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<document id="4B78C80729577F43AFBDDE1BD7B0E7BA" ID-CLB-Dataset="22179" ID-DOI="10.5852/ejt.2018.479" ID-GBIF-Dataset="73087d02-3f6a-4672-a86f-44cba6cfe5cf" ID-Zenodo-Dep="3825259" ID-ZooBank="EA678C5B-2E5B-4F73-B254-B9A524307E70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589392305789" checkinUser="valdenar" docAuthor="Tchesunov, Alexei V." docDate="2018" docId="039387D61803FFF1253DFE87FB51F944" docLanguage="en" docName="ejt-479_tchesunov.pdf.imf" docOrigin="European Journal of Taxonomy 479" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Neostygarctus grossmeteori Tchesunov 2018, sp. nov." docType="treatment" docUuid="0047D8B7-BEAE-4751-B098-86EA728FABA3" docUuidSource="ZooBank" docVersion="8" lastPageNumber="12" masterDocId="FFAAFFAE1807FFFA277FFF8BFFEBFF97" masterDocTitle="A new tardigrade species of the genus Neostygarctus Grimaldi de Zio et al., 1982 (Tardigrada, Arthrotardigrada) from the Great Meteor Seamount, Northeast Atlantic" masterLastPageNumber="17" masterPageNumber="1" pageNumber="5" updateTime="1698839498575" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="9782841222BE6DE97F365344094617E2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="19300AB6234F217A3B232F88499F0BC8">
<mods:title id="E50E1ECE1E640B14BBF20ABAB449E358">A new tardigrade species of the genus Neostygarctus Grimaldi de Zio et al., 1982 (Tardigrada, Arthrotardigrada) from the Great Meteor Seamount, Northeast Atlantic</mods:title>
</mods:titleInfo>
<mods:name id="4BD59392E3BCC9909627DD9935AC5052" type="personal">
<mods:role id="883C4D0FA80348952AE8F69A1A997AB5">
<mods:roleTerm id="36245AE5DA37A49A2C68C46D4DB2191A">Author</mods:roleTerm>
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<mods:namePart id="55B66CCDD716B7CA904211C284C0FC25">Tchesunov, Alexei V.</mods:namePart>
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<mods:title id="94752F8D4696BE86971B3C660A5CD1A1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3C3424056E0E272D670713A15FF37CB3">
<mods:date id="E52A605072725CD46FC40B64B56E019E">2018</mods:date>
<mods:detail id="873EBD5D63CFD607FF4C335B8D994F39" type="pubDate">
<mods:number id="A3459C13EBEE7B09E1E3D21155D9382E">2018-11-27</mods:number>
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<mods:number id="FC336C641024FD7E7047A615667733F7">479</mods:number>
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<mods:start id="0EF70E8FB78905731E22FFC7C84CF574">1</mods:start>
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<mods:classification id="D27D5C0C96115B093C0A737043C63C7A">journal article</mods:classification>
<mods:identifier id="932218586E2E30DD68377F47C45F4D05" type="CLB-Dataset">22179</mods:identifier>
<mods:identifier id="744FC0BF200E55D86BC340F6795B1E5D" type="DOI">10.5852/ejt.2018.479</mods:identifier>
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<mods:identifier id="E91D089D3D9FE95B076B745892962240" type="Zenodo-Dep">3825259</mods:identifier>
<mods:identifier id="2037FB641F2FFA5C9D8E7E6C247901EF" type="ZooBank">EA678C5B-2E5B-4F73-B254-B9A524307E70</mods:identifier>
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<treatment id="039387D61803FFF1253DFE87FB51F944" ID-DOI="http://doi.org/10.5281/zenodo.3845837" ID-GBIF-Taxon="164131226" ID-Zenodo-Dep="3845837" LSID="urn:lsid:zoobank.org:act:0047D8B7-BEAE-4751-B098-86EA728FABA3" httpUri="http://treatment.plazi.org/id/039387D61803FFF1253DFE87FB51F944" lastPageId="11" lastPageNumber="12" pageId="4" pageNumber="5">
<subSubSection id="C320654B1803FFFE253DFE87FB22FEDE" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B8536C01803FFFE253DFE87FC19FEB1" blockId="4.[362,1225,267,365]" box="[578,1010,267,294]" pageId="4" pageNumber="5">
<heading id="D0CD81AC1803FFFE253DFE87FC19FEB1" box="[578,1010,267,294]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C3A4D431803FFFE253DFE87FC7BFEB2" ID-CoL="6647c26c-fb2e-41da-8806-f377c47a6e42" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[578,912,267,294]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD21803FFFE253DFE87FC7BFEB2" bold="true" box="[578,912,267,294]" italics="true" pageId="4" pageNumber="5">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A91803FFFE24E8FE87FC19FEB1" box="[919,1010,268,294]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FFAB3AC21803FFFE2615FEA5FB22FEDE" box="[362,1225,302,329]" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C320654B1803FFFE2531FED9FC0DFEFA" box="[590,998,338,365]" pageId="4" pageNumber="5" type="description">
<paragraph id="8B8536C01803FFFE2531FED9FC0DFEFA" blockId="4.[362,1225,267,365]" box="[590,998,338,365]" pageId="4" pageNumber="5">
<figureCitation id="13012A451803FFFE2531FED9FD5CFEFB" box="[590,695,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="6.[188,231,1705,1733]" captionStartId-1="7.[189,232,1939,1965]" captionStartId-2="8.[189,232,1879,1905]" captionStartId-3="9.[189,232,1924,1950]" captionStartId-4="10.[189,232,1809,1835]" captionTargetBox-0="[222,1898,580,1610]" captionTargetBox-1="[201,1368,303,1824]" captionTargetBox-2="[195,1393,314,1784]" captionTargetBox-3="[212,1352,277,1849]" captionTargetBox-4="[232,1358,272,1771]" captionTargetId-0="figure@6.[188,1927,550,1654]" captionTargetId-1="figure@7.[189,1398,262,1892]" captionTargetId-2="figure@8.[189,1399,308,1794]" captionTargetId-3="figure@9.[195,1380,262,1867]" captionTargetId-4="figure@10.[219,1368,262,1772]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." captionText-1="Fig. 3. Neostygarctus grossmeteori sp. nov., heads. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm." captionText-2="Fig. 4. Neostygarctus grossmeteori sp. nov., optical photopictures. A. Paratype, ♀ (SMF 54), entire body, dorsal view. B. Holotype, ♀ (SMF 51), entire body, ventral view. C–E. Paratype of obscure gender (SMF 59). C–D. Areas of dorsal surface of body with spines. E. Lateral body projections. F. Holotype, ♀ (SMF 51), posterior body with female gonopore and anus. Scale bars: A–B = 50 μm; C–F = 20 μm." captionText-3="Fig. 5. Neostygarctus grossmeteori sp. nov. Paratype, ♀ (SMF 52), details. A. Lateral body processes, ventral view. B–D. Legs I (left and right) and IV. E. Genital area. Scale bars: A = 50 μm; B–E = 20 μm." captionText-4="Fig. 6. Neostygarctus grossmeteori sp. nov., details, SEM. A. Female, entire body, ventral view. B. Right secondary clava and outer cirrus on the head, ventral view. C. Ventral conical spikes on the basal part of the lateral body process. D. Right lateral body processes. E. Lateral fan of spines with membrane on the posteriormost body segment. F. Toes with claws of the leg IV ventrally, dorsal tendon detached in some toes. G. Inner and outer claws, dorsal view. H. Accordion-like joint of the cirrus E. Scale bars: A = 30 μm; B, D, F = 10 μm; C, E, G–H = 3 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3825263" figureDoi-1="http://doi.org/10.5281/zenodo.3825265" figureDoi-2="http://doi.org/10.5281/zenodo.3825269" figureDoi-3="http://doi.org/10.5281/zenodo.3825271" figureDoi-4="http://doi.org/10.5281/zenodo.3825273" httpUri-0="https://zenodo.org/record/3825263/files/figure.png" httpUri-1="https://zenodo.org/record/3825265/files/figure.png" httpUri-2="https://zenodo.org/record/3825269/files/figure.png" httpUri-3="https://zenodo.org/record/3825271/files/figure.png" httpUri-4="https://zenodo.org/record/3825273/files/figure.png" pageId="4" pageNumber="5">Figs 2–6</figureCitation>
,
<tableCitation id="C6B8037B1803FFFE25BDFED9FCF2FEFB" box="[706,793,338,364]" captionStart="Table 1" captionStartId="5.[189,258,268,294]" captionTargetPageId="5" captionText="Table 1. Measurements (in μm) of Neostygarctus grossmeteori sp. nov." httpUri="http://table.plazi.org/id/DF4566481802FFFF27C2FE87FBEAFEB1" pageId="4" pageNumber="5" tableUuid="DF4566481802FFFF27C2FE87FBEAFEB1">Table 1</tableCitation>
(morphometrics)
</paragraph>
</subSubSection>
<subSubSection id="C320654B1803FFFE27C2FE18FCF4FDFF" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8B8536C01803FFFE27C2FE18FEDDFE3A" blockId="4.[189,310,403,429]" box="[189,310,403,429]" pageId="4" pageNumber="5">
<heading id="D0CD81AC1803FFFE27C2FE18FEDDFE3A" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B94EEAD21803FFFE27C2FE18FEDDFE3A" bold="true" box="[189,310,403,429]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FE48FCF4FDFF" blockId="4.[189,1399,450,617]" pageId="4" pageNumber="5">
<taxonomicName id="4C3A4D431803FFFE27C2FE48FE8FFE4B" authorityName="Grimaldi de Zio, D'Addabbo Gallo &amp; Morone de Lucia" authorityYear="1982" box="[189,356,451,476]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
<emphasis id="B94EEAD21803FFFE27C2FE48FE8FFE4B" box="[189,356,451,476]" italics="true" pageId="4" pageNumber="5">Neostygarctus</emphasis>
</taxonomicName>
with five middorsal spines on the cephalic and each body plate and on the caudal plate, the spines decreasing in length backwards. Ocelli present. One or two pairs of ventral cervical spines present. Each lateral body proсess provided basally on the ventral side with a transversal row of two to five short but strong spikes. Only inner claws of each foot provided with a fine dorsal accessory spine; external claws may carry vestigial accessory spines.
