treatments-xml/data/03/84/87/0384879CF543FF9E67E5FC8BFA1F90D1.xml

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<mods:title id="0464117317C4FF1B16418234F5F8E84C">A new species of Saurida (Pisces: Synodontidae) from the Mascarene Plateau, Western Indian Ocean</mods:title>
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<mods:date id="D5AB3F11268E8928917B7264FF9DAB47">2015</mods:date>
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<taxonomicName id="4C2D4D09F543FF9A67E5FC8BFEEA9028" box="[151,353,834,860]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="1" pageNumber="441" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F543FF9A67E5FC8BFEEA9028" bold="true" box="[151,353,834,860]" italics="true" pageId="1" pageNumber="441">Saurida tweddlei</emphasis>
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</emphasis>
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<paragraph id="8B92368AF543FF9A67E5FCAFFE0E90D6" blockId="1.[151,606,834,930]" pageId="1" pageNumber="441">
New English name: Tweddle’s lizardfish 
<figureCitation id="13162A0FF543FF9A67E5FC40FE9490D6" box="[151,287,905,930]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1493,1515]" captionStartId-1="3.[151,250,1981,2003]" captionStartId-2="4.[151,250,676,698]" captionTargetBox-0="[343,1231,193,1471]" captionTargetBox-1="[233,1353,1596,1959]" captionTargetBox-2="[176,1411,193,654]" captionTargetId-0="figure@3.[334,1242,183,1481]" captionTargetId-1="figure@3.[233,1354,1596,1960]" captionTargetId-2="figure@4.[176,1412,193,654]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Map of Mascarene Plateau showing location and station numbers at which Saurida tweddlei n. sp. were caught: + = RV Dr Fridtjof Nansen stations, with station number; ★ = FV Ryuyo-Maru No. 2 stations." captionText-1="FIGURE 2. Holotype of Saurida tweddlei n. sp., SAIAB 185941, 354.0 mm SL, Saya da Malha Bank (photo: O. Alvheim)" captionText-2="FIGURE 3. Photograph of ‘ Saurida undosquamis’ from Saya da Malha Bank (Amaoka et al. 1976) now identified as S. tweddlei n. sp." httpUri-0="https://zenodo.org/record/242542/files/figure.png" httpUri-1="https://zenodo.org/record/242543/files/figure.png" httpUri-2="https://zenodo.org/record/242544/files/figure.png" pageId="1" pageNumber="441">Figures 1–3</figureCitation>
; 
<tableCitation id="C6AF0331F543FF9A6659FC40FEF590D6" box="[299,382,905,930]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[151,1354,201,2023]" captionTargetPageId="2" captionText="TABLE 1. Morphometric and meristic data of the holotype and 11 paratype specimens of Saurida tweddlei n. sp." httpUri="http://table.plazi.org/id/DF526602F540FF9967E5FF51FAC193DB" pageId="1" pageNumber="441" tableUuid="DF526602F540FF9967E5FF51FAC193DB">Table 1</tableCitation>
.
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<treatmentCitationGroup id="AB3D11A4F543FF9A67E5FC18FB039093" box="[151,1160,977,999]" pageId="1" pageNumber="441">
<taxonomicName id="4C2D4D09F543FF9A67E5FC18FEF89093" ID-CoL="6XQ8D" box="[151,371,977,999]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="1" pageNumber="441" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F543FF9A67E5FC18FEF89093" box="[151,371,977,999]" italics="true" pageId="1" pageNumber="441">Saurida undosquamis</emphasis>
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(
<emphasis id="B959EA98F543FF9A66F0FC1BFE239093" box="[386,424,978,999]" italics="true" pageId="1" pageNumber="441">non</emphasis>
<bibRefCitation id="EFBC4B7BF543FF9A66DCFC18FDD59093" author="Richardson" box="[430,606,977,999]" pageId="1" pageNumber="441" refString="Richardson, J. (1848) Ichthyology of the voyage of H. M. S. Erebus &amp; Terror. In: Richardson, J. &amp; Gray, J. E. (Eds.), The zoology of the voyage of H. H. S. &quot; Erebus &amp; Terror, &quot; under the command of Captain Sir J. C. Ross ... during ... 1839 - 43, 2 (2), i - viii + 1 - 139, pls. 1 - 60. [London]" type="journal article" year="1848">Richardson, 1848</bibRefCitation>
)— 
<treatmentCitation id="0A8C109BF543FF9A650DFC18FCF59093" author="Amaoka" box="[639,894,977,999]" page="115" pageId="1" pageNumber="441" year="1976">
<bibRefCitation id="EFBC4B7BF543FF9A650DFC18FCC69093" author="Amaoka" box="[639,845,977,999]" pageId="1" pageNumber="441" refString="Amaoka, K., Hatanaka, H., Ida, H., Ikeda, I., Inada, T., Iwai, T., Kawahara, S., Kono, H., Kuronuma, K., Matsuura, K., Miyake, S., Nakamura, I., Ochiai, A., Okamura, O., Okutani, T., Sasaki, T., Sato, T. (Tetsuya), Sato, T. (Torao), Shimizu, T., Takeda, M., Taniuchi, T., Tominaga, Y. &amp; Uyeno, T. (Eds.) (1976) Colored Illustrations of Bottomfishes Collected by Japanese Trawlers. Vol. II. Japan Deep Sea Trawlers Association, Tokyo, 188 pp." type="book" year="1976">
Amaoka 
<emphasis id="B959EA98F543FF9A65ACFC1BFC9A9093" box="[734,785,977,999]" italics="true" pageId="1" pageNumber="441">et al.</emphasis>
1976
</bibRefCitation>
: 115
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, Plate Sey-1 (
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).
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</paragraph>
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<emphasis id="B959EA98F543FF9A67E5FBDDFE8F9759" bold="true" box="[151,260,1044,1069]" pageId="1" pageNumber="441">Holotype</emphasis>
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. 
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185941 (tissue voucher 
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08-1710), 354.0 mm 
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, Indian Ocean, Saya da Malha Bank, Mascarene Trawl Survey Station 19, 
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, 
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, 
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, D. Tweddle and 
<collectingCountry id="F33A761AF543FF9A633CFBF1FBEE9724" box="[1102,1125,1080,1104]" name="Iceland" pageId="1" pageNumber="441">O</collectingCountry>
. Alvheim, RV 
<emphasis id="B959EA98F543FF9A6268FBF0FF609700" italics="true" pageId="1" pageNumber="441">Dr Fridtjof Nansen</emphasis>
, 
<date id="FF93104AF543FF9A678BFB94FE0B9700" box="[249,384,1116,1141]" pageId="1" pageNumber="441" value="2009-07-07">7 July 2009</date>
.
</paragraph>
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<emphasis id="B959EA98F543FF9A67B5FBB6FEB497EC" bold="true" box="[199,319,1151,1176]" pageId="1" pageNumber="441">Paratypes</emphasis>
</typeStatus>
(11: 247.9–401.0 mm 
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). 
