treatments-xml/data/03/93/3E/03933E752657FFEECBFE34B60C3FFA15.xml

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<mods:title id="1C34D10B3AD44678CC38BBEC8E9C3A3B">Revision of Taurocerus (Heteroptera, Pentatomidae, Pentatomini)</mods:title>
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<mods:namePart id="693C367165ACAE8019F52C828D71E238">Jocélia Grazia</mods:namePart>
<mods:affiliation id="8B5AC338B2C8505C63469172F6ECFE4B">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Bloco IV, Prédio 43435, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:namePart id="5C02788E2E733F2916AC025C23CC5E45">Aline Barcellos</mods:namePart>
<mods:affiliation id="5C25CC76484A9B9E37000D369D574797">Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Caixa Postal 1188, 90001 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:title id="B57709D2BA62432593436E9F613F9FCC">Iheringia, Série Zoologia</mods:title>
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<taxonomicName id="4C3AF4E02657FFEDCBFE34B60D68FF1E" authority="Amyot &amp; Serville, 1843" authorityName="edessoides Amyot &amp; Serville" authorityYear="1843" box="[231,623,218,242]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712657FFEDCBFE34B60D68FF1E" bold="true" box="[231,623,218,242]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDCBFE34B60E63FF1E" bold="true" box="[231,356,218,242]" italics="true" pageId="1" pageNumber="174">Taurocerus</emphasis>
<bibRefCitation id="EFABF2922657FFEDCA7134B60D68FF1E" author="Amyot &amp; Serville" bookContentInfo="681 p." box="[360,623,218,242]" firstAuthor="Amyot" journalOrPublisher="Paris, Librairie Encyclopedique de Roret" pageId="1" pageNumber="174" refId="ref6933" refString="AMYOT, J. B. &amp; SERVILLE, A. 1843. Histoire Naturelle des Insectes. Hemipteres. Paris, Librairie Encyclopedique de Roret. 681 p." title="Histoire Naturelle des Insectes. Hemipteres" type="book" year="1843">Amyot &amp; Serville, 1843</bibRefCitation>
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<emphasis id="B94E53712657FFEDCB9835790FE5FEC4" box="[129,226,277,296]" italics="true" pageId="1" pageNumber="174">Taurocerus</emphasis>
<bibRefCitation id="EFABF2922657FFEDCBF235790EF3FEC4" author="AMYOT &amp; SERVILLE" bookContentInfo="681 p." box="[235,500,277,296]" firstAuthor="AMYOT" journalOrPublisher="Paris, Librairie Encyclopedique de Roret" pageId="1" pageNumber="174" refId="ref6933" refString="AMYOT, J. B. &amp; SERVILLE, A. 1843. Histoire Naturelle des Insectes. Hemipteres. Paris, Librairie Encyclopedique de Roret. 681 p." title="Histoire Naturelle des Insectes. Hemipteres" type="book" year="1843">AMYOT &amp; SERVILLE, 1843:151</bibRefCitation>
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<bibRefCitation id="EFABF2922657FFEDC91835790D99FEC4" author="DALLAS" bookContentInfo="390 p." box="[513,670,277,296]" firstAuthor="DALLAS" journalOrPublisher="London, British Museum" pageId="1" pageNumber="174" refId="ref7078" refString="DALLAS, W. S. 1851. List of the specimens of Hemipterous insects of the British Museum I. London, British Museum. 390 p." title="List of the specimens of Hemipterous insects of the British Museum I" type="book" year="1851">DALLAS, 1851:302</bibRefCitation>
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(list);
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<bibRefCitation id="EFABF2922657FFEDCB8735410E54FEAC" author="WALKER" bookContentInfo="parts 1 - 2, p. 1 - 417" box="[158,339,301,320]" firstAuthor="WALKER" journalOrPublisher="London, British Museum" pageId="1" pageNumber="174" refId="ref7450" refString="WALKER, F. 1867. Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum. London, British Museum. parts 1 - 2, p. 1 - 417." title="Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum" type="book" year="1867">WALKER, 1867:391</bibRefCitation>
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(list);
<treatmentCitation id="0A9BA9722657FFEDCA8735410D33FEAC" box="[414,564,301,320]" pageId="1" pageNumber="174">
<bibRefCitation id="EFABF2922657FFEDCA8735410D33FEAC" author="STAL, C." box="[414,564,301,320]" journalOrPublisher="Ofversigt af Kungliga Vetenskakademiens forhandlingar" pageId="1" pageNumber="174" pagination="491 - 560" part="24" refId="ref7405" refString="___. 1867. Bidrag till hemipternas systematik. Ofversigt af Kungliga Vetenskakademiens forhandlingar 24 (7): 491 - 560." title="Bidrag till hemipternas systematik" type="journal article" year="1867">STÅL, 1867:531</bibRefCitation>
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(key);
