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<document id="C90E79DEA2BB09DEBCEC97BFB675A8C1" ID-CLB-Dataset="29184" ID-DOI="10.11646/zootaxa.4482.1.1" ID-GBIF-Dataset="72a80b13-e18f-4dc3-bec2-bb20d17f0e5f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454415" ID-ZooBank="79FECD6E-5192-45C3-9B46-4E6817366F64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539175145001" checkinUser="plazi" docAuthor="Deeming, John C." docDate="2018" docId="038487A77A4BFFF5FF1CB0A5EB0C12F0" docLanguage="en" docName="zootaxa.4482.1.1.pdf.imf" docOrigin="Zootaxa 4482 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyladelphus Becker 1910" docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="FFBDFFDF7A5CFFEDFF8BB638EF2D1377" masterDocTitle="A revision of the Afrotropical species of Pachylophus Loew (Diptera: Chloropidae) and some related Ovoviviparous genera" masterLastPageNumber="51" masterPageNumber="1" pageNumber="24" updateTime="1698571395358" updateUser="ExternalLinkService">
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<mods:title id="8FFD120433359424EA8A347A7369B3F3">A revision of the Afrotropical species of Pachylophus Loew (Diptera: Chloropidae) and some related Ovoviviparous genera</mods:title>
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<mods:namePart id="53ABE838801421E529E9955A34E9425F">Deeming, John C.</mods:namePart>
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<treatment id="038487A77A4BFFF5FF1CB0A5EB0C12F0" ID-DOI="http://doi.org/10.5281/zenodo.5966150" ID-GBIF-Taxon="148676296" ID-Zenodo-Dep="5966150" LSID="urn:lsid:plazi:treatment:038487A77A4BFFF5FF1CB0A5EB0C12F0" httpUri="http://treatment.plazi.org/id/038487A77A4BFFF5FF1CB0A5EB0C12F0" lastPageId="24" lastPageNumber="24" pageId="23" pageNumber="24">
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<paragraph id="8B9236B17A4BFFFAFF1CB0A5EECC15C0" blockId="23.[151,481,1693,1719]" box="[151,481,1693,1719]" pageId="23" pageNumber="24">
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<taxonomicName id="4C2D4D327A4BFFFAFF1CB0A5EECC15C0" ID-CoL="6PH4" authority="Becker, 1910" authorityName="Becker" authorityYear="1910" box="[151,481,1693,1719]" class="Insecta" family="Chloropidae" genus="Phyladelphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A4BFFFAFF1CB0A5EECC15C0" bold="true" box="[151,481,1693,1719]" pageId="23" pageNumber="24">
<emphasis id="B959EAA37A4BFFFAFF1CB0A5EE1615C0" bold="true" box="[151,315,1693,1719]" italics="true" pageId="23" pageNumber="24">Phyladelphus</emphasis>
<bibRefCitation id="EFBC4B407A4BFFFAFEC8B0A5EECC15C0" box="[323,481,1693,1719]" pageId="23" pageNumber="24" refId="ref41190" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. 2. Teil. Aethiopische Region. Annales historico-naturales Musei nationalis hungarici Budapest, 8, 377 - 443, tab. XIII." type="journal article">Becker, 1910</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="8B9236B17A4BFFFAFF1CB0DDEC58158D" blockId="23.[151,885,1764,1786]" box="[151,885,1764,1786]" pageId="23" pageNumber="24">
Type species
<taxonomicName id="4C2D4D327A4BFFFAFE94B0DCEDCE158D" authority="Becker, 1910: 54" authorityName="Becker" authorityPageNumber="54" authorityYear="1910" box="[287,739,1764,1786]" class="Insecta" family="Chloropidae" genus="Phyladelphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="thalhammeri">
<emphasis id="B959EAA37A4BFFFAFE94B0DCED1D158D" box="[287,560,1764,1786]" italics="true" pageId="23" pageNumber="24">Phyladelphus thalhammeri</emphasis>
<bibRefCitation id="EFBC4B407A4BFFFAFDBDB0DCEDED158D" box="[566,704,1764,1786]" pageId="23" pageNumber="24" refId="ref41190" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. 2. Teil. Aethiopische Region. Annales historico-naturales Musei nationalis hungarici Budapest, 8, 377 - 443, tab. XIII." type="journal article">Becker, 1910</bibRefCitation>
: 54
</taxonomicName>
by monotypy.