<collectionCode id="ED3CAE4FF543FF9A65FCFB49FD6897E3" LSID="urn:lsid:biocol.org:col:34988" box="[654,739,1152,1175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="441">MNHN</collectionCode>
2012-0113, 
<quantity id="4CD59B6FF543FF9A6408FBB6FC7A97E3" box="[890,1009,1151,1176]" metricMagnitude="-1" metricUnit="m" metricValue="3.553" pageId="1" pageNumber="441" unit="mm" value="355.3">355.3 mm</quantity>
<collectionCode id="ED3CAE4FF543FF9A648FFB49FBAB97EC" box="[1021,1056,1152,1176]" pageId="1" pageNumber="441">SL</collectionCode>
, same data as 
<typeStatus id="54968828F543FF9A63A1FBB6FAB297EC" box="[1235,1337,1151,1176]" pageId="1" pageNumber="441" type="holotype">holotype</typeStatus>
. NSMT- P 
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<date id="FF93104AF543FF9A67D5FB6DFF7697C8" box="[167,253,1188,1212]" pageId="1" pageNumber="441" value="34-12-13">121334</date>
, 358.2 mm
</quantity>
<collectionCode id="ED3CAE4FF543FF9A66F4FB6DFE2297C8" box="[390,425,1188,1212]" pageId="1" pageNumber="441">SL</collectionCode>
, station 
<collectionCode id="ED3CAE4FF543FF9A6578FB6DFDDA97C8" box="[522,593,1188,1212]" pageId="1" pageNumber="441">SKSK</collectionCode>
7745, Indian Ocean, Saya da Malha Bank, 
<geoCoordinate id="EE19504DF543FF9A633AFB6DFB2D97C8" box="[1096,1190,1188,1212]" direction="south" orientation="latitude" pageId="1" pageNumber="441" precision="925" value="-10.733334">10°44’S</geoCoordinate>
, 
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, FV 
<emphasis id="B959EA98F543FF9A623FFB6CFE929794" italics="true" pageId="1" pageNumber="441">Ryuyo- Maru No. 2</emphasis>
, 
<date id="FF93104AF543FF9A6655FB0EFE7A9794" box="[295,497,1223,1248]" pageId="1" pageNumber="441" value="1978-12-06">6 December 1978</date>
. NSMT-P 
<quantity id="4CD59B6FF543FF9A6517FB0EFCBE97AB" box="[613,821,1223,1248]" metricMagnitude="5" metricUnit="m" metricValue="1.2133532739999997" pageId="1" pageNumber="441" unit="mm" value="1.213353274E8">
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, 327.4 mm
</quantity>
<collectionCode id="ED3CAE4FF543FF9A644EFB01FCD49794" box="[828,863,1224,1248]" pageId="1" pageNumber="441">SL</collectionCode>
, station 
<collectionCode id="ED3CAE4FF543FF9A64CEFB01FB889794" box="[956,1027,1224,1248]" pageId="1" pageNumber="441">SKSK</collectionCode>
8512, Indian Ocean, Saya da Malha Bank, 
<geoCoordinate id="EE19504DF543FF9A6792FB25FEB59671" box="[224,318,1260,1285]" direction="south" orientation="latitude" pageId="1" pageNumber="441" precision="925" value="-10.6">10°36’S</geoCoordinate>
, 
<geoCoordinate id="EE19504DF543FF9A663BFB25FE229671" box="[329,425,1260,1285]" direction="east" orientation="longitude" pageId="1" pageNumber="441" precision="925" value="61.566666">61°34’E</geoCoordinate>
, FV 
<emphasis id="B959EA98F543FF9A66ADFB24FD249670" box="[479,687,1260,1284]" italics="true" pageId="1" pageNumber="441">Ryuyo-Maru No. 2</emphasis>
, 
<date id="FF93104AF543FF9A65CFFB25FC139670" box="[701,920,1260,1285]" pageId="1" pageNumber="441" value="1978-11-30">30 November 1978</date>
. NSMT-P 
<date id="FF93104AF543FF9A6379FB25FBD59671" box="[1035,1118,1260,1285]" pageId="1" pageNumber="441" value="36-12-13">121336</date>
, 349.0 mm 
<collectionCode id="ED3CAE4FF543FF9A6392FB25FA889670" box="[1248,1283,1260,1284]" pageId="1" pageNumber="441">SL</collectionCode>
, 
<collectionCode id="ED3CAE4FF543FF9A627DFB25FADD9670" box="[1295,1366,1260,1284]" pageId="1" pageNumber="441">SKSK</collectionCode>
8513, same data as NSMT-P 
<date id="FF93104AF543FF9A66E1FAC6FE6D965C" box="[403,486,1295,1320]" pageId="1" pageNumber="441" value="35-12-13">121335</date>
. NSMT-P 
<date id="FF93104AF543FF9A652AFAD9FD21965C" box="[600,682,1296,1320]" pageId="1" pageNumber="441" value="37-12-13">121337</date>
, 365.0 mm 
<collectionCode id="ED3CAE4FF543FF9A645FFAD9FCDB965C" box="[813,848,1296,1320]" pageId="1" pageNumber="441">SL</collectionCode>
, 
<collectionCode id="ED3CAE4FF543FF9A6429FAD9FC28965C" box="[859,931,1296,1320]" pageId="1" pageNumber="441">SKSK</collectionCode>
8514, same data as NSMT-P 
<date id="FF93104AF543FF9A6399FAC6FAB5965C" box="[1259,1342,1295,1320]" pageId="1" pageNumber="441" value="35-12-13">121335</date>
. NSMT- P 
<quantity id="4CD59B6FF543FF9A67D5FAFDFEF09638" box="[167,379,1332,1357]" metricMagnitude="5" metricUnit="m" metricValue="1.2133833859999998" pageId="1" pageNumber="441" unit="mm" value="1.213383386E8">
<date id="FF93104AF543FF9A67D5FAFDFF769638" box="[167,253,1332,1356]" pageId="1" pageNumber="441" value="38-12-13">121338</date>
, 338.6 mm
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<collectionCode id="ED3CAE4FF543FF9A66F3FAFDFE2F9638" box="[385,420,1332,1356]" pageId="1" pageNumber="441">SL</collectionCode>
, 
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8515, same data as NSMT-P 
<date id="FF93104AF543FF9A6431FAFDFC1D9639" box="[835,918,1332,1357]" pageId="1" pageNumber="441" value="35-12-13">121335</date>
. NSMT-P 
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, 352.0 mm 
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, 
<collectionCode id="ED3CAE4FF543FF9A627CFAFDFADD9638" box="[1294,1366,1332,1356]" pageId="1" pageNumber="441">SKSK</collectionCode>
8516, same data as NSMT-P 
<date id="FF93104AF543FF9A66E7FA9EFE639604" box="[405,488,1367,1392]" pageId="1" pageNumber="441" value="35-12-13">121335</date>
. 
<collectionCode id="ED3CAE4FF543FF9A6680FA91FDBB961B" box="[498,560,1368,1391]" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="1" pageNumber="441">NTM</collectionCode>
S.17300-001, 
<quantity id="4CD59B6FF543FF9A65A4FA9EFCC3961B" box="[726,840,1367,1392]" metricMagnitude="-1" metricUnit="m" metricValue="3.453" pageId="1" pageNumber="441" unit="mm" value="345.3">345.3 mm</quantity>
<collectionCode id="ED3CAE4FF543FF9A643DFA91FCF99604" box="[847,882,1368,1392]" pageId="1" pageNumber="441">SL</collectionCode>
, same data as 
<typeStatus id="54968828F543FF9A6361FA9EFBF29604" box="[1043,1145,1367,1392]" pageId="1" pageNumber="441" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED3CAE4FF543FF9A63F1FA91FB5C9604" LSID="urn:lsid:biocol.org:col:34984" box="[1155,1239,1368,1392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="1" pageNumber="441">SAIAB</collectionCode>
84181, 3: 247.9– 401.0 mm 
<collectionCode id="ED3CAE4FF543FF9A667DFAB5FEB996E0" box="[271,306,1404,1428]" pageId="1" pageNumber="441">SL</collectionCode>
, same data as 
<typeStatus id="54968828F543FF9A66A1FAB5FDB296E1" box="[467,569,1404,1429]" pageId="1" pageNumber="441" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376501F543FF9A67B5FA56FAA9953C" pageId="1" pageNumber="441" type="diagnosis">
<paragraph id="8B92368AF543FF9A67B5FA56FAA9953C" blockId="1.[151,1437,1044,2005]" pageId="1" pageNumber="441">
<emphasis id="B959EA98F543FF9A67B5FA56FEB196CC" bold="true" box="[199,314,1439,1464]" pageId="1" pageNumber="441">Diagnosis</emphasis>
. A large species of 
<taxonomicName id="4C2D4D09F543FF9A6556FA69FDF696CC" box="[548,637,1440,1464]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="1" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="B959EA98F543FF9A6556FA69FDF696CC" box="[548,637,1440,1464]" italics="true" pageId="1" pageNumber="441">Saurida</emphasis>
</taxonomicName>
, with the following combination of characters: dorsal fin with 12–13 rays; pectoral fins with 14–15 rays; lateral-line scales 53–55; transverse scale rows above lateral line 4½, below lateral line 5½; pectoral fins moderately long (extending to or just beyond a line from origin of pelvic fins to origin of dorsal fin); 2 rows of teeth on outer palatines; 0–3 teeth on vomer; tongue with about 4–5 rows of teeth posteriorly; caudal peduncle compressed (depth greater than width); stomach and intestine pale whitish.
</paragraph>
</subSubSection>
<subSubSection id="C3376501F543FF9F67B5F99DFBAA960A" lastPageId="4" lastPageNumber="444" pageId="1" pageNumber="441" type="description">
<paragraph id="8B92368AF543FF9A67B5F99DFDA29589" blockId="1.[151,1437,1044,2005]" pageId="1" pageNumber="441">
<emphasis id="B959EA98F543FF9A67B5F99DFEDA9519" bold="true" box="[199,337,1620,1645]" pageId="1" pageNumber="441">Description</emphasis>
. Counts and proportional measurements of the 
<typeStatus id="54968828F543FF9A6417F99CFC1E9519" box="[869,917,1621,1645]" pageId="1" pageNumber="441">type</typeStatus>
series are provided in 
<tableCitation id="C6AF0331F543FF9A63E1F99DFB6C9518" box="[1171,1255,1620,1645]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[151,1354,201,2023]" captionTargetPageId="2" captionText="TABLE 1. Morphometric and meristic data of the holotype and 11 paratype specimens of Saurida tweddlei n. sp." httpUri="http://table.plazi.org/id/DF526602F540FF9967E5FF51FAC193DB" pageId="1" pageNumber="441" tableUuid="DF526602F540FF9967E5FF51FAC193DB">Table 1</tableCitation>
. Dorsal-fin rays 13 (12–13); pectoral-fin rays 14 (14–15); pelvic-fin rays 9; anal-fin rays 11 (10–12). Pored lateral-line scales 54 (53–55); transverse scale rows above lateral-line 4½, transverse scale rows below lateral-line 5½; scale rows below lateral line with melanophores 2 (1–2); pre-dorsal scales 19 (16–20); pre-adipose scales 16 (15–18); post-adipose scales 10 (11–12). Vertebrae 49–50.