</paragraph>
</subSubSection>
<subSubSection id="C320654B1803FFFE27C2FD1BFB6BFD4E" box="[189,1152,656,729]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8B8536C01803FFFE27C2FD1BFEAFFD3D" blockId="4.[189,324,656,682]" box="[189,324,656,682]" pageId="4" pageNumber="5">
<heading id="D0CD81AC1803FFFE27C2FD1BFEAFFD3D" bold="true" box="[189,324,656,682]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B94EEAD21803FFFE27C2FD1BFEAFFD3D" bold="true" box="[189,324,656,682]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FD35FB6BFD4E" blockId="4.[189,1152,702,729]" box="[189,1152,702,729]" pageId="4" pageNumber="5">The species name is derived from the name of locality (Gross Meteor in German).</paragraph>
</subSubSection>
<subSubSection id="C320654B1803FFF127C2FC8BFB51F944" lastPageId="11" lastPageNumber="12" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8B8536C01803FFFE27C2FC8BFE87FC8D" blockId="4.[189,364,768,794]" box="[189,364,768,794]" pageId="4" pageNumber="5">
<heading id="D0CD81AC1803FFFE27C2FC8BFE87FC8D" bold="true" box="[189,364,768,794]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B94EEAD21803FFFE27C2FC8BFE87FC8D" bold="true" box="[189,364,768,794]" pageId="4" pageNumber="5">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B523C9D1803FFFE2795FCA4FB09FC07" ID-GBIF-Occurrence="2618260301" collectingDate="2010-03-01" collectionCode="RV" elevation="299" latitude="29.816233" longLatPrecision="1" longitude="-28.43235" pageId="4" pageNumber="5" specimenCode="SMF 51" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="8B8536C01803FFFE2795FCA4FEB7FCDE" blockId="4.[189,1399,815,912]" box="[234,348,815,841]" pageId="4" pageNumber="5">
<emphasis id="B94EEAD21803FFFE2795FCA4FEB7FCDE" bold="true" box="[234,348,815,841]" pageId="4" pageNumber="5">
<typeStatus id="548188621803FFFE2795FCA4FEB7FCDE" box="[234,348,815,841]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FCD9FB09FC07" blockId="4.[189,1399,815,912]" pageId="4" pageNumber="5">
ATLANTIC OCEAN:
<specimenCount id="9D3CFD491803FFFE26B1FCD9FE0FFCFB" box="[462,484,850,876]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
, Great Meteor Seamount plateau,
<geoCoordinate id="EE0E50071803FFFE24FBFCD9FBF6FCFB" box="[900,1053,850,876]" degrees="29" direction="north" minutes="48.974" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="29.816233">
29º48.974
<collectionCode id="ED2BAE051803FFFE237AFCD9FBF6FCFB" box="[1029,1053,850,876]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="4" pageNumber="5" type="Herbarium">N</collectionCode>
</geoCoordinate>
,
<geoCoordinate id="EE0E50071803FFFE2353FCDAFB22FCFB" box="[1068,1225,849,876]" degrees="28" direction="west" minutes="25.941" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-28.43235">
28º25.941
<collectionCode id="ED2BAE051803FFFE23D2FCD9FB22FCFB" box="[1197,1225,850,876]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="5" type="Herbarium">W</collectionCode>
</geoCoordinate>
, depth
<quantity id="4CC29B251803FFFE225DFCD9FA9AFCFB" box="[1314,1393,850,876]" metricMagnitude="2" metricUnit="m" metricValue="2.9899999999999998" pageId="4" pageNumber="5" unit="m" value="299.0">
<elevation id="0017D1F31803FFFE225DFCD9FA9AFCFB" box="[1314,1393,850,876]" metricMagnitude="2" metricUnit="m" metricValue="2.9899999999999998" pageId="4" pageNumber="5" unit="m" value="299.0">299 m</elevation>
</quantity>
,
<collectionCode id="ED2BAE051803FFFE27C2FCFEFF0AFC18" box="[189,225,885,911]" country="Italy" name="Collection of Leptospira Strains" pageId="4" pageNumber="5">RV</collectionCode>
<taxonomicName id="4C3A4D431803FFFE2795FCFEFEBCFC18" box="[234,343,885,911]" class="Pilidiophora" family="Lineidae" genus="Poseidon" higherTaxonomySource="CoL" kingdom="Animalia" order="Heteronemertea" pageId="4" pageNumber="5" phylum="Nemertea" rank="genus">
<emphasis id="B94EEAD21803FFFE2795FCFEFEBCFC18" box="[234,343,885,911]" italics="true" pageId="4" pageNumber="5">Poseidon</emphasis>
</taxonomicName>
, expedition P397 GROMET, location #17, stn 106-6,
<date id="FF8410001803FFFE24ADFCFEFB8AFC18" box="[978,1121,885,911]" pageId="4" pageNumber="5" value="2010-03-01">
<collectingDate id="EFC0E9E81803FFFE24ADFCFEFB8AFC18" box="[978,1121,885,911]" pageId="4" pageNumber="5" value="2010-03-01">
<date id="FF8410001803FFFE24ADFCFEFBF4FC18" box="[978,1055,885,911]" day="01" month="03" monthName="Mar" pageId="4" pageNumber="5">1 Mar.</date>
<date id="FF8410001803FFFE2359FCFEFB8AFC18" box="[1062,1121,885,911]" pageId="4" pageNumber="5" value="2010" year="2010">2010</date>
</collectingDate>
</date>
(
<specimenCode id="DB9C9EBB1803FFFE230FFCFEFB3EFC07" box="[1136,1237,885,912]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="5">SMF 51</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8B8536C01803FFFE2795FC3CFE8CFC46" blockId="4.[189,1399,951,1082]" box="[234,359,951,977]" pageId="4" pageNumber="5">
<emphasis id="B94EEAD21803FFFE2795FC3CFE8CFC46" bold="true" box="[234,359,951,977]" pageId="4" pageNumber="5">
<typeStatus id="548188621803FFFE2795FC3CFE8CFC46" box="[234,359,951,977]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FC51FE84FBAD" blockId="4.[189,1399,951,1082]" pageId="4" pageNumber="5">
ATLANTIC OCEAN:
<specimenCount id="9D3CFD491803FFFE26BBFC51FDEEFC63" box="[452,517,986,1012]" pageId="4" pageNumber="5" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="ED2BAE051803FFFE256DFC51FDA5FC63" box="[530,590,986,1012]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="5">SMF</collectionCode>
52-54), 1 Ƌ (
<collectionCode id="ED2BAE051803FFFE2598FC51FCC8FC63" box="[743,803,986,1012]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="5">SMF</collectionCode>
58), one specimen of unidentified gender (
<collectionCode id="ED2BAE051803FFFE2272FC51FAA2FC63" box="[1293,1353,986,1012]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="5">SMF</collectionCode>
60) (
<tableCitation id="C6B8037B1803FFFE27BAFC76FEF0FB80" box="[197,283,1021,1047]" captionStart="Table 1" captionStartId="5.[189,258,268,294]" captionTargetPageId="5" captionText="Table 1. Measurements (in μm) of Neostygarctus grossmeteori sp. nov." httpUri="http://table.plazi.org/id/DF4566481802FFFF27C2FE87FBEAFEB1" pageId="4" pageNumber="5" tableUuid="DF4566481802FFFF27C2FE87FBEAFEB1">Table 1</tableCitation>
), same data as for the
<typeStatus id="548188621803FFFE255BFC76FD64FB80" box="[548,655,1021,1047]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
. The
<typeStatus id="548188621803FFFE25B2FC76FCDFFB80" box="[717,820,1021,1047]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
and the
<typeStatus id="548188621803FFFE24E8FC76FBECFB80" box="[919,1031,1021,1047]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
are all embedded in permanent glycerin slides.
