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<document id="8B9B3EDF2BFED83EA4A71A0094B61263" ID-CLB-Dataset="5940" ID-DOI="10.11646/zootaxa.3761.1.1" ID-GBIF-Dataset="5444e27e-3e2a-429b-931b-823a2086e3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909057" ID-ZooBank="82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623107451891" checkinUser="felipe" docAuthor="Luk, Stephen P. L. &amp; Marshall, Stephen A." docDate="2014" docId="038487F1FFB7FFA0FDC7FD6FFB9B0ED0" docLanguage="en" docName="zt03761p156.pdf" docOrigin="Zootaxa 3761 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aptilotella germana Luk &amp; Marshall 2014, sp. n." docType="treatment" docVersion="6" lastPageNumber="20" masterDocId="FFBDFF89FFA5FFB3FD50FE2BFFC70D18" masterDocTitle="A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="19" updateTime="1699002622211" updateUser="ExternalLinkService">
<mods:mods id="7B6C0E92C7D4F673B8EA0F78893D5D74" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4473AD86FEC504E78F579D779C0DE44">
<mods:title id="F4671A42238D9C6DD96D8A9A3AEE4497">A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)</mods:title>
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<mods:namePart id="D18C2F6956B3F06A65C84B53B65B9AED">Luk, Stephen P. L.</mods:namePart>
<mods:nameIdentifier id="75A0B367DBC58A9E0D1D6B682CBC6366" type="email">luk@uoguelph.ca</mods:nameIdentifier>
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<mods:namePart id="7872BC2BBDD25FF2C1CC7433722C1737">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="6A4B437E94DA23199773741EB84698BC" type="email">luk@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="3DE30E28BC6B4430B0659F41D2903507">Zootaxa</mods:title>
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<mods:part id="676D247EE42FF2FB470B2BFE018BAB2A">
<mods:date id="434147F11461B0436A779D7CFBD63325">2014</mods:date>
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<mods:number id="868F664DA1F43E48811C5520C98D2080">2014-02-05</mods:number>
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<mods:number id="C8B5B6FE8C8705168F453F872D39AEAF">3761</mods:number>
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<mods:classification id="DBA1A70BF41B705AFB1DA4B877C84693">journal article</mods:classification>
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<mods:identifier id="A67232C4164A396971561C99155A8A16" type="DOI">10.11646/zootaxa.3761.1.1</mods:identifier>
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<treatment id="038487F1FFB7FFA0FDC7FD6FFB9B0ED0" ID-DOI="http://doi.org/10.5281/zenodo.4909075" ID-GBIF-Taxon="183203076" ID-Zenodo-Dep="4909075" LSID="urn:lsid:plazi:treatment:038487F1FFB7FFA0FDC7FD6FFB9B0ED0" httpUri="http://treatment.plazi.org/id/038487F1FFB7FFA0FDC7FD6FFB9B0ED0" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C337656CFFB7FFA1FDC7FD6FFD520E45" box="[151,661,836,861]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B9236E7FFB7FFA1FDC7FD6FFD520E45" blockId="18.[151,661,836,896]" box="[151,661,836,861]" pageId="18" pageNumber="19">
<heading id="D0DA818BFFB7FFA1FDC7FD6FFD520E45" bold="true" box="[151,661,836,861]" fontSize="10" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B959EAF5FFB7FFA1FDC7FD6FFD520E45" bold="true" box="[151,661,836,861]" pageId="18" pageNumber="19">
<taxonomicName id="4C2D4D64FFB7FFA1FDC7FD6FFD8E0E45" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,585,836,861]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="germana" status="sp. nov.">
<emphasis id="B959EAF5FFB7FFA1FDC7FD6FFEBB0E43" bold="true" box="[151,380,836,861]" italics="true" pageId="18" pageNumber="19">Aptilotella germana</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<taxonomicNameLabel id="A26A578EFFB7FFA1FF05FD6FFD520E45" box="[597,661,836,861]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337656CFFB7FFA0FDC7FD4CFE290CC8" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8B9236E7FFB7FFA1FDC7FD4CFE6B0E98" blockId="18.[151,661,836,896]" box="[151,428,871,896]" pageId="18" pageNumber="19">