<bibRefCitation id="EFABF2922657FFEDC99A35410DD4FEAC" author="STAL, C." box="[643,723,301,320]" journalOrPublisher="Kongliga Svenska Vetenskakademiens handlingar" pageId="1" pageNumber="174" pagination="1 - 159" part="10" refId="ref7428" refString="___. 1872. Enumeratio Hemipterorum. 2. Kongliga Svenska Vetenskakademiens handlingar 10 (4): 1 - 159." title="Enumeratio Hemipterorum" type="journal article" year="1872">1872:46</bibRefCitation>
;
<treatmentCitation id="0A9BA9722657FFEDCB8735290EB7FEB4" box="[158,432,325,344]" pageId="1" pageNumber="174">
<bibRefCitation id="EFABF2922657FFEDCB8735290EB7FEB4" author="LETHIERRY &amp; SEVERIN" bookContentInfo="x + 286 p." box="[158,432,325,344]" firstAuthor="LETHIERRY" journalOrPublisher="Bruxelles, F. Hayez" pageId="1" pageNumber="174" refId="ref7251" refString="LETHIERRY, L. &amp; SEVERIN, G. 1893. Catalogue General des Hemipteres - Pentatomidae 1. Bruxelles, F. Hayez. x + 286 p." title="Catalogue General des Hemipteres - Pentatomidae 1" type="book" year="1893">LETHIERRY &amp; SEVERIN, 1893:185</bibRefCitation>
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(cat.);
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<bibRefCitation id="EFABF2922657FFEDCAF635290D9AFEB4" author="KIRKALDY" bookContentInfo="xl + 392 p." box="[495,669,325,344]" firstAuthor="KIRKALDY" journalOrPublisher="Berlin, Felix L. Dames" pageId="1" pageNumber="174" refId="ref7219" refString="KIRKALDY, G. W. 1909. Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae. Berlin, Felix L. Dames. xl + 392 p." title="Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae" type="book" year="1909">KIRKALDY, 1909:150</bibRefCitation>
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(cat.);
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<bibRefCitation id="EFABF2922657FFEDCB8735310E79FE9C" author="ROLSTON" box="[158,382,349,368]" etAl="et al." firstAuthor="ROLSTON" journalOrPublisher="Journal of New York Entomological Society" pageId="1" pageNumber="174" pagination="120 - 132" part="88" refId="ref7281" refString="ROLSTON, L. H.; MCDONALD, F. J. D. &amp; THOMAS, D. B., JR. 1980. A conspectus of Pentatomini genera of the Western Hemisphere - Part 1 (Hemiptera: Pentatomidae). Journal of New York Entomological Society 88 (2): 120 - 132." title="A conspectus of Pentatomini genera of the Western Hemisphere - Part 1 (Hemiptera: Pentatomidae)" type="journal article" year="1980">
ROLSTON
<emphasis id="B94E53712657FFEDCBE935310E27FE9C" box="[240,288,349,368]" italics="true" pageId="1" pageNumber="174">et al.</emphasis>
, 1980:121
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(key).
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</paragraph>
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<treatmentCitationGroup id="AB2AA84D2657FFEDCB9835190E87FE64" box="[129,384,373,392]" pageId="1" pageNumber="174">
<treatmentCitation id="0A9BA9722657FFEDCB9835190E7CFE64" box="[129,379,373,392]" pageId="1" pageNumber="174">
<taxonomicName id="4C3AF4E02657FFEDCB9835190E7CFE64" ID-CoL="8WCXY" authority="WALKER, 1867: 404" authorityName="abr upta Walker, 1 8 6 7" authorityYear="1867" box="[129,379,373,392]" class="Magnoliopsida" family="Pentatomidae" genus="Canaca" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="174" phylum="Tracheophyta" rank="genus">
<emphasis id="B94E53712657FFEDCB9835190FC3FE64" box="[129,196,373,392]" italics="true" pageId="1" pageNumber="174">Canaca</emphasis>
<bibRefCitation id="EFABF2922657FFEDCBD635190E7CFE64" author="WALKER" bookContentInfo="parts 1 - 2, p. 1 - 417" box="[207,379,373,392]" firstAuthor="WALKER" journalOrPublisher="London, British Museum" pageId="1" pageNumber="174" refId="ref7450" refString="WALKER, F. 1867. Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum. London, British Museum. parts 1 - 2, p. 1 - 417." title="Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum" type="book" year="1867">WALKER, 1867:404</bibRefCitation>
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.
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<emphasis id="B94E53712657FFEDCB9835E10FC4FE4C" box="[129,195,397,416]" italics="true" pageId="1" pageNumber="174">Canoca</emphasis>
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(
<emphasis id="B94E53712657FFEDCBCC35E10FEAFE4C" box="[213,237,397,416]" italics="true" pageId="1" pageNumber="174">sic</emphasis>
);
<bibRefCitation id="EFABF2922657FFEDCA1D35E10EBFFE4C" author="KIRKALDY" bookContentInfo="xl + 392 p." box="[260,440,397,416]" firstAuthor="KIRKALDY" journalOrPublisher="Berlin, Felix L. Dames" pageId="1" pageNumber="174" refId="ref7219" refString="KIRKALDY, G. W. 1909. Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae. Berlin, Felix L. Dames. xl + 392 p." title="Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae" type="book" year="1909">KIRKALDY, 1909:150</bibRefCitation>
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(cat.).
