDFE00" bold="true" box="[199,562,402,428]" pageId="10" pageNumber="351">Additional material examined</emphasis>
(
<specimenCount id="9D3BA58F9909FFDDFDF8FE0DFD02FE01" box="[580,653,403,429]" count="12" pageId="10" pageNumber="351" type="male">12♂♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFD20FE0CFD5BFE01" box="[668,724,402,429]" count="5" pageId="10" pageNumber="351" type="female">5♀♀</specimenCount>
). 
<emphasis id="B949B2149909FFDDFD51FE0CFCEBFE00" bold="true" box="[749,868,402,428]" pageId="10" pageNumber="351">
<collectingCountry id="F32A2E969909FFDDFD51FE0CFCD1FE00" box="[749,862,402,428]" name="Thailand" pageId="10" pageNumber="351">Thailand</collectingCountry>
:
</emphasis>
<materialsCitation id="3B55645B9909FFDDFCD1FE0DFD37FE7D" collectingDate="2022-04-15" collectionCode="WPC" collectorName="Wuttipon Pathomwattananurak" country="Thailand" location="Boon Rawd" pageId="10" pageNumber="351" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai">
<specimenCount id="9D3BA58F9909FFDDFCD1FE0DFC1FFE01" box="[877,912,403,429]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFC1EFE0DFC6BFE01" box="[930,996,403,429]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFC46FE0DFB4DFE01" box="[1018,1218,403,429]" country="Thailand" name="Chiang Rai" pageId="10" pageNumber="351">Chiang Rai Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFB73FE0DFAD9FE01" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFB73FE0DFAD9FE01" box="[1231,1366,403,429]" country="Thailand" name="Boon Rawd" pageId="10" pageNumber="351" stateProvince="Chiang Rai">Boon Rawd</location>
farm, 
<date id="FF8348C69909FFDDFF2BFE28FE99FE7D" box="[151,278,438,465]" pageId="10" pageNumber="351" value="2022-04-15">
<collectingDate id="EFC7B12E9909FFDDFF2BFE28FE99FE7D" box="[151,278,438,465]" pageId="10" pageNumber="351" value="2022-04-15">15.IV.2022</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFE9FFE29FD0DFE7D" box="[291,642,439,465]" pageId="10" pageNumber="351">Wuttipon Pathomwattananurak</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B55645B9909FFDDFD7BFE29FEA2FE59" collectionCode="WPC" country="Thailand" location="Wiang Pa Pao Dist." pageId="10" pageNumber="351" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Chiang Rai">
<specimenCount id="9D3BA58F9909FFDDFD7BFE29FC8EFE7D" box="[711,769,439,465]" count="3" pageId="10" pageNumber="351" type="male">3♂♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFCACFE29FCBCFE7D" box="[784,819,439,465]" count="1" pageId="10" pageNumber="351" type="female">1♀</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFCF8FE29FC09FE7D" box="[836,902,439,465]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFC21FE29FBEAFE7D" box="[925,1125,439,465]" country="Thailand" name="Chiang Rai" pageId="10" pageNumber="351">Chiang Rai Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFBCEFE29FAD7FE7D" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFBCEFE29FAD7FE7D" box="[1138,1368,439,465]" country="Thailand" name="Wiang Pa Pao Dist." pageId="10" pageNumber="351" stateProvince="Chiang Rai">Wiang Pa Pao Dist.</location>
, local collector leg
</materialsCitation>
.; 
<materialsCitation id="3B55645B9909FFDDFEFDFE45FBB0FE59" box="[321,1087,474,501]" collectionCode="WPC" country="Thailand" location="Bo Kluea Dist." pageId="10" pageNumber="351" specimenCount="1" specimenCount-male="1" stateProvince="Nan">
<specimenCount id="9D3BA58F9909FFDDFEFDFE45FEEBFE59" box="[321,356,475,501]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFEC4FE45FE35FE59" box="[376,442,475,501]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFE6EFE45FDC5FE59" box="[466,586,475,501]" country="Thailand" name="Nan" pageId="10" pageNumber="351">Nan Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFDE5FE45FC82FE59" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFDE5FE45FC82FE59" box="[601,781,475,501]" country="Thailand" name="Bo Kluea Dist." pageId="10" pageNumber="351" stateProvince="Nan">Bo Kluea Dist.</location>
, 2015, local collector leg
</materialsCitation>
.; 
<materialsCitation id="3B55645B9909FFDDFBEFFE45FC92FDB5" collectingDate="2023-11-28" collectionCode="WPC" collectorName="Uraiwan" country="Thailand" location="Bua Yai Dist." pageId="10" pageNumber="351" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Nakhon Ratchasima">