<figureCitation id="13162A62FFB7FFA1FDC7FD4CFEEC0E98" box="[151,299,871,896]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURE 11" captionStartId-0="53.[151,264,1585,1608]" captionStartId-1="54.[151,249,1121,1143]" captionTargetBox-0="[340,1247,193,1565]" captionTargetBox-1="[340,1247,193,1100]" captionTargetId-0="figure-39@53.[340,1247,193,1565]" captionTargetId-1="figure-45@54.[340,1247,193,1100]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 9–10. Aptilotella germana Luk &amp; Marshall, sp. n., male: 9—habitus, left lateral; 10—habitus, dorsal. Abbreviation: WP = left wing pad." captionText-1="FIGURE 11. Aptilotella germana Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, DS5 = division of sternite 5, PG = postgonite, SUR = surstylus." figureDoi-0="http://doi.org/10.5281/zenodo.4909063" figureDoi-1="http://doi.org/10.5281/zenodo.4909065" httpUri-0="https://zenodo.org/record/4909063/files/figure.png" httpUri-1="https://zenodo.org/record/4909065/files/figure.png" pageId="18" pageNumber="19">Figures 9–11</figureCitation>
, and 75–85
</paragraph>
<paragraph id="8B9236E7FFB7FFA1FDC7FD84FE870885" blockId="18.[151,1437,943,2013]" pageId="18" pageNumber="19">
<emphasis id="B959EAF5FFB7FFA1FDC7FD84FEE40ED0" bold="true" box="[151,291,943,968]" italics="true" pageId="18" pageNumber="19">Description.</emphasis>
Habitus as in
<figureCitation id="13162A62FFB7FFA1FC95FD84FD420ED0" box="[453,645,943,968]" captionStart="FIGURES 9–10" captionStartId="53.[151,264,1585,1608]" captionTargetBox="[340,1247,193,1565]" captionTargetId="figure-39@53.[340,1247,193,1565]" captionTargetPageId="53" captionText="FIGURES 9–10. Aptilotella germana Luk &amp; Marshall, sp. n., male: 9—habitus, left lateral; 10—habitus, dorsal. Abbreviation: WP = left wing pad." figureDoi="http://doi.org/10.5281/zenodo.4909063" httpUri="https://zenodo.org/record/4909063/files/figure.png" pageId="18" pageNumber="19">Figures 9 and 10</figureCitation>
. Body length 1.0 mm. Head ground color yellow. Frons finely rugose; pale areas silvery-blue; brown medial stripes each about one-fourth the width of frons, diffuse along the anterior margin; brown orbital stripes each half the width of medial stripe; ocular emargination with small pale spot. Ocellar tubercle scarcely raised; ocelli present; ocellar bristle two-fifths the length of frons. Orbital bristle present; orbital setulae minute, in five pairs. Interfrontal setae in two pairs. Lunule polished; face shining; facial excavation with a silvery-white band continuing onto anterior half of gena; gena weakly shining, setaceous. Antenna brown. Scutum and scutellum dark reddish-brown, shining. Scutum creased along posterolateral margin; uniformly setose. Scutellum bare; flat, 2.4 times wider than long, 0.6 times the width of scutum. Apical scutellar bristles 1.4 times as long as basal. Pleuron dark reddish-brown; with two pale pruinose stripes, the first on upper half of the anepisternum and anepimeron, the second fainter, at margin with katepisternum. Legs yellow ochre; mid and hind coxae brown; mid tibia with two anterodorsal bristles, in male with row of five stout setae in ventrodistal third. Wing pad (
<figureCitation id="13162A62FFB7FFA1FC43FB17FE90084D" box="[275,343,1340,1365]" captionStart="FIGURES 9–10" captionStartId="53.[151,264,1585,1608]" captionTargetBox="[340,1247,193,1565]" captionTargetId="figure-39@53.[340,1247,193,1565]" captionTargetPageId="53" captionText="FIGURES 9–10. Aptilotella germana Luk &amp; Marshall, sp. n., male: 9—habitus, left lateral; 10—habitus, dorsal. Abbreviation: WP = left wing pad." figureDoi="http://doi.org/10.5281/zenodo.4909063" httpUri="https://zenodo.org/record/4909063/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
) small, dark brown. Abdomen black, shining; tergites each with two rows of setae, three rows on syntergite; sternites finely microtrichose. Epandrium and synsternite 6+7 dark reddish-brown; cercus and surstylus yellow-orange.