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Type-species:
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<emphasis id="B94E53712657FFEDCA7A35AE0D66FE35" box="[355,609,450,473]" italics="true" pageId="1" pageNumber="174">Taurocerus edessoides</emphasis>
Amyot &amp; Serville
</taxonomicName>
(=
<taxonomicName id="4C3AF4E02657FFEDCA0C35B30D98FE1A" authority="achilles Stal, 1862" authorityName="achilles Stal" authorityYear="1862" box="[277,671,479,502]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712657FFEDCA0C35B30D0BFE1A" box="[277,524,479,502]" italics="true" pageId="1" pageNumber="174">Taurocerus achilles</emphasis>
<bibRefCitation id="EFABF2922657FFEDC90135B30D98FE1A" author="Stal" box="[536,671,479,502]" firstAuthor="Stal" journalOrPublisher="Stettiner entomologische Zeitung" pageId="1" pageNumber="174" pagination="81 - 118" part="23" refId="ref7382" refString="STAL, C. 1862. Hemiptera Mexicana. Enumeravit speciesque novas descripsit. Stettiner entomologische Zeitung 23: 81 - 118." title="Hemiptera Mexicana. Enumeravit speciesque novas descripsit" type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
), by monotypy.
</paragraph>
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<subSubSection id="C320DCE82657FFEDCBD8365B0E71FBB2" pageId="1" pageNumber="174" type="diagnosis">
<paragraph id="8B858F632657FFEDCBD8365B0E71FBB2" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">
Diagnosis. Medium to large-sized species. First antennal segment not surpassing apex of head (
<figureCitation id="130193E62657FFEDC99336380DC1FD87" box="[650,710,596,619]" captionStart="Fig" captionStartId="1.[763,794,1022,1041]" captionTargetBox="[773,1355,216,982]" captionTargetId="figure@1.[763,1355,216,1013]" captionTargetPageId="1" captionText="Fig. 1. Taurocerus amazonensis sp. nov., holotype. Scale bar, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3630714" httpUri="https://zenodo.org/record/3630714/files/figure.png" pageId="1" pageNumber="174">fig. 1</figureCitation>
). Jugae shorter than clypeus. Rostrum reaching third to fourth abdominal sternite; first rostral segment projected beyond bucculae. Bucculae anteriorly truncate and posteriorly lobed. Anterior half of pronotum strongly declivent. Humeri strongly produced laterad or anterolaterad, apices not-spinose. Femora distally armed with one dorsal and 1+1 lateral blackish spines. Tibiae dorsally sulcated. Ostiolar rugae short, digitiform. Mesosternum slightly more elevated than metasternum, nearly continuous in profile with it. Metasternum sligthly elevated, compressed. Abdominal venter with a mesial, longitudinal keel; third sternite with a stout median tubercle, apposed to posterior margin of metasternum. Ventral surface of pygophore with 1 + 1 lateral, concave areas. Genital plates tumid. Spiracles of gonocoxites 8 visible. Laterotergites 9 surpassing transverse band linking laterotergites 8.
</paragraph>
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<paragraph id="8B858F632657FFEDC9E237920A56FBFD" blockId="1.[763,1361,1022,1041]" box="[763,1361,1022,1041]" pageId="1" pageNumber="174">
Fig. 1.
<taxonomicName id="4C3AF4E02657FFEDC82537920B0AFBFD" authority="Jocélia Grazia &amp; Aline Barcellos, 2005" authorityName="Jocélia Grazia &amp; Aline Barcellos" authorityYear="2005" box="[828,1037,1022,1041]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="species" species="amazonensis" status="sp. nov.">
<emphasis id="B94E53712657FFEDC82537920B0AFBFD" box="[828,1037,1022,1041]" italics="true" pageId="1" pageNumber="174">Taurocerus amazonensis</emphasis>
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<taxonomicNameLabel id="A27DEE0A2657FFEDCF0D37920B5DFBFD" box="[1044,1114,1022,1041]" pageId="1" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Scale bar, 2 mm.