</paragraph>
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Left unattributed to a genus group by
<bibRefCitation id="EFBC4B407A4BFFFAFDB6B11EEC6F1448" box="[573,834,1830,1855]" pageId="23" pageNumber="24" refId="ref41065" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. In: Entomologica scandinavica. 8 (Supplement). Scandinavian Science Press, Klampenborg, pp. 1 - 200." type="book chapter">Andersson (1977: 180)</bibRefCitation>
, this genus is known from two oriental, one European and one
<collectingRegion id="49E9F8537A4BFFFAFF7CB173EE0A1415" box="[247,295,1867,1890]" country="Cameroon" name="East" pageId="23" pageNumber="24">East</collectingRegion>
African species. Notes on it are given by
<bibRefCitation id="EFBC4B407A4BFFFAFC8AB172EB001414" box="[769,1069,1866,1891]" pageId="23" pageNumber="24" refId="ref41720" refString="Deeming, J. C. (1981) A contribution to the study of afrotropical Chloropine Diptera. Journal of Natural History, 15 (5), 789 - 828. https: // doi. org / 10.1080 / 00222938100770581" type="journal article">Deeming (1981: 811812)</bibRefCitation>
, who was only able to liken it to
<taxonomicName id="4C2D4D327A4BFFFAFF1CB156EECC14F0" authority="Speiser, 1910" authorityName="Speiser" authorityYear="1910" box="[151,481,1902,1927]" class="Insecta" family="Chloropidae" genus="Mepachymerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A4BFFFAFF1CB156EE6F14F0" box="[151,322,1902,1927]" italics="true" pageId="23" pageNumber="24">Mepachymerus</emphasis>
<bibRefCitation id="EFBC4B407A4BFFFAFEC2B156EECC14F0" box="[329,481,1902,1927]" pageId="23" pageNumber="24" refId="ref43150" refString="Speiser, P. (1910) [Abteilung] 10. Diptera. 5. Cyclorhapha [sic!], Aschiza. Cyclorhapha, Schizophora, Schizometopa. (Auswahl); Holometopa. (Auswahl). In: Koniglich schwedische Akademie der Wissenschaften (Ed.), Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906 unter Leitung von Prof. Dr. Yngve Sjostedt. 2. Band, Abteilung 8 - 14. P. Palmquists Aktiebolag, Stockholm, pp. 113 - 202 incl. index." type="book chapter">Speiser, 1910</bibRefCitation>
</taxonomicName>
which latter
<bibRefCitation id="EFBC4B407A4BFFFAFDFDB156EC5414F0" box="[630,889,1902,1927]" pageId="23" pageNumber="24" refId="ref41065" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. In: Entomologica scandinavica. 8 (Supplement). Scandinavian Science Press, Klampenborg, pp. 1 - 200." type="book chapter">Andersson (1977: 132)</bibRefCitation>
placed along with
<taxonomicName id="4C2D4D327A4BFFFAFBDBB156EA4714F0" authority="Frey, 1961" authorityName="Frey" authorityYear="1961" box="[1104,1386,1902,1927]" class="Insecta" family="Chloropidae" genus="Steleocerellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A4BFFFAFBDBB156EBC614F0" box="[1104,1259,1902,1927]" italics="true" pageId="23" pageNumber="24">Steleocerellus</emphasis>
Frey,
<quantity id="4CD59B547A4BFFFAFAB9B156EA4714F0" box="[1330,1386,1902,1927]" metricMagnitude="1" metricUnit="m" metricValue="4.98094" pageId="23" pageNumber="24" unit="in" value="1961.0">1961</quantity>
</taxonomicName>
in a discrete genus group.
<taxonomicName id="4C2D4D327A4BFFFAFE18B1ACED8014DC" authority="Becker, 1910" authorityName="Becker" authorityYear="1910" box="[403,685,1938,1963]" class="Insecta" family="Chloropidae" genus="Phyladelphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B959EAA37A4BFFFAFE18B1ACED2014DC" box="[403,525,1940,1963]" italics="true" pageId="23" pageNumber="24">P. geminus</emphasis>
<bibRefCitation id="EFBC4B407A4BFFFAFD9DB1AAED8014DC" box="[534,685,1938,1963]" pageId="23" pageNumber="24" refId="ref41190" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. 2. Teil. Aethiopische Region. Annales historico-naturales Musei nationalis hungarici Budapest, 8, 377 - 443, tab. XIII." type="journal article">Becker, 1910</bibRefCitation>
</taxonomicName>
and the new species here described as
<taxonomicName id="4C2D4D327A4BFFFAFBF9B1ACEBFE14DC" authorityName="Deeming" authorityYear="2018" box="[1138,1235,1938,1963]" class="Insecta" family="Chloropidae" genus="Phyladelphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="woodi">
<emphasis id="B959EAA37A4BFFFAFBF9B1ACEBFE14DC" box="[1138,1235,1938,1963]" italics="true" pageId="23" pageNumber="24">P. woodi</emphasis>
</taxonomicName>