</paragraph>
<paragraph id="8B9236E7FFB7FFA0FD97FB8CFF230C05" blockId="18.[151,1437,943,2013]" lastBlockId="19.[151,1436,151,968]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B959EAF5FFB7FFA1FD97FB8CFE4B08D8" bold="true" box="[199,396,1447,1472]" italics="true" pageId="18" pageNumber="19">Male terminalia.</emphasis>
Sternite 5 (
<figureCitation id="13162A62FFB7FFA1FF4CFB8CFDB908D8" box="[540,638,1447,1472]" captionStart="FIGURE 11" captionStartId="54.[151,249,1121,1143]" captionTargetBox="[340,1247,193,1100]" captionTargetId="figure-45@54.[340,1247,193,1100]" captionTargetPageId="54" captionText="FIGURE 11. Aptilotella germana Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, DS5 = division of sternite 5, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909065" httpUri="https://zenodo.org/record/4909065/files/figure.png" pageId="18" pageNumber="19">Figs. 11</figureCitation>
,
<figureCitation id="13162A62FFB7FFA1FFDEFB8CFD6C08D8" box="[654,683,1447,1472]" captionStart="FIGURES 75–78" captionStartId="90.[151,264,1266,1289]" captionTargetBox="[390,1200,208,1237]" captionTargetId="figure-44@90.[378,1210,193,1245]" captionTargetPageId="90" captionText="FIGURES 75–78. Aptilotella germana Luk &amp; Marshall, sp. n.: 75—male terminalia, posterior; 76—male terminalia, left lateral; 77—male synsternite 6+7; 78—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909187" httpUri="https://zenodo.org/record/4909187/files/figure.png" pageId="18" pageNumber="19">78</figureCitation>
) completely divided, the inner margins concave posteriorly and flanked by setae. Synsternite 6+7 as in
<figureCitation id="13162A62FFB7FFA1FF01FBE7FD0608FB" box="[593,705,1484,1509]" captionStart="FIGURES 75–78" captionStartId="90.[151,264,1266,1289]" captionTargetBox="[390,1200,208,1237]" captionTargetId="figure-44@90.[378,1210,193,1245]" captionTargetPageId="90" captionText="FIGURES 75–78. Aptilotella germana Luk &amp; Marshall, sp. n.: 75—male terminalia, posterior; 76—male terminalia, left lateral; 77—male synsternite 6+7; 78—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909187" httpUri="https://zenodo.org/record/4909187/files/figure.png" pageId="18" pageNumber="19">Figure 77</figureCitation>
. Cercus (
<figureCitation id="13162A62FFB7FFA1FE7AFBE7FC4108FD" box="[810,902,1484,1509]" captionStart="FIGURE 11" captionStartId="54.[151,249,1121,1143]" captionTargetBox="[340,1247,193,1100]" captionTargetId="figure-45@54.[340,1247,193,1100]" captionTargetPageId="54" captionText="FIGURE 11. Aptilotella germana Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, DS5 = division of sternite 5, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909065" httpUri="https://zenodo.org/record/4909065/files/figure.png" pageId="18" pageNumber="19">Figs. 11</figureCitation>
,
<figureCitation id="13162A62FFB7FFA1FEC3FBE7FC1D08FD" box="[915,986,1484,1509]" captionStart="FIGURES 75–78" captionStartId="90.[151,264,1266,1289]" captionTargetBox="[390,1200,208,1237]" captionTargetId="figure-44@90.[378,1210,193,1245]" captionTargetPageId="90" captionText="FIGURES 75–78. Aptilotella germana Luk &amp; Marshall, sp. n.: 75—male terminalia, posterior; 76—male terminalia, left lateral; 77—male synsternite 6+7; 78—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909187" httpUri="https://zenodo.org/record/4909187/files/figure.png" pageId="18" pageNumber="19">75, 76</figureCitation>
) compressed, 3.5 times as long as basal width; base triangular, clothed in setulae; apex rounded, thickened; midlength with one long seta reaching apex, distally with several evenly spaced marginal setae and preapical sensory setae. Surstylus (
<figureCitation id="13162A62FFB7FFA1F9C0F83FFB2A0B35" box="[1168,1261,1556,1581]" captionStart="FIGURE 11" captionStartId="54.[151,249,1121,1143]" captionTargetBox="[340,1247,193,1100]" captionTargetId="figure-45@54.[340,1247,193,1100]" captionTargetPageId="54" captionText="FIGURE 11. Aptilotella germana Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, DS5 = division of sternite 5, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909065" httpUri="https://zenodo.org/record/4909065/files/figure.png" pageId="18" pageNumber="19">Figs. 11</figureCitation>
,
<figureCitation id="13162A62FFB7FFA1F9AAF83FFA850B35" box="[1274,1346,1556,1581]" captionStart="FIGURES 75–78" captionStartId="90.[151,264,1266,1289]" captionTargetBox="[390,1200,208,1237]" captionTargetId="figure-44@90.[378,1210,193,1245]" captionTargetPageId="90" captionText="FIGURES 75–78. Aptilotella germana Luk &amp; Marshall, sp. n.: 75—male terminalia, posterior; 76—male terminalia, left lateral; 77—male synsternite 6+7; 78—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909187" httpUri="https://zenodo.org/record/4909187/files/figure.png" pageId="18" pageNumber="19">75, 76</figureCitation>
