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<document id="04649D830A05DD12C97037D4128A822E" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629612136760" checkinUser="felipe" docAuthor="Ballantyne, Lesley A. &amp; Lambkin, Christine" docDate="2009" docId="0394D665BE31FFA3FF3C510626BDEB22" docLanguage="en" docName="zt01997p188.pdf" docOrigin="Zootaxa 1997" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Asymmetricata Ballantyne &amp; Lambkin 2009, gen. n." docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFADAE1DBE2CFFBCFFAB57492227EE30" masterDocTitle="Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)" masterLastPageNumber="188" masterPageNumber="1" pageNumber="30" updateTime="1699083011144" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1C7077CA5D0672BF9C93DB83950A0534">Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="EA319058E982B8A9BB5F2EB4A6A543A9">Ballantyne, Lesley A.</mods:namePart>
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<mods:namePart id="B60313498B3EDA9C74898465388BC5EA">Lambkin, Christine</mods:namePart>
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<treatment id="0394D665BE31FFA3FF3C510626BDEB22" ID-DOI="http://doi.org/10.5281/zenodo.5324230" ID-Zenodo-Dep="5324230" LSID="urn:lsid:plazi:treatment:0394D665BE31FFA3FF3C510626BDEB22" httpUri="http://treatment.plazi.org/id/0394D665BE31FFA3FF3C510626BDEB22" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="C32734F8BE31FFA1FF3C51062382E859" box="[151,421,1615,1641]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8B826773BE31FFA1FF3C51062382E859" blockId="29.[151,421,1615,1676]" box="[151,421,1615,1641]" pageId="29" pageNumber="30">
<heading id="D0CAD01FBE31FFA1FF3C51062382E859" bold="true" box="[151,421,1615,1641]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B949BB61BE31FFA1FF3C51062382E859" bold="true" box="[151,421,1615,1641]" pageId="29" pageNumber="30">
<taxonomicName id="4C3D1CF0BE31FFA1FF3C5106236BE859" ID-CoL="34RG" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,332,1615,1641]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE31FFA1FF3C5106236BE859" bold="true" box="[151,332,1615,1641]" italics="true" pageId="29" pageNumber="30">Asymmetricata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A061ABE31FFA1FEF951062382E859" box="[338,421,1615,1641]" pageId="29" pageNumber="30" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE31FFA1FF3C513B23A3E8BC" box="[151,388,1650,1676]" pageId="29" pageNumber="30" type="description">
<paragraph id="8B826773BE31FFA1FF3C513B23A3E8BC" blockId="29.[151,421,1615,1676]" box="[151,388,1650,1676]" pageId="29" pageNumber="30">
(
<figureCitation id="13067BF6BE31FFA1FF34513B232EE8BC" box="[159,265,1650,1676]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="29" pageNumber="30">Figs 58</figureCitation>
, 108114)
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE31FFA1FF3C51F42085E8E7" box="[151,674,1725,1751]" pageId="29" pageNumber="30" type="type_taxon">
<paragraph id="8B826773BE31FFA1FF3C51F42085E8E7" blockId="29.[151,1436,1725,2031]" box="[151,674,1725,1751]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF3C51F4231CE8E7" bold="true" box="[151,315,1725,1751]" pageId="29" pageNumber="30">
<typeStatus id="5486D9D1BE31FFA1FF3C51F422F3E8E7" box="[151,212,1725,1751]" pageId="29" pageNumber="30">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3D1CF0BE31FFA1FEE851F42085E8E7" ID-CoL="3WCPN" authority="(Motsch.)" baseAuthorityName="Motsch." box="[323,674,1725,1751]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="circumdata">
<emphasis id="B949BB61BE31FFA1FEE851F42000E8E6" box="[323,551,1725,1750]" italics="true" pageId="29" pageNumber="30">Luciola circumdata</emphasis>
(Motsch.)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE31FFA3FF6D51AC23E7EDB2" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8B826773BE31FFA1FF6D51AC2395E97F" blockId="29.[151,1436,1725,2031]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6D51AC2319E8CF" bold="true" box="[198,318,1765,1791]" pageId="29" pageNumber="30">Diagnosis</emphasis>
