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<figureCitation id="13012A75B832FFE4FB8250C7FB0FFC1A" box="[1090,1178,920,943]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="4.[151,186,2055,2076]" captionStartId-1="5.[152,187,2026,2047]" captionStartId-2="6.[151,186,1996,2017]" captionTargetBox-0="[151,1431,1166,2014]" captionTargetBox-1="[155,1435,1130,1985]" captionTargetBox-2="[175,1409,257,1953]" captionTargetId-0="figure-335@4.[151,1431,1166,2014]" captionTargetId-1="figure-411@5.[155,1435,1130,1985]" captionTargetId-2="figure-6@6.[175,1409,257,1953]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 1. The new record of Lycodon pictus (VNMN 011227) from Vietnam in life. Photograph by Tao Thien Nguyen." captionText-1="Fig. 2. New country record of Lycodon pictus (CIB 115609) from China (Nonggang, Guangxi), adult female in life. Photograph by Jin-Long Ren." captionText-2="Fig. 3. Adult female of Lycodon pictus (CIB 115609; Guangxi, China) in preservative. (A) Dorsal general view; (B) Ventral general view; (C) Lateral head view, right side; (D) Lateral head view, left side; (E) Dorsal view of head; (F) Ventral view of head; (G) Lateral body view; (H) Ventral scale view. Scale bar 5 mm.Photographs by Jin-Long Ren." figureDoi-0="http://doi.org/10.5281/zenodo.5743938" figureDoi-1="http://doi.org/10.5281/zenodo.5743940" figureDoi-2="http://doi.org/10.5281/zenodo.5743942" httpUri-0="https://zenodo.org/record/5743938/files/figure.png" httpUri-1="https://zenodo.org/record/5743940/files/figure.png" httpUri-2="https://zenodo.org/record/5743942/files/figure.png" pageId="4" pageNumber="416">Figs 1-3</figureCitation>
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New record for 
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(n = 1):
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), Ha Lang, 
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, 
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22°42’50’’ N, 106°40’74’’ E, 
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a.s.l., collected by 
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on 
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Fig. 1. The new record of 
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(VNMN 011227) from Vietnam in life. Photograph by Tao Thien Nguyen.
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New record for 
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(n = 1):
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(field no. GX2019069; 
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<location id="8EE5602BB833FFE5FDCC527DFEDDFEEF" LSID="urn:lsid:plazi:treatment:039387E6B832FFE8FC8F5009FAB1FD4C:8EE5602BB833FFE5FDCC527DFEDDFEEF" country="China" county="Longzhou County" latitude="22.467756" longLatPrecision="1" longitude="106.95521" name="Nonggang National Nature Reserve" pageId="5" pageNumber="417" stateProvince="Guangxi Zhuang">Nonggang National Nature Reserve</location>
, 
<collectingCounty id="62E44E7CB833FFE5FE9A521CFDB0FEEF" box="[346,549,323,346]" pageId="5" pageNumber="417">Longzhou County</collectingCounty>
, 
<collectingRegion id="49FEF812B833FFE5FDF7521CFD61FEEF" box="[567,756,323,346]" country="China" name="Guangxi" pageId="5" pageNumber="417">Guangxi Zhuang</collectingRegion>
<location id="8EE5602BB833FFE5FF57523CFE01FECF" LSID="urn:lsid:plazi:treatment:039387E6B832FFE8FC8F5009FAB1FD4C:8EE5602BB833FFE5FF57523CFE01FECF" box="[151,404,355,378]" country="China" county="Longzhou County" latitude="22.467756" longLatPrecision="1" longitude="106.95521" municipality="Nonggang National Nature Reserve" name="Autonomous Region" pageId="5" pageNumber="417" stateProvince="Guangxi Zhuang">Autonomous Region</location>
, China, 
<geoCoordinate id="EE0E5037B833FFE5FDDC523CFD65FECF" box="[540,752,355,378]" degrees="106" direction="east" minutes="57" orientation="longitude" pageId="5" pageNumber="417" precision="1" seconds="18.77" value="106.95521">106°57’18.77” E</geoCoordinate>
, 
<geoCoordinate id="EE0E5037B833FFE5FF5752DCFEACFE2F" box="[151,313,387,410]" degrees="22" direction="north" minutes="28" orientation="latitude" pageId="5" pageNumber="417" precision="1" seconds="3.92" value="22.467756">22°28’3.92” N</geoCoordinate>
, 
<quantity id="4CC29B15B833FFE5FE8652DCFE1EFE2F" box="[326,395,387,410]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="5" pageNumber="417" unit="m" value="233.0">
<elevation id="0017D1C3B833FFE5FE8652DCFE1EFE2F" box="[326,395,387,410]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="5" pageNumber="417" unit="m" value="233.0">233 m</elevation>
</quantity>