have in common with this genus group a flattened laceolate arista that is often rather rounded apically and which is clothed in very short setae. However, such aristal development does occur elsewhere. Duda (1933: 142) described a
<taxonomicName id="4C2D4D327A4BFFF5FA97B1E2EFD113C7" class="Insecta" family="Chloropidae" genus="Steleocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="24" lastPageNumber="25" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="lindbergi">
<emphasis id="B959EAA37A4BFFF5FA97B1E2EFD113C7" italics="true" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Steleocerus lindbergi</emphasis>
</taxonomicName>
from
<collectingCountry id="F33A76217A44FFF5FECDB6A0EE8613D8" box="[326,427,152,175]" name="Morocco" pageId="24" pageNumber="25">Morocco</collectingCountry>
with just such aristal development (
<taxonomicName id="4C2D4D327A44FFF5FCDBB6AFEB5D13C7" authority="Becker, 1910" authorityName="Becker" authorityYear="1910" box="[848,1136,151,176]" class="Insecta" family="Chloropidae" genus="Steleocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A44FFF5FCDBB6AFECFD13C7" box="[848,976,151,176]" italics="true" pageId="24" pageNumber="25">Steleocerus</emphasis>
<bibRefCitation id="EFBC4B407A44FFF5FC51B6AFEB5D13C7" box="[986,1136,151,176]" pageId="24" pageNumber="25" refId="ref41190" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. 2. Teil. Aethiopische Region. Annales historico-naturales Musei nationalis hungarici Budapest, 8, 377 - 443, tab. XIII." type="journal article">Becker, 1910</bibRefCitation>
</taxonomicName>
is now placed as a junior synonym of
<taxonomicName id="4C2D4D327A44FFF5FEA3B684ED0313A2" authority="Speiser" authorityName="Speiser" authorityYear="1910" box="[296,558,188,213]" class="Insecta" family="Chloropidae" genus="Mepachymerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A44FFF5FEA3B684EEFE13A2" box="[296,467,188,213]" italics="true" pageId="24" pageNumber="25">Mepachymerus</emphasis>
Speiser
</taxonomicName>
of the same year). This species was recorded from
<collectingCountry id="F33A76217A44FFF5FB0CB684EBCA13A2" box="[1159,1255,188,213]" name="Bulgaria" pageId="24" pageNumber="25">Bulgaria</collectingCountry>
and transferred into
<taxonomicName id="4C2D4D327A44FFF5FF46B6D9EEB8138F" authority="Becker" authorityName="Becker" authorityYear="1910" box="[205,405,223,248]" class="Insecta" family="Chloropidae" genus="Lasiosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAA37A44FFF5FF46B6D9EE17138F" box="[205,314,225,248]" italics="true" pageId="24" pageNumber="25">Lasiosina</emphasis>
Becker
</taxonomicName>
by
<bibRefCitation id="EFBC4B407A44FFF5FE4CB6E7EDD6138F" box="[455,763,223,248]" pageId="24" pageNumber="25" refId="ref41334" refString="Beschovski, V. (1983) On the systematic status of Steleocerus lindbergi (Duda, 1933) (Diptera, Chloropidae). Acta zoologica bulgarica, 21, 78 - 82. [Zoological Record translation of title in Bulgarian]" type="journal article">Beschovski (1983: 7880)</bibRefCitation>
. My dissection of the abdomen of a female from
<collectingCountry id="F33A76217A44FFF5FAC7B6E7EAB1138F" box="[1356,1436,223,248]" name="Turkey" pageId="24" pageNumber="25">Turkey</collectingCountry>
(Ödemis, Lake Gölcük,
<date id="FF9310717A44FFF5FE2EB73CED35126A" box="[421,536,260,285]" pageId="24" pageNumber="25" value="1997-07-09">9.vii.1997</date>
, J.C. Deeming, in
<collectionCode id="ED3CAE747A44FFF5FD63B73CEC69126B" box="[744,836,260,284]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="24" pageNumber="25">NMWC</collectionCode>
) confirms this. It has narrow cerci and the ovipositor, when extended, is nearly as long as the abdomen. Thus, in the
<taxonomicName id="4C2D4D327A44FFF5FCFAB71FECD21237" box="[881,1023,295,320]" class="Insecta" family="Chloropidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
, the character of a blade-like arista, which is undoubtedly apomorphic and cannot be regarded as being of diagnostic significance in the attributing of species to genera, although in the
<taxonomicName id="4C2D4D327A44FFF5FD98B757EDB312FF" box="[531,670,367,392]" class="Insecta" family="Lauxaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="family">Lauxaniidae</taxonomicName>
and
<taxonomicName id="4C2D4D327A44FFF5FD5FB757EC7E12FF" box="[724,851,367,392]" class="Insecta" family="Celyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="family">Celyphidae</taxonomicName>
it may well be so.
</paragraph>
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