. Dorsal surface, especially shape of pronotum, similar to many
<taxonomicName id="4C3D1CF0BE31FFA1FB9051AC268CE8CE" authorityName="Olliff, Madagascan" authorityYear="1890" box="[1083,1195,1765,1790]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE31FFA1FB9051AC268CE8CE" box="[1083,1195,1765,1790]" italics="true" pageId="29" pageNumber="30">Atyphella</emphasis>
</taxonomicName>
; colour very like
<taxonomicName id="4C3D1CF0BE31FFA1FA2A51AC22F5E916" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="wolfi">
<emphasis id="B949BB61BE31FFA1FA2A51AC22F5E916" italics="true" pageId="29" pageNumber="30">C. wolfi</emphasis>
</taxonomicName>
from above; distinguished from all other
<taxonomicName id="4C3D1CF0BE31FFA1FD6F50442166E917" authorityName="sensu McDermott" authorityYear="1966" box="[708,833,1805,1831]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="3" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
by the asymmetrical tergite 8 which is emarginate on its right side (
<figureCitation id="13067BF6BE31FFA1FEF7507C2383E97F" box="[348,420,1845,1871]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="29" pageNumber="30">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8B826773BE31FFA1FF6C501421A8E9AF" blockId="29.[151,1436,1725,2031]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6C5014232AE947" bold="true" box="[199,269,1885,1911]" pageId="29" pageNumber="30">Male.</emphasis>
Pronotum (
<figureCitation id="13067BF6BE31FFA1FE37501423C4E947" box="[412,483,1885,1911]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="29" pageNumber="30">Fig. 5</figureCitation>
) dorsal surface lacking irregularities in posterolateral areas, longitudinal groove in lateral areas; punctation dense. Anterior margin not explanate.
</paragraph>
<paragraph id="8B826773BE31FFA2FF6C50E4219BEF1A" blockId="29.[151,1436,1725,2031]" lastBlockId="30.[151,1436,152,2018]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Pronotum wider across posterior area than rest, pronotal width greater than humeral width. Anterolateral corners rounded obtuse; lateral margins in anterior half divergent; lateral margins in posterior half diverge then converge with rounded convergence; lacking indentation at mid-point, or sinuousity in either horizontal or vertical plane; lacking indentation in lateral margin near posterolateral corner, and irregularities at corner; posterolateral corners rounded; rounded corners obtuse, posterolateral corners project as far as or beyond median posterior margin, separated from it by shallow emargination.</paragraph>
<paragraph id="8B826773BE32FFA2FF6C567127B1EF4A" blockId="30.[151,1436,152,2018]" pageId="30" pageNumber="31">Hypomera closed. Median area of hypomeron not elevated vertically; anterior area of hypomeron not flattened, posterior area of hypomeron widely flattened and strongly adpressed; pronotal width/ GHW index 1.6.</paragraph>
<paragraph id="8B826773BE32FFA2FF6C56C123FFECA2" blockId="30.[151,1436,152,2018]" pageId="30" pageNumber="31">
Elytron (
<figureCitation id="13067BF6BE32FFA2FE8556C12353EF92" box="[302,372,392,418]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="30" pageNumber="31">Fig. 5</figureCitation>
) punctation dense, not linear, not as large as pronotum, nor widely and evenly spaced; apices not deflexed; epipleuron and suture extending beyond mid-point almost to apex, not extending as ridge around apex; no expansion of epipleuron or sutural ridge in apical half; 2 interstitial lines, inner two do not exceed suture; elytral carina absent; in horizontal specimen viewed from beneath epipleuron at elytral base wide, covers humerus; viewed from above arises anterior to posterior margin of MS; epipleuron developed as lateral ridge along most of length; sutural margins approximate along most of their length in closed elytra; lateral margins convexsided.