<specimenCount id="9D3BA58F9909FFDDFBEFFE45FB02FE59" box="[1107,1165,475,501]" count="4" pageId="10" pageNumber="351" type="male">4♂♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFB22FE45FB59FE59" box="[1182,1238,475,501]" count="2" pageId="10" pageNumber="351" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFB55FE45FAA4FE59" box="[1257,1323,475,501]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFAFFFE45FED3FDB5" country="Thailand" name="Nakhon Ratchasima" pageId="10" pageNumber="351">Nakhon Ratchasima Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFED8FE61FE77FDB5" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFED8FE61FE77FDB5" box="[356,504,511,537]" country="Thailand" name="Bua Yai Dist." pageId="10" pageNumber="351" stateProvince="Nakhon Ratchasima">Bua Yai Dist.</location>
, 
<date id="FF8348C69909FFDDFDBDFE61FD0CFDB5" box="[513,643,511,537]" pageId="10" pageNumber="351" value="2023-11-28">
<collectingDate id="EFC7B12E9909FFDDFDBDFE61FD0CFDB5" box="[513,643,511,537]" pageId="10" pageNumber="351" value="2023-11-28">28.XI.2023</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFD30FE61FD63FDB5" box="[652,748,511,537]" pageId="10" pageNumber="351">Uraiwan</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B55645B9909FFDDFC9BFE61FDBEFD91" collectingDate="2022-07" collectionCode="WPC" collectorName="Qiao-Zhi Yang" country="Thailand" location="Khong Chiam Dist." pageId="10" pageNumber="351" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Ubon Ratchathani">
<specimenCount id="9D3BA58F9909FFDDFC9BFE61FCC4FDB5" box="[807,843,511,537]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFCEAFE61FC01FDB5" box="[854,910,511,537]" count="2" pageId="10" pageNumber="351" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFC26FE61FC53FDB5" box="[922,988,511,537]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFC51FE61FB77FDB5" box="[1005,1272,511,537]" country="Thailand" name="Ubon Ratchathani" pageId="10" pageNumber="351">Ubon Ratchathani Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFABDFE61FF5FFD91" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFABDFE61FF5FFD91" country="Thailand" name="Khong Chiam Dist." pageId="10" pageNumber="351" stateProvince="Ubon Ratchathani">Khong Chiam Dist.</location>
, 
<date id="FF8348C69909FFDDFF61FDBDFECAFD91" box="[221,325,547,573]" pageId="10" pageNumber="351" value="2022-07">
<collectingDate id="EFC7B12E9909FFDDFF61FDBDFECAFD91" box="[221,325,547,573]" pageId="10" pageNumber="351" value="2022-07">VII.2022</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFEEFFDBDFE73FD91" box="[339,508,547,573]" pageId="10" pageNumber="351">Qiao-Zhi Yang</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B55645B9909FFDDFD83FDBDFDA5FDCD" collectingDate="2020-12-04" collectionCode="WPC" collectorName="Suchatree Tangcharoenkitsakul" country="Thailand" location="Song Phi Nong" municipality="Tha Mai Dist." pageId="10" pageNumber="351" specimenCount="1" specimenCount-male="1" stateProvince="Chanthaburi">
<specimenCount id="9D3BA58F9909FFDDFD83FDBDFDEDFD91" box="[575,610,547,573]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFDCFFDBDFD3AFD91" box="[627,693,547,573]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFD77FDBDFC13FD91" box="[715,924,547,573]" country="Thailand" name="Chanthaburi" pageId="10" pageNumber="351">Chanthaburi Prov.</collectingRegion>
, 
<collectingMunicipality id="6BE6F47C9909FFDDFC15FDBDFBC2FD91" box="[937,1101,547,573]" pageId="10" pageNumber="351">Tha Mai Dist.</collectingMunicipality>
, 
<location id="8EE238DD9909FFDDFBE6FDBDFA82FD91" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFBE6FDBDFA82FD91" box="[1114,1293,547,573]" country="Thailand" municipality="Tha Mai Dist." name="Song Phi Nong" pageId="10" pageNumber="351" stateProvince="Chanthaburi">Song Phi Nong</location>
, 
<date id="FF8348C69909FFDDFAA6FDBDFA18FD91" box="[1306,1431,547,573]" pageId="10" pageNumber="351" value="2020-12-04">
<collectingDate id="EFC7B12E9909FFDDFAA6FDBDFA18FD91" box="[1306,1431,547,573]" pageId="10" pageNumber="351" value="2020-12-04">4.XII.2020</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFF2BFDD9FE79FDCD" box="[151,502,583,609]" pageId="10" pageNumber="351">Suchatree Tangcharoenkitsakul</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B55645B9909FFDDFD8BFDD9FEBDFD29" collectingDate="2022-08" collectionCode="WPC" collectorName="Qiao-Zhi Yang" country="Thailand" location="Pong Nam Ron Dist." pageId="10" pageNumber="351" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Chanthaburi">
<specimenCount id="9D3BA58F9909FFDDFD8BFDD9FDFEFDCD" box="[567,625,583,609]" count="3" pageId="10" pageNumber="351" type="male">3♂♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFDC2FDD9FD2EFDCD" box="[638,673,583,609]" count="1" pageId="10" pageNumber="351" type="female">1♀</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFD0DFDD9FD7CFDCD" box="[689,755,583,609]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFCB4FDD9FC58FDCD" box="[776,983,583,609]" country="Thailand" name="Chanthaburi" pageId="10" pageNumber="351">Chanthaburi Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFC5FFDD9FB5CFDCD" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFC5FFDD9FB5CFDCD" box="[995,1235,583,609]" country="Thailand" name="Pong Nam Ron Dist." pageId="10" pageNumber="351" stateProvince="Chanthaburi">Pong Nam Ron Dist.</location>