</paragraph>
</caption>
<subSubSection id="C320DCE82657FFEECBD830080C3EFE6E" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174" type="description">
<paragraph id="8B858F632657FFEDCBD830080DA4FAE2" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">Description. Head. Jugae shorter than clypeus, apices rounded; outer margins sinuated, convergent toward clypeus. First antennal segment not surpassing apex of head. Rostrum reaching third to fourth abdominal sternite; first rostral segment projected beyond bucculae. Bucculae anteriorly truncate and posteriorly lobed.</paragraph>
<paragraph id="8B858F632657FFEDCBD831780E88F9DF" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">
Pronotum. Anterior half of the disc strongly declivent. Anterior border slightly concave. Antero-lateral angles toothed. Antero-lateral borders crenulated, more strongly on anterior half. Humeri strongly produced laterad or antero-laterad, apex posteriorly directed (laterally in
<taxonomicName id="4C3AF4E02657FFEDCA1931CB0E71FA52" box="[256,374,1447,1470]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDCA1931CB0E71FA52" box="[256,374,1447,1470]" italics="true" pageId="1" pageNumber="174">T. abruptus</emphasis>
</taxonomicName>
). Posterolateral borders sinuated, concave on the anterior half and slightly convex posterad. Posterior border rectilinear. Medium-sized punctures distributed more densely around cicatrices and on posterior half of the disc.
</paragraph>
<paragraph id="8B858F632657FFEDCBD832550D04F8D7" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">
Scutellum. Coarse punctures irregularly distributed over basal half, decreasing in size toward apex. Apex with a V-shaped, yellowish callous, variable intraspecifically. In most of the specimens of
<taxonomicName id="4C3AF4E02657FFEDCAA232FE0D4EF945" box="[443,585,1682,1705]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDCAA232FE0D4EF945" box="[443,585,1682,1705]" italics="true" pageId="1" pageNumber="174">T. edessoides</emphasis>
</taxonomicName>
, the arms of “V” are broad and closer to each other, making it cordiform in appearance; in some specimens, however, the “V” is faded. In
<taxonomicName id="4C3AF4E02657FFEDCBFF32850E54F8EC" box="[230,339,1769,1792]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDCBFF32850E54F8EC" box="[230,339,1769,1792]" italics="true" pageId="1" pageNumber="174">T. achilles</emphasis>
</taxonomicName>
and the holotype of
<taxonomicName id="4C3AF4E02657FFEDC93632850DAFF8EC" box="[559,680,1769,1792]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDC93632850DAFF8EC" box="[559,680,1769,1792]" italics="true" pageId="1" pageNumber="174">T. abruptus</emphasis>
</taxonomicName>
, the callus is thinner than in
<taxonomicName id="4C3AF4E02657FFEDCA67336B0D0DF8F2" box="[382,522,1799,1822]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDCA67336B0D0DF8F2" box="[382,522,1799,1822]" italics="true" pageId="1" pageNumber="174">T. edessoides</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C3AF4E02657FFEDC92D336B0DD2F8F2" authority="Jocélia Grazia &amp; Aline Barcellos, 2005" authorityName="Jocélia Grazia &amp; Aline Barcellos" authorityYear="2005" box="[564,725,1799,1822]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="species" species="amazonensis" status="sp. nov.">
<emphasis id="B94E53712657FFEDC92D336B0DD2F8F2" box="[564,725,1799,1822]" italics="true" pageId="1" pageNumber="174">T. amazonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DEE0A2657FFEDCB9833480FD1F8D7" box="[129,214,1828,1851]" pageId="1" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
, the callous is also variable.
</paragraph>
<paragraph id="8B858F632657FFEDCBD8332D0D92F85C" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">Hemelytra. Area between claval suture and R + M vein dull, with small punctures homogeneously distributed. Clavus and costal area shiny, coarsely punctured. Membrane with 12 to 15 longitudinal, parallel veins.</paragraph>
<paragraph id="8B858F632657FFEDCBD833DA0DAAF807" blockId="1.[129,727,567,2027]" pageId="1" pageNumber="174">Legs. Femora distally armed with one dorsal and 1+1 lateral blackish spines. Tibiae dorsally sulcated.</paragraph>
<paragraph id="8B858F632657FFEDC822302A0A4DFAA4" blockId="1.[763,1361,1094,2026]" pageId="1" pageNumber="174">Ventral surface of thorax castaneous, more coarsely and densely puntured on ventral surface of prothorax, mesepimeron and metepisternum. Prosternum flat; mesosternum slightly more elevated, almost continuous with metasternum in profile. Metasternum sligthly elevated, laterally compressed; anterior margin entire, posterior margin slightly concave. Ostiolar rugae short, digitiform, reaching about 1/7 to 1/8 of the distance from inner margin of ostiole to lateral margins of metapleura.</paragraph>
<paragraph id="8B858F632657FFEDC82231220B95FA14" blockId="1.[763,1361,1094,2026]" pageId="1" pageNumber="174">Abdominal venter with a mesial, longitudinal keel; surface rarely punctured. Third sternite with a stout median tubercle, apposed to posterior margin of the metasternum. Tricobothria distributed on a longitudinal imaginary line across the middle of spiracles. Apical angles of connexivum acute, blackish.</paragraph>
<paragraph id="8B858F632657FFEDC82231920B62F8B4" blockId="1.[763,1361,1094,2026]" pageId="1" pageNumber="174">
Male genitalia. Pygophore subtrapezoidal to quadrangular. Dorsal rim slightly sinuous. X segment trapezoidal, wider basally. Parameres well-developed. Ventral rim excavated medially, with a median process, varying from an inconspicuous to acute tubercle; lateromedially, 1 + 1 processes, variable in shape and size. Posterolateral angles of pygophore mutic or with a small, acute tubercle.