</paragraph>
<paragraph id="8B826773BE32FFA2FF6C55E92639EA12" blockId="30.[151,1436,152,2018]" pageId="30" pageNumber="31">
Head (
<figureCitation id="13067BF6BE32FFA2FEBE55E9237BEC8A" box="[277,348,672,698]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="30" pageNumber="31">Fig. 6</figureCitation>
) moderately depressed between eyes; moderately exposed in front of pronotum, not capable of complete retraction within prothoracic cavity; eyes moderately separated beneath at level of posterior margin of mouthpart complex; eyes above labrum close to moderately separated; frons-vertex junction rounded, with no median elevation; posterolateral eye excavation not strongly developed, not visible in resting head position; antennal sockets on head between eyes, not contiguous and separated by &lt;ASW; clypeolabral suture present, flexible, not in front of anterior eye margin when head viewed with labrum horizontal; outer edges of labrum reach inner edges of closed mandibles. Mouthparts functional; apical segment of labial palpi nonlunate, strongly flattened, wide triangular shape with inner edge dentate. Antennae 11 segmented; length&gt; GHW to twice GHW; no segments flattened, shortened, or expanded; pedicel not produced; FS1 not shorter than pedicel; in very short antennae FS may be subequal in length and width.
</paragraph>
<paragraph id="8B826773BE32FFA2FF6C53792033EA42" blockId="30.[151,1436,152,2018]" pageId="30" pageNumber="31">
Legs (
<figureCitation id="13067BF6BE32FFA2FEB85379237CEA7A" box="[275,347,1072,1098]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="30" pageNumber="31">Fig. 6</figureCitation>
) with inner tarsal claw not split; lacking MFC; no femora or tibiae swollen or curved; no basitarsi expanded or excavated.
</paragraph>
<paragraph id="8B826773BE32FFA2FF6C53C9208FE972" blockId="30.[151,1436,152,2018]" pageId="30" pageNumber="31">
Abdomen (
<figureCitation id="13067BF6BE32FFA2FEE653C9239CEAAA" box="[333,443,1152,1178]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="30" pageNumber="31">Figs 6, 7</figureCitation>
, 108111) lacking cuticular remnants in association with aedeagal sheath; no ventrites with curved posterior margins nor extending anteriorly into emarginated posterior margin of anterior segment; LO in V7 entire or bipartite, reaching to sides but not to posterior margin; both entire and bipartite LO occupying most of V7; anterior margin of entire LO in V7 narrowly medially emarginate; posterior half of V7 not arched or swollen, muscle impressions not visible here; LO present in V6, occupying almost all V6. MPP present, symmetrical or asymmetrical; medially shallowly emarginate or not; non-emarginate MPP apex rounded, entire; not laterally compressed, short or L=W, slightly inclined dorsally, sometimes engulfed by T8 apex, lacking dorsal ridge, dorsal median longitudinal trough. V7 lacking median carina, PLP, incurving lobes or pointed projections, median dimple, or reflexed lobes; V7 with median longitudinal trough between halves of bipartite LO, anteromedian face of entire LO with narrow longitudinal depression. T7 lacking prolonged posterolateral corners. T8 strongly sclerotised, asymmetrical, emarginated on right side, L&gt;W, visible posterior area not narrowing abruptly, lacking prolonged posterolateral corners, median posterior emargination, median posterior projections; inclining ventrally, engulfing MPP, not extending conspicuously beyond posterior margin of V7. T8 ventral surface lacking flanges, lateral depressed troughs, median longitudinal trough, asymmetrical projections, median posterior ridge; T8 with concealed anterolateral arms as long as visible posterior portion of T8, narrow in horizontal plane and expanded dorsoventrally, not laterally emarginated before their origins, apices lacking bifurcation of inner margin and bases lacking ventrally directed pieces; lateral margins of T8 not enfolding sides of V7.
