treatments-xml/data/03/84/87/038487C2EE123246FF69FD6BFD74F937.xml
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<document id="E907D5AE4FF7B499966DA9FD7208579B" ID-DOI="10.1649/0010-065X-74.1.175" ID-GBIF-Dataset="28046c1b-405b-4b6e-b291-e88a770321e5" ID-ISSN="1938-4394" ID-Zenodo-Dep="3746907" ID-ZooBank="urn:lsid:zoobank.org:pub:F36FA2BB-C7F0-4A34-972A-C34572F9ED94" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586467962798" checkinUser="felipe" docAuthor="Powell, Gareth S." docDate="2020" docId="038487C2EE123246FF69FD6BFD74F937" docLanguage="en" docName="TheColeopteristsBulletin.74.1.175-180.pdf.imf" docOrigin="The Coleopterists Bulletin 74 (1)" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Carpophilus mellarius Powell 2020, new species" docType="treatment" docVersion="5" lastPageNumber="178" masterDocId="FFBDFFBAEE163242FFDDFFC1FF85FFF6" masterDocTitle="Four New Species Of Carpophilus (Ecnomorphus) Motschulsky (Coleoptera: Nitidulidae: Carpophilinae) From The New World" masterLastPageNumber="180" masterPageNumber="175" pageNumber="178" updateTime="1698736860956" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="727E47A92A8B86B5C0AF30815A21BB8C">Four New Species Of Carpophilus (Ecnomorphus) Motschulsky (Coleoptera: Nitidulidae: Carpophilinae) From The New World</mods:title>
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<mods:namePart id="888400E75052156105BB43279D1144B7">Powell, Gareth S.</mods:namePart>
<mods:affiliation id="AFC589663E6B6585B632C2AB4CE1A74A">Department of Biology Brigham Young University</mods:affiliation>
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<mods:title id="2C1A5FC93D946F661DFA57D67FDDDC8E">The Coleopterists Bulletin</mods:title>
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<mods:date id="01E7B30F84D32D8FEFD86956C1F2F1BA">2020</mods:date>
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<taxonomicName id="4C2D4D57EE123246FF69FD6BFE5BFD36" authority="Powell" authorityName="Powell" authorityYear="2020" box="[180,478,682,704]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius" status="sp. nov.">
<emphasis id="B959EAC6EE123246FF69FD6BFE16FD49" bold="true" box="[180,403,682,703]" italics="true" pageId="4" pageNumber="178">Carpophilus mellarius</emphasis>
Powell
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,
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Zoobank.org/
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<figureCitation id="13162A51EE123246FE91FD3BFE24FCF9" box="[332,417,762,783]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="4" pageNumber="178">Figs. 1B</figureCitation>
,
<figureCitation id="13162A51EE123246FE70FD3BFE4CFCF9" box="[429,457,762,783]" captionStart="Fig" captionStartId="3.[197,228,1687,1706]" captionTargetBox="[177,1188,993,1668]" captionTargetId="figure@3.[175,1190,991,1670]" captionTargetPageId="3" captionText="Fig. 2. Carpophilus (Ecnomorphus) species, lateral view of male genitalia. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. marginatus, E) C. discoideus, F) C. brachypterus." figureDoi="http://doi.org/10.5281/zenodo.3746911" httpUri="https://zenodo.org/record/3746911/files/figure.png" pageId="4" pageNumber="178">2B</figureCitation>
)
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<emphasis id="B959EAC6EE123246FF72FCE4FECDFCCC" bold="true" box="[175,328,805,826]" pageId="4" pageNumber="178">Type Material.</emphasis>
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male
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labeled “
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:
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; Mpio: | Coappilla,
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</quantity>
. NW Jet Rds. | Tapalapa/Ocotepec
<quantity id="4CD59B31EE123246FE41FC9BFE64FC99" box="[412,481,858,879]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="4" pageNumber="178" unit="m" value="1960.0">
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.
