treatments-xml/data/03/84/87/038487AF1F3FFFDF116FC54E74BFF97C.xml

<document id="B6BB3834212E21A4907A3BB73F0EE437" ID-DOI="10.5281/zenodo.5401510" ID-ISSN="1638-9387" ID-Zenodo-Dep="5401510" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630334724433" checkinUser="marcus" docAuthor="Cumberlidge, Neil &amp; Sternberg, Richard V." docDate="2002" docId="038487AF1F3FFFDF116FC54E74BFF97C" docLanguage="en" docName="Zoosystema.2002.24.1.41-79.pdf" docOrigin="Zoosystema 24 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Marojejy longimerus Cumberlidge, Boyko &amp; Harvey 2002" docType="treatment" docVersion="3" lastPageNumber="73" masterDocId="FFBDFFD71F21FFFF120AC105702FFFF9" masterDocTitle="The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)" masterLastPageNumber="79" masterPageNumber="41" pageNumber="71" updateTime="1699232587643" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="584A64D5DECECCAF4CAF188AFCE815A2">The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)</mods:title>
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<mods:namePart id="F66EB4AB19BF34BE19BAD87C10AA3458">Cumberlidge, Neil</mods:namePart>
<mods:affiliation id="7BDACE52563B62AA68B81F2583918063">Department of Biology, Northern Michigan University, Marquette, MI, 49855 (USA) ncumberl @ nmu. edu</mods:affiliation>
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<mods:namePart id="77B052A6BB9519E26FF2E9254578A70E">Sternberg, Richard V.</mods:namePart>
<mods:affiliation id="D954FA455B16B5E280593B43DA8B5279">NHB- 163, Department of Invertebrate Zoology, Smithsonian Institution, Washington D. C., 20560 (USA) Sternberg. Richard @ nmnh. si. edu.</mods:affiliation>
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<mods:title id="1130E39226369A0742691B84D1F1D3D6">Zoosystema</mods:title>
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<mods:date id="A80C554D23FF093C8BABC9B24E28A747">2002</mods:date>
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<taxonomicName id="4C2D4D3A1F3FFFE1116FC54E748DFB7C" authority="Cumberlidge, Boyko &amp; Harvey, 2002" authorityName="Cumberlidge, Boyko &amp; Harvey" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="B959EAAB1F3FFFE1116FC54E746BFB9C" bold="true" box="[869,1092,1099,1125]" italics="true" pageId="30" pageNumber="71">Marojejy longimerus</emphasis>
Cumberlidge, Boyko &amp; Harvey, 2002
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</paragraph>
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<subSubSection id="C33765321F3FFFE110C0C58C74F0FB5A" box="[714,1247,1161,1187]" pageId="30" pageNumber="71" type="description">
<paragraph id="8B9236B91F3FFFE110C0C58C74F0FB5A" blockId="30.[714,1247,1099,1187]" box="[714,1247,1161,1187]" pageId="30" pageNumber="71">
(
<figureCitation id="13162A3C1F3FFFE110DBC58C730CFB5A" box="[721,803,1161,1187]" captionStart="FIG" captionStartId="11.[151,162,1354,1371]" captionTargetBox="[192,1245,168,1311]" captionTargetId="figure-26@11.[187,1212,162,1321]" captionTargetPageId="11" captionText="FIG. 2. — Dorsal view of the carapace of adult males of Madagascan freshwater crabs (A-C based on Ng &amp; Takeda 1994); A, Skeleosophusa gollhardi (Bott, 1965), holotype (MNHN B-5165); B, S. prolixa Ng &amp; Takeda, 1994, holotype, (MRSN); C, S. eumeces Ng &amp; Takeda, 1994, holotype, (MRSN); D, Boreas uglowi n. gen., n. sp., holotype (MNHN B- 11292); E, Madagapotamon humberti Bott, 1965; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002, holotype, (AMNH 17833). Scale bar: A, 7.2 mm; B, 7.1 mm; C, 6.5 mm; D, 9 mm; E, 9.4 mm; F, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401514" httpUri="https://zenodo.org/record/5401514/files/figure.png" pageId="30" pageNumber="71">Figs 2F</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE11127C58C737CFB5A" box="[813,851,1161,1187]" captionStart="FIG" captionStartId="14.