</paragraph>
<paragraph id="8B92368AF543FF9A67B5F8CEFDFD94A0" blockId="1.[151,1437,1044,2005]" pageId="1" pageNumber="441">Body elongate and cylindrical, somewhat depressed on head and compressed on caudal peduncle, the latter with slightly raised lateral line scales, forming a slight ridge; scales large, cycloid, not very deciduous; scales present on cheek and opercle. Snout rounded when viewed from above. Nostrils located about midway between snout tip and anterior margin of orbit; anterior nostrils a little larger than posterior, anterior one with a broad dermal flap posteriorly. Eye circular, directed laterally, covered with fleshy adipose eyelid anteriorly and posteriorly. Interorbital region broad, slightly concave.</paragraph>
<caption id="DF526602F540FF9967E5FF51FAC193DB" ID-Table-UUID="DF526602F540FF9967E5FF51FAC193DB" box="[151,1354,151,175]" httpUri="http://table.plazi.org/id/DF526602F540FF9967E5FF51FAC193DB" pageId="2" pageNumber="442" targetBox="[151,1354,201,2023]" targetIsTable="true" targetPageId="2">
<paragraph id="8B92368AF540FF9967E5FF51FAC193DB" blockId="2.[151,1354,151,175]" box="[151,1354,151,175]" pageId="2" pageNumber="442">
<emphasis id="B959EA98F540FF9967E5FF51FE8193DB" bold="true" box="[151,266,152,175]" pageId="2" pageNumber="442">TABLE 1.</emphasis>
Morphometric and meristic data of the holotype and 11 paratype specimens of 
<taxonomicName id="4C2D4D09F540FF99632AFF5EFA8693DB" box="[1112,1293,151,175]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="2" pageNumber="442" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F540FF99632AFF5EFA8693DB" box="[1112,1293,151,175]" italics="true" pageId="2" pageNumber="442">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F540FF996266FF51FAC193DB" box="[1300,1354,151,175]" pageId="2" pageNumber="442" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B92368AF540FF9967EDFF21FB889493" pageId="2" pageNumber="442">
<table id="F92DC42AF540006467E5FF00FAC19493" box="[151,1354,201,2023]" gridcols="3" gridrows="44" pageId="2" pageNumber="442">
<tr id="351D34C8F540006467E5FF00FAC19251" box="[151,1354,201,293]" gridrow="0" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FF00FD009251" box="[151,651,201,293]" gridcol="0" gridrow="0" pageId="2" pageNumber="442">
Standard length (mm) 
<emphasis id="B959EA98F540FF9967EDFEC6FE1C9251" bold="true" box="[159,407,270,293]" pageId="2" pageNumber="442">Morphometrics (%SL)</emphasis>
</th>
<th id="76CC5DB4F540006465C3FF00FC849251" box="[689,783,201,293]" gridcol="1" gridrow="0" pageId="2" pageNumber="442">Holotype 354.0</th>
<th id="76CC5DB4F540006464AFFF00FAC19251" box="[989,1354,201,293]" gridcol="2" gridrow="0" pageId="2" pageNumber="442">Paratypes 247.9–401.0 (n = 11) Mean (range)</th>
</tr>
<tr id="351D34C8F540006467E5FEFFFAC19238" box="[151,1354,310,332]" gridrow="1" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FEFFFD009238" box="[151,651,310,332]" gridcol="0" gridrow="1" pageId="2" pageNumber="442">Predorsal length</th>
<td id="76CC5DB4F540006465C3FEFFFC849238" box="[689,783,310,332]" gridcol="1" gridrow="1" pageId="2" pageNumber="442">40.5</td>
<td id="76CC5DB4F540006464AFFEFFFAC19238" box="[989,1354,310,332]" gridcol="2" gridrow="1" pageId="2" pageNumber="442">42.8 (39.3–46.3)</td>
</tr>
<tr id="351D34C8F540006467E5FE94FAC19207" box="[151,1354,349,371]" gridrow="2" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FE94FD009207" box="[151,651,349,371]" gridcol="0" gridrow="2" pageId="2" pageNumber="442">Preadipose length</th>
<td id="76CC5DB4F540006465C3FE94FC849207" box="[689,783,349,371]" gridcol="1" gridrow="2" pageId="2" pageNumber="442">79.5</td>
<td id="76CC5DB4F540006464AFFE94FAC19207" box="[989,1354,349,371]" gridcol="2" gridrow="2" pageId="2" pageNumber="442">80.8 (78.1–88.2)</td>
</tr>
<tr id="351D34C8F540006467E5FE4CFAC192EF" box="[151,1354,389,411]" gridrow="3" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FE4CFD0092EF" box="[151,651,389,411]" gridcol="0" gridrow="3" pageId="2" pageNumber="442">Preanal length</th>
<td id="76CC5DB4F540006465C3FE4CFC8492EF" box="[689,783,389,411]" gridcol="1" gridrow="3" pageId="2" pageNumber="442">72.0</td>
<td id="76CC5DB4F540006464AFFE4CFAC192EF" box="[989,1354,389,411]" gridcol="2" gridrow="3" pageId="2" pageNumber="442">74.9 (70.7–79.2)</td>
</tr>
<tr id="351D34C8F540006467E5FE65FAC192B6" box="[151,1354,428,450]" gridrow="4" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FE65FD0092B6" box="[151,651,428,450]" gridcol="0" gridrow="4" pageId="2" pageNumber="442">Preanal-fin length</th>
<td id="76CC5DB4F540006465C3FE65FC8492B6" box="[689,783,428,450]" gridcol="1" gridrow="4" pageId="2" pageNumber="442">74.6</td>
<td id="76CC5DB4F540006464AFFE65FAC192B6" box="[989,1354,428,450]" gridcol="2" gridrow="4" pageId="2" pageNumber="442">77.9 (73.2–82.7)</td>
</tr>
<tr id="351D34C8F540006467E5FE1AFAC1929D" box="[151,1354,467,489]" gridrow="5" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FE1AFD00929D" box="[151,651,467,489]" gridcol="0" gridrow="5" pageId="2" pageNumber="442">Prepectoral length</th>
<td id="76CC5DB4F540006465C3FE1AFC84929D" box="[689,783,467,489]" gridcol="1" gridrow="5" pageId="2" pageNumber="442">25.8</td>
<td id="76CC5DB4F540006464AFFE1AFAC1929D" box="[989,1354,467,489]" gridcol="2" gridrow="5" pageId="2" pageNumber="442">27.6 (24.6–29.8)</td>
</tr>
<tr id="351D34C8F540006467E5FE33FAC19164" box="[151,1354,506,528]" gridrow="6" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FE33FD009164" box="[151,651,506,528]" gridcol="0" gridrow="6" pageId="2" pageNumber="442">Prepelvic length</th>
<td id="76CC5DB4F540006465C3FE33FC849164" box="[689,783,506,528]" gridcol="1" gridrow="6" pageId="2" pageNumber="442">36.9</td>
<td id="76CC5DB4F540006464AFFE33FAC19164" box="[989,1354,506,528]" gridcol="2" gridrow="6" pageId="2" pageNumber="442">39.5 (36.7–42.5)</td>
</tr>
<tr id="351D34C8F540006467E5FDEBFAC1914C" box="[151,1354,546,568]" gridrow="7" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FDEBFD00914C" box="[151,651,546,568]" gridcol="0" gridrow="7" pageId="2" pageNumber="442">Head length (HL)</th>
<td id="76CC5DB4F540006465C3FDEBFC84914C" box="[689,783,546,568]" gridcol="1" gridrow="7" pageId="2" pageNumber="442">24.2</td>
<td id="76CC5DB4F540006464AFFDEBFAC1914C" box="[989,1354,546,568]" gridcol="2" gridrow="7" pageId="2" pageNumber="442">26.1 (24.1–27.9)</td>
</tr>
<tr id="351D34C8F540006467E5FD80FAC1912B" box="[151,1354,585,607]" gridrow="8" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FD80FD00912B" box="[151,651,585,607]" gridcol="0" gridrow="8" pageId="2" pageNumber="442">Body depth</th>
<td id="76CC5DB4F540006465C3FD80FC84912B" box="[689,783,585,607]" gridcol="1" gridrow="8" pageId="2" pageNumber="442">11.5</td>