a.s.l., collected by 
<collectorName id="26CF5326B833FFE5FDA052DCFD61FE2F" box="[608,756,387,410]" pageId="5" pageNumber="417">Ren Jin-Long</collectorName>
and 
<collectorName id="26CF5326B833FFE5FF0452FBFEFDFE0E" box="[196,360,420,443]" pageId="5" pageNumber="417">Zeng Fan-Xing</collectorName>
on 
<date id="FF841030B833FFE5FE5052FBFD85FE0E" box="[400,528,420,443]" pageId="5" pageNumber="417" value="2019-07-08">
<collectingDate id="EFC0E9D8B833FFE5FE5052FBFD85FE0E" box="[400,528,420,443]" pageId="5" pageNumber="417" value="2019-07-08">8 July 2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320657BB833FFE8FF57528CFDBBFE6C" lastPageId="8" lastPageNumber="420" pageId="5" pageNumber="417" type="description">
<paragraph id="8B8536F0B833FFE5FF57528CFAC2FCAB" blockId="5.[151,756,467,1072]" lastBlockId="5.[831,1437,258,960]" pageId="5" pageNumber="417">
<emphasis id="B94EEAE2B833FFE5FF57528CFDDDFE5F" bold="true" box="[151,584,467,490]" pageId="5" pageNumber="417">Description of the Chinese specimen:</emphasis>
Head elongate, moderately distinct from neck, rather flattened, longer than wide, snout narrow, distinctly shorter than head width; nostril large, lateral, located in the middle of the nasal; eye large, pupils vertically elliptic; rostral triangular, much broader than high, hardly visible from above; nasal divided, narrowed medially; two internasals, anteriorly round, slightly wider than high, bordered by two large, hexagonal prefrontals posteriorly; frontal single, enlarged, pentagonal to hexagonal, narrowed posteriorly; parietals large, longer than wide, in contact with each other medially, with upper anterior and posterior temporals, paraparietal laterally and four nuchal scales posteriorly; paraparietals elongated, anterior part widened, about equal in length or slightly longer than posterior temporals; loreal 1/1, elongate, not entering orbit on right side, whereas the posterior corner entering orbit on the left side (
<figureCitation id="13012A75B833FFE5FD6050A6FF77FB85" captionStart="Fig" captionStartId="6.[151,186,1996,2017]" captionTargetBox="[175,1409,257,1953]" captionTargetId="figure-6@6.[175,1409,257,1953]" captionTargetPageId="6" captionText="Fig. 3. Adult female of Lycodon pictus (CIB 115609; Guangxi, China) in preservative. (A) Dorsal general view; (B) Ventral general view; (C) Lateral head view, right side; (D) Lateral head view, left side; (E) Dorsal view of head; (F) Ventral view of head; (G) Lateral body view; (H) Ventral scale view. Scale bar 5 mm.Photographs by Jin-Long Ren." figureDoi="http://doi.org/10.5281/zenodo.5743942" httpUri="https://zenodo.org/record/5743942/files/figure.png" pageId="5" pageNumber="417">Fig. 3C and D</figureCitation>
); supralabials 8/8, first and second in contact with nasal, third to fifth entering orbit, sixth largest, slightly higher than third on left side; infralabials 10/9, first pair in broad contact with each other, first to fifth/first to fourth in contact with anterior pair of chin shields; anterior and posterior pairs of chin shields elongate, second pair not meeting in midline; preocular 1/1, located on antero-upper part of eye; postoculars 2/2, lowermost smaller, bordering anterior temporals; anterior temporal 1/1, in contact with sixth supralabial; posterior temporals 3/3, upper one thinner than lower one. Left maxilla arched, with an angular apex, distinctly bent inwards anteriorly. A total of 13 maxillary teeth or teeth alveola, with the following formula: five small anterior teeth, slightly enlarged posteriorly + three strongly enlarged teeth, thick, and not much curved + a distinctly wide gap + three small teeth + a small gap + two enlarged posterior teeth.
</paragraph>
<paragraph id="8B8536F0B833FFE5FC805078FBE2FC75" blockId="5.[831,1437,258,960]" pageId="5" pageNumber="417">
Body elongate, TL 
<quantity id="4CC29B15B833FFE5FBCC5078FBF2FC8B" box="[1036,1127,807,830]" metricMagnitude="-1" metricUnit="m" metricValue="5.91" pageId="5" pageNumber="417" unit="mm" value="591.0">591 mm</quantity>
; SVL 
<quantity id="4CC29B15B833FFE5FB695078FA91FC8B" box="[1193,1284,807,830]" metricMagnitude="-1" metricUnit="m" metricValue="4.67" pageId="5" pageNumber="417" unit="mm" value="467.0">467 mm</quantity>
; TaL 
<quantity id="4CC29B15B833FFE5FAFD5078FA0DFC8B" box="[1341,1432,807,830]" metricMagnitude="-1" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="417" unit="mm" value="124.0">124 mm</quantity>
; preventral 1, ventrals 218, from behind neck region distinctly notched laterally; subcaudals 90, paired; precloacal plate single; DSR 17-17-15, all smooth; the vertebral scales not enlarged.