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,
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<collectingDate id="EFD7E9FCEE123246FEF7FCB5FE15FC7F" box="[298,400,884,905]" pageId="4" pageNumber="178" value="2008-05-26">26-V-2008</collectingDate>
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<collectorName id="26D85302EE123246FE46FCB5FD92FC7F" box="[411,535,884,905]" pageId="4" pageNumber="178">R. Anderson</collectorName>
| dry oak forest litter, 2008-031 || HOLOTYPE: |
<taxonomicName id="4C2D4D57EE123246FDDAFC4FFDFBFC52" authorityName="Stephens" authorityYear="1829" box="[519,638,910,932]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAC6EE123246FDDAFC4FFDFBFC52" box="[519,638,910,932]" italics="true" pageId="4" pageNumber="178">Carpophilus</emphasis>
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|
<taxonomicName id="4C2D4D57EE123246FF7CFC68FF7FFC49" authorityName="Powell" authorityYear="2020" box="[161,250,937,959]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius">
<emphasis id="B959EAC6EE123246FF7CFC68FF7FFC49" box="[161,250,937,959]" italics="true" pageId="4" pageNumber="178">mellarius</emphasis>
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| Des.
<bibRefCitation id="EFBC4B25EE123246FEE4FC6BFE35FC49" author="Powell" box="[313,432,938,959]" firstAuthor="Powell" pageId="4" pageNumber="178" pagination="297 - 300" refId="ref4716" refString="Powell, G. S., and G. J. Martin. 2019. Type designations for sap beetles in the subfamily Carpophilinae Erichson (Coleoptera: Nitidulidae) housed in the Natural History Museum, London. Zootaxa 4590 (2): 297 - 300." type="journal article" year="2019">Powell 2019</bibRefCitation>
” (deposited in
<collectionCode id="ED3CAE11EE123246FDE3FC6BFDF1FC49" box="[574,628,938,959]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="4" pageNumber="178" type="University or college">BYU</collectionCode>
)
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.
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<typeStatus id="54968876EE123246FF4AFC05FF79FC2F" box="[151,252,964,985]" pageId="4" pageNumber="178" type="paratype">Paratypes</typeStatus>
(6):
</emphasis>
<collectingCountry id="F33A7644EE123246FE9BFC05FE17FC2F" box="[326,402,964,985]" name="Mexico" pageId="4" pageNumber="178">Mexico</collectingCountry>
:
<collectingRegion id="49E9F836EE123246FE7EFC05FE77FC2F" box="[419,498,964,985]" country="Mexico" name="Chiapas" pageId="4" pageNumber="178">Chiapas</collectingRegion>
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;
<materialsCitation id="3B453C89EE123246FDDEFC05FDFAFBB2" ID-GBIF-Occurrence="2597530328" collectingDate="2008-05-14" collectingDateMax="2008-05-25" collectingDateMin="2008-05-14" collectionCode="CSCA" country="Mexico" county="Mpio" elevation="1776" latitude="17.18273" location="Corso" longLatPrecision="1" longitude="-93.15168" municipality="Villa" pageId="4" pageNumber="178" specimenCode="LLAMA08" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<collectingCounty id="62F34E58EE123246FDDEFC05FDBFFC2F" box="[515,570,964,985]" pageId="4" pageNumber="178">Mpio</collectingCounty>
:
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|
<location id="8EF2600FEE123246FF4AFC1EFF57FC02" LSID="urn:lsid:plazi:treatment:038487C2EE123246FF69FD6BFD74F937:8EF2600FEE123246FF4AFC1EFF57FC02" box="[151,210,991,1012]" country="Mexico" county="Mpio" latitude="17.18273" longLatPrecision="1" longitude="-93.15168" municipality="Villa" name="Corso" pageId="4" pageNumber="178" stateProvince="Chiapas">Corso</location>
,
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, |
<location id="8EF2600FEE123246FE3EFC1EFE05FBF8" LSID="urn:lsid:plazi:treatment:038487C2EE123246FF69FD6BFD74F937:8EF2600FEE123246FE3EFC1EFE05FBF8" country="Mexico" county="Mpio" latitude="17.18273" longLatPrecision="1" longitude="-93.15168" municipality="Villa" name="R. Biosfera La Sepaltura Cerro Bola" pageId="4" pageNumber="178" stateProvince="Chiapas">R. Biosfera La Sepaltura Cerro Bola</location>
| base
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||
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<collectingDate id="EFD7E9FCEE123246FEE8FBD5FE1EFBDF" box="[309,411,1044,1065]" pageId="4" pageNumber="178" value="2008-05-14">14-V-2008</collectingDate>
</date>
R. Anderson,
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| dry oak forest litter, 2008-006” (1,
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).