, 
<date id="FF8348C69909FFDDFB62FDD9FADEFDCD" box="[1246,1361,583,609]" pageId="10" pageNumber="351" value="2022-08">
<collectingDate id="EFC7B12E9909FFDDFB62FDD9FADEFDCD" box="[1246,1361,583,609]" pageId="10" pageNumber="351" value="2022-08">VIII.2022</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFAE1FDD9FF71FD29" pageId="10" pageNumber="351">Qiao-Zhi Yang</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B55645B9909FFDDFE82FDF5FF4EFD05" collectingDate="2023-10-20" collectionCode="WPC" collectorName="Chayaphol Phothaworn" country="Thailand" location="Nopphitam Dist." pageId="10" pageNumber="351" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nakhon Si Thammarat">
<specimenCount id="9D3BA58F9909FFDDFE82FDF5FEEDFD29" box="[318,354,619,645]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
, 
<specimenCount id="9D3BA58F9909FFDDFED3FDF5FE1DFD29" box="[367,402,619,645]" count="1" pageId="10" pageNumber="351" type="female">1♀</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFE1DFDF5FE6CFD29" box="[417,483,619,645]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingRegion id="49F9A0E49909FFDDFE4BFDF5FCB4FD29" box="[503,827,619,645]" country="Thailand" name="Nakhon Si Thammarat" pageId="10" pageNumber="351">Nakhon Si Thammarat Prov.</collectingRegion>
, 
<location id="8EE238DD9909FFDDFCFAFDF5FB8BFD29" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFCFAFDF5FB8BFD29" box="[838,1028,619,645]" country="Thailand" name="Nopphitam Dist." pageId="10" pageNumber="351" stateProvince="Nakhon Si Thammarat">Nopphitam Dist.</location>
, 
<date id="FF8348C69909FFDDFBACFDF5FB07FD29" box="[1040,1160,619,645]" pageId="10" pageNumber="351" value="2023-10-20">
<collectingDate id="EFC7B12E9909FFDDFBACFDF5FB07FD29" box="[1040,1160,619,645]" pageId="10" pageNumber="351" value="2023-10-20">20.X.2023</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFB28FDF5FA13FD29" box="[1172,1436,619,645]" pageId="10" pageNumber="351">Chayaphol Phothaworn</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B55645B9909FFDDFF7BFD11FC73FD05" box="[199,1020,655,681]" collectingDate="2023-10-11" collectionCode="WPC" collectorName="Somkit Boonmak" country="Myanmar" location="Ye" municipality="Mon State" pageId="10" pageNumber="351" specimenCount="1" specimenCount-male="1">
<emphasis id="B949B2149909FFDDFF7BFD11FEC7FD05" bold="true" box="[199,328,655,681]" pageId="10" pageNumber="351">
<collectingCountry id="F32A2E969909FFDDFF7BFD11FECCFD05" box="[199,323,655,681]" name="Myanmar" pageId="10" pageNumber="351">Myanmar</collectingCountry>
:
</emphasis>
<specimenCount id="9D3BA58F9909FFDDFEF3FD11FEFDFD05" box="[335,370,655,681]" count="1" pageId="10" pageNumber="351" type="male">1♂</specimenCount>
(
<collectionCode id="ED2CF6C39909FFDDFE3DFD11FE4CFD05" box="[385,451,655,681]" country="USA" httpUri="http://grbio.org/cool/hnai-3bfy" name="World Phytophthora Genetic Resource Collection" pageId="10" pageNumber="351">WPC</collectionCode>
), 
<collectingMunicipality id="6BE6F47C9909FFDDFE6BFD11FDC1FD05" box="[471,590,655,681]" pageId="10" pageNumber="351">Mon State</collectingMunicipality>
, 
<location id="8EE238DD9909FFDDFDE4FD11FDF6FD05" LSID="urn:lsid:plazi:treatment:0394DF10990AFFDDFF2BFA6BFA09F935:8EE238DD9909FFDDFDE4FD11FDF6FD05" box="[600,633,655,681]" country="Myanmar" municipality="Mon State" name="Ye" pageId="10" pageNumber="351">Ye</location>
, 
<date id="FF8348C69909FFDDFD3FFD11FD74FD05" box="[643,763,655,681]" pageId="10" pageNumber="351" value="2023-10-11">
<collectingDate id="EFC7B12E9909FFDDFD3FFD11FD74FD05" box="[643,763,655,681]" pageId="10" pageNumber="351" value="2023-10-11">11.X.2023</collectingDate>
</date>
, 
<collectorName id="26C80BD09909FFDDFCBBFD11FC43FD05" box="[775,972,655,681]" pageId="10" pageNumber="351">Somkit Boonmak</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3273D8D9909FFDDFF7BFD2CFA1DFD5D" pageId="10" pageNumber="351" type="distribution">
<paragraph id="8B826E069909FFDDFF7BFD2CFA1DFD5D" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
<emphasis id="B949B2149909FFDDFF7BFD2CFEEEFD60" bold="true" box="[199,353,690,716]" pageId="10" pageNumber="351">Distribution.</emphasis>