</paragraph>
<paragraph id="8B8536F0B833FFE7FC805087FEDAFD8C" blockId="5.[832,1437,984,1072]" lastBlockId="7.[151,756,258,889]" lastPageId="7" lastPageNumber="419" pageId="5" pageNumber="417">
<emphasis id="B94EEAE2B833FFE5FC805087FBE0FC5A" bold="true" box="[832,1141,984,1007]" pageId="5" pageNumber="417">Coloration in preservative:</emphasis>
Dorsal surface of head and neck with pigmentation, paler on lateral side of head, the lower part of supralabials yellowish cream, speckled with pale brownish blotches. Body brownish black, light body bands beginning after 1.5 times the head length behind the head, in total 29 transverse light bands on body and 14 light bands on tail; the first four body bands yellowish cream, and distinctly widened towards the venter, increased in size posteriorly; a dark mottling in the vertebral region more prominent posteriorly; the subsequent light body bands with two distinct indentations on each side, fused in the middle in the last third of the body.
</paragraph>
<caption id="DF456678B833FFE5FF5854B5FEF8F7AE" ID-DOI="http://doi.org/10.5281/zenodo.5743940" ID-Zenodo-Dep="5743940" httpUri="https://zenodo.org/record/5743940/files/figure.png" pageId="5" pageNumber="417" startId="5.[152,187,2026,2047]" targetBox="[155,1435,1130,1985]" targetPageId="5">
<paragraph id="8B8536F0B833FFE5FF5854B5FEF8F7AE" blockId="5.[152,1436,2026,2075]" pageId="5" pageNumber="417">
Fig. 2. New country record of 
<taxonomicName id="4C3A4D73B833FFE5FE0854B5FDCFF84A" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[456,602,2026,2047]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="417" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B833FFE5FE0854B5FDCFF84A" box="[456,602,2026,2047]" italics="true" pageId="5" pageNumber="417">Lycodon pictus</emphasis>
</taxonomicName>
(CIB 115609) from China (Nonggang, Guangxi), adult female in life. Photograph by Jin-Long Ren.
</paragraph>
</caption>
<caption id="DF456678B830FFE6FF575493FBDDF7AE" ID-DOI="http://doi.org/10.5281/zenodo.5743942" ID-Zenodo-Dep="5743942" httpUri="https://zenodo.org/record/5743942/files/figure.png" pageId="6" pageNumber="418" startId="6.[151,186,1996,2017]" targetBox="[175,1409,257,1953]" targetPageId="6">
<paragraph id="8B8536F0B830FFE6FF575493FBDDF7AE" blockId="6.[151,1437,1996,2075]" pageId="6" pageNumber="418">
Fig. 3. Adult female of 
<taxonomicName id="4C3A4D73B830FFE6FE435493FD80F854" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[387,533,1996,2017]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="418" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B830FFE6FE435493FD80F854" box="[387,533,1996,2017]" italics="true" pageId="6" pageNumber="418">Lycodon pictus</emphasis>
</taxonomicName>
(CIB 115609; Guangxi, China) in preservative. (A) Dorsal general view; (B) Ventral general view; (C) Lateral head view, right side; (D) Lateral head view, left side; (E) Dorsal view of head; (F) Ventral view of head; (G) Lateral body view; (H) Ventral scale view. Scale bar 5 mm.Photographs by Jin-Long Ren.
