treatments-xml/data/03/84/87/038487E62F3C2424FF12B3EDFE8FFC7D.xml
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<document id="82E5A58AB194BA11C74E4A08E0F6E73F" ID-CLB-Dataset="6241" ID-DOI="10.11646/zootaxa.4973.1.1" ID-GBIF-Dataset="d0967b08-98d1-4ccd-b472-a678e0e57eda" ID-ISSN="1175-5326" ID-Zenodo-Dep="4771958" ID-ZooBank="42EB4BF2-A571-4894-9EEF-783649A27E4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1621409551899" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander &amp; Auffenberg, Kurt" docDate="2021" docId="038487E62F3C2424FF12B3EDFE8FFC7D" docLanguage="en" docName="zootaxa.4973.1.1.pdf" docOrigin="Zootaxa 4973 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dicharax gnomus Kobelt &amp; Mollendorff 1900, n. sp." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FFBDFF9E2F20243AFF85B51EFF9BFFD8" masterDocTitle="Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)" masterLastPageNumber="61" masterPageNumber="1" pageNumber="29" updateTime="1698940034724" updateUser="ExternalLinkService">
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<mods:title id="907AFBA5C307CCDD0643680EEDE94668">Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)</mods:title>
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<mods:namePart id="F4F05334CD5E75F7B77A0AEF2EC63A3F">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="CFB6450E31D9EBD48FE235AF07D7AF13">Hunyadi, András</mods:namePart>
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<mods:namePart id="DFC98F7E8F43B47E60693217ED7AD661">Grego, Jozef</mods:namePart>
<mods:nameIdentifier id="C3DC600066532FBEEFF2E3C6EFCBBBCF" type="ORCID">0000-0002-4977-0415</mods:nameIdentifier>
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<mods:namePart id="9D6B802DD01544FA2621CD42FDA80912">Reischütz, Alexander</mods:namePart>
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<mods:nameIdentifier id="85553F4C9EC9A0ED50B6203F762D26E8" type="email">alexander.reischuetz@gmx.at</mods:nameIdentifier>
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<mods:namePart id="1F82949309BFD7079981FC1272EA7F51">Auffenberg, Kurt</mods:namePart>
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<mods:nameIdentifier id="BBED1C16CA388309EA6715D6E9C2CBCE" type="email">kurtauffenberg@gmail.com</mods:nameIdentifier>
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<mods:title id="B3B1C16C790904A213753CBA96AB3EEF">Zootaxa</mods:title>
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<mods:part id="1F6DD7E5F31143585BD8516DD3FE4382">
<mods:date id="AD7E161CC5494CC577CB34FCB0D4878F">2021</mods:date>
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<mods:number id="03411EB1E8B28943CAD790302963A841">2021-05-18</mods:number>
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<mods:number id="7ABADF7C3B2A18C74D873CA6403A397F">4973</mods:number>
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<mods:identifier id="8F6CC0EDF4A7490B74E2311AE10493BB" type="DOI">10.11646/zootaxa.4973.1.1</mods:identifier>
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<treatment id="038487E62F3C2424FF12B3EDFE8FFC7D" ID-DOI="http://doi.org/10.5281/zenodo.4916575" ID-GBIF-Taxon="182640502" ID-Zenodo-Dep="4916575" LSID="urn:lsid:plazi:treatment:038487E62F3C2424FF12B3EDFE8FFC7D" httpUri="http://treatment.plazi.org/id/038487E62F3C2424FF12B3EDFE8FFC7D" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C337657B2F3C2426FF12B3EDFD12F8D6" box="[151,649,1778,1806]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B9236F02F3C2426FF12B3EDFD12F8D6" blockId="28.[151,649,1778,1842]" box="[151,649,1778,1806]" pageId="28" pageNumber="29">
<heading id="D0DA819C2F3C2426FF12B3EDFD12F8D6" bold="true" box="[151,649,1778,1806]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B959EAE22F3C2426FF12B3EDFD12F8D6" bold="true" box="[151,649,1778,1806]" pageId="28" pageNumber="29">
<taxonomicName id="4C2D4D732F3C2426FF12B3EDFDDBF8D5" authority="Pall-Gergely" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[151,576,1778,1806]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B959EAE22F3C2426FF12B3EDFE92F8D6" bold="true" box="[151,265,1779,1806]" italics="true" pageId="28" pageNumber="29">Dicharax</emphasis>
(?)