<td id="76CC5DB4F540006464AFFD80FAC1912B" box="[989,1354,585,607]" gridcol="2" gridrow="8" pageId="2" pageNumber="442">13.5 (12.5–15.2)</td>
</tr>
<tr id="351D34C8F540006467E5FDB9FAC191F2" box="[151,1354,624,646]" gridrow="9" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FDB9FD0091F2" box="[151,651,624,646]" gridcol="0" gridrow="9" pageId="2" pageNumber="442">Body width</th>
<td id="76CC5DB4F540006465C3FDB9FC8491F2" box="[689,783,624,646]" gridcol="1" gridrow="9" pageId="2" pageNumber="442">13.5</td>
<td id="76CC5DB4F540006464AFFDB9FAC191F2" box="[989,1354,624,646]" gridcol="2" gridrow="9" pageId="2" pageNumber="442">13.8 (12.2–14.9)</td>
</tr>
<tr id="351D34C8F540006467E5FD5EFAC191D9" box="[151,1354,663,685]" gridrow="10" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FD5EFD0091D9" box="[151,651,663,685]" gridcol="0" gridrow="10" pageId="2" pageNumber="442">Interpelvic width</th>
<td id="76CC5DB4F540006465C3FD5EFC8491D9" box="[689,783,663,685]" gridcol="1" gridrow="10" pageId="2" pageNumber="442">8.5</td>
<td id="76CC5DB4F540006464AFFD5EFAC191D9" box="[989,1354,663,685]" gridcol="2" gridrow="10" pageId="2" pageNumber="442">9.3 (8.6–10.4)</td>
</tr>
<tr id="351D34C8F540006467E5FD77FAC191A0" box="[151,1354,702,724]" gridrow="11" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FD77FD0091A0" box="[151,651,702,724]" gridcol="0" gridrow="11" pageId="2" pageNumber="442">Pectoral-fin length</th>
<td id="76CC5DB4F540006465C3FD77FC8491A0" box="[689,783,702,724]" gridcol="1" gridrow="11" pageId="2" pageNumber="442">13.0</td>
<td id="76CC5DB4F540006464AFFD77FAC191A0" box="[989,1354,702,724]" gridcol="2" gridrow="11" pageId="2" pageNumber="442">14.9 (12.6–17.4)</td>
</tr>
<tr id="351D34C8F540006467E5FD2CFAC1918F" box="[151,1354,741,763]" gridrow="12" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FD2CFD00918F" box="[151,651,741,763]" gridcol="0" gridrow="12" pageId="2" pageNumber="442">Pelvic-fin length</th>
<td id="76CC5DB4F540006465C3FD2CFC84918F" box="[689,783,741,763]" gridcol="1" gridrow="12" pageId="2" pageNumber="442">13.8</td>
<td id="76CC5DB4F540006464AFFD2CFAC1918F" box="[989,1354,741,763]" gridcol="2" gridrow="12" pageId="2" pageNumber="442">15.6 (14.0–17.2)</td>
</tr>
<tr id="351D34C8F540006467E5FCC5FAC19051" box="[151,1354,780,805]" gridrow="13" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FCC5FD009051" box="[151,651,780,805]" gridcol="0" gridrow="13" pageId="2" pageNumber="442">Length of 2nd dorsal ray</th>
<td id="76CC5DB4F540006465C3FCC5FC849051" box="[689,783,780,805]" gridcol="1" gridrow="13" pageId="2" pageNumber="442">18.5</td>
<td id="76CC5DB4F540006464AFFCC5FAC19051" box="[989,1354,780,805]" gridcol="2" gridrow="13" pageId="2" pageNumber="442">19.5 (16.4–22.9)</td>
</tr>
<tr id="351D34C8F540006467E5FCFFFAC19038" box="[151,1354,822,844]" gridrow="14" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FCFFFD009038" box="[151,651,822,844]" gridcol="0" gridrow="14" pageId="2" pageNumber="442">Length of last dorsal ray</th>
<td id="76CC5DB4F540006465C3FCFFFC849038" box="[689,783,822,844]" gridcol="1" gridrow="14" pageId="2" pageNumber="442">4.8</td>
<td id="76CC5DB4F540006464AFFCFFFAC19038" box="[989,1354,822,844]" gridcol="2" gridrow="14" pageId="2" pageNumber="442">5.6 (4.7–6.9)</td>
</tr>
<tr id="351D34C8F540006467E5FC97FAC190E9" box="[151,1354,862,925]" gridrow="15" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FC97FD0090E9" box="[151,651,862,925]" gridcol="0" gridrow="15" pageId="2" pageNumber="442">Length dorsal-fin base Length of 2nd anal ray</th>
<td id="76CC5DB4F540006465C3FC97FC8490E9" box="[689,783,862,925]" gridcol="1" gridrow="15" pageId="2" pageNumber="442">12.9 7.1</td>
<td id="76CC5DB4F540006464AFFC97FAC190E9" box="[989,1354,862,925]" gridcol="2" gridrow="15" pageId="2" pageNumber="442">14.1 (13.2–15.7) 8.9 (8.1–10.1)</td>
</tr>
<tr id="351D34C8F540006467E5FC67FAC190B0" box="[151,1354,942,964]" gridrow="16" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FC67FD0090B0" box="[151,651,942,964]" gridcol="0" gridrow="16" pageId="2" pageNumber="442">Length of last anal ray</th>
<td id="76CC5DB4F540006465C3FC67FC8490B0" box="[689,783,942,964]" gridcol="1" gridrow="16" pageId="2" pageNumber="442">5.6</td>
<td id="76CC5DB4F540006464AFFC67FAC190B0" box="[989,1354,942,964]" gridcol="2" gridrow="16" pageId="2" pageNumber="442">5.4 (2.6–7.2)</td>
</tr>
<tr id="351D34C8F540006467E5FC1FFAC19098" box="[151,1354,982,1004]" gridrow="17" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FC1FFD009098" box="[151,651,982,1004]" gridcol="0" gridrow="17" pageId="2" pageNumber="442">Length of anal-fin base</th>
<td id="76CC5DB4F540006465C3FC1FFC849098" box="[689,783,982,1004]" gridcol="1" gridrow="17" pageId="2" pageNumber="442">9.4</td>
<td id="76CC5DB4F540006464AFFC1FFAC19098" box="[989,1354,982,1004]" gridcol="2" gridrow="17" pageId="2" pageNumber="442">10.0 (8.3–12.0)</td>
</tr>
<tr id="351D34C8F540006467E5FC34FAC19767" box="[151,1354,1021,1043]" gridrow="18" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FC34FD009767" box="[151,651,1021,1043]" gridcol="0" gridrow="18" pageId="2" pageNumber="442">Length of caudal peduncle</th>
<td id="76CC5DB4F540006465C3FC34FC849767" box="[689,783,1021,1043]" gridcol="1" gridrow="18" pageId="2" pageNumber="442">15.8</td>
<td id="76CC5DB4F540006464AFFC34FAC19767" box="[989,1354,1021,1043]" gridcol="2" gridrow="18" pageId="2" pageNumber="442">16.3 (14.9–17.7)</td>
</tr>
<tr id="351D34C8F540006467E5FBEDFAC1974E" box="[151,1354,1060,1082]" gridrow="19" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FBEDFD00974E" box="[151,651,1060,1082]" gridcol="0" gridrow="19" pageId="2" pageNumber="442">Depth of caudal peduncle</th>
<td id="76CC5DB4F540006465C3FBEDFC84974E" box="[689,783,1060,1082]" gridcol="1" gridrow="19" pageId="2" pageNumber="442">6.3</td>
<td id="76CC5DB4F540006464AFFBEDFAC1974E" box="[989,1354,1060,1082]" gridcol="2" gridrow="19" pageId="2" pageNumber="442">6.6 (6.0–7.4)</td>
</tr>
<tr id="351D34C8F540006467E5FB82FAC19715" box="[151,1354,1099,1121]" gridrow="20" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FB82FD009715" box="[151,651,1099,1121]" gridcol="0" gridrow="20" pageId="2" pageNumber="442">Width of caudal peduncle</th>
<td id="76CC5DB4F540006465C3FB82FC849715" box="[689,783,1099,1121]" gridcol="1" gridrow="20" pageId="2" pageNumber="442">4.0</td>
<td id="76CC5DB4F540006464AFFB82FAC19715" box="[989,1354,1099,1121]" gridcol="2" gridrow="20" pageId="2" pageNumber="442">4.9 (4.0–6.0)</td>
</tr>
<tr id="351D34C8F540006467E5FBBBFAC197FD" box="[151,1354,1138,1161]" gridrow="21" pageId="2" pageNumber="442" rowspan-1="1">