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FBEAFE5EFBEC" blockId="4.[189,437,1121,1147]" box="[189,437,1121,1147]" pageId="4" pageNumber="5">
<heading id="D0CD81AC1803FFFE27C2FBEAFE5EFBEC" bold="true" box="[189,437,1121,1147]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B94EEAD21803FFFE27C2FBEAFE5EFBEC" bold="true" box="[189,437,1121,1147]" pageId="4" pageNumber="5">General description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8536C01803FFFE27C2FB1BFD31F9BA" blockId="4.[189,1399,1168,1581]" pageId="4" pageNumber="5">
<collectionCode id="ED2BAE051803FFFE27C2FB1BFDCFFB3F" box="[189,548,1168,1194]" pageId="4" pageNumber="5">BODY SURFACE AND SCULPTURES</collectionCode>
. Dorsal cuticle consists of five transversal dorsal plates (one cephalic, three body and one caudal plates) and intermediate areas between them (
<figureCitation id="13012A451803FFFE2348FB38FB72FB5A" box="[1079,1177,1203,1229]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
). Ventral cuticular plates not developed as such; the only transversal cuticular folds may be discernible (
<figureCitation id="13012A451803FFFE23E0FB5DFB10FB67" box="[1183,1275,1238,1264]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
). Edges of dorsal plates discernible as discontinuous cuticular folds which may be obscured here and there. Cuticle of plates hardly differs from that of intermediate areas – but sparse and irregular coarse punctuations on dorsal plates (
<figureCitation id="13012A451803FFFE26F5FACBFE02FACD" box="[394,489,1344,1370]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
), which are, in fact, seemingly epicuticular pillars not detectable in SEM on surface but visible in optical microscope laterally as short projecting bars. Five robust middorsal spines present, which are elongated and conical, with acute or slightly rounded tips. Each plate carries one median spine attached in pit in middle on cephalic and on body plates I and II, close to posterior edge on body plate III, and close to anterior edge of caudal plate. Middorsal spines tend to be reducing in length from cephalic to caudal plate (
<figureCitation id="13012A451803FFFE2509FA7BFD3FF99D" box="[630,724,1520,1546]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
, ds). Set of middorsal spines may be incomplete in some specimens (
<tableCitation id="C6B8037B1803FFFE2639F998FE76F9BA" box="[326,413,1555,1581]" captionStart="Table 1" captionStartId="5.[189,258,268,294]" captionTargetPageId="5" captionText="Table 1. Measurements (in μm) of Neostygarctus grossmeteori sp. nov." httpUri="http://table.plazi.org/id/DF4566481802FFFF27C2FE87FBEAFEB1" pageId="4" pageNumber="5" tableUuid="DF4566481802FFFF27C2FE87FBEAFEB1">Table 1</tableCitation>
), possibly because of loss.
</paragraph>
<paragraph id="8B8536C01803FFFF27C2F9DEFE76F9C8" blockId="4.[189,1399,1620,1999]" lastBlockId="5.[189,1399,1394,1631]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
CEPHALIC REGION (
<figureCitation id="13012A451803FFFE26EEF9DFFE37F9F8" box="[401,476,1620,1647]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
A–B, 3A–B). Anteriorly subdivided into two pairs of lobes densely feathered by fine hollow cuticular spines: two frontal semicircular lobes (fl), two fronto-lateral lobes (fll), also semicircular but less prominent, and two long lateral common cirrophores of cirrus A and primary clava (cc). Pair of not very prominent but distinct latero-dorsal wart-like cervical lobes (cl) on narrowed neck region just posterior to lateral cirrophores. Dorsal cuticle of cephalic region marked by sparse and irregularly distributed coarse punctuations which are actually projections of short rod-like epicuticular pillars (
<figureCitation id="13012A451803FFFE2668F8A3FE9DF8D5" box="[279,374,1832,1858]" captionStart="Fig" captionStartId="7.[189,232,1939,1965]" captionTargetBox="[201,1368,303,1824]" captionTargetId="figure@7.[189,1398,262,1892]" captionTargetPageId="7" captionText="Fig. 3. Neostygarctus grossmeteori sp. nov., heads. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3825265" httpUri="https://zenodo.org/record/3825265/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
). Long middorsal spine at level of cervical lobes. Anterior edges of frontal and frontolateral lobes as well as both anterior and posterior edges of lateral cirrophores on head flanked by tight rows of finest hollow spines: about 15 spines on frontal lobe, 10-12 spines on fronto-lateral lobe, 9-11 spines on anterior and about 15 spines on posterior edge of lateral lobe. Spines of anterior frontal lobes very fine, spines of fronto-lateral lobes and then to anterior and posterior edges of body lateral processes become gradually more and more robust. Cervical lobes bear about eight medium-sized, slightly curved spines each. Fine spines arranged in tight rows on all head and body processes joined with thin film or membrane (
<figureCitation id="13012A451802FFFF2638FA33FE67FA45" box="[327,396,1464,1490]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A–B); rags of the membranes observable in SEM (
<figureCitation id="13012A451802FFFF24ABFA33FBC5FA44" box="[980,1070,1464,1491]" captionStart="Fig" captionStartId="10.[189,232,1809,1835]" captionTargetBox="[232,1358,272,1771]" captionTargetId="figure@10.[219,1368,262,1772]" captionTargetPageId="10" captionText="Fig. 6. Neostygarctus grossmeteori sp. nov., details, SEM. A. Female, entire body, ventral view. B. Right secondary clava and outer cirrus on the head, ventral view. C. Ventral conical spikes on the basal part of the lateral body process. D. Right lateral body processes. E. Lateral fan of spines with membrane on the posteriormost body segment. F. Toes with claws of the leg IV ventrally, dorsal tendon detached in some toes. G. Inner and outer claws, dorsal view. H. Accordion-like joint of the cirrus E. Scale bars: A = 30 μm; B, D, F = 10 μm; C, E, G–H = 3 μm." figureDoi="http://doi.org/10.5281/zenodo.3825273" httpUri="https://zenodo.org/record/3825273/files/figure.png" pageId="5" pageNumber="6">Fig. 6E</figureCitation>
). About two pairs of straight medium-sized ventro-lateral cervical spines (cs) on neck region just posterior to cervical lobes. Anterior cervical spines a bit smaller and located more antero-ventrally than posterior cervical spines (the latters are paired on either side in one specimen). Soft mouth cone on ventral side situated a bit posterior to anterior head edge.
