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<document id="FF3A23A01E1D1E2AA0C6D2525FA25FAE" ID-CLB-Dataset="46451" ID-DOI="10.11646/zootaxa.3764.5.6" ID-GBIF-Dataset="1526b43e-9524-4978-b6de-12ddc404b3eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="229022" ID-ZooBank="B69CCDF7-53BC-4FE7-BE19-B75C5D780CC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460383928317" checkinUser="plazi" docAuthor="Chen, Jun, Wang, Bo, Zhang, Haichun &amp; Wang, Xiaoli" docDate="2014" docId="039387B67047FFFBFF13AF9A4B75B1CF" docLanguage="en" docName="zt03764p586.pdf" docOrigin="Zootaxa 3764 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tianyuprosbole Chen, Wang, Zhang &amp; Wang, 2014, gen. nov." docType="treatment" docVersion="8" lastPageNumber="582" masterDocId="FFAAFFCE7046FFFAFF84AC16480EB608" masterDocTitle="A remarkable new genus of Tettigarctidae (Insecta, Hemiptera, Cicadoidea) from the Middle Jurassic of northeastern China" masterLastPageNumber="586" masterPageNumber="581" pageNumber="582" updateTime="1698297166485" updateUser="ExternalLinkService">
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<mods:title id="C6A6DE9D18328EFB7ED1DD5A14BD4D7D">A remarkable new genus of Tettigarctidae (Insecta, Hemiptera, Cicadoidea) from the Middle Jurassic of northeastern China</mods:title>
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<mods:namePart id="9B177491371BB14B82D39A2BBD289D17">Chen, Jun</mods:namePart>
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<mods:namePart id="28FDE1AD60E6C184CDE46DE96F1CED28">Wang, Bo</mods:namePart>
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<mods:namePart id="5C2C0DBA3920A7639CE1ECE56D6BF6FE">Zhang, Haichun</mods:namePart>
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<mods:namePart id="A2DD45870BD0E5073B68A97F14BA77AF">Wang, Xiaoli</mods:namePart>
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<treatment id="039387B67047FFFBFF13AF9A4B75B1CF" ID-DOI="http://doi.org/10.5281/zenodo.6125918" ID-GBIF-Taxon="119414808" ID-Zenodo-Dep="6125918" LSID="urn:lsid:plazi:treatment:039387B67047FFFBFF13AF9A4B75B1CF" httpUri="http://treatment.plazi.org/id/039387B67047FFFBFF13AF9A4B75B1CF" lastPageNumber="582" pageId="1" pageNumber="582">
<subSubSection id="C320652B7047FFFBFF13AF9A49B4B5AD" box="[151,442,908,933]" pageId="1" pageNumber="582" type="nomenclature">
<paragraph id="8B8536A07047FFFBFF13AF9A49B4B5AD" blockId="1.[151,442,908,933]" box="[151,442,908,933]" pageId="1" pageNumber="582">
<heading id="D0CD81CC7047FFFBFF13AF9A49B4B5AD" bold="true" box="[151,442,908,933]" fontSize="10" level="2" pageId="1" pageNumber="582" reason="2">
<emphasis id="B94EEAB27047FFFBFF13AF9A49B4B5AD" bold="true" box="[151,442,908,933]" pageId="1" pageNumber="582">
<taxonomicName id="4C3A4D237047FFFBFF13AF9A4945B5AD" ID-CoL="8HS5P" box="[151,331,908,933]" class="Insecta" family="Tettigarctidae" genus="Tianyuprosbole" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94EEAB27047FFFBFF13AF9A4945B5AD" bold="true" box="[151,331,908,933]" italics="true" pageId="1" pageNumber="582">Tianyuprosbole</emphasis>
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<taxonomicNameLabel id="A27D57C97047FFFBFED5AF9A49B4B5AD" box="[337,442,908,933]" pageId="1" pageNumber="582" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C320652B7047FFFBFF13AFC24BBEB5E5" box="[151,944,980,1005]" pageId="1" pageNumber="582" type="materials_examined">
<paragraph id="8B8536A07047FFFBFF13AFC24BBEB5E5" blockId="1.[151,1437,980,1991]" box="[151,944,980,1005]" pageId="1" pageNumber="582">
<emphasis id="B94EEAB27047FFFBFF13AFC2493CB5E5" bold="true" box="[151,306,980,1005]" pageId="1" pageNumber="582">
<typeStatus id="548188027047FFFBFF13AFC248DCB5E5" box="[151,210,980,1005]" pageId="1" pageNumber="582">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3A4D237047FFFBFEBDAFC24A38B5E5" box="[313,566,980,1005]" class="Insecta" family="Tettigarctidae" genus="Tianyuprosbole" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B94EEAB27047FFFBFEBDAFC24A38B5E5" box="[313,566,980,1005]" italics="true" pageId="1" pageNumber="582">Tianyuprosbole zhengi</emphasis>
</taxonomicName>
<emphasis id="B94EEAB27047FFFBFDB9AFC24AE5B5E5" bold="true" box="[573,747,980,1005]" pageId="1" pageNumber="582">gen. et sp. nov.</emphasis>
, here designated.