[151,162,1356,1373]" captionTargetBox="[156,1239,168,1322]" captionTargetId="figure-22@14.[146,1253,158,1327]" captionTargetPageId="14" captionText="FIG. 3. — Inferior view of the sternum of adult males of Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb., G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Boreas uglowi n. gen., n. sp. Scale bar: A, 11.7 mm; B, 17.2 mm; C, 15.9 mm; D, 14.5 mm; E, 10.2 mm; F, 7 mm; G, 9.6 mm; H, 6.4 mm; I, 9.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5401516" httpUri="https://zenodo.org/record/5401516/files/figure.png" pageId="30" pageNumber="71">3H</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE11154C58C73ACFB5A" box="[862,899,1161,1187]" captionStart="FIG" captionStartId="16.[151,162,1071,1088]" captionTargetBox="[155,1241,171,1034]" captionTargetId="figure-282@16.[138,1257,160,1041]" captionTargetPageId="16" captionText="FIG. 4. — Left mandible of Madagascan freshwater crabs, frontal view showing the palp (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2000) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2000; H, Boreas uglowi n. gen., n. sp.; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Madagapotamon humberti Bott, 1965. Scale bar: A-I, K, L, 1.0 mm; J, 1.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5401518" httpUri="https://zenodo.org/record/5401518/files/figure.png" pageId="30" pageNumber="71">4G</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE11187C58C739DFB5A" box="[909,946,1161,1187]" captionStart="FIG" captionStartId="21.[151,162,1262,1279]" captionTargetBox="[248,1159,165,1192]" captionTargetId="figure-109@21.[228,1169,158,1197]" captionTargetPageId="21" captionText="FIG. 5. — Left third maxilliped of Madagascan freshwater crabs, frontal view (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.0 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401520" httpUri="https://zenodo.org/record/5401520/files/figure.png" pageId="30" pageNumber="71">5G</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE111B6C58C73CDFB5A" box="[956,994,1161,1187]" captionStart="FIG" captionStartId="23.[151,162,768,785]" captionTargetBox="[258,1160,169,728]" captionTargetId="figure-337@23.[233,1163,157,745]" captionTargetPageId="23" captionText="FIG. 6. — Superior view of the carpus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs (J, K based on Ng &amp; Takeda 1994); A Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.5 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401522" httpUri="https://zenodo.org/record/5401522/files/figure.png" pageId="30" pageNumber="71">6H</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE111E7C58C7400FB5A" box="[1005,1071,1161,1187]" captionStart="FIG" captionStartId="25.[151,162,1567,1584]" captionTargetBox="[244,1126,230,1497]" captionTargetId="figure-22@25.[232,1187,219,1521]" captionTargetPageId="25" captionText="FIG. 7. — Superior and inferior views of the merus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs; A, B, Hydrothelphusa agilis A. Milne-Edwards, 1872; C, D, H. madagascariensis (A. Milne-Edwards, 1872); E, F, H. bombetokensis (Rathbun, 1904); G, H, H. goudoti (H. Milne Edwards, 1853) n. comb.; I, J, Malagasya antongilensis (Rathbun, 1905) n. comb.; K, L, Madagapotamon humberti Bott, 1965; M, N, Skelosophusa gollhardi (Bott, 1965); O, P, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; Q, R, Boreas uglowi n. gen., n. sp. Scale bar: A-H, 5.0 mm; I-T, 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401524" httpUri="https://zenodo.org/record/5401524/files/figure.png" pageId="30" pageNumber="71">7O, P</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE11633C58C7477FB5A" box="[1081,1112,1161,1187]" captionStart="FIG" captionStartId="29.