<collectingCountry id="F32A2E969909FFDDFED7FD2DFE28FD61" box="[363,423,691,717]" name="India" pageId="10" pageNumber="351">India</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFE0AFD2DFDB3FD61" box="[438,572,691,717]" name="Bangladesh" pageId="10" pageNumber="351">Bangladesh</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFDF6FD2DFD1FFD61" box="[586,656,691,717]" name="China" pageId="10" pageNumber="351">China</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFD22FD2DFD7CFD61" box="[670,755,691,717]" name="Taiwan" pageId="10" pageNumber="351">Taiwan</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFCBDFD2DFCFFFD61" box="[769,880,691,717]" name="Myanmar" pageId="10" pageNumber="351">Myanmar</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFCC2FD2DFC6FFD61" box="[894,992,691,717]" name="Vietnam" pageId="10" pageNumber="351">Vietnam</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFC51FD2DFBDCFD61" box="[1005,1107,691,717]" name="Thailand" pageId="10" pageNumber="351">Thailand</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFBDEFD2DFB14FD61" box="[1122,1179,691,717]" name="Laos" pageId="10" pageNumber="351">Laos</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFB16FD2DFAAFFD61" box="[1194,1312,691,717]" name="Cambodia" pageId="10" pageNumber="351">Cambodia</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFA93FD2DFA17FD61" box="[1327,1432,691,717]" name="Malaysia" pageId="10" pageNumber="351">Malaysia</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFF2BFD49FE84FD5D" box="[151,267,727,753]" name="Singapore" pageId="10" pageNumber="351">Singapore</collectingCountry>
, and 
<collectingCountry id="F32A2E969909FFDDFEFAFD49FE3CFD5D" box="[326,435,727,753]" name="Indonesia" pageId="10" pageNumber="351">Indonesia</collectingCountry>
(
<bibRefCitation id="EFAC13F79909FFDDFE7DFD49FD6AFD5D" author="Mizunuma, T. &amp; Nagai, T." box="[449,741,727,753]" pageId="10" pageNumber="351" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai 1994</bibRefCitation>
; 
<bibRefCitation id="EFAC13F79909FFDDFD4DFD49FC72FD5D" author="Pinratana, A. &amp; Maes, J. M." box="[753,1021,727,753]" pageId="10" pageNumber="351" refId="ref10072" refString="Pinratana, A. &amp; Maes, J. M. (2003) Lucanidae of Thailand. Sunprinting, Bangkok, 248 pp." type="book" year="2003">Pinratana &amp; Maes 2003</bibRefCitation>
; 
<bibRefCitation id="EFAC13F79909FFDDFBB5FD49FB02FD5D" author="Fujita, H." box="[1033,1165,727,753]" pageId="10" pageNumber="351" refId="ref9102" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
; 
<bibRefCitation id="EFAC13F79909FFDDFB24FD49FA09FD5D" author="Huang, H. &amp; Chen, C. C." box="[1176,1414,727,753]" pageId="10" pageNumber="351" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3273D8D9909FFDDFF7BFD65FA09F935" pageId="10" pageNumber="351" type="discussion">
<paragraph id="8B826E069909FFDDFF7BFD65FE88FCF1" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
<emphasis id="B949B2149909FFDDFF7BFD65FEB4FCB9" bold="true" box="[199,315,763,789]" pageId="10" pageNumber="351">Remarks.</emphasis>
<taxonomicName id="4C3D15859909FFDDFEFFFD65FE6AFCB9" authorityName="Macleay" authorityYear="1819" box="[323,485,763,789]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFEFFFD65FE6AFCB9" box="[323,485,763,789]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="EFAC13F79909FFDDFD0AFD65FCE6FCB9" author="MacLeay, W. S." box="[694,873,763,789]" pageId="10" pageNumber="351" refId="ref9658" refString="MacLeay, W. S. (1819) Horae entomologicae: or essays on the annulose animals. Vol. 1. Part 1. S. Bagster, London, 160 pp. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">Macleay (1819)</bibRefCitation>
from “Australasia” without more precise locality. The 
<typeStatus id="5486D0A49909FFDDFF71FC81FEABFC95" box="[205,292,799,825]" pageId="10" pageNumber="351" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="4C3D15859909FFDDFEF3FC81FE7BFC95" authorityName="Macleay" authorityYear="1819" box="[335,500,799,825]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFEF3FC81FE7BFC95" box="[335,500,799,825]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
(
<figureCitation id="130672839909FFDDFDBAFC81FD00FC95" box="[518,655,798,825]" captionStart="FIGURES 45–47" captionStartId="11.[151,264,943,967]" captionTargetBox="[154,1435,184,916]" captionTargetId="figure-17@11.[151,1435,181,919]" captionTargetPageId="11" captionText="FIGURES 45–47. Syntype of Aegus chelifer Macleay, 1819 (image ©BMNH/Keita Matsumoto). 45, dorsal habitus; 46, ventral habitus; 47, labels." figureDoi="http://doi.org/10.5281/zenodo.11234142" httpUri="https://zenodo.org/record/11234142/files/figure.png" pageId="10" pageNumber="351">Figs. 45–47</figureCitation>
) is a small size male that does not present the middle tooth on the mandible.