</paragraph>
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<subSubSection id="C320652B7047FFFBFF43AFE14C6AB3C4" pageId="1" pageNumber="582" type="diagnosis">
<paragraph id="8B8536A07047FFFBFF43AFE14A65B275" blockId="1.[151,1437,980,1991]" pageId="1" pageNumber="582">
<emphasis id="B94EEAB27047FFFBFF43AFE1494FB218" bold="true" box="[199,321,1015,1040]" pageId="1" pageNumber="582">Diagnosis.</emphasis>
Body small and stout. Antennae well-developed; scape thick; pedicel slightly thinner and longer than scape; flagellum aristiform. Eyes large, semi-circular in lateral view. Postclypeus greatly inflated. Pronotum exceedingly expanded and concealing mesonotum, with posterior two-thirds transversely rugose. Fore femur slightly thick; hind tibia slender and long.
</paragraph>
<paragraph id="8B8536A07047FFFBFF43A89E4C6AB3C4" blockId="1.[151,1437,980,1991]" pageId="1" pageNumber="582">
Tegmen short, nearly trapezoid. Clavus and costal area long and broad. Postnodal area ornamented with ruffles, short, almost half as long as prenodal area. Nodal line wide but not well-defined. Nodus distinct. Vein ScP fused with R+M at wing base, dividing from ScP + RA at nodal line, and ending at nodus. Stem R+M branching at basal 0.21 wing length. Vein R about 0.10 wing length and two-branched; vein RA two-branched; vein
<collectingCountry id="F32D76307047FFFBFAACA8E34D44B304" box="[1320,1354,1269,1292]" name="Poland" pageId="1" pageNumber="582">RP</collectingCountry>
single, strongly curved at nodal line. Vein M fused with CuA for a distance about 0.10 wing length, branching into M1+2 and M3+4 at basal 0.48 wing length; Veins M1+2 and M3+4 branching into M1, M2, M3, and M4 nearly at the same level. Vein CuA nearly straight, but curved just beyond nodal line and then branching into CuA1 and CuA2; branch CuA1 curved at crossvein m-cua and recurved near tegmen margin; branch CuA2 long and sinuous. Vein CuP straight, ending at nodal line. Vein Pcu ending at about midpoint of wing. Vein A1 longitudinal.
</paragraph>
</subSubSection>
<subSubSection id="C320652B7047FFFBFF43A9C14C44B030" pageId="1" pageNumber="582" type="etymology">
<paragraph id="8B8536A07047FFFBFF43A9C14D50B01B" blockId="1.[151,1437,980,1991]" pageId="1" pageNumber="582">
<emphasis id="B94EEAB27047FFFBFF43A9C14940B3F8" bold="true" box="[199,334,1495,1520]" pageId="1" pageNumber="582">Etymology.</emphasis>
The generic name refers to Shandong Tianyu Museum of Nature, where the specimen is housed, and prosbole, a common suffix for the genera of the subfamily Cicadoprosbolinae
<bibRefCitation id="EFAB4B517047FFFBFBC5A9ED4D54B01B" author="Becker-Migdisova" box="[1089,1370,1531,1556]" pageId="1" pageNumber="582" refString="Becker-Migdisova, E. E. (1947) Cicadoprosbole sogutensis gen. n. sp. n., a transitional form between Permian" type="book" year="1947">Becker-Migdisova, 1947</bibRefCitation>
.