<emphasis id="B959EAE22F3C2426FEBFB3EAFE01F8D6" bold="true" box="[314,410,1780,1806]" italics="true" pageId="28" pageNumber="29">gnomus</emphasis>
Páll-Gergely
</taxonomicName>
<taxonomicNameLabel id="A26A57992F3C2426FDC2B3EAFD12F8D6" box="[583,649,1780,1806]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3C2426FF12B209FF7CF8EA" box="[151,231,1815,1842]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9236F02F3C2426FF12B209FF7CF8EA" blockId="28.[151,649,1778,1842]" box="[151,231,1815,1842]" pageId="28" pageNumber="29">
<figureCitation id="13162A752F3C2426FF12B209FF7CF8EA" box="[151,231,1815,1842]" captionStart="FIGURE 22" captionStartId="29.[151,250,1776,1800]" captionTargetBox="[151,1436,181,1752]" captionTargetId="figure-18@29.[151,1436,181,1752]" captionTargetPageId="29" captionText="FIGURE 22. Dicharax (?) gnomus Páll-Gergely, n. sp., holotype (UF 380401)." figureDoi="http://doi.org/10.5281/zenodo.4772008" httpUri="https://zenodo.org/record/4772008/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3C2426FF12B241FDBCF81A" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B9236F02F3C2426FF12B241FDBCF81A" blockId="28.[151,1437,1887,1986]" pageId="28" pageNumber="29">
<emphasis id="B959EAE22F3C2426FF12B241FE1AF8A1" bold="true" box="[151,385,1887,1913]" pageId="28" pageNumber="29">Material examined.</emphasis>
<materialsCitation id="3B453CAD2F3C2426FE0CB27EFDBCF81A" ID-GBIF-Occurrence="3128117308" accessionNumber="UF 380401" collectingDate="1988-05-16" collectionCode="UF" collectorName="F. G. Thompson" country="Thailand" elevation="200" latitude="18.168884" location="Ban Nim Rin" longLatPrecision="1" longitude="99.93722" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Phrae" typeStatus="holotype">
<emphasis id="B959EAE22F3C2426FE0CB27EFE64F8A2" bold="true" box="[393,511,1888,1914]" pageId="28" pageNumber="29">
<typeStatus id="549688522F3C2426FE0CB27EFE61F8A2" box="[393,506,1888,1914]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F33A76602F3C2426FD82B241FDF6F8A1" box="[519,621,1887,1913]" name="Thailand" pageId="28" pageNumber="29">Thailand</collectingCountry>
,
<collectingRegion id="49E9F8122F3C2426FDFCB241FCBCF8A1" box="[633,807,1887,1913]" country="Thailand" name="Phrae" pageId="28" pageNumber="29">Phrae Province</collectingRegion>
,
<quantity id="4CD59B152F3C2426FCB6B27EFCF6F8A1" box="[819,877,1887,1914]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="km" value="4.0">4 km</quantity>
N of
<location id="8EF2602B2F3C2426FC35B27EFBD1F8A1" LSID="urn:lsid:plazi:treatment:038487E62F3C2424FF12B3EDFE8FFC7D:8EF2602B2F3C2426FC35B27EFBD1F8A1" box="[944,1098,1887,1914]" country="Thailand" latitude="18.168884" longLatPrecision="1" longitude="99.93722" name="Ban Nim Rin" pageId="28" pageNumber="29" stateProvince="Phrae">Ban Nim Rin</location>
, valley with limestone ridges,
<geoCoordinate id="EE1950372F3C2426FF12B29AFEB7F846" box="[151,300,1924,1950]" degrees="18" direction="north" minutes="10.133" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="18.168884">18°10.133N</geoCoordinate>
,
<geoCoordinate id="EE1950372F3C2426FEBFB29DFE50F846" box="[314,459,1923,1950]" degrees="99" direction="east" minutes="56.233" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="99.93722">99°56.233E</geoCoordinate>
,
<elevation id="0000D1C32F3C2426FE5FB29AFDFFF845" box="[474,612,1923,1950]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="200.0">
<quantity id="4CD59B152F3C2426FE5FB29AFDBFF846" box="[474,548,1924,1950]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9310302F3C2426FDF1B29DFC88F846" box="[628,787,1923,1950]" pageId="28" pageNumber="29" value="1988-05-16">
<collectingDate id="EFD7E9D82F3C2426FDF1B29DFC88F846" box="[628,787,1923,1950]" pageId="28" pageNumber="29" value="1988-05-16">16 May 1988</collectingDate>
</date>
,
<collectorName id="26D853262F3C2426FCA4B29AFC48F845" box="[801,979,1923,1950]" pageId="28" pageNumber="29">F.G. Thompson</collectorName>
leg., leaf litter at base of boulders,
<collectionCode id="ED3CAE352F3C2426FAFCB29AFA06F846" box="[1401,1437,1924,1950]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="28" pageNumber="29" type="Museum">UF</collectionCode>
380401 (D:
<quantity id="4CD59B152F3C2426FE9AB2B9FE1EF81A" box="[287,389,1959,1986]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="28" pageNumber="29" unit="mm" value="1.65">1.65 mm</quantity>
, H:
<quantity id="4CD59B152F3C2426FE36B2B9FD80F81A" box="[435,539,1959,1986]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="28" pageNumber="29" unit="mm" value="1.15">1.15 mm</quantity>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF5266782F3D2427FF12B3EEFC27F8D0" ID-DOI="http://doi.org/10.5281/zenodo.4772008" ID-Zenodo-Dep="4772008" box="[151,956,1776,1800]" httpUri="https://zenodo.org/record/4772008/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1776,1800]" targetBox="[151,1436,181,1752]" targetPageId="29">
<paragraph id="8B9236F02F3D2427FF12B3EEFC27F8D0" blockId="29.[151,956,1776,1800]" box="[151,956,1776,1800]" pageId="29" pageNumber="30">
<emphasis id="B959EAE22F3D2427FF12B3EEFEBBF8D0" bold="true" box="[151,288,1776,1800]" pageId="29" pageNumber="30">FIGURE 22.</emphasis>
<taxonomicName id="4C2D4D732F3D2427FEA3B3EEFE1DF8D0" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[294,390,1776,1800]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F3D2427FEA3B3EEFE1DF8D0" box="[294,390,1776,1800]" italics="true" pageId="29" pageNumber="30">Dicharax</emphasis>
</taxonomicName>
(?)
<taxonomicName id="4C2D4D732F3D2427FE2AB3EFFD1DF8D0" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2021" box="[431,646,1776,1800]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="gnomus" status="sp. nov.">
<emphasis id="B959EAE22F3D2427FE2AB3EFFE65F8D0" box="[431,510,1777,1800]" italics="true" pageId="29" pageNumber="30">gnomus</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="A26A57992F3D2427FD15B3EEFD53F8DF" box="[656,712,1776,1799]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
, holotype (UF 380401).
</paragraph>
</caption>
<subSubSection id="C337657B2F3D2427FF42B228FBD0F8AD" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B9236F02F3D2427FF42B228FBD0F8AD" blockId="29.[151,1436,1846,2017]" pageId="29" pageNumber="30">
<emphasis id="B959EAE22F3D2427FF42B228FED5F888" bold="true" box="[199,334,1846,1872]" pageId="29" pageNumber="30">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2D4D732F3D2427FDBBB229FD0EF888" authorityName="Pall-Gergely" authorityYear="2021" box="[574,661,1847,1872]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="gnomus">
<emphasis id="B959EAE22F3D2427FDBBB229FD0EF888" box="[574,661,1847,1872]" italics="true" pageId="29" pageNumber="30">gnomus</emphasis>
</taxonomicName>
(Medieval Latin for gnome) refers to the tiny shell that has unusual shell area ratios compared to most other species (to be used as a noun in apposition).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3D2427FF42B260FDE3F865" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8B9236F02F3D2427FF42B260FDE3F865" blockId="29.[151,1436,1846,2017]" pageId="29" pageNumber="30">
<emphasis id="B959EAE22F3D2427FF42B260FED9F840" bold="true" box="[199,322,1918,1944]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
Tiny
<taxonomicName id="4C2D4D732F3D2427FE0FB260FDCBF840" box="[394,592,1918,1944]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B959EAE22F3D2427FE0FB260FE6FF840" box="[394,500,1918,1944]" italics="true" pageId="29" pageNumber="30">Dicharax</emphasis>
species
</taxonomicName>
with wide, flat R1 ribs and fine spiral striation, short R2 with flat ribs, strongly reduced R3, and circular aperture.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3D2424FF42B2D8FE4FFDE5" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8B9236F02F3D2424FF42B2D8FF44FE2D" blockId="29.[151,1436,1846,2017]" lastBlockId="30.[151,1437,150,933]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B959EAE22F3D2427FF42B2D8FEC2F838" bold="true" box="[199,345,1990,2016]" pageId="29" pageNumber="30">Description.</emphasis>
Shell light grey, slightly semi-translucent; shell outline rounded in dorsal view; spire slightly elevated, body whorl rounded; protoconch low, finely granular, glossy, 1.5 whorls; R1 of 1.25 whorls, beginning with low ribs that appear curved anteriorly, developing into wide, flattened ribs separated by grooves, forming a smoothish, glossy surface; first whorl of R1 with some fine spiral striation; boundary between R1 and R2 not conspicuous due to similar rib morphology, although ribs become denser, ca. 30 R2 ribs present; R2+R3 less than quarter whorl (ca. 7080˚); R3 considerably shorter than R2; boundary between R2 and R3 distinct due to sculpture change and rather deep constriction; R3 strongly reduced, represented only as a slight swelling just behind peristome; aperture almost parallel with shell axis, circular, round; peristome weak, very slightly thickened; boundary between inner and outer peristomes barely discernible; inner peristome weak, somewhat protruding and very slightly expanded, outer peristome very weakly developed, only indicated; umbilicus relatively narrow, less than one third of shell width.