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:
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: |
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NNW Coapilla
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|
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|
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,
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<collectingDate id="EFD7E9FCEE123246FE37FBA5FDD5FB8F" box="[490,592,1124,1145]" pageId="4" pageNumber="178" value="2008-05-25">25-V-2008</collectingDate>
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| ex.
<location id="8EF2600FEE123246FF4AFBBFFF4AFB65" LSID="urn:lsid:plazi:treatment:038487C2EE123246FF69FD6BFD74F937:8EF2600FEE123246FF4AFBBFFF4AFB65" box="[151,207,1150,1171]" country="Mexico" county="Mpio" latitude="17.18273" longLatPrecision="1" longitude="-93.15168" municipality="Villa" name="Sifted" pageId="4" pageNumber="178" stateProvince="Chiapas">Sifted</location>
leaf litter, | 2° mesophil forest |
<specimenCode id="DB8B9EAFEE123246FDCCFBBFFDF8FB65" box="[529,637,1150,1171]" collectionCode="LLAMA" pageId="4" pageNumber="178">LLAMA08</specimenCode>
Ma-A-04-2-all” (1,
<collectionCode id="ED3CAE11EE123246FE82FB58FE1AFB58" box="[351,415,1177,1198]" pageId="4" pageNumber="178">GSPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B453C89EE123246FE69FB59FF66FAEE" ID-GBIF-Occurrence="2597530330" collectingDate="2008-05-25" collectingDateMax="2008-05-25" collectingDateMin="2008-05-14" collectionCode="SEMC" country="Mexico" county="Mpio" elevation="1915" latitude="17.18273" location="Chiapas" longLatPrecision="1" longitude="-93.15168" municipality="Villa" pageId="4" pageNumber="178" specimenCode="LLAMA08" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<collectingCountry id="F33A7644EE123246FE62FB58FD8EFB58" box="[447,523,1177,1198]" name="Mexico" pageId="4" pageNumber="178">Mexico</collectingCountry>
:
<collectingRegion id="49E9F836EE123246FDC7FB58FDEFFB58" box="[538,618,1177,1198]" country="Mexico" name="Chiapas" pageId="4" pageNumber="178">Chiapas</collectingRegion>
: |
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<quantity id="4CD59B31EE123246FF4AFB72FF47FB3E" box="[151,194,1203,1224]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="178" unit="km" value="5.0">5km</quantity>
NNW Coapilla
</locationDeviation>
|
<geoCoordinate id="EE195013EE123246FEAAFB72FE69FB3E" box="[375,492,1203,1224]" degrees="17.18273" direction="north" orientation="latitude" pageId="4" pageNumber="178" precision="1" value="17.18273">17.18273°N</geoCoordinate>
<geoCoordinate id="EE195013EE123246FE28FB72FDEAFB3E" box="[501,623,1203,1224]" degrees="93.15168" direction="west" orientation="longitude" pageId="4" pageNumber="178" precision="1" value="-93.15168">93.15168°W</geoCoordinate>
|
<quantity id="4CD59B31EE123246FF4AFB0FFF59FB15" box="[151,220,1230,1251]" metricMagnitude="3" metricUnit="m" metricValue="1.9149999999999998" pageId="4" pageNumber="178" unit="m" value="1915.0">
<elevation id="0000D1E7EE123246FF4AFB0FFF59FB15" box="[151,220,1230,1251]" metricMagnitude="3" metricUnit="m" metricValue="1.9149999999999998" pageId="4" pageNumber="178" unit="m" value="1915.0">1915m</elevation>
</quantity>
,
<date id="FF931014EE123246FF33FB0FFED6FB15" box="[238,339,1230,1251]" pageId="4" pageNumber="178" value="2008-05-25">
<collectingDate id="EFD7E9FCEE123246FF33FB0FFED6FB15" box="[238,339,1230,1251]" pageId="4" pageNumber="178" value="2008-05-25">25-V-2008</collectingDate>
</date>
| ex. Sifted leaf litter, | 2° mesophil forest |
<specimenCode id="DB8B9EAFEE123246FE96FB29FE32FB0B" box="[331,439,1256,1277]" collectionCode="LLAMA" pageId="4" pageNumber="178">LLAMA08</specimenCode>
Ma-A-04-2-all” (4,
<collectionCode id="ED3CAE11EE123246FF4AFAC2FF59FAEE" box="[151,220,1283,1304]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="178" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE123246FF72FADFFE78F9CA" pageId="4" pageNumber="178" type="diagnosis">
