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<document id="B6577EE42D78A7323FEA9ABB27D85AB7" ID-CLB-Dataset="26473" ID-DOI="10.11646/zootaxa.4616.1.1" ID-GBIF-Dataset="324b030a-18a2-4dac-9ddd-28c468aea7df" ID-ISSN="1175-5326" ID-Zenodo-Dep="3248018" ID-ZooBank="1B9DFBC9-2BA8-4619-8F70-E372DCBD7411" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560832298691" checkinUser="plazi" docAuthor="Normark, Benjamin B., Okusu, Akiko, Morse, Geoffrey E., Peterson, Daniel A., Itioka, Takao &amp; Schneider, Scott A." docDate="2019" docId="038487C687589632A2A1F90DEAA6FB59" docLanguage="en" docName="zootaxa.4616.1.1.pdf" docOrigin="Zootaxa 4616 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Diaspididae Bacteriodetes A 1096" docType="key" docVersion="11" lastPageNumber="47" masterDocId="FFBDFFBE8775961CA236FFB9EF3AFFCB" masterDocTitle="Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="46" updateTime="1698734953532" updateUser="ExternalLinkService">
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<mods:title id="446D7A383580608CE386C34A5AE66993">Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae)</mods:title>
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<mods:namePart id="7932D93B47EA85C5EF557DD90AE1887E">Normark, Benjamin B.</mods:namePart>
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<mods:namePart id="FED7121F3446B04364242D66D81DD222">Okusu, Akiko</mods:namePart>
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<mods:namePart id="AF7782215A9CB85DD50D31F8D76F3FF3">Morse, Geoffrey E.</mods:namePart>
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<mods:namePart id="EA32866144169B1E9B22B83B94B7ABED">Peterson, Daniel A.</mods:namePart>
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<mods:namePart id="EE94A3FC457F44413D7C01C222FD07AB">Itioka, Takao</mods:namePart>
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<mods:namePart id="34FE6933E6B6391BA641F647418A8472">Schneider, Scott A.</mods:namePart>
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<mods:date id="60A16B9D60EBF596721898BCDA32CBFA">2019</mods:date>
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<mods:number id="A8CE52A6B5683BE9E10D2DEBE73C7F4E">4616</mods:number>
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<mods:classification id="6B3E5A688A25A5D3BD7D02FCDED5281A">journal article</mods:classification>
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<mods:identifier id="24BC4F1DEDB8206F79F38E3D744752A8" type="DOI">10.11646/zootaxa.4616.1.1</mods:identifier>
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<treatment id="038487C687589632A2A1F90DEAA6FB59" ID-DOI="http://doi.org/10.5281/zenodo.5925392" ID-GBIF-Taxon="158289975" ID-Zenodo-Dep="5925392" LSID="urn:lsid:plazi:treatment:038487C687589632A2A1F90DEAA6FB59" httpUri="http://treatment.plazi.org/id/038487C687589632A2A1F90DEAA6FB59" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection id="C337655B87589631A2A1F90DED03F905" box="[151,569,1716,1742]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8B9236D087589631A2A1F90DED03F905" blockId="45.[151,569,1716,1742]" box="[151,569,1716,1742]" pageId="45" pageNumber="46">
<heading id="D0DA81BC87589631A2A1F90DED03F905" bold="true" box="[151,569,1716,1742]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="B959EAC287589631A2A1F90DED03F905" bold="true" box="[151,569,1716,1742]" pageId="45" pageNumber="46">
Key to subfamilies of
<taxonomicName id="4C2D4D5387589631A393F90DED03F905" ID-CoL="96X" ID-ENA="30080" authorityName="Bacteriodetes A" authorityYear="1096" box="[421,569,1716,1742]" class="Insecta" family="Diaspididae" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="family">Diaspididae</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337655B87589632A2A1F943EAA6FB59" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="key">
<paragraph id="8B9236D087589632A2A1F943EAA1FE5C" blockId="45.[151,1436,1786,2012]" lastBlockId="46.[151,1436,151,1170]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<keyStep id="30D92E7587589632A2A1F943EAA1FE5C" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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1
<emphasis id="B959EAC287589631A2D7F943EE38F8DB" bold="true" box="[225,258,1786,1808]" pageId="45" pageNumber="46">AF</emphasis>
permanently enclosed within sclerotized cuticle of 2F (pupillarial); without plates or gland spines; ducts absent or minute; perivulvar pores, when present, forming a horizontal band anterior to vulva; antenna multisetose (rarely unisetose); with 03 (usually 0) pairs of pygidial lobes; median lobes, when present, roughly quadrate, with serrate to notched apical margin (sometimes appearing bilobed); body size minute (