[151,162,1545,1562]" captionTargetBox="[151,1247,251,1506]" captionTargetId="figure-43@29.[153,1248,238,1515]" captionTargetPageId="29" captionText="FIG. 8. — Last walking leg (pereiopod 5) of adult males of Madagascan freshwater crabs (H based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Boreas uglowi n. gen., n. sp.; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; G, Madagapotamon humberti Bott, 1965; H, Skelosophusa prolixa Ng &amp; Takeda, 1994; I, Malagasya antongilensis (Rathbun, 1905) n. comb.; J, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb. Scale bar: A-D, 5.0 mm; E-L, 2.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401526" httpUri="https://zenodo.org/record/5401526/files/figure.png" pageId="30" pageNumber="71">8F</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE11669C58C748FFB5A" box="[1123,1184,1161,1187]" captionStart="FIG" captionStartId="32.[151,162,1363,1380]" captionTargetBox="[149,1247,169,1337]" captionTargetId="figure-94@32.[141,1252,160,1342]" captionTargetPageId="32" captionText="FIG. 10. — Right first gonopod (dorsal and ventral views) of adult males of Madagascan freshwater crabs; A, B, Boreas uglowi n. gen., n. sp.; C, D, Malagasya antongilensis (Rathbun, 1905) n. comb.; E, F, Skeleosophusa gollhardi (Bott, 1965); G, H, Madagapotamon humberti Bott, 1965; I, J, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002. Scale bar: 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401530" httpUri="https://zenodo.org/record/5401530/files/figure.png" pageId="30" pageNumber="71">10I, J</figureCitation>
; 
<figureCitation id="13162A3C1F3FFFE116A1C58C74F8FB5A" box="[1195,1239,1161,1187]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="30" pageNumber="71">11E</figureCitation>
)
</paragraph>
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<paragraph id="8B9236B91F3FFFE110C0C5C2738FFB00" blockId="30.[714,1248,1223,1669]" pageId="30" pageNumber="71">
<taxonomicName id="4C2D4D3A1F3FFFE110C0C5C27369FB00" authority="Cumberlidge, Boyko &amp; Harvey, 2002: 67 - 71" authorityName="Cumberlidge, Boyko &amp; Harvey" authorityPageNumber="67 - 71" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="B959EAAB1F3FFFE110C0C5C273A0FB19" box="[714,911,1223,1248]" italics="true" pageId="30" pageNumber="71">Marojejy longimerus</emphasis>
Cumberlidge, Boyko &amp; Harvey, 2002: 67-71
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, figs 1-3.
</paragraph>
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<paragraph id="8B9236B91F3FFFE110C0C40C740FF97C" blockId="30.[714,1248,1223,1669]" pageId="30" pageNumber="71">
MATERIAL EXAMINED. — 
<emphasis id="B959EAAB1F3FFFE111E3C40C745BFAD8" bold="true" box="[1001,1140,1289,1313]" pageId="30" pageNumber="71">
<collectingCountry id="F33A76291F3FFFE111E3C40C7440FAD8" box="[1001,1135,1289,1313]" name="Madagascar" pageId="30" pageNumber="71">Madagascar</collectingCountry>
.
</emphasis>
Province d’Antsiranana, Réserve naturelle intégrale de 
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river, 
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northwest of Manantenina, 
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cw 23, cl 15.1, ch 7.45, fw 
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17833), 
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, 
<specimenCount id="9D2BFD301F3FFFE116AFC4A774CFFA43" box="[1189,1248,1442,1466]" pageId="30" pageNumber="71" type="generic">1 juv.</specimenCount>
, 
<typeStatus id="5496881B1F3FFFE110F8C4BF7365FA2B" box="[754,842,1466,1490]" pageId="30" pageNumber="71" type="paratype">paratype</typeStatus>
cw 11.1, cl 8.4, ch 4.5, fw 
<quantity id="4CD59B5C1F3FFFE11680C4BE74F0FA28" box="[1162,1247,1466,1491]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="30" pageNumber="71" unit="mm" value="3.1">3.1 mm</quantity>
(