<paragraph id="8B9236D4EE123246FF72FADFFE78F9CA" blockId="4.[151,640,804,1730]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FF72FADFFE92FAC5" bold="true" box="[175,279,1310,1331]" pageId="4" pageNumber="178">Diagnosis.</emphasis>
<taxonomicName id="4C2D4D57EE123246FEF7FADCFD89FAC5" authorityName="Powell" authorityYear="2020" box="[298,524,1309,1331]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius">
<emphasis id="B959EAC6EE123246FEF7FADCFD89FAC5" box="[298,524,1309,1331]" italics="true" pageId="4" pageNumber="178">Carpophilus mellarius</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C2D4D57EE123246FF4AFAF6FE67FABB" authority="LeConte" authorityName="LeConte" authorityYear="1858" box="[151,482,1335,1357]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B959EAC6EE123246FF4AFAF6FE04FABB" box="[151,385,1335,1357]" italics="true" pageId="4" pageNumber="178">Carpophilus discoideus</emphasis>
LeConte
</taxonomicName>
(
<figureCitation id="13162A51EE123246FE2AFAF9FDCCFABB" box="[503,585,1336,1357]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="4" pageNumber="178">Fig. 1G</figureCitation>
) but differs in the following ways:
<taxonomicName id="4C2D4D57EE123246FE1DFA93FDB2FA9E" authorityName="Powell" authorityYear="2020" box="[448,567,1362,1384]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius">
<emphasis id="B959EAC6EE123246FE1DFA93FDB2FA9E" box="[448,567,1362,1384]" italics="true" pageId="4" pageNumber="178">C. mellarius</emphasis>
</taxonomicName>
is unicolorous, while
<taxonomicName id="4C2D4D57EE123246FEEFFAADFE36FA74" authorityName="LeConte" authorityYear="1858" box="[306,435,1388,1410]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B959EAC6EE123246FEEFFAADFE36FA74" box="[306,435,1388,1410]" italics="true" pageId="4" pageNumber="178">C. discoideus</emphasis>
</taxonomicName>
has distinctly lighter patches in the center of each elytron, surrounded by a much darker habitus; the scutellar shield in
<taxonomicName id="4C2D4D57EE123246FDB5FA63FF75FA24" authorityName="Powell" authorityYear="2020" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius">
<emphasis id="B959EAC6EE123246FDB5FA63FF75FA24" italics="true" pageId="4" pageNumber="178">C. mellarius</emphasis>
</taxonomicName>
is much shorter than in
<taxonomicName id="4C2D4D57EE123246FE08FA7DFDD1FA24" authorityName="LeConte" authorityYear="1858" box="[469,596,1468,1490]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B959EAC6EE123246FE08FA7DFDD1FA24" box="[469,596,1468,1490]" italics="true" pageId="4" pageNumber="178">C. discoideus</emphasis>
</taxonomicName>
; the elytral apices are clearly fimbriate in
<taxonomicName id="4C2D4D57EE123246FE20FA16FDF8FA1B" authorityName="LeConte" authorityYear="1858" box="[509,637,1495,1517]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B959EAC6EE123246FE20FA16FDF8FA1B" box="[509,637,1495,1517]" italics="true" pageId="4" pageNumber="178">C. discoideus</emphasis>
</taxonomicName>
but bare in
<taxonomicName id="4C2D4D57EE123246FF23FA30FEF7F9F1" authorityName="Powell" authorityYear="2020" box="[254,370,1521,1543]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="mellarius">
<emphasis id="B959EAC6EE123246FF23FA30FEF7F9F1" box="[254,370,1521,1543]" italics="true" pageId="4" pageNumber="178">C. mellarius</emphasis>
</taxonomicName>
. The arrow-shaped tip of the tegmen is also diagnostic in comparison with other members in this group (