</paragraph>
</caption>
<paragraph id="8B8536F0B831FFE7FF57511DFE0EFCCC" blockId="7.[151,756,258,889]" pageId="7" pageNumber="419">Ventral surface of head and neck yellowish cream, belly cream and greyish cream in the last third part of body and on lower tail surface; the dark dorsal bands in part extending towards the venter, forming complete dark bands around the posterior body and on the tail; lateral side of the head cream below, with the lighter pattern beginning in the supralabial region; tip of lower jaw and infralabial region in part greyish brown; dorsal surface of the head and upper head sides a bit paler than the remaining head dorsum.</paragraph>
<paragraph id="8B8536F0B831FFE7FF5750CEFC42FE2C" blockId="7.[151,756,913,1160]" lastBlockId="7.[831,1437,258,825]" pageId="7" pageNumber="419">
<emphasis id="B94EEAE2B831FFE7FF5750CEFDD3FC1D" bold="true" box="[151,582,913,936]" pageId="7" pageNumber="419">Variation of the new specimens:</emphasis>
The Chinese specimen (CIB 115609) largely matches the morphological data of the original description of 
<taxonomicName id="4C3A4D73B831FFE7FD58508EFF42FBBD" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="419" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B831FFE7FD58508EFF42FBBD" italics="true" pageId="7" pageNumber="419">Lycodon pictus</emphasis>
</taxonomicName>
, except for having (1) a slightly shorter tail, TaL/TL 0.210 vs 
<quantity id="4CC29B15B831FFE7FE9E574EFE2FFB9D" box="[350,442,1041,1064]" metricMagnitude="-3" metricUnit="m" metricValue="5.461" pageId="7" pageNumber="419" unit="in" value="0.215">0.215 in</quantity>
the single female 
<typeStatus id="54818852B831FFE7FD50574EFD65FB9D" box="[656,752,1041,1064]" pageId="7" pageNumber="419" type="paratype">paratype</typeStatus>
; (2) fewer infralabials 9/10 vs 10; (3) fewer anterior temporals 1 vs 2; however, 
<bibRefCitation id="EFAB4B01B831FFE7FE01570EFD0BFBDD" author="Janssen H. Y. &amp; Pham C. T. &amp; Ngo H. T. &amp; Le M. D. &amp; Nguyen T. Q. &amp; Ziegler T." box="[449,670,1105,1128]" pageId="7" pageNumber="419" pagination="1 - 29" refId="ref5787" refString="Janssen H. Y., Pham C. T., Ngo H. T., Le M. D., Nguyen T. Q., Ziegler T. 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. Zookeys 875: 1 - 29." type="journal article" year="2019">
Janssen 
<emphasis id="B94EEAE2B831FFE7FDD9570EFDC5FBDD" box="[537,592,1105,1128]" italics="true" pageId="7" pageNumber="419">et al.</emphasis>
(2019)
</bibRefCitation>
already documented the condition “2*” each for one side of both 
<typeStatus id="54818852B831FFE7FCBC525DFC73FEAC" box="[892,998,258,281]" pageId="7" pageNumber="419" type="paratype">paratypes</typeStatus>
, meaning two anterior temporals with the lowermost not touching the postocular, thus referred to as posterior temporal herein, resulting in only one anterior temporal in afore mentioned cases in the type series already.
</paragraph>
<paragraph id="8B8536F0B831FFE7FCFF52FDFBCEFD2C" blockId="7.[831,1437,258,825]" pageId="7" pageNumber="419">
The Vietnamese specimen (VNMN011227) generally agrees with the morphological data of the original description of 
<taxonomicName id="4C3A4D73B831FFE7FC2252BDFB1DFE4C" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[994,1160,482,505]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="419" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B831FFE7FC2252BDFB1DFE4C" box="[994,1160,482,505]" italics="true" pageId="7" pageNumber="419">Lycodon pictus</emphasis>
</taxonomicName>
, except for having (1) a slightly longer tail, TaL/TL 0.219 vs 
<quantity id="4CC29B15B831FFE7FB1E515DFA08FDAC" box="[1246,1437,514,537]" metricMagnitude="-3" metricUnit="m" metricValue="5.4102" metricValueMax="5.461" metricValueMin="5.3594" pageId="7" pageNumber="419" unit="in" value="0.213" valueMax="0.215" valueMin="0.211">0.211 to 0.215 in</quantity>
the single juvenile and the single female from the 
<typeStatus id="54818852B831FFE7FAAE517DFA09FD8C" box="[1390,1436,546,569]" pageId="7" pageNumber="419">type</typeStatus>
series for which TaL/TL ratio was available, (2) slightly fewer maxillary teeth (12 vs 13 or 14) and (3) fewer subcaudals 85 vs 90 or 91.