<collectionCode id="ED3CAE7C1F3FFFE110DBC4D07307FA14" box="[721,808,1493,1517]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="30" pageNumber="71">AMNH</collectionCode>
17833). — 
<quantity id="4CD59B5C1F3FFFE111D1C4D07401FA14" box="[987,1070,1493,1517]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="30" pageNumber="71" unit="km" value="11.0">11 km</quantity>
northwest of Manantenina, Province d’Antsiranana, Réserve naturelle intégrale de 
<taxonomicName id="4C2D4D3A1F3FFFE111E9C7027462F9E6" box="[995,1101,1543,1567]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
, 
<geoCoordinate id="EE19507E1F3FFFE11669C70274F5F9E6" box="[1123,1242,1543,1567]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="30" pageNumber="71" precision="15" seconds="8" value="-14.435555">14°26’8”S</geoCoordinate>
, 
<geoCoordinate id="EE19507E1F3FFFE110C0C724731CF9C0" box="[714,819,1569,1593]" degrees="49" direction="east" minutes="44" orientation="longitude" pageId="30" pageNumber="71" precision="15" seconds="1" value="49.733612">49°44’1”E</geoCoordinate>
, source of Andranomifototra river, under rocks, 
<quantity id="4CD59B5C1F3FFFE1111AC73F734BF9A9" box="[784,868,1593,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="30" pageNumber="71" unit="m" value="1875.0">1875 m</quantity>
, 
<date id="FF9310791F3FFFE1117EC73F7424F9AB" box="[884,1035,1594,1618]" pageId="30" pageNumber="71" value="1996-11-13" valueMax="1996-11-19" valueMin="1996-11-13">13-19.XI.1996</date>
, coll. E. Quinter, 4, 
<typeStatus id="5496881B1F3FFFE11106C7577347F993" box="[780,872,1618,1642]" pageId="30" pageNumber="71" type="paratype">paratypes</typeStatus>
cl 13.0- 
<quantity id="4CD59B5C1F3FFFE111B5C7567431F990" box="[959,1054,1618,1643]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="30" pageNumber="71" unit="mm" value="14.7">14.7 mm</quantity>
, 5, 
<typeStatus id="5496881B1F3FFFE11689C75774F0F993" box="[1155,1247,1618,1642]" pageId="30" pageNumber="71" type="paratype">paratypes</typeStatus>
cl 
<quantity id="4CD59B5C1F3FFFE110EEC7687359F97A" box="[740,886,1644,1669]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.42" metricValueMin="1.22" pageId="30" pageNumber="71" unit="mm" value="13.2" valueMax="14.2" valueMin="12.2">12.2-14.2 mm</quantity>
(
<collectionCode id="ED3CAE7C1F3FFFE1118DC76873FAF97C" box="[903,981,1645,1669]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="30" pageNumber="71" type="Museum">FMNH</collectionCode>
4656).
</paragraph>
<caption id="DF5266311F3EFFE0129DC74C74E6F97C" ID-DOI="http://doi.org/10.5281/zenodo.5401528" ID-Zenodo-Dep="5401528" httpUri="https://zenodo.org/record/5401528/files/figure.png" pageId="31" pageNumber="72" startId="31.[151,162,1609,1626]" targetBox="[224,1179,165,1574]" targetPageId="31">
<paragraph id="8B9236B91F3EFFE0129DC74C74E6F97C" blockId="31.[151,1247,1609,1669]" pageId="31" pageNumber="72">
FIG. 9. — Right first gonopod (dorsal view; superior view; ventral view) and right second gonopod (dorsal view) of adult males of species of 
<taxonomicName id="4C2D4D3A1F3EFFE012FCC75A727CF989" authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" box="[246,595,1631,1648]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAAB1F3EFFE012FCC75A7154F989" box="[246,379,1631,1648]" italics="true" pageId="31" pageNumber="72">Hydrothelphusa</emphasis>
A. Milne-Edwards, 1872
</taxonomicName>
; 
<emphasis id="B959EAAB1F3EFFE01056C75A7251F989" bold="true" box="[604,638,1631,1648]" pageId="31" pageNumber="72">A -D</emphasis>
, 
<taxonomicName id="4C2D4D3A1F3EFFE01081C75A7388F989" authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" box="[651,935,1631,1648]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B959EAAB1F3EFFE01081C75A72E0F989" box="[651,719,1631,1648]" italics="true" pageId="31" pageNumber="72">H. agilis</emphasis>
A. Milne-Edwards, 1872
</taxonomicName>
; 