</paragraph>
<paragraph id="8B826E069909FFDDFF7BFCF9FF6DFBF5" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
There are three junior synonyms of 
<taxonomicName id="4C3D15859909FFDDFDE0FCF9FD5DFC2D" authorityName="Macleay" authorityYear="1819" box="[604,722,871,897]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFDE0FCF9FD5DFC2D" box="[604,722,871,897]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
, two of which have been treated as subspecies in some studies. 
<bibRefCitation id="EFAC13F79909FFDDFF2BFC15FEB0FC09" author="Boileau, H." box="[151,319,907,933]" pageId="10" pageNumber="351" pagination="319 - 322" refId="ref8526" refString="Boileau, H. (1899) Descriptions sommaires d'Aegus nouveaux. Bulletin de la Societe entomologique de France, 48, 319 - 322. https: // doi. org / 10.3406 / bsef. 1899.22441" type="journal article" year="1899">Boileau (1899)</bibRefCitation>
described 
<taxonomicName id="4C3D15859909FFDDFE0AFC15FDC3FC09" authorityName="Boileaui" authorityYear="1899" box="[438,588,907,933]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B949B2149909FFDDFE0AFC15FDC3FC09" box="[438,588,907,933]" italics="true" pageId="10" pageNumber="351">Aegus nitidus</emphasis>
</taxonomicName>
from Borneo which he later synonymized to 
<taxonomicName id="4C3D15859909FFDDFBFDFC15FA17FC09" authority="(Boileau, 1913)" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1089,1432,907,933]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFBFDFC15FB6EFC09" box="[1089,1249,907,933]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
(
<bibRefCitation id="EFAC13F79909FFDDFB52FC15FA1FFC09" author="Boileau, H." box="[1262,1424,907,933]" pageId="10" pageNumber="351" pagination="213 - 272" refId="ref8564" refString="Boileau, H. (1913) Note sur Lucanides conserves dans les collections de l'Universite d'Oxford et du British Museum. Transactions of the entomological Society London, 1913, 213 - 272. https: // doi. org / 10.1111 / j. 1365 - 2311.1913. tb 01776. x" type="journal article" year="1913">Boileau, 1913</bibRefCitation>
)
</taxonomicName>
. Subsequently, 
<bibRefCitation id="EFAC13F79909FFDDFE85FC31FDE7FC65" author="Mizunuma, T. &amp; Nagai, T." box="[313,616,943,969]" pageId="10" pageNumber="351" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai (1994)</bibRefCitation>
treated taxon 
<taxonomicName id="4C3D15859909FFDDFCB8FC31FCDFFC65" authorityName="Boileaui" authorityYear="1899" box="[772,848,943,969]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B949B2149909FFDDFCB8FC31FCDFFC65" box="[772,848,943,969]" italics="true" pageId="10" pageNumber="351">nitidus</emphasis>
</taxonomicName>
as a subspecies of 
<taxonomicName id="4C3D15859909FFDDFB9EFC31FB4DFC65" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1058,1218,943,969]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFB9EFC31FB4DFC65" box="[1058,1218,943,969]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
without any formal status revise or explanatory remark. Another junior synonym, 
<taxonomicName id="4C3D15859909FFDDFCDCFC4DFAD2FC41" authority="Kriesche, 1920" authorityName="Kriesche" authorityYear="1920" box="[864,1373,979,1005]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B2149909FFDDFCDCFC4DFB27FC41" box="[864,1192,979,1005]" italics="true" pageId="10" pageNumber="351">Aegus specularis tonkinensis</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFB0DFC4DFAD2FC41" author="Kriesche, R." box="[1201,1373,979,1005]" pageId="10" pageNumber="351" pagination="92 - 107" refId="ref9544" refString="Kriesche, R. (1920) Zur Kenntnis der Lucaniden. Archiv fur Naturgeschichte, 86 A (8), 92 - 107." type="journal article" year="1920">Kriesche, 1920</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F32A2E969909FFDDFF2BFC69FF66FBBD" box="[151,233,1015,1041]" name="Vietnam" pageId="10" pageNumber="351">Tonkin</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFF4BFC69FED6FBBD" box="[247,345,1015,1041]" name="Vietnam" pageId="10" pageNumber="351">Vietnam</collectingCountry>
, was synonymized as 
<taxonomicName id="4C3D15859909FFDDFDE1FC69FC8DFBBD" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[605,770,1015,1041]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFDE1FC69FC8DFBBD" box="[605,770,1015,1041]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFAC13F79909FFDDFC8DFC69FC54FBBD" author="Benesh, B." box="[817,987,1015,1041]" pageId="10" pageNumber="351" pagination="1 - 178" refId="ref8502" refString="Benesh, B. (1960) Lucanidae. Coleopterorum Catalogus, Supplementa, 2 nd Edition, 8, 1 - 178." type="journal article" year="1960">Benesh (1960)</bibRefCitation>
. 