</paragraph>
<caption id="DF4566481802FFFF27C2FE87FBEAFEB1" ID-Table-UUID="DF4566481802FFFF27C2FE87FBEAFEB1" box="[189,1025,267,294]" httpUri="http://table.plazi.org/id/DF4566481802FFFF27C2FE87FBEAFEB1" pageId="5" pageNumber="6" targetBox="[200,1367,357,1318]" targetIsTable="true" targetPageId="5">
<paragraph id="8B8536C01802FFFF27C2FE87FBEAFEB1" blockId="5.[189,1025,267,294]" box="[189,1025,267,294]" pageId="5" pageNumber="6">
<emphasis id="B94EEAD21802FFFF27C2FE87FECBFEB2" bold="true" box="[189,288,267,294]" pageId="5" pageNumber="6">Table 1.</emphasis>
Measurements (in μm) of
<taxonomicName id="4C3A4D431802FFFF2527FE87FC75FEB2" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[600,926,268,293]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD21802FFFF2527FE87FC75FEB2" box="[600,926,268,293]" italics="true" pageId="5" pageNumber="6">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A91802FFFF24D9FE87FBEAFEB1" box="[934,1025,268,294]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8536C01802FFFF27B7FE0EFACCFAB1" pageId="5" pageNumber="6">
<table id="F93AC4601802000527B7FEEEFABCFAB1" box="[200,1367,357,1318]" gridcols="7" gridrows="18" pageId="5" pageNumber="6">
<tr id="350A34821802000527B7FEEEFABCFE2A" box="[200,1367,357,445]" gridrow="0" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FEEEFE7CFE2A" box="[200,407,357,445]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B94EEAD21802FFFF27B7FE0EFED6FE0A" bold="true" box="[200,317,389,413]" pageId="5" pageNumber="6">Character</emphasis>
</th>
<th id="76DB5DFE18020005268AFEEEFC0DFE2A" box="[501,998,357,445]" colspan="4" colspanRight="3" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B94EEAD21802FFFF2506FEFEFC70FE3A" bold="true" pageId="5" pageNumber="6">Holotype female (51) and paratype females (52─54)</emphasis>
</th>
<th id="76DB5DFE18020005236DFEEEFB5BFE2A" box="[1042,1200,357,445]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B94EEAD21802FFFF2352FEFEFB97FE3A" bold="true" box="[1069,1173,373,429]" pageId="5" pageNumber="6">Paratype male</emphasis>
</th>
<th id="76DB5DFE1802000523A4FEEEFABCFE2A" box="[1243,1367,357,445]" gridcol="6" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B94EEAD21802FFFF2399FEEEFAAAFE2A" bold="true" pageId="5" pageNumber="6">Paratype of obscure gender</emphasis>
</th>
</tr>
<tr id="350A34821802000527B7FE5BFABCFE7E" box="[200,1367,464,489]" gridrow="1" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FE5BFE7CFE7E" box="[200,407,464,489]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Inventory number</th>
<td id="76DB5DFE18020005268AFE5BFDA7FE7E" box="[501,588,464,489]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">SMF 51</td>
<td id="76DB5DFE180200052506FE5BFD3BFE7E" box="[633,720,464,489]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">SMF 52</td>
<td id="76DB5DFE180200052582FE5BFCBFFE7E" box="[765,852,464,489]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">SMF 53</td>
<td id="76DB5DFE1802000524F8FE5BFC0DFE7E" box="[903,998,464,489]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">SMF 54</td>
<td id="76DB5DFE18020005236DFE5BFB5BFE7E" box="[1042,1200,464,489]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">SMF 58</td>
<td id="76DB5DFE1802000523A4FE5BFABCFE7E" box="[1243,1367,464,489]" gridcol="6" gridrow="1" pageId="5" pageNumber="6">SMF 60</td>
</tr>
<tr id="350A34821802000527B7FE73FABCFD87" box="[200,1367,504,528]" gridrow="2" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FE73FE7CFD87" box="[200,407,504,528]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Body length</th>
<td id="76DB5DFE18020005268AFE73FDA7FD87" box="[501,588,504,528]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">167</td>
<td id="76DB5DFE180200052506FE73FD3BFD87" box="[633,720,504,528]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">181</td>
<td id="76DB5DFE180200052582FE73FCBFFD87" box="[765,852,504,528]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">195</td>
<td id="76DB5DFE1802000524F8FE73FC0DFD87" box="[903,998,504,528]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">177</td>
<td id="76DB5DFE18020005236DFE73FB5BFD87" box="[1042,1200,504,528]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">166</td>
<td id="76DB5DFE1802000523A4FE73FABCFD87" box="[1243,1367,504,528]" gridcol="6" gridrow="2" pageId="5" pageNumber="6">161</td>
</tr>
<tr id="350A34821802000527B7FDABFABCFDAF" box="[200,1367,544,568]" gridrow="3" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FDABFE7CFDAF" box="[200,407,544,568]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Median cirrus</th>
<td id="76DB5DFE18020005268AFDABFDA7FDAF" box="[501,588,544,568]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">42</td>
<td id="76DB5DFE180200052506FDABFD3BFDAF" box="[633,720,544,568]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">57</td>
<td id="76DB5DFE180200052582FDABFCBFFDAF" box="[765,852,544,568]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">67</td>
<td id="76DB5DFE1802000524F8FDABFC0DFDAF" box="[903,998,544,568]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">45</td>
<td id="76DB5DFE18020005236DFDABFB5BFDAF" box="[1042,1200,544,568]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">63</td>
<td id="76DB5DFE1802000523A4FDABFABCFDAF" box="[1243,1367,544,568]" gridcol="6" gridrow="3" pageId="5" pageNumber="6">39</td>
</tr>
<tr id="350A34821802000527B7FDC3FABCFDF7" box="[200,1367,584,608]" gridrow="4" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FDC3FE7CFDF7" box="[200,407,584,608]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Internal cirrus</th>
<td id="76DB5DFE18020005268AFDC3FDA7FDF7" box="[501,588,584,608]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">26</td>
<td id="76DB5DFE180200052506FDC3FD3BFDF7" box="[633,720,584,608]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">29</td>
<td id="76DB5DFE180200052582FDC3FCBFFDF7" box="[765,852,584,608]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">18</td>
<td id="76DB5DFE1802000524F8FDC3FC0DFDF7" box="[903,998,584,608]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">23</td>
<td id="76DB5DFE18020005236DFDC3FB5BFDF7" box="[1042,1200,584,608]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">25</td>
<td id="76DB5DFE1802000523A4FDC3FABCFDF7" box="[1243,1367,584,608]" gridcol="6" gridrow="4" pageId="5" pageNumber="6">17.5</td>
</tr>
<tr id="350A34821802000527B7FDE4FABCFD1F" box="[200,1367,623,648]" gridrow="5" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FDE4FE7CFD1F" box="[200,407,623,648]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">External cirrus</th>
<td id="76DB5DFE18020005268AFDE4FDA7FD1F" box="[501,588,623,648]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">47</td>
<td id="76DB5DFE180200052506FDE4FD3BFD1F" box="[633,720,623,648]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">50</td>
<td id="76DB5DFE180200052582FDE4FCBFFD1F" box="[765,852,623,648]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">54</td>
<td id="76DB5DFE1802000524F8FDE4FC0DFD1F" box="[903,998,623,648]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">47</td>
<td id="76DB5DFE18020005236DFDE4FB5BFD1F" box="[1042,1200,623,648]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">43</td>
<td id="76DB5DFE1802000523A4FDE4FABCFD1F" box="[1243,1367,623,648]" gridcol="6" gridrow="5" pageId="5" pageNumber="6">45</td>
</tr>
<tr id="350A34821802000527B7FD1CFABCFD27" box="[200,1367,663,688]" gridrow="6" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FD1CFE7CFD27" box="[200,407,663,688]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Cirrus A</th>
<td id="76DB5DFE18020005268AFD1CFDA7FD27" box="[501,588,663,688]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">62</td>
<td id="76DB5DFE180200052506FD1CFD3BFD27" box="[633,720,663,688]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">54</td>
<td id="76DB5DFE180200052582FD1CFCBFFD27" box="[765,852,663,688]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">61</td>
<td id="76DB5DFE1802000524F8FD1CFC0DFD27" box="[903,998,663,688]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">65</td>
<td id="76DB5DFE18020005236DFD1CFB5BFD27" box="[1042,1200,663,688]" gridcol="5" gridrow="6" pageId="5" pageNumber="6">62</td>
<td id="76DB5DFE1802000523A4FD1CFABCFD27" box="[1243,1367,663,688]" gridcol="6" gridrow="6" pageId="5" pageNumber="6">50</td>
</tr>
<tr id="350A34821802000527B7FD34FABCFD40" box="[200,1367,703,727]" gridrow="7" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FD34FE7CFD40" box="[200,407,703,727]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Primary clava</th>
<td id="76DB5DFE18020005268AFD34FDA7FD40" box="[501,588,703,727]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">6.5</td>
<td id="76DB5DFE180200052506FD34FD3BFD40" box="[633,720,703,727]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">6</td>
<td id="76DB5DFE180200052582FD34FCBFFD40" box="[765,852,703,727]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">7</td>