<emphasis id="B959EAAB1F3EFFE011BAC75A73E5F989" bold="true" box="[944,970,1631,1648]" pageId="31" pageNumber="72">E -I</emphasis>
, 
<taxonomicName id="4C2D4D3A1F3EFFE011DFC75A74A9F989" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1872" box="[981,1158,1631,1648]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B959EAAB1F3EFFE011DFC75A74A9F989" box="[981,1158,1631,1648]" italics="true" pageId="31" pageNumber="72">H. madagascariensis</emphasis>
</taxonomicName>
(A. Milne- 
<bibRefCitation id="EFBC4B481F3EFFE0129DC7717139F97C" author="EDWARDS A." box="[151,278,1652,1669]" pageId="31" pageNumber="72" pagination="1 - 3" refId="ref26376" refString="MILNE- EDWARDS A. 1872. - Note sur les Crabes d'eau douce de Madagascar. Bibliotheque de l'Ecole des hautes Etudes, Section Sciences naturelles 5 (8): 1 - 3." type="journal article" year="1872">Edwards, 1872</bibRefCitation>
); 
<emphasis id="B959EAAB1F3EFFE01329C7717169F97C" bold="true" box="[291,326,1652,1669]" pageId="31" pageNumber="72">J -M</emphasis>
, 
<taxonomicName id="4C2D4D3A1F3EFFE0135AC771725CF97C" authority="(Rathbun, 1904)" baseAuthorityName="Rathbun" baseAuthorityYear="1904" box="[336,627,1652,1669]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B959EAAB1F3EFFE0135AC77171C8F97C" box="[336,487,1652,1669]" italics="true" pageId="31" pageNumber="72">H. bombetokensis</emphasis>
(
<bibRefCitation id="EFBC4B481F3EFFE013FAC7717241F97C" author="RATHBUN M. J." box="[496,622,1652,1669]" pageId="31" pageNumber="72" pagination="255 - 312" refId="ref26930" refString="RATHBUN M. J. 1904. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 6 (4): 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
)
</taxonomicName>
; 
<emphasis id="B959EAAB1F3EFFE01071C77172B0F97C" bold="true" box="[635,671,1652,1669]" pageId="31" pageNumber="72">N -Q</emphasis>
, 
<taxonomicName id="4C2D4D3A1F3EFFE010A3C77173F9F97C" authority="(H. Milne Edwards, 1853)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[681,982,1652,1669]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="goudoti" status="comb. nov.">
<emphasis id="B959EAAB1F3EFFE010A3C771732FF97C" box="[681,768,1652,1669]" italics="true" pageId="31" pageNumber="72">H. goudoti</emphasis>
(H. 
<bibRefCitation id="EFBC4B481F3EFFE01128C77173FDF97C" author="MILNE EDWARDS H." box="[802,978,1652,1669]" pageId="31" pageNumber="72" pagination="20" refId="ref26442" refString="MILNE EDWARDS H. 1853. - Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences naturelles, Zoologie, Serie 3, 20: 163 - 28." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F3EFFE011D1C7717408F97C" box="[987,1063,1652,1669]" pageId="31" pageNumber="72" rank="species">n. comb.</taxonomicNameLabel>
Scale bar: 4.5 mm.
</paragraph>
</caption>
<caption id="DF5266311F01FFDF129DC456749BFA76" ID-DOI="http://doi.org/10.5281/zenodo.5401530" ID-Zenodo-Dep="5401530" httpUri="https://zenodo.org/record/5401530/files/figure.png" pageId="32" pageNumber="73" startId="32.[151,162,1363,1380]" targetBox="[149,1247,169,1337]" targetPageId="32">
<paragraph id="8B9236B91F01FFDF129DC456749BFA76" blockId="32.[151,1248,1363,1423]" pageId="32" pageNumber="73">
FIG. 10. — Right first gonopod (dorsal and ventral views) of adult males of Madagascan freshwater crabs; 
<emphasis id="B959EAAB1F01FFDF1639C456746FFA9D" bold="true" box="[1075,1088,1363,1380]" pageId="32" pageNumber="73">A</emphasis>
, 
<emphasis id="B959EAAB1F01FFDF1646C4567476FA9D" bold="true" box="[1100,1113,1363,1380]" pageId="32" pageNumber="73">B</emphasis>
, 
<taxonomicName id="4C2D4D3A1F01FFDF166FC45674F0FA9D" authority="Cumberlidge &amp; Sternberg, 2002" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[1125,1247,1363,1380]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="uglowi" status="n. gen., n. sp.">