<bibRefCitation id="EFAC13F79909FFDDFC56FC69FB61FBBD" author="Huang, H. &amp; Chen, C. C." box="[1002,1262,1015,1041]" pageId="10" pageNumber="351" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang &amp; Chen (2017)</bibRefCitation>
illustrated 
<specimenCount id="9D3BA58F9909FFDDFACEFC69FF76FB99" count="2" pageId="10" pageNumber="351" type="generic" typeStatus="syntype">two syntypes</specimenCount>
of 
<taxonomicName id="4C3D15859909FFDDFE9CFB85FDEBFB99" authorityName="Kriesche" authorityYear="1920" box="[288,612,1051,1077]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B2149909FFDDFE9CFB85FDEBFB99" box="[288,612,1051,1077]" italics="true" pageId="10" pageNumber="351">Aegus specularis tonkinensis</emphasis>
</taxonomicName>
and revised it to subspecies level of 
<taxonomicName id="4C3D15859909FFDDFBB8FB85FB29FB99" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1028,1190,1051,1077]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFBB8FB85FB29FB99" box="[1028,1190,1051,1077]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
without formal status revise.
</paragraph>
<paragraph id="8B826E069909FFDDFF7BFBFDFC8BFA9D" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
<bibRefCitation id="EFAC13F79909FFDDFF7BFBFDFEFAFBD1" author="Bomans, H. E." box="[199,373,1123,1149]" pageId="10" pageNumber="351" pagination="179 - 196" refId="ref8841" refString="Bomans, H. E. (1992) Considerations sur le genre Aegus Macleay et description d'especes nouvelles 3 eme note. (77 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 92 (2), 179 - 196." type="journal article" year="1992">Bomans (1992)</bibRefCitation>
later described 
<taxonomicName id="4C3D15859909FFDDFD94FBFDFCEBFBD1" authorityName="Pathomwattananurak &amp; Jiaranaisakul" authorityYear="2024" box="[552,868,1123,1149]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B2149909FFDDFD94FBFDFCEBFBD1" box="[552,868,1123,1149]" italics="true" pageId="10" pageNumber="351">Aegus chelifer crassodontus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F32A2E969909FFDDFC1BFBFDFC73FBD1" box="[935,1020,1123,1149]" name="Vietnam" pageId="10" pageNumber="351">Annam</collectingCountry>
, 
<collectingCountry id="F32A2E969909FFDDFBBBFBFDFBE9FBD1" box="[1031,1126,1123,1149]" name="Vietnam" pageId="10" pageNumber="351">Vietnam</collectingCountry>
with some of the 
<typeStatus id="5486D0A49909FFDDFA8DFBFDFA13FBD1" box="[1329,1436,1123,1149]" pageId="10" pageNumber="351" type="paratype">paratypes</typeStatus>
from northern 
<collectingCountry id="F32A2E969909FFDDFE82FB19FE2AFB0D" box="[318,421,1159,1185]" name="Thailand" pageId="10" pageNumber="351">Thailand</collectingCountry>
. However, the 
<typeStatus id="5486D0A49909FFDDFDEEFB19FD3BFB0D" box="[594,692,1159,1185]" pageId="10" pageNumber="351" type="holotype">holotype</typeStatus>
(
<figureCitation id="130672839909FFDDFD79FB19FCC1FB0D" box="[709,846,1158,1185]" captionStart="FIGURES 48–50" captionStartId="11.[151,264,1877,1901]" captionTargetBox="[168,1435,1027,1853]" captionTargetId="figure-39@11.[151,1435,1027,1853]" captionTargetPageId="11" captionText="FIGURES 48–50. Holotype of Aegus chelifer crassodontus Bomans, 1992 (image ©BMNH/Keita Matsumoto). 48, dorsal habitus; 49, ventral habitus; 50, labels." figureDoi="http://doi.org/10.5281/zenodo.11234146" httpUri="https://zenodo.org/record/11234146/files/figure.png" pageId="10" pageNumber="351">Figs. 48–50</figureCitation>
) shows high similarity to one of the 
<typeStatus id="5486D0A49909FFDDFB45FB19FAD4FB0D" box="[1273,1371,1159,1185]" pageId="10" pageNumber="351" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C3D15859909FFDDFA38FB19FE1FFB69" authorityName="Kriesche" authorityYear="1920" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B2149909FFDDFA38FB19FE1FFB69" italics="true" pageId="10" pageNumber="351">
<emphasis id="B949B2149909FFDDFA38FB19FA13FB0D" bold="true" box="[1412,1436,1159,1185]" italics="true" pageId="10" pageNumber="351">A.</emphasis>
specularis tonkinensis
</emphasis>
</taxonomicName>
, by the short-thickness of mandible with the large middle tooth. This character was used by 
<bibRefCitation id="EFAC13F79909FFDDFF2BFB51FECBFB45" author="Bomans, H. E." box="[151,324,1231,1257]" pageId="10" pageNumber="351" pagination="179 - 196" refId="ref8841" refString="Bomans, H. E. (1992) Considerations sur le genre Aegus Macleay et description d'especes nouvelles 3 eme note. (77 eme contribution a l'etude des Coleopteres Lucanides). Lambillionea, 92 (2), 179 - 196." type="journal article" year="1992">Bomans (1992)</bibRefCitation>