) a halfcone; posteromedially weakly humped and bearing several long setae; apex round and flexed up. Postgonite (
<figureCitation id="13162A62FFB7FFA1F834F81CFF730B6D" captionStart="FIGURE 11" captionStartId="54.[151,249,1121,1143]" captionTargetBox="[340,1247,193,1100]" captionTargetId="figure-45@54.[340,1247,193,1100]" captionTargetPageId="54" captionText="FIGURE 11. Aptilotella germana Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, DS5 = division of sternite 5, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909065" httpUri="https://zenodo.org/record/4909065/files/figure.png" pageId="18" pageNumber="19">Figs. 11</figureCitation>
,
<figureCitation id="13162A62FFB7FFA1FD91F877FF190B6D" box="[193,222,1628,1653]" captionStart="FIGURES 79–81" captionStartId="91.[151,264,1237,1260]" captionTargetBox="[416,1185,205,1216]" captionTargetId="figure-70@91.[403,1185,193,1216]" captionTargetPageId="91" captionText="FIGURES 79–81. Aptilotella germana Luk &amp; Marshall, sp. n.: 79—aedeagus with postgonites removed; 80—left postgonite; 81—hypandrium. Abbreviations: BP = basiphallus, DTP = descending tab of paired arched sclerites, LFS = lateral flanking sclerite, PAS = paired arched sclerites, PDS = paired dorsal sclerites, TBS = tooth-bearing stubs, VBS = ventrobasal sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909189" httpUri="https://zenodo.org/record/4909189/files/figure.png" pageId="18" pageNumber="19">80</figureCitation>
) broad; descending arm short, slender and curved forward; articulatory process for pregonite undeveloped, truncate; lower portion before descending arm acutely angled; articulatory process for basiphallus knobbed with blunt anterior and posterior teeth. Hypandrium (
<figureCitation id="13162A62FFB7FFA1FFE1F88FFCC50BA5" box="[689,770,1700,1725]" captionStart="FIGURES 79–81" captionStartId="91.[151,264,1237,1260]" captionTargetBox="[416,1185,205,1216]" captionTargetId="figure-70@91.[403,1185,193,1216]" captionTargetPageId="91" captionText="FIGURES 79–81. Aptilotella germana Luk &amp; Marshall, sp. n.: 79—aedeagus with postgonites removed; 80—left postgonite; 81—hypandrium. Abbreviations: BP = basiphallus, DTP = descending tab of paired arched sclerites, LFS = lateral flanking sclerite, PAS = paired arched sclerites, PDS = paired dorsal sclerites, TBS = tooth-bearing stubs, VBS = ventrobasal sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909189" httpUri="https://zenodo.org/record/4909189/files/figure.png" pageId="18" pageNumber="19">Fig. 81</figureCitation>
) broad; medial rod apically spatulate, basal margin dilated; posteromedial fork thick and divergent; hypandrial arms slender, distal half weakly recurved; pregonite short, conical. Aedeagus as in
<figureCitation id="13162A62FFB7FFA1FCF8F8C7FDDF0A1D" box="[424,536,1772,1797]" captionStart="FIGURES 79–81" captionStartId="91.[151,264,1237,1260]" captionTargetBox="[416,1185,205,1216]" captionTargetId="figure-70@91.[403,1185,193,1216]" captionTargetPageId="91" captionText="FIGURES 79–81. Aptilotella germana Luk &amp; Marshall, sp. n.: 79—aedeagus with postgonites removed; 80—left postgonite; 81—hypandrium. Abbreviations: BP = basiphallus, DTP = descending tab of paired arched sclerites, LFS = lateral flanking sclerite, PAS = paired arched sclerites, PDS = paired dorsal sclerites, TBS = tooth-bearing stubs, VBS = ventrobasal sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909189" httpUri="https://zenodo.org/record/4909189/files/figure.png" pageId="18" pageNumber="19">Figure 79</figureCitation>
. Basiphallus stout, cylindrical; posterodorsally humped; articulatory process for postgonite truncate, curved upward. Ejaculatory apodeme pale, with the disc embedded midway. Ventrobasal sclerite divided. Lateral flanking sclerite broadly fused posteriorly, with a narrow descending tab; deeply indented at level of basiphallus; fused dorsally until distal third, where it darkens and tapers to a blunt apex. Paired arched sclerites very dark, originating at descending tab of lateral flanking sclerite; their trunks bearing a truncate lateral process, ascending between lateral flanking sclerites and descending ventrodistally, becoming depressed before merging into a prominent process; this process lunate, broadly rounded, basally formed into a sharp thorn. Paired dorsal sclerites slender, originating within dorsal division of lateral flanking sclerite, apically bearing a club of denticles. Lateral flanking sclerite giving rise to a pair of stubs beneath paired dorsal sclerites, capped in trifid denticles, which descend in a line next to lunate process, broadening and curving posteriorly to the level of its thorns.