<emphasis id="B959EAAB1F01FFDF166FC45674F0FA9D" box="[1125,1247,1363,1380]" italics="true" pageId="32" pageNumber="73">Boreas uglowi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F01FFDF129DC46D7131FA80" box="[151,286,1384,1401]" pageId="32" pageNumber="73" rank="species">n. gen., n. sp.</taxonomicNameLabel>
; 
<emphasis id="B959EAAB1F01FFDF1323C46D7118FA80" bold="true" box="[297,311,1384,1401]" pageId="32" pageNumber="73">C</emphasis>
, 
<emphasis id="B959EAAB1F01FFDF134FC46D717CFA80" bold="true" box="[325,339,1384,1401]" pageId="32" pageNumber="73">D</emphasis>
, 
<taxonomicName id="4C2D4D3A1F01FFDF136BC46D72CDFA80" authority="(Rathbun, 1905)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="Rathbun" baseAuthorityYear="1905" box="[353,738,1384,1401]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="antongilensis" status="comb. nov.">
<emphasis id="B959EAAB1F01FFDF136BC46D726AFA80" box="[353,581,1384,1401]" italics="true" pageId="32" pageNumber="73">Malagasya antongilensis</emphasis>
(
<bibRefCitation id="EFBC4B481F01FFDF105BC46D72F2FA80" author="RATHBUN M. J." box="[593,733,1384,1401]" pageId="32" pageNumber="73" pagination="159 - 322" refId="ref26961" refString="RATHBUN M. J. 1905. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 7 (4): 159 - 322." type="journal article" year="1905">Rathbun, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F01FFDF10E0C46D736EFA80" box="[746,833,1384,1401]" pageId="32" pageNumber="73" rank="species">n. comb.</taxonomicNameLabel>
; 
<emphasis id="B959EAAB1F01FFDF1140C46D7379FA80" bold="true" box="[842,854,1384,1401]" pageId="32" pageNumber="73">E</emphasis>
, 
<emphasis id="B959EAAB1F01FFDF116EC46D7340FA80" bold="true" box="[868,879,1384,1401]" pageId="32" pageNumber="73">F</emphasis>
, 
<taxonomicName id="4C2D4D3A1F01FFDF1174C46D74F3FA80" authority="(Bott, 1965)" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[894,1244,1384,1401]" class="Malacostraca" family="Potamonautidae" genus="Skeleosophusa" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F01FFDF1174C46D744CFA80" box="[894,1123,1384,1401]" italics="true" pageId="32" pageNumber="73">Skeleosophusa gollhardi</emphasis>
(
<bibRefCitation id="EFBC4B481F01FFDF167AC46D74F7FA80" author="BOTT R." box="[1136,1240,1384,1401]" pageId="32" pageNumber="73" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
)
</taxonomicName>
; 
<emphasis id="B959EAAB1F01FFDF129DC47B708AFA76" bold="true" box="[151,165,1406,1423]" pageId="32" pageNumber="73">G</emphasis>
, 
<emphasis id="B959EAAB1F01FFDF12A5C47B7092FA76" bold="true" box="[175,189,1406,1423]" pageId="32" pageNumber="73">H</emphasis>
, 
<taxonomicName id="4C2D4D3A1F01FFDF12C2C47B722CFA76" authority="Bott, 1965" authorityName="Bott" authorityYear="1965" box="[200,515,1406,1423]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="humberti">
<emphasis id="B959EAAB1F01FFDF12C2C47B718BFA76" box="[200,420,1406,1423]" italics="true" pageId="32" pageNumber="73">Madagapotamon humberti</emphasis>
<bibRefCitation id="EFBC4B481F01FFDF13A3C47B722CFA76" author="BOTT R." box="[425,515,1406,1423]" pageId="32" pageNumber="73" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B959EAAB1F01FFDF1001C47B723EFA76" bold="true" box="[523,529,1406,1423]" pageId="32" pageNumber="73">I</emphasis>
, 
<emphasis id="B959EAAB1F01FFDF1011C47B720AFA76" bold="true" box="[539,549,1406,1423]" pageId="32" pageNumber="73">J</emphasis>
, 
<taxonomicName id="4C2D4D3A1F01FFDF103AC47B7421FA76" authority="Cumberlidge, Boyko &amp; Harvey, 2002" authorityName="Cumberlidge, Boyko &amp; Harvey" authorityYear="2002" box="[560,1038,1406,1423]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="B959EAAB1F01FFDF103AC47B72F8FA76" box="[560,727,1406,1423]" italics="true" pageId="32" pageNumber="73">Marojejy longimerus</emphasis>
Cumberlidge, Boyko &amp; Harvey, 2002
</taxonomicName>
. Scale bar: 4.0 mm.