to distinguish 
<taxonomicName id="4C3D15859909FFDDFE5BFB51FD7AFB45" authorityName="Pathomwattananurak &amp; Jiaranaisakul" authorityYear="2024" box="[487,757,1231,1257]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B2149909FFDDFE5BFB51FD7AFB45" box="[487,757,1231,1257]" italics="true" pageId="10" pageNumber="351">A. chelifer crassodontus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C3D15859909FFDDFC8BFB51FC24FB45" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[823,939,1231,1257]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFC8BFB51FC24FB45" box="[823,939,1231,1257]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
. Based on the specimens examined from our collection and the series of 
<taxonomicName id="4C3D15859909FFDDFE78FB6DFDB7FAA1" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[452,568,1267,1293]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFE78FB6DFDB7FAA1" box="[452,568,1267,1293]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFAC13F79909FFDDFDE5FB6DFC08FAA1" author="Mizunuma, T. &amp; Nagai, T." box="[601,903,1267,1293]" pageId="10" pageNumber="351" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai (1994)</bibRefCitation>
and 
<bibRefCitation id="EFAC13F79909FFDDFC07FB6DFBC2FAA1" author="Fujita, H." box="[955,1101,1267,1293]" pageId="10" pageNumber="351" refId="ref9102" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita (2010)</bibRefCitation>
these characters are not stable and can be found in the sympatric area with 
<taxonomicName id="4C3D15859909FFDDFD34FA89FD72FA9D" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[648,765,1303,1329]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFD34FA89FD72FA9D" box="[648,765,1303,1329]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B826E069909FFDDFF7BFAA5FA09F935" blockId="10.[151,1437,150,1689]" pageId="10" pageNumber="351">
Overall, the morphological characters of the 
<typeStatus id="5486D0A49909FFDDFD04FAA5FC80FAF9" box="[696,783,1339,1365]" pageId="10" pageNumber="351" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="4C3D15859909FFDDFC8FFAA5FC5CFAF9" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[819,979,1339,1365]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFC8FFAA5FC5CFAF9" box="[819,979,1339,1365]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
</taxonomicName>
, the 
<typeStatus id="5486D0A49909FFDDFBB5FAA5FBE4FAF9" box="[1033,1131,1339,1365]" pageId="10" pageNumber="351" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C3D15859909FFDDFB32FAA5FA13FAF9" authorityName="Pathomwattananurak &amp; Jiaranaisakul" authorityYear="2024" box="[1166,1436,1339,1365]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B2149909FFDDFB32FAA5FA13FAF9" box="[1166,1436,1339,1365]" italics="true" pageId="10" pageNumber="351">A. chelifer crassodontus</emphasis>
</taxonomicName>
and the 
<typeStatus id="5486D0A49909FFDDFF49FAC1FEC3FAD5" box="[245,332,1375,1401]" pageId="10" pageNumber="351" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="4C3D15859909FFDDFECBFAC1FD1CFAD5" authorityName="Kriesche" authorityYear="1920" box="[375,659,1375,1401]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B2149909FFDDFECBFAC1FD1CFAD5" box="[375,659,1375,1401]" italics="true" pageId="10" pageNumber="351">A. specularis tonkinensis</emphasis>
</taxonomicName>
can be placed in between the variations of 
<taxonomicName id="4C3D15859909FFDDFB32FAC1FA89FAD5" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1166,1286,1375,1401]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFB32FAC1FA89FAD5" box="[1166,1286,1375,1401]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
, even within the variation of the populations distributed in 
<collectingCountry id="F32A2E969909FFDDFD1AFA1DFC85FA31" box="[678,778,1411,1437]" name="Thailand" pageId="10" pageNumber="351">Thailand</collectingCountry>
(figs. 1–5; 
<bibRefCitation id="EFAC13F79909FFDDFC2DFA1DFB4FFA31" author="Mizunuma, T. &amp; Nagai, T." box="[913,1216,1411,1437]" pageId="10" pageNumber="351" refId="ref9822" refString="Mizunuma, T. &amp; Nagai, T. (1994) The Lucanid beetles of the world. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai, 1994</bibRefCitation>