<emphasis id="B94E53712657FFEDC8E432A00B54F90F" box="[1021,1107,1740,1763]" italics="true" pageId="1" pageNumber="174">Phallus</emphasis>
. Conjunctiva present, bearing or not processes.
<emphasis id="B94E53712657FFEDCF0E32850B9FF8EC" box="[1047,1176,1769,1792]" italics="true" pageId="1" pageNumber="174">Phallotheca</emphasis>
with 1 + 1 long and slender ventral processes.
<emphasis id="B94E53712657FFEDCF48336A0A57F8F1" box="[1105,1360,1798,1821]" italics="true" pageId="1" pageNumber="174">Ductus seminis distalis</emphasis>
surpassing or not conjunctiva. Vesica with a ventral process, not visible in
<taxonomicName id="4C3AF4E02657FFEDC8E9332D0B59F8B4" box="[1008,1118,1857,1880]" pageId="1" pageNumber="174">
<emphasis id="B94E53712657FFEDC8E9332D0B59F8B4" box="[1008,1118,1857,1880]" italics="true" pageId="1" pageNumber="174">T. achilles</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B858F632657FFEEC82233320C3EFE6E" blockId="1.[763,1361,1094,2026]" lastBlockId="2.[227,825,216,884]" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174">
Female genitalia. Genital plates tumid. Sutural borders of gonocoxites 8 convergent toward apex; posterior borders sligthly to conspicuously concave. Spiracles on laterotergites 8 present. Laterotergites 8 with acute, blackish apex. Laterotergites 9 much shorter than laterotergites 8, rounded at apex, surpassing transversal band linking laterotergites 8. Gonapophyses 9 with a mesial and 1 + 1 lateral, secondary thickenings variable interespecifically in shape.
<emphasis id="B94E53712654FFEEC900355D0DE0FEA4" box="[537,743,305,328]" italics="true" pageId="2" pageNumber="175">Pars intermedialis</emphasis>
with a median constriction; annular crests convergent to each other;
<emphasis id="B94E53712654FFEECA3C35070ED8FE6E" box="[293,479,363,386]" italics="true" pageId="2" pageNumber="175">capsula seminalis</emphasis>
with three tooth-like projections.
</paragraph>
</subSubSection>
<subSubSection id="C320DCE82654FFEECA3A35E40C3FFD81" pageId="2" pageNumber="175" type="distribution">
<paragraph id="8B858F632654FFEECA3A35E40C3FFD81" blockId="2.[227,825,216,884]" pageId="2" pageNumber="175">Distribution. MEXICO (Veracruz), GUATEMALA (Alta Verapaz), COSTA RICA (Guanacaste), PANAMA (Panama, Canal Zone), GUYANA (Cuyuni-Mazaruni), BRAZIL (Amazonas, Pará, Acre, Rondônia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo), COLOMBIA (Distrito Capital, Meta, Amazonas), ECUADOR (Napo), PERU (Loreto, Amazonas, San Martin, Ucayali, Huánuco, Madre de Dios, Junín, Cuzco), BOLIVIA (Cochabamba).</paragraph>
</subSubSection>
<subSubSection id="C320DCE82654FFEECA3A361F0EAEFC98" pageId="2" pageNumber="175" type="discussion">
<paragraph id="8B858F632654FFEECA3A361F0EAEFC98" blockId="2.[227,825,216,884]" pageId="2" pageNumber="175">
Comments. The apical spines at femora, the mesial tubercle on third sternite, apposed to metasternal carena, and the morphological pattern of the
<emphasis id="B94E53712654FFEEC98136C20DECFD29" box="[664,747,686,709]" italics="true" pageId="2" pageNumber="175">phallus</emphasis>
, make
<taxonomicName id="4C3AF4E02654FFEECBFA36A70E5DFD0E" authorityName="achilles Stal" authorityYear="1862" box="[227,346,715,738]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEECBFA36A70E5DFD0E" box="[227,346,715,738]" italics="true" pageId="2" pageNumber="175">Taurocerus</emphasis>
</taxonomicName>
similar to
<taxonomicName id="4C3AF4E02654FFEECAD636A70D84FD0E" authority="Spinola" authorityName="Spinola" authorityYear="1837" box="[463,643,715,738]" class="Insecta" family="Pentatomidae" genus="Arvelius" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEECAD636A70D2FFD0E" box="[463,552,715,738]" italics="true" pageId="2" pageNumber="175">Arvelius</emphasis>
Spinola
</taxonomicName>
, from which can be distinguished by the darker general color, jugae apically rounded, shorter than clypeus; bucculae lobed posteriorly; obtuse apex of humeri, and, on ventral rim of the pygophore, the presence of a median and two lateromedian processes.