</paragraph>
</caption>
<paragraph id="8B9236B91F01FFDF129DC4C67266F9D1" blockId="32.[151,684,1475,1670]" pageId="32" pageNumber="73">
TYPELOCALITY. — Andranomifototra river, 
<geoCoordinate id="EE19507E1F01FFDF129DC4D870D0FA0C" box="[151,255,1501,1525]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="32" pageNumber="73" precision="15" seconds="8" value="-14.435555">14°26’8”S</geoCoordinate>
, 
<geoCoordinate id="EE19507E1F01FFDF1306C4D87158FA0C" box="[268,375,1501,1525]" degrees="49" direction="east" minutes="44" orientation="longitude" pageId="32" pageNumber="73" precision="15" seconds="1" value="49.733612">49°44’1”E</geoCoordinate>
, 
<quantity id="4CD59B5C1F01FFDF1389C4D871FAFA0A" box="[387,469,1500,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="32" pageNumber="73" unit="m" value="1875.0">1875 m</quantity>
, 
<quantity id="4CD59B5C1F01FFDF13E8C4D8720BFA0C" box="[482,548,1501,1525]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="32" pageNumber="73" unit="km" value="11.0">11 km</quantity>
northwest of Manantenina, Province d’Antsiranana, Réserve naturelle intégrale de 
<taxonomicName id="4C2D4D3A1F01FFDF1360C71571EAF9D1" box="[362,453,1552,1576]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
, 
<collectingCountry id="F33A76291F01FFDF13C5C715726AF9D1" box="[463,581,1552,1576]" name="Madagascar" pageId="32" pageNumber="73">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F01FFDF129DC7347181F97C" pageId="32" pageNumber="73" type="distribution">
<paragraph id="8B9236B91F01FFDF129DC7347181F97C" blockId="32.[151,684,1475,1670]" pageId="32" pageNumber="73">
DISTRIBUTION. — 
<collectingCountry id="F33A76291F01FFDF136AC73771F5F9B3" box="[352,474,1586,1610]" name="Madagascar" pageId="32" pageNumber="73">Madagascar</collectingCountry>
(
<figureCitation id="13162A3C1F01FFDF13E0C7377265F9B3" box="[490,586,1586,1610]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="32" pageNumber="73">Fig. 11E</figureCitation>
). Known only from the Réserve naturelle intégrale de 
<taxonomicName id="4C2D4D3A1F01FFDF104DC74E728CF99A" box="[583,675,1611,1635]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="73" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
. DIAGNOSIS. — As for genus.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F01FFDF10C0C4C674BFF97C" pageId="32" pageNumber="73" type="description">
<paragraph id="8B9236B91F01FFDF10C0C4C67374FA22" blockId="32.[714,1045,1475,1533]" box="[714,859,1475,1501]" pageId="32" pageNumber="73">DESCRIPTION</paragraph>
<paragraph id="8B9236B91F01FFDF10C0C4E6743AFA04" blockId="32.[714,1045,1475,1533]" box="[714,1045,1506,1533]" pageId="32" pageNumber="73">
See Cumberlidge 
<emphasis id="B959EAAB1F01FFDF1180C4E77399FA04" box="[906,950,1506,1533]" italics="true" pageId="32" pageNumber="73">et al</emphasis>
. (2002).
</paragraph>
<paragraph id="8B9236B91F01FFDF10C0C72F72DBF9BD" blockId="32.[714,1247,1578,1669]" box="[714,756,1578,1604]" pageId="32" pageNumber="73">
<emphasis id="B959EAAB1F01FFDF10C0C72F72DBF9BD" box="[714,756,1578,1604]" italics="true" pageId="32" pageNumber="73">Size</emphasis>
</paragraph>
<paragraph id="8B9236B91F01FFDF10C0C74E74BFF97C" blockId="32.[714,1247,1578,1669]" pageId="32" pageNumber="73">
The adult size range is from cw 
<quantity id="4CD59B5C1F01FFDF1640C74E748BF99D" box="[1098,1188,1610,1637]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="32" pageNumber="73" unit="mm" value="24.0">24 mm</quantity>
. The carapace proportions are given in 
<tableCitation id="C6AF03021F01FFDF163CC76E74A4F97C" box="[1078,1163,1643,1669]" captionStart="TABLE" captionStartId="9.[151,162,169,186]" captionText="TABLE 3. — Carapace and other proportions of Madagascan freshwater crabs. Abbreviations: cw, carapace width; cl, carapace length; ch, carapace height; fw, front width; p5m, length of merus of right pereiopod 5." pageId="32" pageNumber="73">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>