<figureCitation id="13162A51EE123246FE5DF9E6FE74F9CA" box="[384,497,1575,1596]" captionStart="Fig" captionStartId="3.[197,228,1687,1706]" captionTargetBox="[177,1188,993,1668]" captionTargetId="figure@3.[175,1190,991,1670]" captionTargetPageId="3" captionText="Fig. 2. Carpophilus (Ecnomorphus) species, lateral view of male genitalia. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. marginatus, E) C. discoideus, F) C. brachypterus." figureDoi="http://doi.org/10.5281/zenodo.3746911" httpUri="https://zenodo.org/record/3746911/files/figure.png" pageId="4" pageNumber="178">Figs. 2B, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE123246FF72F982FC4EF9F1" pageId="4" pageNumber="178" type="description">
<paragraph id="8B9236D4EE123246FF72F982FCEFFA24" blockId="4.[151,640,804,1730]" lastBlockId="4.[685,1172,194,1729]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FF72F982FEAFF9AE" bold="true" box="[175,298,1603,1624]" pageId="4" pageNumber="178">Description.</emphasis>
Body parallel-sided, weakly dorsoventrally flattened. Length
<quantity id="4CD59B31EE123246FE50F99CFE64F984" box="[397,481,1629,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="4" pageNumber="178" unit="mm" value="1.36">1.36 mm</quantity>
, width
<quantity id="4CD59B31EE123246FDFBF99CFDFFF984" box="[550,634,1629,1650]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="4" pageNumber="178" unit="mm" value="0.59">0.59 mm</quantity>
, color light brown (
<figureCitation id="13162A51EE123246FE8BF9B9FE26F97B" box="[342,419,1656,1677]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="4" pageNumber="178">Fig. 1B</figureCitation>
). Ventral surface light brown. Dorsal sculpturing moderately shiny; dorsum with fine, golden pubescence.
<emphasis id="B959EAC6EE123246FE69F96CFE6AF934" bold="true" box="[436,495,1709,1730]" pageId="4" pageNumber="178">Head:</emphasis>
Narrower than pronotum, finely punctured. Punctures uniform in size, becoming sparser at midline. Frontoclypeal region truncate, labrum deeply emarginate, mandibles toothed, light brown. Palpi fusiform, reaching about 2/3 length of mandibles. Eyes small, finely faceted, interocular distance
<quantity id="4CD59B31EE123246FD22FE87FCD6FEAD" box="[767,851,326,347]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="178" unit="mm" value="0.29">0.29 mm</quantity>
(
<emphasis id="B959EAC6EE123246FCBDFE86FCE9FEAA" box="[864,876,327,348]" italics="true" pageId="4" pageNumber="178">n</emphasis>
= 2). Antenna 1.5× longer than head; antennomere 1 robust and curved, antennomere 2 as long as basal antennomere, antennomeres 2 and 3 subequal in length, antennomeres 28 expanding apically, with each apical expansion conspicuously darker in coloration, antennomeres 48 each about half length of antennomere 2, antennomeres 911 forming strong, compact club, slightly longer than wide, space between antennomeres 9 and 10 larger than that of 10 and 11.
<emphasis id="B959EAC6EE123246FCA6FDF4FC63FDBC" bold="true" box="[891,998,565,586]" pageId="4" pageNumber="178">Pronotum:</emphasis>
1.95 times as wide as long, sides weakly arcuate with anterior third increasingly rounded, anterior angles and posterior angles broadly obtuse. Posterior margin with welldeveloped marginal line, particularly at middle. Disc flattened, moderately shiny, evenly punctured except at midline.
<emphasis id="B959EAC6EE123246FCFBFD14FC48FD1C" bold="true" box="[806,973,725,746]" pageId="4" pageNumber="178">Scutellar shield:</emphasis>
Feebly pentagonal, reaching obtuse point posteriorly; anteriorly finely granulate, gradually becoming glabrous.
<emphasis id="B959EAC6EE123246FB90FCCBFB16FCE9" bold="true" box="[1101,1171,778,799]" pageId="4" pageNumber="178">Elytra:</emphasis>
Slightly wider than pronotum, quadrate, exactly as long as wide. Humeri not pronounced. Anterior angles almost at right angles, sides mildly arcuate, posterior angles slightly acute, apices truncate, feebly converging anteriorly.
<emphasis id="B959EAC6EE123246FC5FFC4EFC6CFC52" bold="true" box="[898,1001,911,932]" pageId="4" pageNumber="178">Abdomen:</emphasis>
Two tergites dorsally visible, golden brown setae denser than on rest of dorsum. Pygidium rounded with distinct, evenly spaced punctures.