</paragraph>
<paragraph id="8B9236E7FFB6FFA0FD97FF0CFE290CC8" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<emphasis id="B959EAF5FFB6FFA0FD97FF0CFE610C58" bold="true" box="[199,422,295,320]" italics="true" pageId="19" pageNumber="20">Female terminalia.</emphasis>
Epiproct (
<figureCitation id="13162A62FFB6FFA0FF71FF0CFD6B0C58" box="[545,684,295,320]" captionStart="FIGURES 82–85" captionStartId="92.[151,264,1220,1243]" captionTargetBox="[410,1184,202,1192]" captionTargetId="figure-49@92.[398,1189,193,1199]" captionTargetPageId="92" captionText="FIGURES 82–85. Aptilotella germana Luk &amp; Marshall, sp. n.: 82—female terminalia, dorsal; 83—female terminalia, left lateral; 84—female terminalia, ventral; 85—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct." figureDoi="http://doi.org/10.5281/zenodo.4909191" httpUri="https://zenodo.org/record/4909191/files/figure.png" pageId="19" pageNumber="20">Figs. 82, 83</figureCitation>
) indistinct, triangular; microtrichose. Each half of tergite 8 (
<figureCitation id="13162A62FFB6FFA0F834FF0CFF190C7D" captionStart="FIGURES 82–85" captionStartId="92.[151,264,1220,1243]" captionTargetBox="[410,1184,202,1192]" captionTargetId="figure-49@92.[398,1189,193,1199]" captionTargetPageId="92" captionText="FIGURES 82–85. Aptilotella germana Luk &amp; Marshall, sp. n.: 82—female terminalia, dorsal; 83—female terminalia, left lateral; 84—female terminalia, ventral; 85—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct." figureDoi="http://doi.org/10.5281/zenodo.4909191" httpUri="https://zenodo.org/record/4909191/files/figure.png" pageId="19" pageNumber="20">Figs. 82–84</figureCitation>
) convex; apex truncate; distal half setose. Cercus three times as long as wide; with one long apical seta and several preapical setae. Hypoproct (
<figureCitation id="13162A62FFB6FFA0FF1CFF44FD220C90" box="[588,741,367,392]" captionStart="FIGURES 82–85" captionStartId="92.[151,264,1220,1243]" captionTargetBox="[410,1184,202,1192]" captionTargetId="figure-49@92.[398,1189,193,1199]" captionTargetPageId="92" captionText="FIGURES 82–85. Aptilotella germana Luk &amp; Marshall, sp. n.: 82—female terminalia, dorsal; 83—female terminalia, left lateral; 84—female terminalia, ventral; 85—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct." figureDoi="http://doi.org/10.5281/zenodo.4909191" httpUri="https://zenodo.org/record/4909191/files/figure.png" pageId="19" pageNumber="20">Figs. 83, 84</figureCitation>
) reduced to a pair of curved, convergent rods; apically microtrichose and with two pairs of setae. Spermathecae (
<figureCitation id="13162A62FFB6FFA0FE7BFFBFFCB90CB5" box="[811,894,404,429]" captionStart="FIGURES 82–85" captionStartId="92.[151,264,1220,1243]" captionTargetBox="[410,1184,202,1192]" captionTargetId="figure-49@92.[398,1189,193,1199]" captionTargetPageId="92" captionText="FIGURES 82–85. Aptilotella germana Luk &amp; Marshall, sp. n.: 82—female terminalia, dorsal; 83—female terminalia, left lateral; 84—female terminalia, ventral; 85—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct." figureDoi="http://doi.org/10.5281/zenodo.4909191" httpUri="https://zenodo.org/record/4909191/files/figure.png" pageId="19" pageNumber="20">Fig. 85</figureCitation>
) simple; sclerotized ducts very long, five times the diameter of a spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFFB6FFA0FD97FFF7FBE90F00" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8B9236E7FFB6FFA0FD97FFF7FBE90F00" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<emphasis id="B959EAF5FFB6FFA0FD97FFF7FE800CED" bold="true" box="[199,327,476,501]" italics="true" pageId="19" pageNumber="20">Etymology.</emphasis>
The species epithet is from the Latin
<emphasis id="B959EAF5FFB6FFA0FFA4FFF6FCA50CEC" box="[756,866,477,500]" italics="true" pageId="19" pageNumber="20">germanus</emphasis>
, “having the same parents,” since this species and its relatives are characterized by unmistakably homologous aedeagal morphology.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFFB6FFA0FD97FC0FFF330FE8" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B9236E7FFB6FFA0FD97FC0FFC7A0F78" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<emphasis id="B959EAF5FFB6FFA0FD97FC0FFEB60F25" bold="true" box="[199,369,548,573]" italics="true" pageId="19" pageNumber="20">Type material.</emphasis>
<materialsCitation id="3B453CBAFFB6FFA0FC2DFC0FFC7A0F78" ID-GBIF-Occurrence="3311767341" collectingDate="2003-07-23" collectionCode="UNAM" collectorName="Cerro El Calvario &amp; R. S. Anderson." country="Mexico" elevation="2200" latitude="17.186666" location="Tapalapa" longLatPrecision="21" longitude="-93.122505" municipality="Cerro El Calvario" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="54968845FFB6FFA0FC2DFC0FFE220F25" box="[381,485,548,573]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2BFD6EFFB6FFA0FCA2FC0FFDCE0F23" box="[498,521,548,571]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FFB6FFA0FF4AFC0FFDB40F25" box="[538,627,548,573]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="19" pageNumber="20">UNAM</collectionCode>
.