</paragraph>
<paragraph id="8B9236F02F3E2424FF42B4E1FD3CFDC1" blockId="30.[151,1437,150,933]" box="[199,679,510,537]" pageId="30" pageNumber="31">
<emphasis id="B959EAE22F3E2424FF42B4E1FEE5FDC1" bold="true" box="[199,382,511,537]" pageId="30" pageNumber="31">Measurements.</emphasis>
D:
<quantity id="4CD59B152F3E2424FE22B4E0FD96FDC1" box="[423,525,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="30" pageNumber="31" unit="mm" value="1.65">1.65 mm</quantity>
, H:
<quantity id="4CD59B152F3E2424FDB9B4E0FD38FDC1" box="[572,675,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="30" pageNumber="31" unit="mm" value="1.15">1.15 mm</quantity>
.
</paragraph>
<paragraph id="8B9236F02F3E2424FF42B73CFE4FFDE5" blockId="30.[151,1437,150,933]" box="[199,468,546,573]" pageId="30" pageNumber="31">
<emphasis id="B959EAE22F3E2424FF42B73CFECCFDE4" bold="true" box="[199,343,546,572]" pageId="30" pageNumber="31">Operculum.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3E2424FF42B758FBADFD71" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8B9236F02F3E2424FF42B758FBADFD71" blockId="30.[151,1437,150,933]" pageId="30" pageNumber="31">
<emphasis id="B959EAE22F3E2424FF42B758FE54FDB8" bold="true" box="[199,463,582,608]" pageId="30" pageNumber="31">Differential diagnosis.</emphasis>
The R3 region practically absent in this species. No other low-spired alycaeid species has this trait in
<collectingCountry id="F33A76602F3E2424FEC2B775FE30FD5D" box="[327,427,619,645]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
or
<collectingCountry id="F33A76602F3E2424FE54B775FDDBFD5D" box="[465,576,619,645]" name="Myanmar" pageId="30" pageNumber="31">Myanmar</collectingCountry>
. The
<taxonomicName id="4C2D4D732F3E2424FDFBB775FCDAFD5D" box="[638,833,619,645]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B959EAE22F3E2424FDFBB775FD73FD5D" box="[638,744,619,645]" italics="true" pageId="30" pageNumber="31">Dicharax</emphasis>
species
</taxonomicName>
described as
<taxonomicName id="4C2D4D732F3E2424FC5EB775FA99FD5D" authority="Kuroda, 1951" authorityName="Kuroda" authorityYear="1951" box="[987,1282,618,645]" class="Gastropoda" family="Cyclophoridae" genus="Awalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F3E2424FC5EB775FBC4FD5D" box="[987,1119,619,645]" italics="true" pageId="30" pageNumber="31">Awalycaeus</emphasis>
<bibRefCitation id="EFBC4B012F3E2424FBE2B775FA99FD5D" author="Kuroda, T." box="[1127,1282,618,645]" pageId="30" pageNumber="31" pagination="73 - 74" refId="ref25526" refString="Kuroda, T. (1951) On Awalycaeus abei gen. et. sp. nov. (Cyclophoridae). Venus, 16 (5 - 8), 73 - 74." type="journal article" year="1951">
Kuroda,
<quantity id="4CD59B152F3E2424FB4FB774FA99FD5D" box="[1226,1282,618,645]" metricMagnitude="1" metricUnit="m" metricValue="4.95554" pageId="30" pageNumber="31" unit="in" value="1951.0">1951</quantity>
</bibRefCitation>
</taxonomicName>
in
<collectingCountry id="F33A76602F3E2424FAADB775FAF3FD5D" box="[1320,1384,619,645]" name="Japan" pageId="30" pageNumber="31">Japan</collectingCountry>
also have a strongly reduced R3, but we consider this is a case of convergent evolution.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3E2424FF42B7ACFBAEFD15" box="[199,1077,690,717]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8B9236F02F3E2424FF42B7ACFBAEFD15" blockId="30.[151,1437,150,933]" box="[199,1077,690,717]" pageId="30" pageNumber="31">
<emphasis id="B959EAE22F3E2424FF42B7ACFEFAFD14" bold="true" box="[199,353,690,716]" pageId="30" pageNumber="31">Distribution.</emphasis>
This new species is known only from the
<typeStatus id="549688522F3E2424FCBFB7ADFCF1FD15" box="[826,874,691,717]" pageId="30" pageNumber="31">type</typeStatus>
locality (