<th id="76CC5DB4F540006467E5FBBBFD0097FD" box="[151,651,1138,1161]" gridcol="0" gridrow="21" pageId="2" pageNumber="442">
<emphasis id="B959EA98F540FF9967EDFBBAFE2897FD" bold="true" box="[159,419,1138,1161]" pageId="2" pageNumber="442">Morphometrics (% HL)</emphasis>
</th>
<td id="76CC5DB4F540006464AFFBBBFAC197FD" box="[989,1354,1138,1161]" gridcol="2" gridrow="21" pageId="2" pageNumber="442">Mean (range)</td>
</tr>
<tr id="351D34C8F540006467E5FB53FAC197C4" box="[151,1354,1178,1200]" gridrow="22" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FB53FD0097C4" box="[151,651,1178,1200]" gridcol="0" gridrow="22" pageId="2" pageNumber="442">Snout length</th>
<td id="76CC5DB4F540006465C3FB53FC8497C4" box="[689,783,1178,1200]" gridcol="1" gridrow="22" pageId="2" pageNumber="442">24.3</td>
<td id="76CC5DB4F540006464AFFB53FAC197C4" box="[989,1354,1178,1200]" gridcol="2" gridrow="22" pageId="2" pageNumber="442">23.0 (21.2–24.1)</td>
</tr>
<tr id="351D34C8F540006467E5FB08FAC197A3" box="[151,1354,1217,1239]" gridrow="23" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FB08FD0097A3" box="[151,651,1217,1239]" gridcol="0" gridrow="23" pageId="2" pageNumber="442">Eye diameter</th>
<td id="76CC5DB4F540006465C3FB08FC8497A3" box="[689,783,1217,1239]" gridcol="1" gridrow="23" pageId="2" pageNumber="442">16.2</td>
<td id="76CC5DB4F540006464AFFB08FAC197A3" box="[989,1354,1217,1239]" gridcol="2" gridrow="23" pageId="2" pageNumber="442">16.8 (15.5–17.7)</td>
</tr>
<tr id="351D34C8F540006467E5FB21FAC1978A" box="[151,1354,1256,1278]" gridrow="24" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FB21FD00978A" box="[151,651,1256,1278]" gridcol="0" gridrow="24" pageId="2" pageNumber="442">Snout width</th>
<td id="76CC5DB4F540006465C3FB21FC84978A" box="[689,783,1256,1278]" gridcol="1" gridrow="24" pageId="2" pageNumber="442">25.6</td>
<td id="76CC5DB4F540006464AFFB21FAC1978A" box="[989,1354,1256,1278]" gridcol="2" gridrow="24" pageId="2" pageNumber="442">26.4 (23.8–28.5)</td>
</tr>
<tr id="351D34C8F540006467E5FAC6FAC19651" box="[151,1354,1295,1317]" gridrow="25" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FAC6FD009651" box="[151,651,1295,1317]" gridcol="0" gridrow="25" pageId="2" pageNumber="442">Interorbital width</th>
<td id="76CC5DB4F540006465C3FAC6FC849651" box="[689,783,1295,1317]" gridcol="1" gridrow="25" pageId="2" pageNumber="442">21.6</td>
<td id="76CC5DB4F540006464AFFAC6FAC19651" box="[989,1354,1295,1317]" gridcol="2" gridrow="25" pageId="2" pageNumber="442">22.5 (21.0–24.3)</td>
</tr>
<tr id="351D34C8F540006467E5FAFEFAC19639" box="[151,1354,1335,1357]" gridrow="26" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FAFEFD009639" box="[151,651,1335,1357]" gridcol="0" gridrow="26" pageId="2" pageNumber="442">Post orbital length</th>
<td id="76CC5DB4F540006465C3FAFEFC849639" box="[689,783,1335,1357]" gridcol="1" gridrow="26" pageId="2" pageNumber="442">63.5</td>
<td id="76CC5DB4F540006464AFFAFEFAC19639" box="[989,1354,1335,1357]" gridcol="2" gridrow="26" pageId="2" pageNumber="442">64.1 (61.6–66.0)</td>
</tr>
<tr id="351D34C8F540006467E5FA97FAC19600" box="[151,1354,1374,1396]" gridrow="27" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FA97FD009600" box="[151,651,1374,1396]" gridcol="0" gridrow="27" pageId="2" pageNumber="442">Upper jaw length</th>
<td id="76CC5DB4F540006465C3FA97FC849600" box="[689,783,1374,1396]" gridcol="1" gridrow="27" pageId="2" pageNumber="442">67.4</td>
<td id="76CC5DB4F540006464AFFA97FAC19600" box="[989,1354,1374,1396]" gridcol="2" gridrow="27" pageId="2" pageNumber="442">67.8 (64.1–70.3)</td>
</tr>
<tr id="351D34C8F540006467E5FA4CFAC196EF" box="[151,1354,1413,1435]" gridrow="28" pageId="2" pageNumber="442" rowspan-1="1">
<th id="76CC5DB4F540006467E5FA4CFD0096EF" box="[151,651,1413,1435]" gridcol="0" gridrow="28" pageId="2" pageNumber="442">
<emphasis id="B959EA98F540FF9967EDFA4CFE8896EF" bold="true" box="[159,259,1413,1435]" pageId="2" pageNumber="442">Meristics</emphasis>
</th>
<td id="76CC5DB4F540006464AFFA4CFAC196EF" box="[989,1354,1413,1435]" gridcol="2" gridrow="28" pageId="2" pageNumber="442">range</td>
</tr>
<tr id="351D34C8F540006467E5FA65FAC196B6" box="[151,1354,1452,1474]" gridrow="29" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FA65FD0096B6" box="[151,651,1452,1474]" gridcol="0" gridrow="29" pageId="2" pageNumber="442">Dorsal fin</th>
<td id="76CC5DB4F540006465C3FA65FC8496B6" box="[689,783,1452,1474]" gridcol="1" gridrow="29" pageId="2" pageNumber="442">13</td>
<td id="76CC5DB4F540006464AFFA65FAC196B6" box="[989,1354,1452,1474]" gridcol="2" gridrow="29" pageId="2" pageNumber="442">12–13</td>
</tr>
<tr id="351D34C8F540006467E5FA1DFAC1969E" box="[151,1354,1492,1514]" gridrow="30" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FA1DFD00969E" box="[151,651,1492,1514]" gridcol="0" gridrow="30" pageId="2" pageNumber="442">Pectoral fin</th>
<td id="76CC5DB4F540006465C3FA1DFC84969E" box="[689,783,1492,1514]" gridcol="1" gridrow="30" pageId="2" pageNumber="442">14</td>
<td id="76CC5DB4F540006464AFFA1DFAC1969E" box="[989,1354,1492,1514]" gridcol="2" gridrow="30" pageId="2" pageNumber="442">14–15</td>
</tr>
<tr id="351D34C8F540006467E5FA32FAC19565" box="[151,1354,1531,1553]" gridrow="31" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5FA32FD009565" box="[151,651,1531,1553]" gridcol="0" gridrow="31" pageId="2" pageNumber="442">Pelvic fin</th>
<td id="76CC5DB4F540006465C3FA32FC849565" box="[689,783,1531,1553]" gridcol="1" gridrow="31" pageId="2" pageNumber="442">9</td>
<td id="76CC5DB4F540006464AFFA32FAC19565" box="[989,1354,1531,1553]" gridcol="2" gridrow="31" pageId="2" pageNumber="442">9</td>
</tr>
<tr id="351D34C8F540006467E5F9EBFAC1954C" box="[151,1354,1570,1592]" gridrow="32" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F9EBFD00954C" box="[151,651,1570,1592]" gridcol="0" gridrow="32" pageId="2" pageNumber="442">Anal fin</th>
<td id="76CC5DB4F540006465C3F9EBFC84954C" box="[689,783,1570,1592]" gridcol="1" gridrow="32" pageId="2" pageNumber="442">11</td>
<td id="76CC5DB4F540006464AFF9EBFAC1954C" box="[989,1354,1570,1592]" gridcol="2" gridrow="32" pageId="2" pageNumber="442">10–12</td>
</tr>
<tr id="351D34C8F540006467E5F980FAC1952B" box="[151,1354,1609,1631]" gridrow="33" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F980FD00952B" box="[151,651,1609,1631]" gridcol="0" gridrow="33" pageId="2" pageNumber="442">Pored lateral-line scales</th>
<td id="76CC5DB4F540006465C3F980FC84952B" box="[689,783,1609,1631]" gridcol="1" gridrow="33" pageId="2" pageNumber="442">54</td>
<td id="76CC5DB4F540006464AFF980FAC1952B" box="[989,1354,1609,1631]" gridcol="2" gridrow="33" pageId="2" pageNumber="442">53–55</td>
</tr>