; 
<bibRefCitation id="EFAC13F79909FFDDFB72FA1DFAD2FA31" author="Fujita, H." box="[1230,1373,1411,1437]" pageId="10" pageNumber="351" refId="ref9102" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita, 2010</bibRefCitation>
). The variations of these type specimens can be found to occur in nearby locations with some intermediates between them. There are no stable morphological characters or distinct, separated populations which could be used to discriminate subspecies. Moreover, we have dissected the genitalia of males from several places in 
<collectingCountry id="F32A2E969909FFDDFB2EFA71FB79F9A5" box="[1170,1270,1519,1545]" name="Thailand" pageId="10" pageNumber="351">Thailand</collectingCountry>
and found no significant difference. In addition, according to 
<bibRefCitation id="EFAC13F79909FFDDFD07F98DFC5CF981" author="Huang, H. &amp; Chen, C. C." box="[699,979,1555,1581]" pageId="10" pageNumber="351" refId="ref9384" refString="Huang, H. &amp; Chen, C. C. (2017) Stag beetles of China III. Formosa Ecological Company, Taipei, 524 pp." type="book" year="2017">Huang and Chen (2017)</bibRefCitation>
, subspecific classification of 
<taxonomicName id="4C3D15859909FFDDFA99F98DFA12F981" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1317,1437,1555,1581]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFA99F98DFA12F981" box="[1317,1437,1555,1581]" italics="true" pageId="10" pageNumber="351">A. chelifer</emphasis>
</taxonomicName>
should not be clarified due to the precise type locality being unknown. From the aforementioned reasons, we thereby confirm the synonym status of 
<taxonomicName id="4C3D15859909FFDDFE4BF9C5FCBAF9D9" authority="Boileau, 1899" authorityName="Boileaui" authorityYear="1899" box="[503,821,1627,1653]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B949B2149909FFDDFE4BF9C5FD00F9D9" box="[503,655,1627,1653]" italics="true" pageId="10" pageNumber="351">Aegus nitidus</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFD2BF9C5FCBAF9D9" author="Boileau, H." box="[663,821,1627,1653]" pageId="10" pageNumber="351" pagination="319 - 322" refId="ref8526" refString="Boileau, H. (1899) Descriptions sommaires d'Aegus nouveaux. Bulletin de la Societe entomologique de France, 48, 319 - 322. https: // doi. org / 10.3406 / bsef. 1899.22441" type="journal article" year="1899">Boileau, 1899</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3D15859909FFDDFCD2F9C5FAE7F9D9" authority="Kriesche, 1920" authorityName="Kriesche" authorityYear="1920" box="[878,1384,1627,1653]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="specularis" subSpecies="tonkinensis">
<emphasis id="B949B2149909FFDDFCD2F9C5FB3CF9D9" box="[878,1203,1627,1653]" italics="true" pageId="10" pageNumber="351">Aegus specularis tonkinensis</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFB07F9C5FAE7F9D9" author="Kriesche, R." box="[1211,1384,1627,1653]" pageId="10" pageNumber="351" pagination="92 - 107" refId="ref9544" refString="Kriesche, R. (1920) Zur Kenntnis der Lucaniden. Archiv fur Naturgeschichte, 86 A (8), 92 - 107." type="journal article" year="1920">Kriesche, 1920</bibRefCitation>
</taxonomicName>
, and treat 
<taxonomicName id="4C3D15859909FFDDFF6CF9E1FD3BF935" authority="Bomans, 1993" authorityName="Bomans" authorityYear="1993" box="[208,692,1663,1689]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="crassodontus">
<emphasis id="B949B2149909FFDDFF6CF9E1FD83F935" box="[208,524,1663,1689]" italics="true" pageId="10" pageNumber="351">Aegus chelifer crassodontus</emphasis>
Bomans, 1993
</taxonomicName>
as new junior synonym of 
<taxonomicName id="4C3D15859909FFDDFC59F9E1FABDF935" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[997,1330,1663,1689]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B949B2149909FFDDFC59F9E1FB08F935" box="[997,1159,1663,1689]" italics="true" pageId="10" pageNumber="351">Aegus chelifer</emphasis>
<bibRefCitation id="EFAC13F79909FFDDFB31F9E1FABDF935" author="MacLeay, W. S." box="[1165,1330,1663,1689]" pageId="10" pageNumber="351" refId="ref9658" refString="MacLeay, W. S. (1819) Horae entomologicae: or essays on the annulose animals. Vol. 1. Part 1. S. Bagster, London, 160 pp. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
herein.
</paragraph>
</subSubSection>
</treatment>
</document>