<collectingCountry id="F33A7677FFB6FFA0FFD3FC0FFD310F25" box="[643,758,548,573]" name="Mexico" pageId="19" pageNumber="20">MEXICO</collectingCountry>
:
<collectingRegion id="49E9F805FFB6FFA0FE55FC0FFCA50F25" box="[773,866,548,573]" country="Mexico" name="Chiapas" pageId="19" pageNumber="20">Chiapas</collectingRegion>
,
<collectorName id="26D85331FFB6FFA0FE23FC0FFB8F0F25" box="[883,1096,548,573]" pageId="19" pageNumber="20">
<collectingMunicipality id="6BF6AC9DFFB6FFA0FE23FC0FFB8F0F25" box="[883,1096,548,573]" pageId="19" pageNumber="20">Cerro El Calvario</collectingMunicipality>
</collectorName>
, near
<location id="8EF2603CFFB6FFA0F9C4FC0FFB3C0F25" LSID="urn:lsid:plazi:treatment:038487F1FFB7FFA0FDC7FD6FFB9B0ED0:8EF2603CFFB6FFA0F9C4FC0FFB3C0F25" box="[1172,1275,548,573]" country="Mexico" latitude="17.186666" longLatPrecision="21" longitude="-93.122505" municipality="Cerro El Calvario" name="Tapalapa" pageId="19" pageNumber="20" stateProvince="Chiapas">Tapalapa</location>
,
<geoCoordinate id="EE195020FFB6FFA0F85CFC0FFA500F25" box="[1292,1431,548,573]" degrees="17" direction="north" minutes="11" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="12" value="17.186666">17°11’12”N</geoCoordinate>
,
<geoCoordinate id="EE195020FFB6FFA0FDC7FC6CFEDF0F78" box="[151,280,583,608]" degrees="93" direction="west" minutes="7" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="21" value="-93.122505">93°7’21”W</geoCoordinate>
,
<quantity id="4CD59B02FFB6FFA0FC74FC6CFEBC0F47" box="[292,379,583,608]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="m" value="2200.0">
<elevation id="0000D1D4FFB6FFA0FC74FC6CFEBC0F47" box="[292,379,583,608]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<date id="FF931027FFB6FFA0FCD6FC6CFDCF0F78" box="[390,520,583,608]" pageId="19" pageNumber="20" value="2003-07-23">
<collectingDate id="EFD7E9CFFFB6FFA0FCD6FC6CFDCF0F78" box="[390,520,583,608]" pageId="19" pageNumber="20" value="2003-07-23">23.vii.2003</collectingDate>
</date>
, wet cloud forest litter,
<collectorName id="26D85331FFB6FFA0FE41FC6CFC7A0F78" box="[785,957,583,608]" pageId="19" pageNumber="20">R.S. Anderson.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9236E7FFB6FFA0FD97FC47FF330FE8" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<materialsCitation id="3B453CBAFFB6FFA0FD97FC47FB7A0F9D" ID-GBIF-Occurrence="3311767410" box="[199,1213,620,645]" collectionCode="DEBU, UNAM" country="Mexico" location="Chiapas" pageId="19" pageNumber="20" specimenCount="19" specimenCount-female="10" specimenCount-male="9" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="B959EAF5FFB6FFA0FD97FC47FEF80F9B" bold="true" box="[199,319,620,643]" italics="true" pageId="19" pageNumber="20">
<typeStatus id="54968845FFB6FFA0FD97FC47FEFC0F9B" box="[199,315,620,643]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F33A7677FFB6FFA0FC16FC47FE7E0F9D" box="[326,441,620,645]" name="Mexico" pageId="19" pageNumber="20">MEXICO</collectingCountry>
:
<collectingRegion id="49E9F805FFB6FFA0FC94FC47FDE60F9D" box="[452,545,620,645]" country="Mexico" name="Chiapas" pageId="19" pageNumber="20">Chiapas</collectingRegion>
, same label as
<typeStatus id="54968845FFB6FFA0FF98FC47FCED0F9D" box="[712,810,620,645]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D2BFD6EFFB6FFA0FE6BFC47FCA70F9D" box="[827,864,620,645]" pageId="19" pageNumber="20" type="male">6♂</specimenCount>
,
<specimenCount id="9D2BFD6EFFB6FFA0FE3CFC47FC570F9D" box="[876,912,620,645]" pageId="19" pageNumber="20" type="female">6♀</specimenCount>
,
<collectionCode id="ED3CAE22FFB6FFA0FECDFC47FC2A0F9B" box="[925,1005,620,643]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="19" pageNumber="20">DEBU</collectionCode>
;
<specimenCount id="9D2BFD6EFFB6FFA0FEA8FC47FBDA0F9D" box="[1016,1053,620,645]" pageId="19" pageNumber="20" type="male">3♂</specimenCount>
,
<specimenCount id="9D2BFD6EFFB6FFA0F979FC47FB890F9D" box="[1065,1102,620,645]" pageId="19" pageNumber="20" type="female">4♀</specimenCount>
,