<td id="76DB5DFE1802000524F8FD34FC0DFD40" box="[903,998,703,727]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">5.4</td>
<td id="76DB5DFE18020005236DFD34FB5BFD40" box="[1042,1200,703,727]" gridcol="5" gridrow="7" pageId="5" pageNumber="6">7</td>
<td id="76DB5DFE1802000523A4FD34FABCFD40" box="[1243,1367,703,727]" gridcol="6" gridrow="7" pageId="5" pageNumber="6">6</td>
</tr>
<tr id="350A34821802000527B7FD6CFABCFC88" box="[200,1367,743,799]" gridrow="8" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FD6CFE7CFC88" box="[200,407,743,799]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">Secondary clava length × width</th>
<td id="76DB5DFE18020005268AFD6CFDA7FC88" box="[501,588,743,799]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">7.5 × 7</td>
<td id="76DB5DFE180200052506FD6CFD3BFC88" box="[633,720,743,799]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">8 × 7</td>
<td id="76DB5DFE180200052582FD6CFCBFFC88" box="[765,852,743,799]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">6.5 × 9</td>
<td id="76DB5DFE1802000524F8FD6CFC0DFC88" box="[903,998,743,799]" gridcol="4" gridrow="8" pageId="5" pageNumber="6">5.6 × 7.3</td>
<td id="76DB5DFE18020005236DFD6CFB5BFC88" box="[1042,1200,743,799]" gridcol="5" gridrow="8" pageId="5" pageNumber="6">6.5 × 7.6</td>
<td id="76DB5DFE1802000523A4FD6CFABCFC88" box="[1243,1367,743,799]" gridcol="6" gridrow="8" pageId="5" pageNumber="6">6 × 7</td>
</tr>
<tr id="350A34821802000527B7FCA4FABCFCF0" box="[200,1367,815,871]" gridrow="9" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FCA4FE7CFCF0" box="[200,407,815,871]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">Dorsal spine I</th>
<td id="76DB5DFE18020005268AFCA4FDA7FCF0" box="[501,588,815,871]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">42</td>
<td id="76DB5DFE180200052506FCA4FD3BFCF0" box="[633,720,815,871]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">?</td>
<td id="76DB5DFE180200052582FCA4FCBFFCF0" box="[765,852,815,871]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">67</td>
<td id="76DB5DFE1802000524F8FCA4FC0DFCF0" box="[903,998,815,871]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE18020005236DFCA4FB5BFCF0" box="[1042,1200,815,871]" gridcol="5" gridrow="9" pageId="5" pageNumber="6">64</td>
<td id="76DB5DFE1802000523A4FCA4FABCFCF0" box="[1243,1367,815,871]" gridcol="6" gridrow="9" pageId="5" pageNumber="6">48</td>
</tr>
<tr id="350A34821802000527B7FCFCFABCFC38" box="[200,1367,887,943]" gridrow="10" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FCFCFE7CFC38" box="[200,407,887,943]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Dorsal spine II</th>
<td id="76DB5DFE18020005268AFCFCFDA7FC38" box="[501,588,887,943]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">26</td>
<td id="76DB5DFE180200052506FCFCFD3BFC38" box="[633,720,887,943]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">44</td>
<td id="76DB5DFE180200052582FCFCFCBFFC38" box="[765,852,887,943]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">20</td>
<td id="76DB5DFE1802000524F8FCFCFC0DFC38" box="[903,998,887,943]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE18020005236DFCFCFB5BFC38" box="[1042,1200,887,943]" gridcol="5" gridrow="10" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE1802000523A4FCFCFABCFC38" box="[1243,1367,887,943]" gridcol="6" gridrow="10" pageId="5" pageNumber="6">47</td>
</tr>
<tr id="350A34821802000527B7FC35FABCFC61" box="[200,1367,958,1014]" gridrow="11" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FC35FE7CFC61" box="[200,407,958,1014]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">Dorsal spine III</th>
<td id="76DB5DFE18020005268AFC35FDA7FC61" box="[501,588,958,1014]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">31</td>
<td id="76DB5DFE180200052506FC35FD3BFC61" box="[633,720,958,1014]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">39</td>
<td id="76DB5DFE180200052582FC35FCBFFC61" box="[765,852,958,1014]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">18</td>
<td id="76DB5DFE1802000524F8FC35FC0DFC61" box="[903,998,958,1014]" gridcol="4" gridrow="11" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE18020005236DFC35FB5BFC61" box="[1042,1200,958,1014]" gridcol="5" gridrow="11" pageId="5" pageNumber="6">29</td>
<td id="76DB5DFE1802000523A4FC35FABCFC61" box="[1243,1367,958,1014]" gridcol="6" gridrow="11" pageId="5" pageNumber="6">46</td>
</tr>
<tr id="350A34821802000527B7FB8DFABCFBA9" box="[200,1367,1030,1086]" gridrow="12" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FB8DFE7CFBA9" box="[200,407,1030,1086]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">Dorsal spine IV</th>
<td id="76DB5DFE18020005268AFB8DFDA7FBA9" box="[501,588,1030,1086]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">16</td>
<td id="76DB5DFE180200052506FB8DFD3BFBA9" box="[633,720,1030,1086]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">29</td>
<td id="76DB5DFE180200052582FB8DFCBFFBA9" box="[765,852,1030,1086]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">16</td>
<td id="76DB5DFE1802000524F8FB8DFC0DFBA9" box="[903,998,1030,1086]" gridcol="4" gridrow="12" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE18020005236DFB8DFB5BFBA9" box="[1042,1200,1030,1086]" gridcol="5" gridrow="12" pageId="5" pageNumber="6">18</td>
<td id="76DB5DFE1802000523A4FB8DFABCFBA9" box="[1243,1367,1030,1086]" gridcol="6" gridrow="12" pageId="5" pageNumber="6">27</td>
</tr>
<tr id="350A34821802000527B7FBC5FABCFB11" box="[200,1367,1102,1158]" gridrow="13" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FBC5FE7CFB11" box="[200,407,1102,1158]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">Dorsal spine V</th>
<td id="76DB5DFE18020005268AFBC5FDA7FB11" box="[501,588,1102,1158]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">6</td>
<td id="76DB5DFE180200052506FBC5FD3BFB11" box="[633,720,1102,1158]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">9</td>
<td id="76DB5DFE180200052582FBC5FCBFFB11" box="[765,852,1102,1158]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">6</td>
<td id="76DB5DFE1802000524F8FBC5FC0DFB11" box="[903,998,1102,1158]" gridcol="4" gridrow="13" pageId="5" pageNumber="6">missing (lost?)</td>
<td id="76DB5DFE18020005236DFBC5FB5BFB11" box="[1042,1200,1102,1158]" gridcol="5" gridrow="13" pageId="5" pageNumber="6">2</td>
<td id="76DB5DFE1802000523A4FBC5FABCFB11" box="[1243,1367,1102,1158]" gridcol="6" gridrow="13" pageId="5" pageNumber="6">3</td>
</tr>
<tr id="350A34821802000527B7FB1DFABCFB39" box="[200,1367,1174,1198]" gridrow="14" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FB1DFE7CFB39" box="[200,407,1174,1198]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">Cirrus E</th>
<td id="76DB5DFE18020005268AFB1DFDA7FB39" box="[501,588,1174,1198]" gridcol="1" gridrow="14" pageId="5" pageNumber="6">112</td>
<td id="76DB5DFE180200052506FB1DFD3BFB39" box="[633,720,1174,1198]" gridcol="2" gridrow="14" pageId="5" pageNumber="6">113</td>
<td id="76DB5DFE180200052582FB1DFCBFFB39" box="[765,852,1174,1198]" gridcol="3" gridrow="14" pageId="5" pageNumber="6">129</td>
<td id="76DB5DFE1802000524F8FB1DFC0DFB39" box="[903,998,1174,1198]" gridcol="4" gridrow="14" pageId="5" pageNumber="6">97</td>
<td id="76DB5DFE18020005236DFB1DFB5BFB39" box="[1042,1200,1174,1198]" gridcol="5" gridrow="14" pageId="5" pageNumber="6">121</td>
<td id="76DB5DFE1802000523A4FB1DFABCFB39" box="[1243,1367,1174,1198]" gridcol="6" gridrow="14" pageId="5" pageNumber="6">118</td>
</tr>
<tr id="350A34821802000527B7FB35FABCFB41" box="[200,1367,1214,1238]" gridrow="15" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FB35FE7CFB41" box="[200,407,1214,1238]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">Leg IV sense organ</th>
<td id="76DB5DFE18020005268AFB35FDA7FB41" box="[501,588,1214,1238]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">5.9</td>
<td id="76DB5DFE180200052506FB35FD3BFB41" box="[633,720,1214,1238]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">5.8</td>
<td id="76DB5DFE180200052582FB35FCBFFB41" box="[765,852,1214,1238]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">6.7</td>
<td id="76DB5DFE1802000524F8FB35FC0DFB41" box="[903,998,1214,1238]" gridcol="4" gridrow="15" pageId="5" pageNumber="6">5.1</td>
<td id="76DB5DFE18020005236DFB35FB5BFB41" box="[1042,1200,1214,1238]" gridcol="5" gridrow="15" pageId="5" pageNumber="6">5.6</td>
<td id="76DB5DFE1802000523A4FB35FABCFB41" box="[1243,1367,1214,1238]" gridcol="6" gridrow="15" pageId="5" pageNumber="6">3.8</td>
</tr>
<tr id="350A34821802000527B7FB6EFABCFB69" box="[200,1367,1253,1278]" gridrow="16" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FB6EFE7CFB69" box="[200,407,1253,1278]" gridcol="0" gridrow="16" pageId="5" pageNumber="6">Leg I digits</th>
<td id="76DB5DFE18020005268AFB6EFDA7FB69" box="[501,588,1253,1278]" gridcol="1" gridrow="16" pageId="5" pageNumber="6">21</td>
<td id="76DB5DFE180200052506FB6EFD3BFB69" box="[633,720,1253,1278]" gridcol="2" gridrow="16" pageId="5" pageNumber="6">22</td>