</paragraph>
<paragraph id="8B8536F0B831FFE7FCFF51FDFCFFFC8C" blockId="7.[831,1437,258,825]" pageId="7" pageNumber="419">Both specimens showed more bands on the tail; 14 dark bands in CIB 115609 and 15 dark bands in VNMN011227 (vs 9 and 13); 14 light bands in CIB 115609 and 16 light bands in VNMN011227 (vs 9+ and 13).</paragraph>
<paragraph id="8B8536F0B831FFE8FCFF500EFDBBFE6C" blockId="7.[831,1437,849,1160]" lastBlockId="8.[151,756,258,473]" lastPageId="8" lastPageNumber="420" pageId="7" pageNumber="419">
<emphasis id="B94EEAE2B831FFE7FCFF500EFBD1FCDD" bold="true" box="[831,1092,849,872]" pageId="7" pageNumber="419">Extended diagnosis:</emphasis>
<taxonomicName id="4C3A4D73B831FFE7FBA2500EFAB5FCDD" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[1122,1312,849,872]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="419" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B831FFE7FBA2500EFAB5FCDD" box="[1122,1312,849,872]" italics="true" pageId="7" pageNumber="419">Lycodon pictus</emphasis>
</taxonomicName>
can be differentiated from its congeners by the following morphological characters: dorsal scales in 17–17–15 rows, all smooth; supralabials eight (rarely nine); infralabials ten (rarely nine); one elongated loreal on each side, either in contact with the eye or separated from it; precloacal plate single; ventral scales 212-218 (plus one or two preventral scales); subcaudals 85-91; a total length of 597+ mm in males and 
<quantity id="4CC29B15B831FFE7FB31570EFAE8FBDD" box="[1265,1405,1105,1128]" metricMagnitude="-1" metricUnit="m" metricValue="5.67" metricValueMax="5.91" metricValueMin="5.43" pageId="7" pageNumber="419" unit="mm" value="567.0" valueMax="591.0" valueMin="543.0">543-591 mm</quantity>
in females; tail / total length ratio 
<quantity id="4CC29B15B831FFE7FB5B572EFAACFB3D" box="[1179,1337,1137,1160]" metricMagnitude="-3" metricUnit="m" metricValue="5.4483" metricValueMax="5.5626" metricValueMin="5.334" pageId="7" pageNumber="419" unit="in" value="0.2145" valueMax="0.219" valueMin="0.21">
<geoCoordinate id="EE0E5037B831FFE7FB5B572EFB4DFB3D" box="[1179,1240,1137,1160]" degrees="0.210" direction="north" orientation="latitude" pageId="7" pageNumber="419" precision="55" value="0.21">0.210</geoCoordinate>
<geoCoordinate id="EE0E5037B831FFE7FB18572EFA89FB3D" box="[1240,1308,1137,1160]" degrees="0.219" direction="west" orientation="longitude" pageId="7" pageNumber="419" precision="55" value="-0.219">-0.219</geoCoordinate>
in
</quantity>
females; maxillary teeth 12 to 14; dorsal surface of body with 28 or 29 light body bands; dorsal surface of tail with 9 to 15 dark bands, and 13 to 16 cream bands forming a distinct blotch in the vertebral region; ventral surface of body and tail mostly cream with the dark body bands in part extending towards the venter, sometimes forming complete dark bands around the body.
</paragraph>
</subSubSection>
<caption id="DF456678B831FFE7FF5754B6FAB1F7AE" ID-DOI="http://doi.org/10.5281/zenodo.5743944" ID-Zenodo-Dep="5743944" httpUri="https://zenodo.org/record/5743944/files/figure.png" pageId="7" pageNumber="419" startId="7.[151,186,2025,2046]" targetBox="[151,1436,1219,1980]" targetPageId="7">
<paragraph id="8B8536F0B831FFE7FF5754B6FAB1F7AE" blockId="7.[151,1437,2025,2075]" pageId="7" pageNumber="419">
Fig. 4. Map showing the new findings of 
<taxonomicName id="4C3A4D73B831FFE7FDF354B6FD53F84B" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[563,710,2025,2046]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="419" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B831FFE7FDF354B6FD53F84B" box="[563,710,2025,2046]" italics="true" pageId="7" pageNumber="419">Lycodon pictus</emphasis>
</taxonomicName>
. 1: VNMN 011227, from closeby the type locality in Cao Bang Province, northern Vietnam. 2: CIB 115609, the new country record of 
<taxonomicName id="4C3A4D73B831FFE7FCEF5B59FC14F7AE" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[815,897,2054,2075]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="419" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B831FFE7FCEF5B59FC14F7AE" box="[815,897,2054,2075]" italics="true" pageId="7" pageNumber="419">L. pictus</emphasis>
</taxonomicName>
from Nanggong, Guangxi, southern China.