<emphasis id="B959EAC6EE123246FCB1FC1EFC32FC02" bold="true" box="[876,951,991,1012]" pageId="4" pageNumber="178">Venter:</emphasis>
Overall lighter, submentum narrow and transverse, antennal grooves well-developed. Prosternum coarsely punctate, punctures with fine golden setae, process slightly convex in lateral profile, apically rounded. Mesothoracic ventrite finely punctured. Metathoracic ventrite setose, finely punctate throughout. Abdominal ventrite 1 almost as long as metathoracic ventrite, abdominal ventrites 23 small, finely, sparsely punctate, ventrites 4 and 5 large, granulate, more densely pubescent.
<emphasis id="B959EAC6EE123246FC70FB29FC66FB0B" bold="true" box="[941,995,1256,1277]" pageId="4" pageNumber="178">Legs:</emphasis>
Somewhat short, femora robust, covered with fine golden pubescence. Tibiae expanded apically, with 57 apical spines.
<emphasis id="B959EAC6EE123246FD20FAF9FC19FABB" bold="true" box="[765,924,1336,1357]" pageId="4" pageNumber="178">Male genitalia:</emphasis>
Well-sclerotized; lateral lobes parallel in dorsal view, forming elongate opening; in lateral view, lobes curved to sharp point, both inner and outer margins curved (
<figureCitation id="13162A51EE123246FBCEFA49FBD8FA6B" box="[1043,1117,1416,1437]" captionStart="Fig" captionStartId="3.[197,228,1687,1706]" captionTargetBox="[177,1188,993,1668]" captionTargetId="figure@3.[175,1190,991,1670]" captionTargetPageId="3" captionText="Fig. 2. Carpophilus (Ecnomorphus) species, lateral view of male genitalia. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. marginatus, E) C. discoideus, F) C. brachypterus." figureDoi="http://doi.org/10.5281/zenodo.3746911" httpUri="https://zenodo.org/record/3746911/files/figure.png" pageId="4" pageNumber="178">Fig. 2B</figureCitation>
), row of long setae directed inward, long sparse setae at arrow-shaped apex.
</paragraph>
<paragraph id="8B9236D4EE123246FD18FA16FC4EF9F1" blockId="4.[685,1172,194,1729]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FD18FA16FCAEFA1A" bold="true" box="[709,811,1495,1516]" pageId="4" pageNumber="178">Variation.</emphasis>
The metatibiae are less expanded apically in female specimens.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE123246FD18F9CDFC50F9CA" pageId="4" pageNumber="178" type="distribution">
<paragraph id="8B9236D4EE123246FD18F9CDFC50F9CA" blockId="4.[685,1172,194,1729]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FD18F9CDFC4FF9D7" bold="true" box="[709,970,1548,1569]" pageId="4" pageNumber="178">Geographic Distribution.</emphasis>
The
<typeStatus id="54968876EE123246FBD6F9CDFBB1F9D7" box="[1035,1076,1548,1569]" pageId="4" pageNumber="178">type</typeStatus>
series is known from
<collectingRegion id="49E9F836EE123246FCF1F9E6FCFEF9CA" box="[812,891,1575,1596]" country="Mexico" name="Chiapas" pageId="4" pageNumber="178">Chiapas</collectingRegion>
,
<collectingCountry id="F33A7644EE123246FC5BF9E6FC57F9CA" box="[902,978,1575,1596]" name="Mexico" pageId="4" pageNumber="178">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE123246FD18F983FB0BF987" pageId="4" pageNumber="178" type="biology_ecology">
<paragraph id="8B9236D4EE123246FD18F983FB0BF987" blockId="4.[685,1172,194,1729]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FD18F983FC92F9A1" bold="true" box="[709,791,1602,1623]" pageId="4" pageNumber="178">Biology.</emphasis>
All specimens studied were captured in leaf litter; no obvious feeding behavior is known.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE123246FD18F9B6FD74F937" pageId="4" pageNumber="178" type="etymology">
<paragraph id="8B9236D4EE123246FD18F9B6FD74F937" blockId="4.[685,1172,194,1729]" pageId="4" pageNumber="178">
<emphasis id="B959EAC6EE123246FD18F9B6FCBDF97A" bold="true" box="[709,824,1655,1676]" pageId="4" pageNumber="178">Etymology.</emphasis>
The specific epithet is Latin derived and refers to the overall coloration appearing honey brown.
</paragraph>
</subSubSection>
</treatment>
</document>