<collectionCode id="ED3CAE22FFB6FFA0F90AFC47FB720F9D" box="[1114,1205,620,645]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="19" pageNumber="20">UNAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFFB6FFA0F998FC47FA970FB0" ID-GBIF-Occurrence="3311767358" collectingDate="2008-05-27" collectionCode="DEBU" collectorName="de Tapalapa &amp; R. S. Anderson" country="Mexico" elevation="2260" latitude="17.187777" location="Cerro de Tapalapa" longLatPrecision="21" longitude="-93.123055" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<location id="8EF2603CFFB6FFA0F998FC47FA500F9D" LSID="urn:lsid:plazi:treatment:038487F1FFB7FFA0FDC7FD6FFB9B0ED0:8EF2603CFFB6FFA0F998FC47FA500F9D" box="[1224,1431,620,645]" country="Mexico" latitude="17.187777" longLatPrecision="21" longitude="-93.123055" name="Cerro de Tapalapa" pageId="19" pageNumber="20" stateProvince="Chiapas">
Cerro
<collectorName id="26D85331FFB6FFA0F85FFC47FA500F9D" box="[1295,1431,620,645]" pageId="19" pageNumber="20">de Tapalapa</collectorName>
</location>
,
<geoCoordinate id="EE195020FFB6FFA0FDC7FCA4FEE60FB0" box="[151,289,655,680]" degrees="17" direction="north" minutes="11" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="16" value="17.187777">17°11’16”N</geoCoordinate>
,
<geoCoordinate id="EE195020FFB6FFA0FC7EFCA4FE770FB0" box="[302,432,655,680]" degrees="93" direction="west" minutes="7" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="23" value="-93.123055">93°7’23”W</geoCoordinate>
,
<quantity id="4CD59B02FFB6FFA0FCEDFCA4FDD20FBF" box="[445,533,655,680]" metricMagnitude="3" metricUnit="m" metricValue="2.26" pageId="19" pageNumber="20" unit="m" value="2260.0">
<elevation id="0000D1D4FFB6FFA0FCEDFCA4FDD20FBF" box="[445,533,655,680]" metricMagnitude="3" metricUnit="m" metricValue="2.26" pageId="19" pageNumber="20" unit="m" value="2260.0">2260 m</elevation>
</quantity>
,
<date id="FF931027FFB6FFA0FF72FCA4FD550FB0" box="[546,658,655,680]" pageId="19" pageNumber="20" value="2008-05-27">
<collectingDate id="EFD7E9CFFFB6FFA0FF72FCA4FD550FB0" box="[546,658,655,680]" pageId="19" pageNumber="20" value="2008-05-27">27.v.2008</collectingDate>
</date>
, cloud forest, ex. sifted leaf litter,
<collectorName id="26D85331FFB6FFA0F94BFCA4FB060FB0" box="[1051,1217,655,680]" pageId="19" pageNumber="20">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFFB6FFA0F982FCA4FB2E0FB0" box="[1234,1257,655,680]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FFB6FFA0F9A6FCA4FA800FB0" box="[1270,1351,655,680]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="19" pageNumber="20">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFFB6FFA0F80CFCA4FF370FE8" ID-GBIF-Occurrence="3311767313" collectingDate="2008-05-27" collectionCode="DEBU" collectorName="de Tapalapa &amp; R. S. Anderson" country="Mexico" elevation="2240" latitude="17.191666" location="Cerro de Tapalapa" longLatPrecision="21" longitude="-93.11778" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas" typeStatus="paratype">
<location id="8EF2603CFFB6FFA0F80CFCA4FEE50FD5" LSID="urn:lsid:plazi:treatment:038487F1FFB7FFA0FDC7FD6FFB9B0ED0:8EF2603CFFB6FFA0F80CFCA4FEE50FD5" country="Mexico" latitude="17.191666" longLatPrecision="21" longitude="-93.11778" name="Cerro de Tapalapa" pageId="19" pageNumber="20" stateProvince="Chiapas">
Cerro
<collectorName id="26D85331FFB6FFA0FDC7FC9FFEE50FD5" box="[151,290,692,717]" pageId="19" pageNumber="20">de Tapalapa</collectorName>
</location>
,
<geoCoordinate id="EE195020FFB6FFA0FC7EFC9FFE7F0FD5" box="[302,440,692,717]" degrees="17" direction="north" minutes="11" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="30" value="17.191666">17°11’30”N</geoCoordinate>
,
<geoCoordinate id="EE195020FFB6FFA0FC95FC9FFDFE0FD5" box="[453,569,692,717]" degrees="93" direction="west" minutes="7" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="4" value="-93.11778">93°7’4”W</geoCoordinate>
,
<quantity id="4CD59B02FFB6FFA0FF15FC9FFD590FD3" box="[581,670,692,717]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="19" pageNumber="20" unit="m" value="2240.0">