</paragraph>
</subSubSection>
<subSubSection id="C320DCE82654FFEECBFA37F40D20FC43" box="[227,551,920,943]" pageId="2" pageNumber="175" type="description">
<paragraph id="8B858F632654FFEECBFA37F40D20FC43" blockId="2.[227,551,920,943]" box="[227,551,920,943]" pageId="2" pageNumber="175">
<heading id="D0CD380F2654FFEECBFA37F40D20FC43" box="[227,551,920,943]" fontSize="10" level="3" pageId="2" pageNumber="175" reason="1">
Key to species of
<emphasis id="B94E53712654FFEECAB137F40D20FC43" box="[424,551,920,943]" italics="true" pageId="2" pageNumber="175">
<taxonomicName id="4C3AF4E02654FFEECAB137F40D24FC43" authorityName="achilles Stal" authorityYear="1862" box="[424,547,920,943]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">Taurocerus</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320DCE82654FFEECBFA37BF0C3FFA15" pageId="2" pageNumber="175" type="key">
<keyStep id="30CE97C62654FFEECBFA37BF0C30FBEB" pageId="2" pageNumber="175">
<paragraph id="8B858F632654FFEECBFA37BF0C30FC06" blockId="2.[227,823,979,1031]" box="[227,823,979,1002]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBFA37BF0C30FC06" box="[227,823,979,1002]" pageId="2" pageNumber="175">1. Apex of humeri black ................................................. 2</keyLead>
</paragraph>
<paragraph id="8B858F632654FFEECBE4379C0C30FBEB" blockId="2.[227,823,979,1031]" box="[253,823,1008,1031]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBE4379C0C30FBEB" box="[253,823,1008,1031]" pageId="2" pageNumber="175">Apex of humeri yellowish ......................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE97C62654FFEECBFA30610C30FAE3" pageId="2" pageNumber="175">
<paragraph id="8B858F632654FFEECBFA30610C3FFB90" blockId="2.[227,825,1037,1295]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBFA30610C3FFB90" pageId="2" pageNumber="175">
2. General color castaneous. Humeral angles somewhat curved, directed antero-laterad. Gonocoxites 8 almost rectangular, two and a half wider than long (
<figureCitation id="130193E62654FFEEC9FD30240C2EFBB3" box="[740,809,1096,1119]" captionStart="Figs" captionStartId="6.[861,900,1932,1951]" captionTargetBox="[870,1443,1026,1900]" captionTargetId="figure@6.[861,1453,1003,1901]" captionTargetPageId="6" captionText="Figs. 25-26. Genital plates, ventral view: 25, T. achilles Stål, 1862; 26, T. edessoides (Spinola, 1837) (gc8, gonocoxite 8; gc9; gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh abdominal sternite; X, tenth segment). Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3630724" httpUri="https://zenodo.org/record/3630724/files/figure.png" pageId="2" pageNumber="175">fig. 25</figureCitation>
)
<emphasis id="B94E53712654FFEECA06300A0EF3FB91" box="[287,500,1126,1149]" italics="true" pageId="2" pageNumber="175">................................</emphasis>
<taxonomicName id="4C3AF4E02654FFEECAEE300A0C3FFB90" authority="achilles Stal, 1862" authorityName="achilles Stal" authorityYear="1862" box="[503,824,1125,1149]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEECAEE300A0DC1FB91" box="[503,710,1126,1149]" italics="true" pageId="2" pageNumber="175">Taurocerus achilles</emphasis>
<bibRefCitation id="EFABF2922654FFEEC9D430090C3FFB90" author="Stal" box="[717,824,1125,1148]" firstAuthor="Stal" journalOrPublisher="Stettiner entomologische Zeitung" pageId="2" pageNumber="175" pagination="81 - 118" part="23" refId="ref7382" refString="STAL, C. 1862. Hemiptera Mexicana. Enumeravit speciesque novas descripsit. Stettiner entomologische Zeitung 23: 81 - 118." title="Hemiptera Mexicana. Enumeravit speciesque novas descripsit" type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B858F632654FFEECBE430EF0C30FAE3" blockId="2.[227,825,1037,1295]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBE430EF0C30FAE3" pageId="2" pageNumber="175">
General color reddish to dark reddish-castaneous. Humeral angles almost straigth, directed laterad (
<figureCitation id="130193E62654FFEEC80A30CC0E20FB38" captionStart="Figs" captionStartId="2.[861,901,1297,1316]" captionTargetBox="[873,1441,580,1229]" captionTargetId="figure@2.[861,1453,557,1237]" captionTargetPageId="2" captionText="Figs. 2-3. Taurocerus abruptus (Walker, 1867), holotype: 2, dorsal aspect; 3, genital plates, ventral view (gc8, gonocoxite 8; gc9; gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh abdominal sternite; X, tenth segment)." figureDoi="http://doi.org/10.5281/zenodo.3630716" httpUri="https://zenodo.org/record/3630716/files/figure.png" pageId="2" pageNumber="175">fig. 2</figureCitation>
). Gonocoxites 8 quadrangular, one and a half wider than long (
<figureCitation id="130193E62654FFEECA9B30B70EB1FB1E" box="[386,438,1243,1266]" captionStart="Figs" captionStartId="2.[861,901,1297,1316]" captionTargetBox="[873,1441,580,1229]" captionTargetId="figure@2.[861,1453,557,1237]" captionTargetPageId="2" captionText="Figs. 2-3. Taurocerus abruptus (Walker, 1867), holotype: 2, dorsal aspect; 3, genital plates, ventral view (gc8, gonocoxite 8; gc9; gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh abdominal sternite; X, tenth segment)." figureDoi="http://doi.org/10.5281/zenodo.3630716" httpUri="https://zenodo.org/record/3630716/files/figure.png" pageId="2" pageNumber="175">fig. 3</figureCitation>
) ............................................................