<td id="76DB5DFE180200052582FB6EFCBFFB69" box="[765,852,1253,1278]" gridcol="3" gridrow="16" pageId="5" pageNumber="6">17</td>
<td id="76DB5DFE1802000524F8FB6EFC0DFB69" box="[903,998,1253,1278]" gridcol="4" gridrow="16" pageId="5" pageNumber="6">18</td>
<td id="76DB5DFE18020005236DFB6EFB5BFB69" box="[1042,1200,1253,1278]" gridcol="5" gridrow="16" pageId="5" pageNumber="6">20</td>
<td id="76DB5DFE1802000523A4FB6EFABCFB69" box="[1243,1367,1253,1278]" gridcol="6" gridrow="16" pageId="5" pageNumber="6">16</td>
</tr>
<tr id="350A34821802000527B7FA86FABCFAB1" box="[200,1367,1293,1318]" gridrow="17" pageId="5" pageNumber="6">
<th id="76DB5DFE1802000527B7FA86FE7CFAB1" box="[200,407,1293,1318]" gridcol="0" gridrow="17" pageId="5" pageNumber="6">Leg IV digits</th>
<td id="76DB5DFE18020005268AFA86FDA7FAB1" box="[501,588,1293,1318]" gridcol="1" gridrow="17" pageId="5" pageNumber="6">26</td>
<td id="76DB5DFE180200052506FA86FD3BFAB1" box="[633,720,1293,1318]" gridcol="2" gridrow="17" pageId="5" pageNumber="6">25</td>
<td id="76DB5DFE180200052582FA86FCBFFAB1" box="[765,852,1293,1318]" gridcol="3" gridrow="17" pageId="5" pageNumber="6">26</td>
<td id="76DB5DFE1802000524F8FA86FC0DFAB1" box="[903,998,1293,1318]" gridcol="4" gridrow="17" pageId="5" pageNumber="6">26</td>
<td id="76DB5DFE18020005236DFA86FB5BFAB1" box="[1042,1200,1293,1318]" gridcol="5" gridrow="17" pageId="5" pageNumber="6">24</td>
<td id="76DB5DFE1802000523A4FA86FABCFAB1" box="[1243,1367,1293,1318]" gridcol="6" gridrow="17" pageId="5" pageNumber="6">23</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8536C01802FFF127C2F90CFA9CFEDF" blockId="5.[189,1399,1670,2014]" lastBlockId="11.[189,1399,267,329]" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6">
<collectionCode id="ED2BAE051802FFFF27C2F90CFE45F936" box="[189,430,1671,1697]" pageId="5" pageNumber="6">BODY SEGMENTS I–III</collectionCode>
. With long lateral processes (lp) almost normal to long body axis, ornamented by rows of long hollow spines on anterior and posterior edges; terminal narrowed region of processes lack spines and slightly curved rearward (
<figureCitation id="13012A451802FFFF2516F946FD5AF970" box="[617,689,1741,1767]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
A–B, 4E, 5A, 6A–B, D). Spines make processes feather-like. Numbers and position of spines on body lateral projections: (1) about 10 medium-sized spines arranged in arch dorso-posteriorly near bases of processes; (2) about 19–22 long marginal spines along anterior edge of process; (3) about 14-16 also long marginal spines along posterior edge of process (both anterior and posterior rows of marginal sets of dorso-lateral process give impression of feather). Two to five short and stout but acute ventral conical spikes (vs) arranged in row parallel to long body axis on ventral side of each lateral body process (
<figureCitation id="13012A451802FFFF2568F82BFD9CF82D" box="[535,631,1952,1978]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="5" pageNumber="6">Figs 2B</figureCitation>
,
<figureCitation id="13012A451802FFFF25FAF82BFD40F82C" box="[645,683,1952,1979]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[212,1352,277,1849]" captionTargetId="figure@9.[195,1380,262,1867]" captionTargetPageId="9" captionText="Fig. 5. Neostygarctus grossmeteori sp. nov. Paratype, ♀ (SMF 52), details. A. Lateral body processes, ventral view. B–D. Legs I (left and right) and IV. E. Genital area. Scale bars: A = 50 μm; B–E = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3825271" httpUri="https://zenodo.org/record/3825271/files/figure.png" pageId="5" pageNumber="6">5A</figureCitation>
,
<figureCitation id="13012A451802FFFF25C7F82BFD35F82C" box="[696,734,1952,1979]" captionStart="Fig" captionStartId="10.[189,232,1809,1835]" captionTargetBox="[232,1358,272,1771]" captionTargetId="figure@10.[219,1368,262,1772]" captionTargetPageId="10" captionText="Fig. 6. Neostygarctus grossmeteori sp. nov., details, SEM. A. Female, entire body, ventral view. B. Right secondary clava and outer cirrus on the head, ventral view. C. Ventral conical spikes on the basal part of the lateral body process. D. Right lateral body processes. E. Lateral fan of spines with membrane on the posteriormost body segment. F. Toes with claws of the leg IV ventrally, dorsal tendon detached in some toes. G. Inner and outer claws, dorsal view. H. Accordion-like joint of the cirrus E. Scale bars: A = 30 μm; B, D, F = 10 μm; C, E, G–H = 3 μm." figureDoi="http://doi.org/10.5281/zenodo.3825273" httpUri="https://zenodo.org/record/3825273/files/figure.png" pageId="5" pageNumber="6">6C</figureCitation>
). Pair of wart-like latero-dorsal lobes (ldl) on each body segment just posterior to long lateral processes on segments
<collectionCode id="ED2BAE051802FFFF24FCF84FFC53F849" box="[899,952,1988,2014]" pageId="5" pageNumber="6">I–III</collectionCode>
and in the middle on caudal segment. In shape, they resemble cervical latero-dorsal lobes. Caudal segment bears another pair of caudal dorsal wart-like lobes (cdl) on posterior edge. All wart-like lobes armed with five to seven medium-sized spines.
</paragraph>
<caption id="DF4566481801FFFC27C3F922F87DF952" ID-DOI="http://doi.org/10.5281/zenodo.3825263" ID-Zenodo-Dep="3825263" box="[188,1942,1705,1733]" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="6" pageNumber="7" startId="6.[188,231,1705,1733]" targetBox="[222,1898,580,1610]" targetPageId="6">
<paragraph id="8B8536C01801FFFC27C3F922F87DF952" blockId="6.[188,1942,1705,1733]" box="[188,1942,1705,1733]" pageId="6" pageNumber="7">
<emphasis id="B94EEAD21801FFFC27C3F922FEE2F952" bold="true" box="[188,265,1705,1733]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
<taxonomicName id="4C3A4D431801FFFC2670F922FDBFF952" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[271,596,1705,1733]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD21801FFFC2670F922FDBFF952" box="[271,596,1705,1733]" italics="true" pageId="6" pageNumber="7">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A91801FFFC2524F922FD5CF952" box="[603,695,1705,1733]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, entire.
<emphasis id="B94EEAD21801FFFC246DF922FCCCF952" bold="true" box="[786,807,1705,1733]" pageId="6" pageNumber="7">A</emphasis>
. Holotype, ♀ (SMF 51), dorsal view.
<emphasis id="B94EEAD21801FFFC23A0F922FB18F952" bold="true" box="[1247,1267,1705,1733]" pageId="6" pageNumber="7">B</emphasis>
. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm.
</paragraph>
</caption>
<caption id="DF4566481800FFFD27C2F818FD50F846" ID-DOI="http://doi.org/10.5281/zenodo.3825265" ID-Zenodo-Dep="3825265" httpUri="https://zenodo.org/record/3825265/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1939,1965]" targetBox="[201,1368,303,1824]" targetPageId="7">
<paragraph id="8B8536C01800FFFD27C2F818FD50F846" blockId="7.[189,1398,1939,2001]" pageId="7" pageNumber="8">
<emphasis id="B94EEAD21800FFFD27C2F818FEE3F83A" bold="true" box="[189,264,1939,1965]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
<taxonomicName id="4C3A4D431800FFFD2672F81FFDBAF83A" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[269,593,1940,1965]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD21800FFFD2672F81FFDBAF83A" box="[269,593,1940,1965]" italics="true" pageId="7" pageNumber="8">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A91800FFFD2529F81FFD5AF839" box="[598,689,1940,1966]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, heads.
<emphasis id="B94EEAD21800FFFD2475F818FCF4F83A" bold="true" box="[778,799,1939,1965]" pageId="7" pageNumber="8">A</emphasis>
. Holotype, ♀ (SMF 51), dorsal view.
<emphasis id="B94EEAD21800FFFD23B2F818FB0AF83A" bold="true" box="[1229,1249,1939,1965]" pageId="7" pageNumber="8">B</emphasis>
. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm.
</paragraph>
</caption>
<caption id="DF456648180FFFF227C2F8DCFAA1F84B" ID-DOI="http://doi.org/10.5281/zenodo.3825269" ID-Zenodo-Dep="3825269" httpUri="https://zenodo.org/record/3825269/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1879,1905]" targetBox="[195,1393,314,1784]" targetPageId="8">
<paragraph id="8B8536C0180FFFF227C2F8DCFAA1F84B" blockId="8.[189,1399,1879,2012]" pageId="8" pageNumber="9">
<emphasis id="B94EEAD2180FFFF227C2F8DCFEE5F8E5" bold="true" box="[189,270,1879,1906]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
<taxonomicName id="4C3A4D43180FFFF22666F8D2FD89F8E5" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[281,610,1881,1906]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="9" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD2180FFFF22666F8D2FD89F8E5" box="[281,610,1881,1906]" italics="true" pageId="8" pageNumber="9">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A9180FFFF22512F8D3FD25F8E5" box="[621,718,1880,1906]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, optical photopictures.
<emphasis id="B94EEAD2180FFFF22495F8DCFC14F8E6" bold="true" box="[1002,1023,1879,1905]" pageId="8" pageNumber="9">A</emphasis>
. Paratype, ♀ (SMF 54), entire body, dorsal view.