<elevation id="0000D1D4FFB6FFA0FF15FC9FFD590FD3" box="[581,670,692,717]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="19" pageNumber="20" unit="m" value="2240.0">2240 m</elevation>
</quantity>
,
<date id="FF931027FFB6FFA0FFFBFC9FFCDC0FD5" box="[683,795,692,717]" pageId="19" pageNumber="20" value="2008-05-27">
<collectingDate id="EFD7E9CFFFB6FFA0FFFBFC9FFCDC0FD5" box="[683,795,692,717]" pageId="19" pageNumber="20" value="2008-05-27">27.v.2008</collectingDate>
</date>
, oak-pine forest, ex. sifted leaf litter,
<collectorName id="26D85331FFB6FFA0F999FC9FFAA90FD5" box="[1225,1390,692,717]" pageId="19" pageNumber="20">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFFB6FFA0F8D2FC9FFA510FD5" box="[1410,1430,692,717]" pageId="19" pageNumber="20" type="female">♀</specimenCount>
,
<collectionCode id="ED3CAE22FFB6FFA0FDC7FCFCFF2F0FE8" box="[151,232,727,752]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="19" pageNumber="20">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFFB6FFA0FD97FCD7FB9B0ED0" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B9236E7FFB6FFA0FD97FCD7FAE50E98" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<emphasis id="B959EAF5FFB6FFA0FD97FCD7FE8F0E0D" bold="true" box="[199,328,764,789]" pageId="19" pageNumber="20">
<emphasis id="B959EAF5FFB6FFA0FD97FCD7FE850E0D" bold="true" box="[199,322,764,789]" italics="true" pageId="19" pageNumber="20">Comments</emphasis>
.
</emphasis>
<taxonomicName id="4C2D4D64FFB6FFA0FC00FCD7FDF50E0C" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[336,562,764,788]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="germana">
<emphasis id="B959EAF5FFB6FFA0FC00FCD7FDF50E0C" box="[336,562,764,788]" italics="true" pageId="19" pageNumber="20">Aptilotella germana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D64FFB6FFA0FF39FCD7FCC60E0C" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[617,769,764,788]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pyropanda">
<emphasis id="B959EAF5FFB6FFA0FF39FCD7FCC60E0C" box="[617,769,764,788]" italics="true" pageId="19" pageNumber="20">A. pyropanda</emphasis>
</taxonomicName>
are the only
<taxonomicName id="4C2D4D64FFB6FFA0FEC5FCD7FBCC0E0C" authorityName="Duda" authorityYear="1924" box="[917,1035,764,788]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FFB6FFA0FEC5FCD7FBCC0E0C" box="[917,1035,764,788]" italics="true" pageId="19" pageNumber="20">Aptilotella</emphasis>
</taxonomicName>
with a nearly or fully divided male sternite 5. The aedeagus is incredibly autapomorphic and character-rich, with paired arched sclerites of the distiphallus, broad postgonite and hypandrium, and squat basiphallus. They also differ from other squat-bodied
<taxonomicName id="4C2D4D64FFB6FFA0FDC7FD43FECA0E98" authorityName="Duda" authorityYear="1924" box="[151,269,872,896]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FFB6FFA0FDC7FD43FECA0E98" box="[151,269,872,896]" italics="true" pageId="19" pageNumber="20">Aptilotella</emphasis>
</taxonomicName>
in possessing a row of 5–6 stout peg-like setae in the ventrodistal third of the male mid tibia.
</paragraph>
<paragraph id="8B9236E7FFB6FFA0FDC7FDA7FB9B0ED0" blockId="19.[151,1436,151,968]" pageId="19" pageNumber="20">
<taxonomicName id="4C2D4D64FFB6FFA0FDC7FDA7FEBD0EBC" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,378,908,932]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="germana">
<emphasis id="B959EAF5FFB6FFA0FDC7FDA7FEBD0EBC" box="[151,378,908,932]" italics="true" pageId="19" pageNumber="20">Aptilotella germana</emphasis>
</taxonomicName>
differs subtly from all other
<taxonomicName id="4C2D4D64FFB6FFA0FF99FDA7FCF80EBC" authorityName="Duda" authorityYear="1924" box="[713,831,908,932]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FFB6FFA0FF99FDA7FCF80EBC" box="[713,831,908,932]" italics="true" pageId="19" pageNumber="20">Aptilotella</emphasis>
</taxonomicName>
in possessing a minute, dark brown, thickened wing “bud.” Several undescribed species have somewhat larger and triangular wing “buds.”
</paragraph>
</subSubSection>
</treatment>
</document>
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