<emphasis id="B94E53712654FFEECA0130940ECCFAE3" box="[280,459,1272,1295]" italics="true" pageId="2" pageNumber="175">............................</emphasis>
<taxonomicName id="4C3AF4E02654FFEECAD230940C30FAE3" authority="abruptus (Walker, 1867)" authorityName="abruptus (Walker" authorityYear="1867" box="[459,823,1272,1295]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEECAD230940D9CFAE3" box="[459,667,1272,1295]" italics="true" pageId="2" pageNumber="175">Taurocerus abruptus</emphasis>
(
<bibRefCitation id="EFABF2922654FFEEC9BF30940C37FAE3" author="Walker" bookContentInfo="parts 1 - 2, p. 1 - 417" box="[678,816,1272,1295]" firstAuthor="Walker" journalOrPublisher="London, British Museum" pageId="2" pageNumber="175" refId="ref7450" refString="WALKER, F. 1867. Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum. London, British Museum. parts 1 - 2, p. 1 - 417." title="Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum" type="book" year="1867">Walker, 1867</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE97C62654FFEECBFA31790C3FFA15" pageId="2" pageNumber="175">
<paragraph id="8B858F632654FFEECBFA31790C3FFA68" blockId="2.[227,824,1301,1530]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBFA31790C3FFA68" pageId="2" pageNumber="175">
3. Ventral rim of pygophore with an inconspicuous median tubercle and lateral-median processes truncate (
<figureCitation id="130193E62654FFEEC80A315E0E33FA8B" captionStart="Figs" captionStartId="3.[129,171,1946,1965]" captionTargetBox="[173,1285,673,1916]" captionTargetId="figure@3.[172,1308,647,1916]" captionTargetPageId="3" captionText="Figs. 4-12. Pygophore, respectively dorsal, caudal and ventral views: 4, 7, 10, Taurocerus achilles Stål, 1862; 5, 8, 11, Taurocerus amazonensis sp. nov.; 6, 9, 12, Taurocerus edessoides (Spinola, 1837) (dr, dorsal rim; lmp, lateral-median process; mp, median process; pa, paramere; vr, ventral rim; X, tenth segment). Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3630718" httpUri="https://zenodo.org/record/3630718/files/figure.png" pageId="2" pageNumber="175">fig. 11</figureCitation>
). Apex of parameres slightly curved laterad (
<figureCitation id="130193E62654FFEEC9E3313C0C28FA8B" box="[762,815,1360,1383]" captionStart="Figs" captionStartId="3.[129,171,1946,1965]" captionTargetBox="[173,1285,673,1916]" captionTargetId="figure@3.[172,1308,647,1916]" captionTargetPageId="3" captionText="Figs. 4-12. Pygophore, respectively dorsal, caudal and ventral views: 4, 7, 10, Taurocerus achilles Stål, 1862; 5, 8, 11, Taurocerus amazonensis sp. nov.; 6, 9, 12, Taurocerus edessoides (Spinola, 1837) (dr, dorsal rim; lmp, lateral-median process; mp, median process; pa, paramere; vr, ventral rim; X, tenth segment). Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3630718" httpUri="https://zenodo.org/record/3630718/files/figure.png" pageId="2" pageNumber="175">fig. 8</figureCitation>
) ..........................
<taxonomicName id="4C3AF4E02654FFEECACD31010DDCFA68" authority="Jocélia Grazia &amp; Aline Barcellos, 2005" authorityName="Jocélia Grazia &amp; Aline Barcellos" authorityYear="2005" box="[468,731,1389,1412]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="amazonensis" status="sp. nov.">
<emphasis id="B94E53712654FFEECACD31010DDCFA68" box="[468,731,1389,1412]" italics="true" pageId="2" pageNumber="175">Taurocerus amazonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DEE0A2654FFEEC9FD31010C3FFA68" box="[740,824,1389,1412]" pageId="2" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<caption id="DF45DFEB2654FFEEC844317D0BC4FA80" ID-DOI="http://doi.org/10.5281/zenodo.3630716" ID-Zenodo-Dep="3630716" httpUri="https://zenodo.org/record/3630716/files/figure.png" pageId="2" pageNumber="175" startId="2.[861,901,1297,1316]" targetBox="[873,1441,580,1229]" targetPageId="2">
<paragraph id="8B858F632654FFEEC844317D0BC4FA80" blockId="2.[861,1459,1297,1388]" pageId="2" pageNumber="175">
Figs. 2-3.