</paragraph>
<paragraph id="8B826773BE32FFA3FF6C501923CBEF32" blockId="30.[151,1436,152,2018]" lastBlockId="31.[151,1436,152,1298]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">Aedeagal sheath (Figs 115117) never&gt; 4 times as long as wide; lacking paraprocts; symmetrical in posterior area where sheath sternite subparallel-sided along length; sternite not angulate on L or R sides, posterior area not evenly emarginated on either side preapically, posterior margin of sternite broadly and slightly asymmetrically emarginated; anterior half of sternite broad, apically rounded; tergite lateral arms extending narrowly anteriorly to sides of sheath sternite; tergite not subdivided, lacking projecting pieces along posterior margin of tergite 9, anterior margin of tergite darker than rest with lateral margins narrowly darker (wider and slightly longer on right side).</paragraph>
<paragraph id="8B826773BE33FFA3FF6C565923DEECA2" blockId="31.[151,1436,152,1298]" pageId="31" pageNumber="32">Aedeagus (Figs 112114) L/W 3/1; LL lacking lateral appendages, visible from beneath at sides of ML, LL/ML wide; LL of equal length, slightly shorter than ML, diverging along inner margins, separated there by&gt; half their length; LL base width not = LL apex width; LL apex width subequal to ML width; LL apices not expanded horizontally; dorsal base of LL asymmetrical, not excavated, anterior margin prolonged to left; LL lacking lateral hairy appendages along outer ventral margins, margins not produced preapically nor narrowly on inner apical margin, not obliquely truncate along preapical inner margins; apices of LL not inturned, or outturned; lacking projection on left LL only; inner margins lacking slender leaf-like projection; ML symmetrical, lacking paired lateral teeth, tooth to left side; not strongly arched, apex not shaped like arrowhead, not bulbous, not inclined ventrally; BP hooded, not very narrow, not strongly sclerotised, not strongly emarginated along anterior margin.</paragraph>
<paragraph id="8B826773BE33FFA3FF6C55D621CBED6A" blockId="31.[151,1436,152,1298]" pageId="31" pageNumber="32">
<emphasis id="B949BB61BE33FFA3FF6C55D6230EEC89" bold="true" box="[199,297,671,697]" pageId="31" pageNumber="32">Female.</emphasis>
Macropterous in
<taxonomicName id="4C3D1CF0BE33FFA3FE5055E92049EC89" box="[507,622,672,697]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="B949BB61BE33FFA3FE5055E92049EC89" box="[507,622,672,697]" italics="true" pageId="31" pageNumber="32">As. ovalis</emphasis>
</taxonomicName>
(
<figureCitation id="13067BF6BE33FFA3FDD455E920F2EC8A" box="[639,725,672,698]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="31" pageNumber="32">Fig. 14</figureCitation>
), assumed capable of flight. Pronotum lacking irregularities in posterolateral areas; punctation moderate to dense; pronotum&gt; humeral width; indentation of lateral margin, irregularities at posterolateral corner, absent; outline similar to male. Elytral punctation not as large as pronotum nor evenly spaced; 2 interstitial lines; elytral carina absent. No legs or parts thereof swollen and/or curved. LO in V6 only, V7 lacking any elevations, depressions or ridges.
</paragraph>
<paragraph id="8B826773BE33FFA3FF6C542123E7EDB2" blockId="31.[151,1436,152,1298]" box="[199,448,872,898]" pageId="31" pageNumber="32">Larva not associated.</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE33FFA3FF6C54C62157EDCA" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8B826773BE33FFA3FF6C54C62157EDCA" blockId="31.[151,1436,152,1298]" pageId="31" pageNumber="32">
<emphasis id="B949BB61BE33FFA3FF6C54C6236BED99" bold="true" box="[199,332,911,937]" pageId="31" pageNumber="32">Etymology</emphasis>
.