<tr id="351D34C8F540006467E5F9B9FAC195F2" box="[151,1354,1648,1670]" gridrow="34" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F9B9FD0095F2" box="[151,651,1648,1670]" gridcol="0" gridrow="34" pageId="2" pageNumber="442">Transverse scales above lateral-line</th>
<td id="76CC5DB4F540006465C3F9B9FC8495F2" box="[689,783,1648,1670]" gridcol="1" gridrow="34" pageId="2" pageNumber="442">4½</td>
<td id="76CC5DB4F540006464AFF9B9FAC195F2" box="[989,1354,1648,1670]" gridcol="2" gridrow="34" pageId="2" pageNumber="442">4½</td>
</tr>
<tr id="351D34C8F540006467E5F95EFAC195D9" box="[151,1354,1687,1709]" gridrow="35" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F95EFD0095D9" box="[151,651,1687,1709]" gridcol="0" gridrow="35" pageId="2" pageNumber="442">Transverse scales below lateral-line</th>
<td id="76CC5DB4F540006465C3F95EFC8495D9" box="[689,783,1687,1709]" gridcol="1" gridrow="35" pageId="2" pageNumber="442">5½</td>
<td id="76CC5DB4F540006464AFF95EFAC195D9" box="[989,1354,1687,1709]" gridcol="2" gridrow="35" pageId="2" pageNumber="442">5½</td>
</tr>
<tr id="351D34C8F540006467E5F976FAC195A1" box="[151,1354,1727,1749]" gridrow="36" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F976FD0095A1" box="[151,651,1727,1749]" gridcol="0" gridrow="36" pageId="2" pageNumber="442">Scale rows below lateral line with melanophores</th>
<td id="76CC5DB4F540006465C3F976FC8495A1" box="[689,783,1727,1749]" gridcol="1" gridrow="36" pageId="2" pageNumber="442">2</td>
<td id="76CC5DB4F540006464AFF976FAC195A1" box="[989,1354,1727,1749]" gridcol="2" gridrow="36" pageId="2" pageNumber="442">1–2</td>
</tr>
<tr id="351D34C8F540006467E5F92FFAC19588" box="[151,1354,1766,1788]" gridrow="37" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F92FFD009588" box="[151,651,1766,1788]" gridcol="0" gridrow="37" pageId="2" pageNumber="442">Pre-dorsal scales</th>
<td id="76CC5DB4F540006465C3F92FFC849588" box="[689,783,1766,1788]" gridcol="1" gridrow="37" pageId="2" pageNumber="442">19</td>
<td id="76CC5DB4F540006464AFF92FFAC19588" box="[989,1354,1766,1788]" gridcol="2" gridrow="37" pageId="2" pageNumber="442">16–20</td>
</tr>
<tr id="351D34C8F540006467E5F8C4FAC19457" box="[151,1354,1805,1827]" gridrow="38" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F8C4FD009457" box="[151,651,1805,1827]" gridcol="0" gridrow="38" pageId="2" pageNumber="442">Pre-adipose scales</th>
<td id="76CC5DB4F540006465C3F8C4FC849457" box="[689,783,1805,1827]" gridcol="1" gridrow="38" pageId="2" pageNumber="442">16</td>
<td id="76CC5DB4F540006464AFF8C4FAC19457" box="[989,1354,1805,1827]" gridcol="2" gridrow="38" pageId="2" pageNumber="442">15–18</td>
</tr>
<tr id="351D34C8F540006467E5F8FDFAC1943E" box="[151,1354,1844,1866]" gridrow="39" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F8FDFD00943E" box="[151,651,1844,1866]" gridcol="0" gridrow="39" pageId="2" pageNumber="442">Post-adipose scales</th>
<td id="76CC5DB4F540006465C3F8FDFC84943E" box="[689,783,1844,1866]" gridcol="1" gridrow="39" pageId="2" pageNumber="442">10</td>
<td id="76CC5DB4F540006464AFF8FDFAC1943E" box="[989,1354,1844,1866]" gridcol="2" gridrow="39" pageId="2" pageNumber="442">11–12</td>
</tr>
<tr id="351D34C8F540006467E5F895FAC19406" box="[151,1354,1884,1906]" gridrow="40" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F895FD009406" box="[151,651,1884,1906]" gridcol="0" gridrow="40" pageId="2" pageNumber="442">No. vertebrae</th>
<td id="76CC5DB4F540006465C3F895FC849406" box="[689,783,1884,1906]" gridcol="1" gridrow="40" pageId="2" pageNumber="442">-</td>
<td id="76CC5DB4F540006464AFF895FAC19406" box="[989,1354,1884,1906]" gridcol="2" gridrow="40" pageId="2" pageNumber="442">49–50</td>
</tr>
<tr id="351D34C8F540006467E5F84AFAC194ED" box="[151,1354,1923,1945]" gridrow="41" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F84AFD0094ED" box="[151,651,1923,1945]" gridcol="0" gridrow="41" pageId="2" pageNumber="442">No. rows of palatine teeth anteriorly</th>
<td id="76CC5DB4F540006465C3F84AFC8494ED" box="[689,783,1923,1945]" gridcol="1" gridrow="41" pageId="2" pageNumber="442">2</td>
<td id="76CC5DB4F540006464AFF84AFAC194ED" box="[989,1354,1923,1945]" gridcol="2" gridrow="41" pageId="2" pageNumber="442">2</td>
</tr>
<tr id="351D34C8F540006467E5F863FAC194B4" box="[151,1354,1962,1984]" gridrow="42" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F863FD0094B4" box="[151,651,1962,1984]" gridcol="0" gridrow="42" pageId="2" pageNumber="442">No. of vomerine teeth</th>
<td id="76CC5DB4F540006465C3F863FC8494B4" box="[689,783,1962,1984]" gridcol="1" gridrow="42" pageId="2" pageNumber="442">2</td>
<td id="76CC5DB4F540006464AFF863FAC194B4" box="[989,1354,1962,1984]" gridcol="2" gridrow="42" pageId="2" pageNumber="442">0–3</td>
</tr>
<tr id="351D34C8F540006467E5F818FAC19493" box="[151,1354,2001,2023]" gridrow="43" pageId="2" pageNumber="442">
<th id="76CC5DB4F540006467E5F818FD009493" box="[151,651,2001,2023]" gridcol="0" gridrow="43" pageId="2" pageNumber="442">No. teeth rows across tongue</th>
<td id="76CC5DB4F540006465C3F818FC849493" box="[689,783,2001,2023]" gridcol="1" gridrow="43" pageId="2" pageNumber="442">4</td>
<td id="76CC5DB4F540006464AFF818FAC19493" box="[989,1354,2001,2023]" gridcol="2" gridrow="43" pageId="2" pageNumber="442">4–5</td>
</tr>
</table>
</paragraph>
<caption id="DF526602F541FF9867E5FA1CFBB5957E" httpUri="https://zenodo.org/record/242542/files/figure.png" pageId="3" pageNumber="443" targetBox="[343,1231,193,1471]" targetPageId="3">
<paragraph id="8B92368AF541FF9867E5FA1CFBB5957E" blockId="3.[151,1435,1493,1547]" pageId="3" pageNumber="443">
<emphasis id="B959EA98F541FF9867E5FA1CFE98969F" bold="true" box="[151,275,1493,1515]" pageId="3" pageNumber="443">FIGURE 1.</emphasis>
Map of Mascarene Plateau showing location and station numbers at which 
<taxonomicName id="4C2D4D09F541FF986363FA1CFB35969F" box="[1041,1214,1493,1515]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="3" pageNumber="443" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F541FF986363FA1CFB35969F" box="[1041,1214,1493,1515]" italics="true" pageId="3" pageNumber="443">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F541FF9863B6FA1FFB7C969F" box="[1220,1271,1494,1515]" pageId="3" pageNumber="443" rank="species">
<emphasis id="B959EA98F541FF9863B6FA1FFB7C969F" bold="true" box="[1220,1271,1494,1515]" pageId="3" pageNumber="443">n.sp.</emphasis>
</taxonomicNameLabel>
were caught: + = RV 
<emphasis id="B959EA98F541FF9867A1FA3CFE1D957E" box="[211,406,1524,1546]" italics="true" pageId="3" pageNumber="443">Dr Fridtjof Nansen</emphasis>
stations, with station number; ★ = FV 
<emphasis id="B959EA98F541FF986455FA3CFC6F957E" box="[807,996,1524,1546]" italics="true" pageId="3" pageNumber="443">Ryuyo-Maru No. 2</emphasis>
stations.