<taxonomicName id="4C3AF4E02654FFEEC8A2317D0BF8FAC8" authority="abruptus (Walker, 1867)" authorityName="abruptus (Walker" authorityYear="1867" box="[955,1279,1297,1316]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEEC8A2317D0B76FAC8" box="[955,1137,1297,1316]" italics="true" pageId="2" pageNumber="175">Taurocerus abruptus</emphasis>
(
<bibRefCitation id="EFABF2922654FFEECF67317D0BFEFAC8" author="Walker" bookContentInfo="parts 1 - 2, p. 1 - 417" box="[1150,1273,1297,1316]" firstAuthor="Walker" journalOrPublisher="London, British Museum" pageId="2" pageNumber="175" refId="ref7450" refString="WALKER, F. 1867. Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum. London, British Museum. parts 1 - 2, p. 1 - 417." title="Catalogue of the specimens of Hemiptera- Heteroptera in the collection of the British Museum" type="book" year="1867">Walker, 1867</bibRefCitation>
)
</taxonomicName>
, holotype: 2, dorsal aspect; 3, genital plates, ventral view (gc8, gonocoxite 8; gc9; gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh abdominal sternite; X, tenth segment).
</paragraph>
</caption>
<paragraph id="8B858F632654FFEECBE431E60C3FFA15" blockId="2.[227,824,1301,1530]" pageId="2" pageNumber="175">
<keyLead id="30CB2C562654FFEECBE431E60C3FFA15" pageId="2" pageNumber="175">
Ventral rim of pygophore with a conspicuous and acute median tubercle; latero-median processes conical (
<figureCitation id="130193E62654FFEEC80D31C40E32FA30" captionStart="Figs" captionStartId="3.[129,171,1946,1965]" captionTargetBox="[173,1285,673,1916]" captionTargetId="figure@3.[172,1308,647,1916]" captionTargetPageId="3" captionText="Figs. 4-12. Pygophore, respectively dorsal, caudal and ventral views: 4, 7, 10, Taurocerus achilles Stål, 1862; 5, 8, 11, Taurocerus amazonensis sp. nov.; 6, 9, 12, Taurocerus edessoides (Spinola, 1837) (dr, dorsal rim; lmp, lateral-median process; mp, median process; pa, paramere; vr, ventral rim; X, tenth segment). Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3630718" httpUri="https://zenodo.org/record/3630718/files/figure.png" pageId="2" pageNumber="175">fig. 12</figureCitation>
). Apex of parameres strongly curved ventrad (
<figureCitation id="130193E62654FFEEC80A31A90E22FA15" captionStart="Figs" captionStartId="3.[129,171,1946,1965]" captionTargetBox="[173,1285,673,1916]" captionTargetId="figure@3.[172,1308,647,1916]" captionTargetPageId="3" captionText="Figs. 4-12. Pygophore, respectively dorsal, caudal and ventral views: 4, 7, 10, Taurocerus achilles Stål, 1862; 5, 8, 11, Taurocerus amazonensis sp. nov.; 6, 9, 12, Taurocerus edessoides (Spinola, 1837) (dr, dorsal rim; lmp, lateral-median process; mp, median process; pa, paramere; vr, ventral rim; X, tenth segment). Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3630718" httpUri="https://zenodo.org/record/3630718/files/figure.png" pageId="2" pageNumber="175">fig. 9</figureCitation>
) ................
<taxonomicName id="4C3AF4E02654FFEECABB318F0C3FFA15" authority="edessoides (Spinola, 1837)" authorityName="edessoides (Spinola" authorityYear="1837" box="[418,824,1506,1530]" class="Insecta" family="Pentatomidae" genus="Taurocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="B94E53712654FFEECABB318F0D88FA16" box="[418,655,1507,1530]" italics="true" pageId="2" pageNumber="175">Taurocerus edessoides</emphasis>
(
<bibRefCitation id="EFABF2922654FFEEC985318E0C37FA15" author="Spinola" bookContentInfo="383 p." box="[668,816,1506,1529]" firstAuthor="Spinola" journalOrPublisher="Genes, Yves Gravier" pageId="2" pageNumber="175" refId="ref7341" refString="SPINOLA, M. 1837. Essai sur les genres d'insectes appartenants a l'ordre des Hemipteres, Lin. ou Rhyngotes, Fab. et a la section des Heteropteres, Dufour. Genes, Yves Gravier. 383 p." title="Essai sur les genres d'insectes appartenants a l'ordre des Hemipteres, Lin. ou Rhyngotes, Fab. et a la section des Heteropteres, Dufour" type="book" year="1837">Spinola, 1837</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>