<quantity id="4CD59B3587589631A0B9F8EBEC3EF8AC" box="[655,772,1874,1895]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="45" pageNumber="46" unit="mm" value="0.6" valueMax="0.8" valueMin="0.4">0.40.8 mm</quantity>
long);
<emphasis id="B959EAC287589631A17DF8EBEC5FF8A3" bold="true" box="[843,869,1874,1896]" pageId="45" pageNumber="46">2F</emphasis>
body margin without plates; with or without gland spines; with or without ducts—when marginal ducts present, these are 2-barred and often short and wide (barrel-shaped);
<emphasis id="B959EAC287589631A71CF8D6EA77F84E" bold="true" box="[1322,1357,1903,1925]" pageId="45" pageNumber="46">CR</emphasis>
abdominal segments IIIVI each with 2 submedian setae (1 dorsal, 1 ventral); head without dorsal ducts; abdominal segment II with a submedian dorsal duct;
<emphasis id="B959EAC287589631A3F3F813EEE6F80B" bold="true" box="[453,476,1962,1984]" pageId="45" pageNumber="46">M</emphasis>
diploid, with half of the chromosomes heterochromatic; antenna 6-segmented, without annulations; Nearctic and Neotropical................................................................ ANCEPASPIDINAE -
<emphasis id="B959EAC2875B9632A2D7FF2EEE38FF66" bold="true" box="[225,258,151,173]" pageId="46" pageNumber="47">AF</emphasis>
permanently enclosed within the sclerotized cuticle of 2F (pupillarial) or not (non-pupillarial); usually with gland spines, or plates, or both; ducts usually present, often large; perivulvar pores, when present, usually in 45 groups, some pores situated posterolaterad of vulva; antenna unisetose or (less commonly) multisetose; with 05 (usually 13) pairs of pygidial lobes; median lobes various, but never appearing bilobed; body size various (
<quantity id="4CD59B35875B9632A1B6FF56ECD8FECF" box="[896,994,239,260]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="3.0" metricValueMin="0.4" pageId="46" pageNumber="47" unit="mm" value="1.7" valueMax="3.0" valueMin="0.4">0.43 mm</quantity>
long);
<emphasis id="B959EAC2875B9632A611FF56EB7AFECE" bold="true" box="[1063,1088,239,261]" pageId="46" pageNumber="47">2F</emphasis>
body margin with or without plates; with or without gland spines; usually with ducts, which may be 1- or 2-barred and of various sizes and shapes;
<emphasis id="B959EAC2875B9632A738FEB5EA0BFEE9" bold="true" box="[1294,1329,268,290]" pageId="46" pageNumber="47">CR</emphasis>
abdominal segments IIIVI each with 1 submedian seta, which may be dorsal or ventral; head with or without dorsal ducts; if submarginal dorsal ducts are absent from head, abdominal segment II lacks a submedial dorsal duct;
<emphasis id="B959EAC2875B9632A60BFEFEEB6EFE96" bold="true" box="[1085,1108,327,349]" pageId="46" pageNumber="47">M</emphasis>
haploid, without heterochromatic chromosomes or (less commonly) diploid, with half of the chromosomes heterochromatic; antenna 5- or 6- segmented, with or without annulations; widespread.......................................................................... 2
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<keyLead id="30DC95E5875B9632A2A1FE26EAA1FDE1" pageId="46" pageNumber="47">
2
<emphasis id="B959EAC2875B9632A2D7FE26EE38FE7E" bold="true" box="[225,258,415,437]" pageId="46" pageNumber="47">AF</emphasis>
antenna with 38 setae; plates distinctive, consisting of a dorsoventrally thickened portion that appears bifurcate or trifurcate and a thinner marginal portion stretched between the tines; without fringing on plates and without gland spines; without pores near posterior spiracle; first 3 pygidial lobes similar in shape, unilobular, without notches; dorsal ducts very thin, resembling ventral microducts;
<emphasis id="B959EAC2875B9632A3EBFE4EEECEFDC6" bold="true" box="[477,500,503,525]" pageId="46" pageNumber="47">M</emphasis>
diploid, with half of the chromosomes heterochromatic; circumtropical, extending to temperate regions of
<collectingCountry id="F33A7640875B9632A371FDACEEF8FDE1" box="[327,450,533,554]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
and
<collectingCountry id="F33A7640875B9632A3C7FDACED76FDE1" box="[497,588,533,554]" name="Australia" pageId="46" pageNumber="47">Australia</collectingCountry>
, and in greenhouses...................................... FURCASPIDINAE
</keyLead>
</paragraph>
<paragraph id="8B9236D0875B9632A2A1FD8BEAA1FD54" blockId="46.[151,1436,151,1170]" pageId="46" pageNumber="47">
<keyLead id="30DC95E5875B9632A2A1FD8BEAA1FD54" pageId="46" pageNumber="47">
-