<figureCitation id="13162A752F3E2424FC51B7ADFBBCFD15" box="[980,1063,691,717]" captionStart="FIGURE 17" captionStartId="23.[151,250,1861,1885]" captionTargetBox="[202,1386,1025,1838]" captionTargetId="figure-32@23.[202,1386,1025,1838]" captionTargetPageId="23" captionText="FIGURE 17. Distribution of alycaeid species in northern Thailand and Myanmar. filled circle: Dicharax admirandus Páll- Gergely &amp; Auffenberg, n. sp., empty square: Dicharax ebrius Páll-Gergely &amp; Auffenberg, n. sp., filled triangle: Dicharax gnomus Páll-Gergely, n. sp., filled square: Dicharax mirounga Páll-Gergely, n. sp., inverted empty triangles: Dicharax verruco- sus Páll-Gergely &amp; Hunyadi n. sp., rhombus: Dicharax stuparum Páll-Gergely &amp; Hunyadi, 2018, empty triangle: Metalycaeus anas Páll-Gergely &amp; A. Reischütz, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4771996" httpUri="https://zenodo.org/record/4771996/files/figure.png" pageId="30" pageNumber="31">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F3E2424FF42B7C9FE8FFC7D" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8B9236F02F3E2424FF42B7C9FE8FFC7D" blockId="30.[151,1437,150,933]" pageId="30" pageNumber="31">
<emphasis id="B959EAE22F3E2424FF42B7C9FEA0FD29" bold="true" box="[199,315,727,753]" pageId="30" pageNumber="31">Remarks.</emphasis>
This species has unusual conchological characters, and therefore, its generic placement is unclear. The spiral striation on R1 and the lack of spiral striation on the protoconch suggests a relationship to
<taxonomicName id="4C2D4D732F3E2424FB75B7E5FA0EFCCD" authorityName="Mollendorff" authorityYear="1897" box="[1264,1429,763,789]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F3E2424FB75B7E5FA0EFCCD" box="[1264,1429,763,789]" italics="true" pageId="30" pageNumber="31">Chamalycaeus</emphasis>
</taxonomicName>
, whereas the glossy shell surface, the flattened R2 ribs, and the low protoconch indicate a position in
<taxonomicName id="4C2D4D732F3E2424FAAEB601FA0EFCE1" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[1323,1429,799,825]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F3E2424FAAEB601FA0EFCE1" box="[1323,1429,799,825]" italics="true" pageId="30" pageNumber="31">Dicharax</emphasis>
</taxonomicName>
. This study revealed other
<taxonomicName id="4C2D4D732F3E2424FE39B65DFDE5FC85" box="[444,638,835,861]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B959EAE22F3E2424FE39B65DFDBDFC85" box="[444,550,835,861]" italics="true" pageId="30" pageNumber="31">Dicharax</emphasis>
species
</taxonomicName>
/populations that possess some spiral striae on R1 (see under
<taxonomicName id="4C2D4D732F3E2424FAB7B65DFE86FC59" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg, 2021" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="admirandus" status="sp. nov.">
<emphasis id="B959EAE22F3E2424FAB7B65DFE86FC59" italics="true" pageId="30" pageNumber="31">Dicharax admirandus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26A57992F3E2424FEA9B679FEF2FC59" box="[300,361,871,897]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C2D4D732F3E2424FE27B679FD85FC58" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[418,542,871,896]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="omissus">
<emphasis id="B959EAE22F3E2424FE27B679FD85FC58" box="[418,542,871,896]" italics="true" pageId="30" pageNumber="31">D. omissus</emphasis>
</taxonomicName>
and Páll-Gergely
<emphasis id="B959EAE22F3E2424FD68B679FCBBFC59" box="[749,800,871,897]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 2017). Thus, the placement of this species in
<taxonomicName id="4C2D4D732F3E2424FAB7B679FA07FC59" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[1330,1436,871,897]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F3E2424FAB7B679FA07FC59" box="[1330,1436,871,897]" italics="true" pageId="30" pageNumber="31">Dicharax</emphasis>
</taxonomicName>
is justified.
</paragraph>
</subSubSection>
</treatment>
</document>