</paragraph>
</caption>
<caption id="DF526602F541FF9867E5F874FAFE94A7" box="[151,1397,1981,2003]" httpUri="https://zenodo.org/record/242543/files/figure.png" pageId="3" pageNumber="443" targetBox="[233,1353,1596,1959]" targetPageId="3">
<paragraph id="8B92368AF541FF9867E5F874FAFE94A7" blockId="3.[151,1397,1981,2003]" box="[151,1397,1981,2003]" pageId="3" pageNumber="443">
<emphasis id="B959EA98F541FF9867E5F874FE9994A7" bold="true" box="[151,274,1981,2003]" pageId="3" pageNumber="443">FIGURE 2.</emphasis>
Holotype of 
<taxonomicName id="4C2D4D09F541FF9866EBF874FDCE94A7" box="[409,581,1981,2003]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="3" pageNumber="443" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F541FF9866EBF874FDCE94A7" box="[409,581,1981,2003]" italics="true" pageId="3" pageNumber="443">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F541FF986539F877FDF594A7" box="[587,638,1982,2003]" pageId="3" pageNumber="443" rank="species">
<emphasis id="B959EA98F541FF986539F877FDF594A7" bold="true" box="[587,638,1982,2003]" pageId="3" pageNumber="443">n.sp.</emphasis>
</taxonomicNameLabel>
, SAIAB 185941, 354.0 mm SL, Saya da Malha Bank (photo: O. Alvheim)
</paragraph>
</caption>
<caption id="DF526602F546FF9F67E5FD6DFEAD91AC" httpUri="https://zenodo.org/record/242544/files/figure.png" pageId="4" pageNumber="444" targetBox="[176,1411,193,654]" targetPageId="4">
<paragraph id="8B92368AF546FF9F67E5FD6DFEAD91AC" blockId="4.[151,1436,676,728]" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67E5FD6DFE9D91CD" bold="true" box="[151,278,676,698]" pageId="4" pageNumber="444">FIGURE 3.</emphasis>
Photograph of ‘ 
<emphasis id="B959EA98F546FF9F66B4FD6DFD2591CE" box="[454,686,676,698]" italics="true" pageId="4" pageNumber="444">
<taxonomicName id="4C2D4D09F546FF9F66B4FD6DFD9D91CE" box="[454,534,676,698]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="4" pageNumber="444" phylum="Chordata" rank="genus">Saurida</taxonomicName>
undosquamis’
</emphasis>
from Saya da Malha Bank (Amaoka 
<emphasis id="B959EA98F546FF9F6336FD6CFBF091CE" box="[1092,1147,676,698]" italics="true" pageId="4" pageNumber="444">et al.</emphasis>
1976) now identified as 
<taxonomicName id="4C2D4D09F546FF9F62FBFD6CFF6691AC" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="4" pageNumber="444" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F546FF9F62FBFD6CFF6691AC" italics="true" pageId="4" pageNumber="444">S. tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F546FF9F6781FD0AFEAD91AC" box="[243,294,707,728]" pageId="4" pageNumber="444" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B92368AF546FF9F67B5FCC8FDFD9762" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">Mouth very large, the gape extending well beyond posterior margin of eye. Teeth on jaws numerous, small, canine-like, generally in four rows, outermost teeth smallest, innermost longest, visible when mouth closed; similar teeth on the palate, tongue and branchial arches; palatine teeth in two separate series, teeth of outer series long, in two rows, with anterior teeth longer, teeth of inner series short, in about six poorly defined rows; outer palatine series converge anteriorly but separated by vomer which is toothless or bears 1–3 teeth; tongue short, spatulate, fleshy, with triangular shaped patch of teeth, in about 4 (4–5) rows posteriorly; lower jaw slightly shorter than upper jaw and fits into groove between teeth at tip of upper jaw; gill rakers absent, branchial arches each with about 4 rows of teeth, those on the inside largest.</paragraph>
<paragraph id="8B92368AF546FF9F67B5FBE8FB1E97BE" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">Origin of dorsal fin just posterior to midpoint of distance from snout to adipose-fin origin; base of dorsal fin longer than that of anal fin; adipose fin above posterior half of anal fin; anus just anterior to origin of anal fin; pectoral fins moderately long, 13.0% (12.6–17.4%) of SL, tip of pectoral fin reaching to or just beyond a line from base of pelvic fin to origin of dorsal fin (P-D line); pectoral and pelvic fins with long and pointed axillary scales, those of pelvic a little shorter; caudal fin deeply forked, upper and lower lobes about equal.</paragraph>
<paragraph id="8B92368AF546FF9F67B5FB1CFA1F979A" blockId="4.[151,1437,769,2018]" box="[199,1428,1237,1262]" pageId="4" pageNumber="444">Peritoneum pale whitish; stomach pale (slightly greyish anteriorly in a few specimens), intestine pale whitish.</paragraph>
<paragraph id="8B92368AF546FF9F67B5FB30FBAA960A" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67B5FB30FE109665" box="[199,411,1273,1297]" italics="true" pageId="4" pageNumber="444">Colour in alcohol.</emphasis>
Brown on back, paler below lateral line, pigment extending to about 2 (1–2) scale rows below lateral line; pectoral fins dusky over outer two-thirds of fin; pelvic fins pale; dorsal, caudal and anal fins pale, anterior margin and posterior edge of dorsal and caudal fins dusky; leading edge of caudal fin with 10 (8–16) blackish spots; adipose fin pale or dusky over much of fin or along outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C3376501F546FF9F67B5FA40FC469546" pageId="4" pageNumber="444" type="materials_examined">
<paragraph id="8B92368AF546FF9F67B5FA40FC469546" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67B5FA40FE7296D5" box="[199,505,1417,1441]" italics="true" pageId="4" pageNumber="444">Colour of fresh specimens</emphasis>
(
<figureCitation id="13162A0FF546FF9F657FFA40FDDD96D6" box="[525,598,1417,1442]" captionStart="FIGURE 2" captionStartId="3.[151,250,1981,2003]" captionTargetBox="[233,1353,1596,1959]" captionTargetId="figure@3.[233,1354,1596,1960]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Saurida tweddlei n. sp., SAIAB 185941, 354.0 mm SL, Saya da Malha Bank (photo: O. Alvheim)" httpUri="https://zenodo.org/record/242543/files/figure.png" pageId="4" pageNumber="444">Fig. 2</figureCitation>
, 
<figureCitation id="13162A0FF546FF9F6514FA43FDFE96D6" box="[614,629,1418,1442]" captionStart="FIGURE 3" captionStartId="4.[151,250,676,698]" captionTargetBox="[176,1411,193,654]" captionTargetId="figure@4.[176,1412,193,654]" captionTargetPageId="4" captionText="FIGURE 3. Photograph of ‘ Saurida undosquamis’ from Saya da Malha Bank (Amaoka et al. 1976) now identified as S. tweddlei n. sp." httpUri="https://zenodo.org/record/242544/files/figure.png" pageId="4" pageNumber="444">3</figureCitation>
). Top of head and back dark brown or coppery brown, some scales narrowly edged with black, forming a variegated pattern; silvery white below lateral line; leading edge and outer margin of dorsal fin light dusky; adipose fin light dusky over most of fin; leading edge of caudal fin with 8–16 blackish spots, posterior edge and lower lobe of caudal fin dusky; pectoral fins dusky, base of fin translucent; pelvic fins pale lemon at base, rest of fin translucent; anal fin translucent.
</paragraph>
</subSubSection>
<subSubSection id="C3376501F546FF9F67B5F9F4FDC9945A" pageId="4" pageNumber="444" type="distribution">
<paragraph id="8B92368AF546FF9F67B5F9F4FDC9945A" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67B5F9F4FDAC9522" bold="true" box="[199,551,1597,1622]" pageId="4" pageNumber="444">Distribution and abundance.</emphasis>
<taxonomicName id="4C2D4D09F546FF9F6540F9F4FD7E9521" box="[562,757,1597,1621]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="4" pageNumber="444" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F546FF9F6540F9F4FD7E9521" box="[562,757,1597,1621]" italics="true" pageId="4" pageNumber="444">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F546FF9F6473F9F4FCB39522" box="[769,824,1597,1622]" pageId="4" pageNumber="444" rank="species">
<emphasis id="B959EA98F546FF9F6473F9F4FCB39522" bold="true" box="[769,824,1597,1622]" pageId="4" pageNumber="444">n.sp.</emphasis>
</taxonomicNameLabel>
is so far known only from the northern part of the Mascarene Plateau in the Indian Ocean, from 
<collectingCountry id="F33A761AF546FF9F65CEF9A8FCBF950E" box="[700,820,1633,1658]" name="Seychelles" pageId="4" pageNumber="444">Seychelles</collectingCountry>
Bank and Saya de Malha Bank (
<figureCitation id="13162A0FF546FF9F63B5F9A8FA85950E" box="[1223,1294,1633,1658]" captionStart="FIGURE 1" captionStartId="3.[151,250,1493,1515]" captionTargetBox="[343,1231,193,1471]" captionTargetId="figure@3.[334,1242,183,1481]" captionTargetPageId="3" captionText="FIGURE 1. Map of Mascarene Plateau showing location and station numbers at which Saurida tweddlei n. sp. were caught: + = RV Dr Fridtjof Nansen stations, with station number; ★ = FV Ryuyo-Maru No. 2 stations." httpUri="https://zenodo.org/record/242542/files/figure.png" pageId="4" pageNumber="444">Fig. 1</figureCitation>
). The typespecimens collected by RV 
<emphasis id="B959EA98F546FF9F66A2F94FFD2195E9" box="[464,682,1669,1693]" italics="true" pageId="4" pageNumber="444">Dr Fridtjof Nansen</emphasis>
at station 19 were trawled in depths of 
<quantity id="4CD59B6FF546FF9F631EF94FFB3D95E9" box="[1132,1206,1670,1694]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="444" unit="m" value="127.0">127 m</quantity>
, but the species was also caught at other stations (
<figureCitation id="13162A0FF546FF9F6691F960FDAF95B6" box="[483,548,1705,1730]" captionStart="FIGURE 1" captionStartId="3.[151,250,1493,1515]" captionTargetBox="[343,1231,193,1471]" captionTargetId="figure@3.[334,1242,183,1481]" captionTargetPageId="3" captionText="FIGURE 1. Map of Mascarene Plateau showing location and station numbers at which Saurida tweddlei n. sp. were caught: + = RV Dr Fridtjof Nansen stations, with station number; ★ = FV Ryuyo-Maru No. 2 stations." httpUri="https://zenodo.org/record/242542/files/figure.png" pageId="4" pageNumber="444">Fig. 1</figureCitation>
) in depths ranging from 
<quantity id="4CD59B6FF546FF9F6431F960FC2195B5" box="[835,938,1705,1730]" metricMagnitude="1" metricUnit="m" metricValue="6.65" metricValueMax="7.6" metricValueMin="5.7" pageId="4" pageNumber="444" unit="m" value="66.5" valueMax="76.0" valueMin="57.0">
57– 
<specimenCount id="9D2BFD03F546FF9F641FF960FC2195B5" box="[877,938,1705,1730]" pageId="4" pageNumber="444" type="generic">76 m</specimenCount>
</quantity>
. This species was one of the most abundant trawl species on the northern Mascarene Plateau, comprising 22.5–61.2% (mean 36%, n = 3) of the total catch weight on the Saya de Malha Bank, and 0.5–41.4% (mean 18.8%, n = 6) of the total catch weight on the 
<collectingCountry id="F33A761AF546FF9F6256F938FA17947E" box="[1316,1436,1777,1802]" name="Seychelles" pageId="4" pageNumber="444">Seychelles</collectingCountry>
Bank (D. Tweddle, unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C3376501F546FF9E67B5F8F0FA1F90D1" lastPageId="5" lastPageNumber="445" pageId="4" pageNumber="444" type="etymology">
<paragraph id="8B92368AF546FF9F67B5F8F0FE3F94CA" blockId="4.[151,1437,769,2018]" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67B5F8F0FECD9426" bold="true" box="[199,326,1849,1874]" pageId="4" pageNumber="444">Etymology</emphasis>
. Named in honour of Denis Tweddle, Honorary Research Associate at SAIAB, who collected this species during an exploratory fishing survey of the Mascarene Plateau by the RV 
<emphasis id="B959EA98F546FF9F6339F897FAA29401" box="[1099,1321,1885,1909]" italics="true" pageId="4" pageNumber="444">Dr Fridtjof Nansen</emphasis>
, and who brought it to the attention of the author during a visit to SAIAB. The common English name Tweddle’s lizardfish is proposed for this species.