<emphasis id="B959EAC2875B9632A2D7FD8BEE38FD83" bold="true" box="[225,258,562,584]" pageId="46" pageNumber="47">AF</emphasis>
antenna unisetose or (less commonly) multisetose; plates, when present, of uniform thickness and often fringed; with or without gland spines; with or without pores near posterior spiracle; lobes various; ducts various, dorsal ducts often wider than ventral microducts;
<emphasis id="B959EAC2875B9632A395FDD5EE80FD49" bold="true" box="[419,442,620,642]" pageId="46" pageNumber="47">M</emphasis>
haploid, without heterochromatic chromosomes or (less commonly) diploid, with half of the chromosomes heterochromatic; widespread....................................................................... 3
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<paragraph id="8B9236D0875B9632A2A1FD1EEAA6FC6C" blockId="46.[151,1436,151,1170]" pageId="46" pageNumber="47">
<keyLead id="30DC95E5875B9632A2A1FD1EEAA6FC6C" pageId="46" pageNumber="47">
3
<emphasis id="B959EAC2875B9632A2D7FD1EEE38FD76" bold="true" box="[225,258,679,701]" pageId="46" pageNumber="47">AF</emphasis>
usually with fringed plates and without gland spines; L2 and L3, when present, unilobular; ducts 1- or 2-barred; orifices of marginal macroducts, if oblong, with long axis usually parallel to body margin; pores by anterior spiracle present or absent, when present usually 5-locular;
<emphasis id="B959EAC2875B9632A020FD5BED0AFD33" bold="true" box="[534,560,738,760]" pageId="46" pageNumber="47">2F</emphasis>
usually with fringed plates; L2 unilobular;
<emphasis id="B959EAC2875B9632A1E2FD5BECCDFD33" bold="true" box="[980,1015,738,760]" pageId="46" pageNumber="47">CR</emphasis>
abdominal segment II with submedian dorsal duct; abdominal segments IIIVI without dorsal submedian setae; abdominal segments IVVI each without ventral submedian seta; head and mesothorax each with submarginal or submedial dorsal duct; terminal segment of antenna with 2 apical setae; tarsus with seta near base of claw; tarsus and tibia fused or separated by a septum;
<emphasis id="B959EAC2875B9632A661FC83EB45FC9B" bold="true" box="[1111,1151,826,848]" pageId="46" pageNumber="47">AM</emphasis>
midcranial ridge with dorsal branches reduced; tubercular ocelli present; pronotal ridges present; abdomen without dorsal setae on segments IVVII;
<emphasis id="B959EAC2875B9632A7B3FCEEEAA6FCA6" bold="true" box="[1413,1436,855,877]" pageId="46" pageNumber="47">M</emphasis>
haploid without heterochromatic chromosomes or diploid with half of the chromosomes heterochromatic; widespread.............................................................................................. ASPIDIOTINAE
</keyLead>
</paragraph>
<paragraph id="8B9236D0875B9632A2A1FC16EAA6FB59" blockId="46.[151,1436,151,1170]" pageId="46" pageNumber="47">
<keyLead id="30DC95E5875B9632A2A1FC16EAA6FB59" pageId="46" pageNumber="47">
-
<emphasis id="B959EAC2875B9632A2D7FC16EE38FC0E" bold="true" box="[225,258,943,965]" pageId="46" pageNumber="47">AF</emphasis>
usually without fringed plates and with gland spines; L2 usually bilobular, L3 bilobular or unilobular; ducts usually 2- barred; orifices of marginal macroducts, if oblong, with long axis oblique or perpendicular to body margin; pores by anterior spiracle usually present, usually 3-locular;
<emphasis id="B959EAC2875B9632A0B6FC53EDA0FBCB" bold="true" box="[640,666,1002,1024]" pageId="46" pageNumber="47">2F</emphasis>
usually without fringed plates; L2 usually bilobular;
<emphasis id="B959EAC2875B9632A696FC53EBF9FBCB" bold="true" box="[1184,1219,1002,1024]" pageId="46" pageNumber="47">CR</emphasis>
abdominal segment II without submedian dorsal duct; abdominal segments IIIVI each with dorsal submedian seta; abdominal segments IVVI each with ventral submedian seta; head and mesothorax without submarginal or submedial dorsal ducts; terminal segment of antenna with 1 apical seta; tarsus with or without seta near base of claw; tarsus and tibia separated by a septum;
<emphasis id="B959EAC2875B9632A73EFBFBEA0AFB93" bold="true" box="[1288,1328,1090,1112]" pageId="46" pageNumber="47">AM</emphasis>
midcranial ridge with dorsal branches present; tubercular ocelli absent; pronotal ridges absent; abdomen with dorsal setae on segments IVVII;
<emphasis id="B959EAC2875B9632A302FBC5EE71FB59" bold="true" box="[308,331,1148,1170]" pageId="46" pageNumber="47">M</emphasis>
haploid, without heterochromatic chromosomes; widespread.............................. DIASPIDINAE
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</paragraph>
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</subSubSection>
</treatment>
</document>