<taxonomicName id="4C3D1CF0BE33FFA3FEF754D9202BED99" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[348,524,912,937]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE33FFA3FEF754D9202BED99" box="[348,524,912,937]" italics="true" pageId="31" pageNumber="32">Asymmetricata</emphasis>
</taxonomicName>
is a feminine noun latinised from the English word asymmetrical. If the last two syllables (at-a) are pronounced with the first a sounded ay (as in hay), then the generic name aptly evokes the distinguishing feature of an asymmetrical tergite 8.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE33FFA3FF6C534E26BDEB22" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8B826773BE33FFA3FF6C534E26BDEB22" blockId="31.[151,1436,152,1298]" pageId="31" pageNumber="32">
<emphasis id="B949BB61BE33FFA3FF6C534E231EEA11" bold="true" box="[199,313,1031,1057]" pageId="31" pageNumber="32">Remarks</emphasis>
. Ballantyne included representatives of all recognised species groups within the
<taxonomicName id="4C3D1CF0BE33FFA3FB5F53412756EA12" authorityName="sensu McDermott" authorityYear="1966" box="[1268,1393,1032,1058]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="3" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
for this analysis, including these species. The
<taxonomicName id="4C3D1CF0BE33FFA3FD3C53792120EA79" authorityName="Olliff, Madagascan" authorityYear="1890" box="[663,775,1072,1097]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE33FFA3FD3C53792120EA79" box="[663,775,1072,1097]" italics="true" pageId="31" pageNumber="32">Atyphella</emphasis>
</taxonomicName>
complex was shown to be paraphyletic, with the clear separation of these two species from
<taxonomicName id="4C3D1CF0BE33FFA3FDFF531120E3EA41" authorityName="Olliff, Madagascan" authorityYear="1890" box="[596,708,1112,1137]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE33FFA3FDFF531120E3EA41" box="[596,708,1112,1137]" italics="true" pageId="31" pageNumber="32">Atyphella</emphasis>
</taxonomicName>
. This new genus is however based on a very restricted treatment and hindered by lack of specimens, and Ballantynes inability to borrow
<typeStatus id="5486D9D1BE33FFA3FB9953C92643EAAA" box="[1074,1124,1152,1178]" pageId="31" pageNumber="32">type</typeStatus>
material of
<taxonomicName id="4C3D1CF0BE33FFA3FB5E53C9277DEAA9" box="[1269,1370,1152,1177]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="B949BB61BE33FFA3FB5E53C9277DEAA9" box="[1269,1370,1152,1177]" italics="true" pageId="31" pageNumber="32">L. ovalis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFAC1A82BE33FFA3FAC253C92374EAF2" author="Ballantyne, L. A." pageId="31" pageNumber="32" pagination="171 - 188" refId="ref102796" refString="Ballantyne, L. A. (1987 b) Lucioline Morphology, Taxonomy and Behaviour: A Reappraisal. (Coleoptera, Lampyridae). Transactions of the American Entomological Society 113, 171 - 188." type="journal article" year="1987">Ballantyne (1987b)</bibRefCitation>
described the asymmetrical T
<quantity id="4CC5CA96BE33FFA3FD1553E120CCEAF2" box="[702,747,1192,1218]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="31" pageNumber="32" unit="in" value="8.0">8 in</quantity>
males in
<taxonomicName id="4C3D1CF0BE33FFA3FCC953E1266EEAF1" box="[866,1097,1192,1217]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="circumdata">
<emphasis id="B949BB61BE33FFA3FCC953E1266EEAF1" box="[866,1097,1192,1217]" italics="true" pageId="31" pageNumber="32">Luciola circumdata</emphasis>
</taxonomicName>
and suggested the aedeagus would have to be extracted dorsally. This feature is doubtless more widespread than this restricted treatment allows, and it is probable that
<taxonomicName id="4C3D1CF0BE33FFA3FE5C53B120FEEB22" authority="Olivier" authorityName="Olivier" box="[503,729,1272,1298]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="impressa">
<emphasis id="B949BB61BE33FFA3FE5C53B12059EB21" box="[503,638,1272,1297]" italics="true" pageId="31" pageNumber="32">L. impressa</emphasis>
Olivier
</taxonomicName>
and
<taxonomicName id="4C3D1CF0BE33FFA3FCB953B12626EB22" authority="Walker" authorityName="Walker" box="[786,1025,1272,1298]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B949BB61BE33FFA3FCB953B12181EB21" box="[786,934,1272,1297]" italics="true" pageId="31" pageNumber="32">L. humeralis</emphasis>
Walker
</taxonomicName>
belong here.
</paragraph>
</subSubSection>
</treatment>
</document>