</paragraph>
</caption>
<subSubSection id="C320657BB83EFFE8FF5752AEFEA1FDDD" pageId="8" pageNumber="420" type="distribution">
<paragraph id="8B8536F0B83EFFE8FF5752AEFEA1FDDD" blockId="8.[151,756,497,616]" pageId="8" pageNumber="420">
<emphasis id="B94EEAE2B83EFFE8FF5752AEFEBAFDBD" bold="true" box="[151,303,497,520]" pageId="8" pageNumber="420">Distribution:</emphasis>
<taxonomicName id="4C3A4D73B83EFFE8FE8452AEFE63FDBD" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[324,502,497,520]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B83EFFE8FE8452AEFE63FDBD" box="[324,502,497,520]" italics="true" pageId="8" pageNumber="420">Lycodon pictus</emphasis>
</taxonomicName>
is currently known from the 
<typeStatus id="54818852B83EFFE8FEC0514EFEBBFD9D" box="[256,302,529,552]" pageId="8" pageNumber="420">type</typeStatus>
locality in 
<collectingRegion id="49FEF812B83EFFE8FE72514EFD1EFD9D" box="[434,651,529,552]" country="Vietnam" name="Cao Bang" pageId="8" pageNumber="420">Cao Bang Province</collectingRegion>
, northern 
<collectingCountry id="F32D7660B83EFFE8FF57516EFF67FDFD" box="[151,242,561,584]" name="Vietnam" pageId="8" pageNumber="420">Vietnam</collectingCountry>
and Longzhou County in 
<collectingRegion id="49FEF812B83EFFE8FDE8516EFD12FDFD" box="[552,647,561,584]" country="China" name="Guangxi" pageId="8" pageNumber="420">Guangxi</collectingRegion>
, southern 
<collectingCountry id="F32D7660B83EFFE8FF57510EFF42FDDD" box="[151,215,593,616]" name="China" pageId="8" pageNumber="420">China</collectingCountry>
(
<figureCitation id="13012A75B83EFFE8FF25510EFEB2FDDD" box="[229,295,593,616]" captionStart="Fig" captionStartId="7.[151,186,2025,2046]" captionTargetBox="[151,1436,1219,1980]" captionTargetId="figure-477@7.[151,1436,1219,1976]" captionTargetPageId="7" captionText="Fig. 4. Map showing the new findings of Lycodon pictus. 1: VNMN 011227, from closeby the type locality in Cao Bang Province, northern Vietnam. 2: CIB 115609, the new country record of L. pictus from Nanggong, Guangxi, southern China." figureDoi="http://doi.org/10.5281/zenodo.5743944" httpUri="https://zenodo.org/record/5743944/files/figure.png" pageId="8" pageNumber="420">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320657BB83EFFE8FF5751DFFDD5FCE2" pageId="8" pageNumber="420" type="etymology">
<paragraph id="8B8536F0B83EFFE8FF5751DFFDD5FCE2" blockId="8.[151,756,640,855]" pageId="8" pageNumber="420">
<emphasis id="B94EEAE2B83EFFE8FF5751DFFE8EFD22" bold="true" box="[151,283,640,663]" pageId="8" pageNumber="420">Etymology:</emphasis>
The specific name of the species, “ 
<taxonomicName id="4C3A4D73B83EFFE8FD6251DFFD77FD22" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[674,738,640,663]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B83EFFE8FD6251DFFD77FD22" box="[674,738,640,663]" italics="true" pageId="8" pageNumber="420">pictus</emphasis>
</taxonomicName>
”, meaning painted or decorated in Latin, refers to the unique dorsal color pattern of this species (
<bibRefCitation id="EFAB4B01B83EFFE8FD42519FFF68FD42" author="Janssen H. Y. &amp; Pham C. T. &amp; Ngo H. T. &amp; Le M. D. &amp; Nguyen T. Q. &amp; Ziegler T." pageId="8" pageNumber="420" pagination="1 - 29" refId="ref5787" refString="Janssen H. Y., Pham C. T., Ngo H. T., Le M. D., Nguyen T. Q., Ziegler T. 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. Zookeys 875: 1 - 29." type="journal article" year="2019">
Janssen 
<emphasis id="B94EEAE2B83EFFE8FD21519FFF26FD42" italics="true" pageId="8" pageNumber="420">et al.</emphasis>
, 2019
</bibRefCitation>
). Because this species is herein reported as a new member of the Chinese snake fauna, we suggest “Jin Bai Huan She (DZ白ẇffi)” as its Chinese common name, deriving from its scientific name.