</paragraph>
<paragraph id="8B8536A07047FFFBFF43AA094C44B030" blockId="1.[151,1437,980,1991]" box="[199,1098,1567,1592]" pageId="1" pageNumber="582">
<emphasis id="B94EEAB27047FFFBFF43AA09499DB030" bold="true" box="[199,403,1567,1592]" pageId="1" pageNumber="582">Species included.</emphasis>
Only the
<typeStatus id="548188027047FFFBFD86AA364A3CB030" box="[514,562,1568,1592]" pageId="1" pageNumber="582">type</typeStatus>
species,
<taxonomicName id="4C3A4D237047FFFBFD1CAA094B9BB030" box="[664,917,1567,1592]" class="Insecta" family="Tettigarctidae" genus="Tianyuprosbole" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B94EEAB27047FFFBFD1CAA094B9BB030" box="[664,917,1567,1592]" italics="true" pageId="1" pageNumber="582">Tianyuprosbole zhengi</emphasis>
</taxonomicName>
<emphasis id="B94EEAB27047FFFBFC18AA094C44B030" bold="true" box="[924,1098,1567,1592]" pageId="1" pageNumber="582">gen. et sp. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C320652B7047FFFBFF43AA554B75B1CF" pageId="1" pageNumber="582" type="discussion">
<paragraph id="8B8536A07047FFFBFF43AA5549E2B0C0" blockId="1.[151,1437,980,1991]" pageId="1" pageNumber="582">
<emphasis id="B94EEAB27047FFFBFF43AA554935B054" bold="true" box="[199,315,1603,1628]" pageId="1" pageNumber="582">Remarks.</emphasis>
The genus undoubtedly belongs to the family
<taxonomicName id="4C3A4D237047FFFBFCD8AA554BF8B054" box="[860,1014,1603,1628]" class="Insecta" family="Tettigarctidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="family">Tettigarctidae</taxonomicName>
<emphasis id="B94EEAB27047FFFBFB84AA554C2EB054" box="[1024,1056,1603,1628]" italics="true" pageId="1" pageNumber="582">s.l.</emphasis>
on the following characteristics: postclypeus greatly inflated; pronotum expanded, with posterior area transversely rugose; nodal line of tegmen visible; branch RA forked and
<collectingCountry id="F32D76307047FFFBFE79AA9A4A11B0AB" box="[509,543,1676,1699]" name="Poland" pageId="1" pageNumber="582">RP</collectingCountry>
single; vein M four-branched, forming a short stalk with two-branched CuA; vein CuP, Pcu, and A1 simple.
</paragraph>
<paragraph id="8B8536A07047FFFBFF43AAC54B75B1CF" blockId="1.[151,1437,980,1991]" pageId="1" pageNumber="582">
The tegmen of
<taxonomicName id="4C3A4D237047FFFBFE03AAC54A3AB0E4" box="[391,564,1747,1772]" class="Insecta" family="Tettigarctidae" genus="Tianyuprosbole" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94EEAB27047FFFBFE03AAC54A3AB0E4" box="[391,564,1747,1772]" italics="true" pageId="1" pageNumber="582">Tianyuprosbole</emphasis>
</taxonomicName>
<emphasis id="B94EEAB27047FFFBFDC6AAC54ABCB0E4" bold="true" box="[578,690,1747,1772]" pageId="1" pageNumber="582">
<taxonomicNameLabel id="A27D57C97047FFFBFDC6AAC54ABCB0E4" box="[578,690,1747,1772]" pageId="1" pageNumber="582" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is similar to that of
<taxonomicName id="4C3A4D237047FFFBFC3BAAC54D92B0E3" authority="Becker-Migdisova, 1947" authorityName="Becker-Migdisova" authorityYear="1947" box="[959,1436,1747,1772]" class="Insecta" family="Cicadoprosbolidae" genus="Cicadoprosbole" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAB27047FFFBFC3BAAC54C7FB0E4" box="[959,1137,1747,1772]" italics="true" pageId="1" pageNumber="582">Cicadoprosbole</emphasis>
<bibRefCitation id="EFAB4B517047FFFBFBFBAAC54D92B0E3" author="Becker-Migdisova" box="[1151,1436,1747,1772]" pageId="1" pageNumber="582" refString="Becker-Migdisova, E. E. (1947) Cicadoprosbole sogutensis gen. n. sp. n., a transitional form between Permian" type="book" year="1947">Becker-Migdisova, 1947</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFAB4B517047FFFBFF1AAAE149B2B107" author="Becker-Migdisova" box="[158,444,1783,1808]" pageId="1" pageNumber="582" refString="Becker-Migdisova, E. E. (1947) Cicadoprosbole sogutensis gen. n. sp. n., a transitional form between Permian" type="book" year="1947">Becker-Migdisova, 1947</bibRefCitation>
;
<bibRefCitation id="EFAB4B517047FFFBFE4EAAEE4A55B118" author="Evans" box="[458,603,1783,1808]" pageId="1" pageNumber="582" refString="Evans, J. W. (1956) Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165 - 258. http: // dx. doi. org / 10.1071 / zo 9560165" type="journal article" year="1956">Evans, 1956</bibRefCitation>
), the
<typeStatus id="548188027047FFFBFD19AAEE4AC3B118" box="[669,717,1784,1808]" pageId="1" pageNumber="582">type</typeStatus>
genus of the subfamily Cicadoprosbolinae. It differs from the latter in possessing vein RA bifurcated, vein M and CuA fused as a short stalk for tegmen. The new genus can be distinguished from other genera within the subfamily Cicadoprosbolinae in the following features of the tegmen: wing shape nearly trapezoid; vein RA having two terminations; vein M forked far away from nodal line, and fourbranched; branch CuA2 long and sinuous. Furthermore, it is distinctly different from the genera of the subfamily Tettigarctinae in having a broad costal area and clavus on tegmen.
</paragraph>
</subSubSection>
</treatment>
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