<emphasis id="B94EEAD2180FFFF226EAF8F0FE42F802" bold="true" box="[405,425,1915,1941]" pageId="8" pageNumber="9">B</emphasis>
. Holotype, ♀ (SMF 51), entire body, ventral view.
<emphasis id="B94EEAD2180FFFF22489F8F0FBC6F802" bold="true" box="[1014,1069,1915,1941]" pageId="8" pageNumber="9">C–E</emphasis>
. Paratype of obscure gender (SMF 59).
<emphasis id="B94EEAD2180FFFF22644F815FE9FF82F" bold="true" box="[315,372,1950,1976]" pageId="8" pageNumber="9">C–D</emphasis>
. Areas of dorsal surface of body with spines.
<emphasis id="B94EEAD2180FFFF22401F815FC79F82F" bold="true" box="[894,914,1950,1976]" pageId="8" pageNumber="9">E</emphasis>
. Lateral body projections.
<emphasis id="B94EEAD2180FFFF223B8F815FB32F82F" bold="true" box="[1223,1241,1950,1976]" pageId="8" pageNumber="9">F</emphasis>
. Holotype, ♀ (SMF 51), posterior body with female gonopore and anus. Scale bars: A–B = 50 μm; C–F = 20 μm.
</paragraph>
</caption>
<caption id="DF456648180EFFF327C2F80FFA9CF855" ID-DOI="http://doi.org/10.5281/zenodo.3825271" ID-Zenodo-Dep="3825271" httpUri="https://zenodo.org/record/3825271/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1924,1950]" targetBox="[212,1352,277,1849]" targetPageId="9">
<paragraph id="8B8536C0180EFFF327C2F80FFA9CF855" blockId="9.[189,1399,1924,1986]" pageId="9" pageNumber="10">
<emphasis id="B94EEAD2180EFFF327C2F80FFEE0F808" bold="true" box="[189,267,1924,1951]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
<taxonomicName id="4C3A4D43180EFFF3266BF80EFDB0F809" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[276,603,1925,1950]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD2180EFFF3266BF80EFDB0F809" box="[276,603,1925,1950]" italics="true" pageId="9" pageNumber="10">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A9180EFFF3251CF80EFD54F808" box="[611,703,1925,1951]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
Paratype, ♀ (SMF 52), details.
<emphasis id="B94EEAD2180EFFF32344F80FFBBBF809" bold="true" box="[1083,1104,1924,1950]" pageId="9" pageNumber="10">A</emphasis>
. Lateral body processes, ventral view.
<emphasis id="B94EEAD2180EFFF32624F823FE79F855" bold="true" box="[347,402,1960,1986]" pageId="9" pageNumber="10">B–D</emphasis>
. Legs I (left and right) and IV.
<emphasis id="B94EEAD2180EFFF3247FF823FCFFF855" bold="true" box="[768,788,1960,1986]" pageId="9" pageNumber="10">E</emphasis>
. Genital area. Scale bars: A = 50 μm; B–E = 20 μm.
</paragraph>
</caption>
<caption id="DF456648180DFFF027C2F89AFD2CF84B" ID-DOI="http://doi.org/10.5281/zenodo.3825273" ID-Zenodo-Dep="3825273" httpUri="https://zenodo.org/record/3825273/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1809,1835]" targetBox="[232,1358,272,1771]" targetPageId="10">
<paragraph id="8B8536C0180DFFF027C2F89AFD2CF84B" blockId="10.[189,1399,1809,2012]" pageId="10" pageNumber="11">
<emphasis id="B94EEAD2180DFFF027C2F89AFEE2F8BC" bold="true" box="[189,265,1809,1835]" pageId="10" pageNumber="11">Fig. 6.</emphasis>
<taxonomicName id="4C3A4D43180DFFF02670F899FDBFF8BC" authority="Tchesunov, 2018" authorityName="Tchesunov" authorityYear="2018" box="[271,596,1810,1835]" class="Heterotardigrada" family="Neostygarctidae" genus="Neostygarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="grossmeteori" status="sp. nov.">
<emphasis id="B94EEAD2180DFFF02670F899FDBFF8BC" box="[271,596,1810,1835]" italics="true" pageId="10" pageNumber="11">Neostygarctus grossmeteori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A9180DFFF02525F899FD5DF8BB" box="[602,694,1810,1836]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, details, SEM.
<emphasis id="B94EEAD2180DFFF0241CF89AFC93F8BC" bold="true" box="[867,888,1809,1835]" pageId="10" pageNumber="11">A</emphasis>
. Female, entire body, ventral view.
<emphasis id="B94EEAD2180DFFF0226BF899FAC3F8BB" bold="true" box="[1300,1320,1810,1836]" pageId="10" pageNumber="11">B</emphasis>
. Right secondary clava and outer cirrus on the head, ventral view.
<emphasis id="B94EEAD2180DFFF024FEF8BFFC7DF8D9" bold="true" box="[897,918,1844,1870]" pageId="10" pageNumber="11">C</emphasis>
. Ventral conical spikes on the basal part of the lateral body process.
<emphasis id="B94EEAD2180DFFF0257DF8D3FDFCF8E5" bold="true" box="[514,535,1880,1906]" pageId="10" pageNumber="11">D</emphasis>
. Right lateral body processes.
<emphasis id="B94EEAD2180DFFF024FFF8D3FC7FF8E5" bold="true" box="[896,916,1880,1906]" pageId="10" pageNumber="11">E</emphasis>
. Lateral fan of spines with membrane on the posteriormost body segment.
<emphasis id="B94EEAD2180DFFF02537F8F0FDB1F802" bold="true" box="[584,602,1915,1941]" pageId="10" pageNumber="11">F</emphasis>
. Toes with claws of the leg IV ventrally, dorsal tendon detached in some toes.
<emphasis id="B94EEAD2180DFFF02641F815FEBEF82F" bold="true" box="[318,341,1950,1976]" pageId="10" pageNumber="11">G</emphasis>
. Inner and outer claws, dorsal view.
<emphasis id="B94EEAD2180DFFF02580F815FCFDF82F" bold="true" box="[767,790,1950,1976]" pageId="10" pageNumber="11">H</emphasis>
. Accordion-like joint of the cirrus E. Scale bars: A = 30 μm; B, D, F = 10 μm; C, E, G–H = 3 μm.
</paragraph>
</caption>
<paragraph id="8B8536C0180CFFF127C2FEFBFF1AFC29" blockId="11.[189,1399,368,958]" pageId="11" pageNumber="12">
SENSORY ORGANS. Long anterior flagellar median cirrus (mc) rooted on prominent cirrophore hanging over anterior edge of head (
<figureCitation id="13012A45180CFFF126BAFE18FDE4FE3A" box="[453,527,403,429]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
A–B, 3A–B). Internal cirri (ic) consist of short stub-shaped cirrophore and whip-like flagellum situated ventrally to anterior frontal lobes (
<figureCitation id="13012A45180CFFF124D6FE3DFC05FE47" box="[937,1006,438,464]" captionStart="Fig" captionStartId="7.[189,232,1939,1965]" captionTargetBox="[201,1368,303,1824]" captionTargetId="figure@7.[189,1398,262,1892]" captionTargetPageId="7" captionText="Fig. 3. Neostygarctus grossmeteori sp. nov., heads. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3825265" httpUri="https://zenodo.org/record/3825265/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
A–B). External cirri (ec) situated ventrally to fronto-lateral lobes and also consist of short stub-shaped cirrophore and longer whip-like flagellum; similar in shape to internal cirri but a bit longer (
<figureCitation id="13012A45180CFFF124F7FE76FC38FD80" box="[904,979,509,535]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
A–B, 3A–B, 6B). All cephalic cirri (flagellar parts) possess minute and sparse irregular lateral sprouts visible in SEM (
<figureCitation id="13012A45180CFFF1230EFDABFB21FDAC" box="[1137,1226,544,571]" captionStart="Fig" captionStartId="10.[189,232,1809,1835]" captionTargetBox="[232,1358,272,1771]" captionTargetId="figure@10.[219,1368,262,1772]" captionTargetPageId="10" captionText="Fig. 6. Neostygarctus grossmeteori sp. nov., details, SEM. A. Female, entire body, ventral view. B. Right secondary clava and outer cirrus on the head, ventral view. C. Ventral conical spikes on the basal part of the lateral body process. D. Right lateral body processes. E. Lateral fan of spines with membrane on the posteriormost body segment. F. Toes with claws of the leg IV ventrally, dorsal tendon detached in some toes. G. Inner and outer claws, dorsal view. H. Accordion-like joint of the cirrus E. Scale bars: A = 30 μm; B, D, F = 10 μm; C, E, G–H = 3 μm." figureDoi="http://doi.org/10.5281/zenodo.3825273" httpUri="https://zenodo.org/record/3825273/files/figure.png" pageId="11" pageNumber="12">Fig. 6B</figureCitation>
). External cirri located close to secondary clava just ventrally of fronto-lateral lobe edge. Globular secondary clava on short narrow stalk situated at inner position to external cirrus (
<figureCitation id="13012A45180CFFF124E6FDEDFC09FD17" box="[921,994,614,640]" captionStart="Fig" captionStartId="7.[189,232,1939,1965]" captionTargetBox="[201,1368,303,1824]" captionTargetId="figure@7.[189,1398,262,1892]" captionTargetPageId="7" captionText="Fig. 3. Neostygarctus grossmeteori sp. nov., heads. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3825265" httpUri="https://zenodo.org/record/3825265/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
A–B, 6B). Cirrus A (lateral cirrus) and small primary clava rooted together on common cylindrical cirrophore (cc) which is continuation of lateral head lobe (
<figureCitation id="13012A45180CFFF126D0FD26FE13FD50" box="[431,504,685,711]" captionStart="Fig" captionStartId="7.[189,232,1939,1965]" captionTargetBox="[201,1368,303,1824]" captionTargetId="figure@7.[189,1398,262,1892]" captionTargetPageId="7" captionText="Fig. 3. Neostygarctus grossmeteori sp. nov., heads. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, ♀ (SMF 52), ventral view. Scale bar = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3825265" httpUri="https://zenodo.org/record/3825265/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
A–B, 6B); primary clava ovoid to lemon-shaped, with pointed terminal end. In some female and male specimens, paired ventral transversal slits (ts) observed just posterior to legs III (
<figureCitation id="13012A45180CFFF12793FD78FEA2FC9A" box="[236,329,755,781]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Fig. 2B</figureCitation>
); they are considered as possible sensory organs. No sensory organs present on legs
<collectionCode id="ED2BAE05180CFFF12245FD7FFA98FC99" box="[1338,1395,756,782]" pageId="11" pageNumber="12">I–III</collectionCode>
. Cirri E (cE) on caudal segment growing on body postero-laterally. Each cirrus E consists of long robust cirrophore without any discernible tubercles, cylindrical accordion-like thickening composed of three to five rings, and long whip-like flagellum (
<figureCitation id="13012A45180CFFF125C2FCD6FCECFCE0" box="[701,775,861,887]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
A–B, 6H). Legs IV provided with latero-dorsal sense organs (se4) on external side of coxae. Sense organs small, ovoid, with tiny apical papillum (
<figureCitation id="13012A45180CFFF12390FC0BFAD3FC0D" box="[1263,1336,896,922]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
A–B, 5D).