</paragraph>
</subSubSection>
<subSubSection id="C320657BB83EFFE8FF575030FAB1FD4C" pageId="8" pageNumber="420" type="biology_ecology">
<paragraph id="8B8536F0B83EFFE8FF575030FAB1FD4C" blockId="8.[151,756,879,966]" lastBlockId="8.[831,1437,258,761]" pageId="8" pageNumber="420">
<emphasis id="B94EEAE2B83EFFE8FF575030FEF3FC33" bold="true" box="[151,358,879,902]" pageId="8" pageNumber="420">Natural history:</emphasis>
<taxonomicName id="4C3A4D73B83EFFE8FEBF5030FDA3FC33" authorityName="Janssen, Pham, Ngo, Le, Nguyen &amp; Ziegler" authorityYear="2019" box="[383,566,879,902]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B94EEAE2B83EFFE8FEBF5030FDA3FC33" box="[383,566,879,902]" italics="true" pageId="8" pageNumber="420">Lycodon pictus</emphasis>
</taxonomicName>
inhabits karst environment in between elevations of 
<quantity id="4CC29B15B83EFFE8FD5C50D0FD66FC13" box="[668,755,911,934]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="8" pageNumber="420" unit="m" value="233.0">
<elevation id="0017D1C3B83EFFE8FD5C50D0FD66FC13" box="[668,755,911,934]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="8" pageNumber="420" unit="m" value="233.0">233 m</elevation>
</quantity>
(
<location id="8EE5602BB83EFFE8FF5F50F0FE86FC73" LSID="urn:lsid:plazi:treatment:039387E6B832FFE8FC8F5009FAB1FD4C:8EE5602BB83EFFE8FF5F50F0FE86FC73" box="[159,275,943,966]" country="China" name="Nonggang" pageId="8" pageNumber="420" stateProvince="Guangxi">Nonggang</location>
, 
<collectingRegion id="49FEF812B83EFFE8FEE550F0FE10FC73" box="[293,389,943,966]" country="China" name="Guangxi" pageId="8" pageNumber="420">Guangxi</collectingRegion>
, 
<collectingCountry id="F32D7660B83EFFE8FE5750F0FE4FFC73" box="[407,474,943,966]" name="China" pageId="8" pageNumber="420">China</collectingCountry>
, this study, 
<figureCitation id="13012A75B83EFFE8FDB050F0FD22FC73" box="[624,695,943,966]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[151,1435,1019,1983]" captionTargetId="figure-365@8.[151,1436,1019,1983]" captionTargetPageId="8" captionText="Fig. 5. Macrohabitat of Lycodon pictus in Nonggang National Nature Reserve, Longzhou, Guangxi, China. Photograph by Jin-Long Ren." figureDoi="http://doi.org/10.5281/zenodo.5743946" httpUri="https://zenodo.org/record/5743946/files/figure.png" pageId="8" pageNumber="420">Fig. 5</figureCitation>
) and 
<quantity id="4CC29B15B83EFFE8FCFF525DFC1FFEAC" box="[831,906,258,281]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="8" pageNumber="420" unit="m" value="701.0">701 m</quantity>
a.s.l. (
<typeStatus id="54818852B83EFFE8FC1F525DFB9AFEAC" box="[991,1039,258,281]" pageId="8" pageNumber="420">type</typeStatus>
locality; 
<bibRefCitation id="EFAB4B01B83EFFE8FB42525DFAF8FEAC" author="Janssen H. Y. &amp; Pham C. T. &amp; Ngo H. T. &amp; Le M. D. &amp; Nguyen T. Q. &amp; Ziegler T." box="[1154,1389,258,281]" pageId="8" pageNumber="420" pagination="1 - 29" refId="ref5787" refString="Janssen H. Y., Pham C. T., Ngo H. T., Le M. D., Nguyen T. Q., Ziegler T. 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. Zookeys 875: 1 - 29." type="journal article" year="2019">
Janssen 
<emphasis id="B94EEAE2B83EFFE8FB22525DFAB3FEAC" box="[1250,1318,258,281]" italics="true" pageId="8" pageNumber="420">et al.,</emphasis>
2019
</bibRefCitation>
). In southern 
<collectingCountry id="F32D7660B83EFFE8FC64527DFC73FE8C" box="[932,998,290,313]" name="China" pageId="8" pageNumber="420">China</collectingCountry>
, the species was found at night between 00:00 and 1:00 am after light rain, active in a drainage ditch along a forest path, covered with dead leaves. The surrounding habitat was secondary karst forest, consisting of short hardwood, shrubs and vines. The snake did not attempt to bite when it was handled, neither aggressive nor defensive behavior was observed. In terms of the herpetofauna, several other amphibians and reptiles were observed during the survey period in 
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the same microhabitat, including 
<taxonomicName id="4C3A4D73B83EFFE8FAF2511DFC5FFDCC" baseAuthorityName="Ye &amp; Fei in Ye, Fei &amp; Hu" baseAuthorityYear="1993" class="Amphibia" family="Rhacophoridae" genus="Kurixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="odontotarsus">
<emphasis id="B94EEAE2B83EFFE8FAF2511DFC5FFDCC" italics="true" pageId="8" pageNumber="420">Kurixalus odontotarsus</emphasis>
</taxonomicName>
(Ye &amp; Fei in 
<bibRefCitation id="EFAB4B01B83EFFE8FB9C513DFB6EFDCC" author="Ye C. &amp; Fei L. &amp; Hu S." box="[1116,1275,610,633]" pageId="8" pageNumber="420" refId="ref6637" refString="Ye C., Fei L., Hu S. 1993. Rare and economic amphibians of China. Sichuan Publishing House of Science &amp; Technology, Chengdu, 412 pp. [in Chinese]" type="book" year="1993">
Ye 
<emphasis id="B94EEAE2B83EFFE8FB40513DFB20FDCC" box="[1152,1205,610,633]" italics="true" pageId="8" pageNumber="420">et al.</emphasis>
, 1993
</bibRefCitation>
), 
<taxonomicName id="4C3A4D73B83EFFE8FACE513DFB12FD2C" authority="(Ohler &amp; Delorme, 2006)" baseAuthorityName="Ohler &amp; Delorme" baseAuthorityYear="2006" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="420" phylum="Arthropoda" rank="species" species="kio">
<emphasis id="B94EEAE2B83EFFE8FACE513DFCF5FD2C" italics="true" pageId="8" pageNumber="420">Rhacophorus kio</emphasis>
(
<bibRefCitation id="EFAB4B01B83EFFE8FCB151DDFBEBFD2C" author="Ohler A. &amp; Delorme M." box="[881,1150,642,665]" pageId="8" pageNumber="420" pagination="86 - 97" refId="ref6180" refString="Ohler A., Delorme M. 2006. Well known does not mean well studied: Morphological and molecular support for existence of sibling species in the Javanese gliding frog Rhacophorus reinwardtii (Amphibia, Anura). Comptes Rendus Biologies 329: 86 - 97." type="journal article" year="2006">Ohler &amp; Delorme, 2006</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C3A4D73B83EFFE8FB5451DDFC14FD0C" authority="(Gray, 1845)" baseAuthorityName="Gray" baseAuthorityYear="1845" class="Reptilia" family="Agamidae" genus="Draco" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B94EEAE2B83EFFE8FB5451DDFAC5FD2C" box="[1172,1360,642,665]" italics="true" pageId="8" pageNumber="420">Draco maculatus</emphasis>
(
<bibRefCitation id="EFAB4B01B83EFFE8FAA051DDFCECFD0C" author="Gray J. E." pageId="8" pageNumber="420" refId="ref5630" refString="Gray J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of the British Museum / Edward Newman, London: xxvii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C3A4D73B83EFFE8FC5B51FDFA0DFD0C" authority="(Cuvier, 1829)" baseAuthorityName="Cuvier" baseAuthorityYear="1829" box="[923,1432,674,697]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="lepidogaster">
<emphasis id="B94EEAE2B83EFFE8FC5B51FDFB47FD0C" box="[923,1234,674,697]" italics="true" pageId="8" pageNumber="420">Acanthosaura lepidogaster</emphasis>
(
<bibRefCitation id="EFAB4B01B83EFFE8FB2F51FDFA05FD0C" author="Cuvier G. L." box="[1263,1424,674,697]" pageId="8" pageNumber="420" pagination="1 - 406" refId="ref5278" refString="Cuvier G. L. 1829. Le Regne Animal Distribue, d'apres son Organisation, pour servir de base a l'Histoire naturelle des Animaux et d'introduction a l'Anatomie Comparee. Nouvelle Edition [second edition]. Vol. 2. Les Reptiles. Deterville, Paris, i-xvi, 1 - 406." type="book chapter" year="1829">Cuvier, 1829</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C3A4D73B83EFFE8FCFF519DFAF2FD6C" authority="Grismer, Viets &amp; Boyle, 1999" authorityName="Grismer, Viets &amp; Boyle" authorityYear="1999" box="[831,1383,706,729]" class="Reptilia" family="Eublepharidae" genus="Goniurosaurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="luii">
<emphasis id="B94EEAE2B83EFFE8FCFF519DFB9AFD6C" box="[831,1039,706,729]" italics="true" pageId="8" pageNumber="420">Goniurosaurus luii</emphasis>
<bibRefCitation id="EFAB4B01B83EFFE8FBD8519DFAF2FD6C" author="Grismer L. &amp; Viets B. E. &amp; Boyle L. J." box="[1048,1383,706,729]" pageId="8" pageNumber="420" pagination="382 - 393" refId="ref5667" refString="Grismer L., Viets B. E., Boyle L. J. 1999. Two new continental species of Goniurosaurus (Squamata: Eublepharidae) with a phylogeny and evolutionary classification of the genus. Journal of Herpetology 33 (3): 382 - 393." type="journal article" year="1999">Grismer, Viets &amp; Boyle, 1999</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3A4D73B83EFFE8FCFF51BDFAB5FD4C" authority="(Boie, 1827)" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[831,1312,738,761]" class="Reptilia" family="Lamprophiidae" genus="Psammodynastes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="420" phylum="Chordata" rank="species" species="pulverulentus">
<emphasis id="B94EEAE2B83EFFE8FCFF51BDFB1AFD4C" box="[831,1167,738,761]" italics="true" pageId="8" pageNumber="420">Psammodynastes pulverulentus</emphasis>
(
<bibRefCitation id="EFAB4B01B83EFFE8FB5D51BDFA82FD4C" author="Boie F." box="[1181,1303,738,761]" pageId="8" pageNumber="420" pagination="508 - 566" refId="ref5116" refString="Boie F. 1827. Bemerkungen uber Merrem's Versuch eines Systems der Amphibien, 1. Lieferung: Ophidier. Isis van Oken 20: 508 - 566." type="journal article" year="1827">Boie, 1827</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>