</paragraph>
<paragraph id="8B8536C0180CFFF127C2FC6EFB2FFB44" blockId="11.[189,1399,997,1235]" pageId="11" pageNumber="12">
<collectionCode id="ED2BAE05180CFFF127C2FC6EFEDFFC68" box="[189,308,997,1023]" pageId="11" pageNumber="12">LEGS I–IV</collectionCode>
. Rather long, consisting of long truncate conical coxa/femur, telescoping cylindrical tibia and palm-shaped (hand-shaped in
<bibRefCitation id="EFAB4B31180CFFF12563FB83FCF6FBB4" author="Kristensen R. M. &amp; Sorensen M. V. &amp; Hansen J. G. &amp; Zeppilli D." box="[540,797,1032,1059]" pageId="11" pageNumber="12" pagination="453 - 467" refId="ref7306" refString="Kristensen R. M., Sorensen M. V., Hansen J. G., Zeppilli D. 2015. A new species of Neostygarctus (Arthrotardigrada) from the Condor Seamount in the Azores, Northeast Atlantic. Marine Biodiversity 45 (3): 453 - 467. https: // doi. org / 10.1007 / s 12526 - 015 - 0323 - y" type="journal article" year="2015">
Kristensen
<emphasis id="B94EEAD2180CFFF125DFFB82FD38FBB5" box="[672,723,1032,1058]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2015
</bibRefCitation>
) tarsus with four equal digits. Border between coxa and femur indistinct. Legs
<collectionCode id="ED2BAE05180CFFF12681FBA7FDDFFBD1" box="[510,564,1068,1094]" pageId="11" pageNumber="12">I–III</collectionCode>
have neither sensory organs nor bristles. Legs IV provided with three groups of spines on coxa: (1) about ten short to medium-sized spines on anterior wart-like projection (awp) on internal side of coxa basally, (2) five to six medium-sized spines on posterior wart-like projection (pwp) on internal side of coxa right opposite to ovoid papilla, (3) about four medium-sized spines on external side of leg just ventrally to ovoid latero-dorsal sense organ (
<figureCitation id="13012A45180CFFF12327FB33FB5CFB44" box="[1112,1207,1208,1235]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[212,1352,277,1849]" captionTargetId="figure@9.[195,1380,262,1867]" captionTargetPageId="9" captionText="Fig. 5. Neostygarctus grossmeteori sp. nov. Paratype, ♀ (SMF 52), details. A. Lateral body processes, ventral view. B–D. Legs I (left and right) and IV. E. Genital area. Scale bars: A = 50 μm; B–E = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3825271" httpUri="https://zenodo.org/record/3825271/files/figure.png" pageId="11" pageNumber="12">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="8B8536C0180CFFF127C2FB71FCD7FA36" blockId="11.[189,1399,1273,1441]" pageId="11" pageNumber="12">
DIGITS. Digits of legs IV bit longer than those of legs
<collectionCode id="ED2BAE05180CFFF1245BFB71FCB6FA83" box="[804,861,1274,1300]" pageId="11" pageNumber="12">I–III</collectionCode>
. All digits of same leg about equal in length to one another. Digits long, slim and flat, with slight subterminal ventral bulge and terminal solid beak-like claws. Only internal claws provided with fine but distinct dorsal spine (accessory spine); external claws may bear very tiny appendage visible only in SEM and probably presenting vestigial spine. Peduncles (cuticular supports) not visible (
<figureCitation id="13012A45180CFFF1254BFA0DFD97FA36" box="[564,636,1414,1441]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[212,1352,277,1849]" captionTargetId="figure@9.[195,1380,262,1867]" captionTargetPageId="9" captionText="Fig. 5. Neostygarctus grossmeteori sp. nov. Paratype, ♀ (SMF 52), details. A. Lateral body processes, ventral view. B–D. Legs I (left and right) and IV. E. Genital area. Scale bars: A = 50 μm; B–E = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3825271" httpUri="https://zenodo.org/record/3825271/files/figure.png" pageId="11" pageNumber="12">Figs 5</figureCitation>
B–D, G, 6F–G).
</paragraph>
<paragraph id="8B8536C0180CFFF127C2FA43FCD1F9DB" blockId="11.[189,1398,1479,1612]" pageId="11" pageNumber="12">
<collectionCode id="ED2BAE05180CFFF127C2FA43FE7AFA48" box="[189,401,1480,1506]" pageId="11" pageNumber="12">GENITO- ANAL AREA</collectionCode>
. In females, gonopore (go) is six-lobed rosette-like structure within circle; two lateral sinuous seminal ducts (du) open posteriorly into spherical seminal receptacles (sr) (
<figureCitation id="13012A45180CFFF123E6FA60FB1FF992" box="[1177,1268,1514,1541]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[212,1352,277,1849]" captionTargetId="figure@9.[195,1380,262,1867]" captionTargetPageId="9" captionText="Fig. 5. Neostygarctus grossmeteori sp. nov. Paratype, ♀ (SMF 52), details. A. Lateral body processes, ventral view. B–D. Legs I (left and right) and IV. E. Genital area. Scale bars: A = 50 μm; B–E = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3825271" httpUri="https://zenodo.org/record/3825271/files/figure.png" pageId="11" pageNumber="12">Fig. 5E</figureCitation>
). In males, transversally-oval gonopore situated more close to anal opening (
<figureCitation id="13012A45180CFFF124CDF985FBE6F9BF" box="[946,1037,1550,1576]" captionStart="Fig" captionStartId="6.[188,231,1705,1733]" captionTargetBox="[222,1898,580,1610]" captionTargetId="figure@6.[188,1927,550,1654]" captionTargetPageId="6" captionText="Fig. 2. Neostygarctus grossmeteori sp. nov., entire. A. Holotype, ♀ (SMF 51), dorsal view. B. Paratype, Ƌ (SMF 58), ventral view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3825263" httpUri="https://zenodo.org/record/3825263/files/figure.png" pageId="11" pageNumber="12">Fig. 2B</figureCitation>
). Anal opening in both genders is longitudinal slit on anal plate posterior to gonopore.
</paragraph>
<paragraph id="8B8536C0180CFFF127C2F9F8FB51F944" blockId="11.[189,1399,1650,1747]" pageId="11" pageNumber="12">INTERNAL ORGANS. Internal organs obscure in some specimens, possibly because of maceration. Pair of wide-set internal spheric ocelli (oc) at level of lateral lobes of head; ocelli pallid and indistinct in some specimens. Pharyngeal bulb obscure. Stylets not seen. Midgut and gonad not observed.</paragraph>
</subSubSection>
</treatment>
</document>
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