</paragraph>
<paragraph id="8B92368AF546FF9E67B5F800FB3E93A1" blockId="4.[151,1437,769,2018]" lastBlockId="5.[151,1437,151,933]" lastPageId="5" lastPageNumber="445" pageId="4" pageNumber="444">
<emphasis id="B959EA98F546FF9F67B5F800FED19496" bold="true" box="[199,346,1993,2018]" pageId="4" pageNumber="444">Comparison</emphasis>
. This species was previously reported from the 
<collectingCountry id="F33A761AF546FF9F6409F800FC789496" box="[891,1011,1993,2018]" name="Seychelles" pageId="4" pageNumber="444">Seychelles</collectingCountry>
by 
<bibRefCitation id="EFBC4B7BF546FF9F636CF800FA809496" author="Amaoka" box="[1054,1291,1993,2018]" pageId="4" pageNumber="444" refString="Amaoka, K., Hatanaka, H., Ida, H., Ikeda, I., Inada, T., Iwai, T., Kawahara, S., Kono, H., Kuronuma, K., Matsuura, K., Miyake, S., Nakamura, I., Ochiai, A., Okamura, O., Okutani, T., Sasaki, T., Sato, T. (Tetsuya), Sato, T. (Torao), Shimizu, T., Takeda, M., Taniuchi, T., Tominaga, Y. &amp; Uyeno, T. (Eds.) (1976) Colored Illustrations of Bottomfishes Collected by Japanese Trawlers. Vol. II. Japan Deep Sea Trawlers Association, Tokyo, 188 pp." type="book" year="1976">
Amaoka 
<emphasis id="B959EA98F546FF9F63F7F803FB359495" box="[1157,1214,1993,2017]" italics="true" pageId="4" pageNumber="444">et al.</emphasis>
(1976
</bibRefCitation>
: p.115, Plate Sey-1) misidentified as 
<taxonomicName id="4C2D4D09F547FF9E66D9FF51FCC993C4" authority="Richardson" authorityName="Richardson" box="[427,834,151,176]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E66D9FF51FD2993C4" box="[427,674,152,176]" italics="true" pageId="5" pageNumber="445">Saurida undosquamis</emphasis>
(Richardson)
</taxonomicName>
. Although the specimen could not be located in the NSMT and is presumed lost, the photograph (
<figureCitation id="13162A0FF547FF9E65E5FF75FD5293A0" box="[663,729,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,676,698]" captionTargetBox="[176,1411,193,654]" captionTargetId="figure@4.[176,1412,193,654]" captionTargetPageId="4" captionText="FIGURE 3. Photograph of ‘ Saurida undosquamis’ from Saya da Malha Bank (Amaoka et al. 1976) now identified as S. tweddlei n. sp." httpUri="https://zenodo.org/record/242544/files/figure.png" pageId="5" pageNumber="445">Fig. 3</figureCitation>
) is clearly identifiable as 
<taxonomicName id="4C2D4D09F547FF9E6489FF75FBFD93A0" box="[1019,1142,188,212]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F547FF9E6489FF75FBFD93A0" box="[1019,1142,188,212]" italics="true" pageId="5" pageNumber="445">S. tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F547FF9E630FFF75FB3E93A1" box="[1149,1205,188,213]" pageId="5" pageNumber="445" rank="species">
<emphasis id="B959EA98F547FF9E630FFF75FB3E93A1" bold="true" box="[1149,1205,188,213]" pageId="5" pageNumber="445">n.sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8B92368AF547FF9E67B5FF29FD0491F1" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="445">
<bibRefCitation id="EFBC4B7BF547FF9E67B5FF29FE55938C" author="Inoue" box="[199,478,223,248]" pageId="5" pageNumber="445" refString="Inoue, T. &amp; Nakabo, T. (2006) The Saurida undosquamis group (Aulopiformes: Synodontidae), with description of a new species from southern Japan. Ichthyological research, 53 (4), 379 - 397. http: // dx. doi. org / 10.1007 / s 10228 - 006 - 0358 - y" type="journal article" year="2006">Inoue &amp; Nakabo (2006)</bibRefCitation>
, in a review of the 
<taxonomicName id="4C2D4D09F547FF9E65B7FF29FC37938C" box="[709,956,224,248]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E65B7FF29FC37938C" box="[709,956,224,248]" italics="true" pageId="5" pageNumber="445">Saurida undosquamis</emphasis>
</taxonomicName>
group, showed the name 
<taxonomicName id="4C2D4D09F547FF9E639BFF29FA17938C" box="[1257,1436,224,248]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E639BFF29FA17938C" box="[1257,1436,224,248]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
(type-locality north-western 
<collectingCountry id="F33A761AF547FF9E66AAFECDFDCA9269" box="[472,577,260,285]" name="Australia" pageId="5" pageNumber="445">Australia</collectingCountry>
) has been applied to a complex of species distributed widely throughout the Indo-West Pacific. They recognised four species in the group (
<taxonomicName id="4C2D4D09F547FF9E6401FEE1FBAD9234" box="[883,1062,296,320]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E6401FEE1FBAD9234" box="[883,1062,296,320]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C2D4D09F547FF9E634AFEE1FF589211" authority="Inoue &amp; Nakabo, 2006" authorityName="Inoue &amp; Nakabo" authorityYear="2006" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="umeyoshii">
<emphasis id="B959EA98F547FF9E634AFEE1FB429234" box="[1080,1225,296,320]" italics="true" pageId="5" pageNumber="445">S. umeyoshii</emphasis>
<bibRefCitation id="EFBC4B7BF547FF9E63A1FEE1FF589211" author="Inoue" pageId="5" pageNumber="445" refString="Inoue, T. &amp; Nakabo, T. (2006) The Saurida undosquamis group (Aulopiformes: Synodontidae), with description of a new species from southern Japan. Ichthyological research, 53 (4), 379 - 397. http: // dx. doi. org / 10.1007 / s 10228 - 006 - 0358 - y" type="journal article" year="2006">Inoue &amp; Nakabo, 2006</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C2D4D09F547FF9E6792FE85FD959210" authority="Tanaka, 1917" authorityName="Tanaka" authorityYear="1917" box="[224,542,332,357]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B959EA98F547FF9E6792FE85FEF29210" box="[224,377,332,356]" italics="true" pageId="5" pageNumber="445">S. macrolepis</emphasis>
<bibRefCitation id="EFBC4B7BF547FF9E66F1FE85FD959210" author="Tanaka" box="[387,542,332,357]" pageId="5" pageNumber="445" refString="Tanaka, S. (1917) Six new species of Japanese fishes. Dobutsugaku Zasshi, 29 (340), 37 - 40. [in Japanese]." type="journal article" year="1917">Tanaka, 1917</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C2D4D09F547FF9E652CFE85FC3B9210" authority="Norman, 1939" authorityName="Norman" authorityYear="1939" box="[606,944,332,356]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="B959EA98F547FF9E652CFE85FD749210" box="[606,767,332,356]" italics="true" pageId="5" pageNumber="445">S. longimanus</emphasis>
<bibRefCitation id="EFBC4B7BF547FF9E647AFE84FC3B9210" author="Norman" box="[776,944,332,356]" pageId="5" pageNumber="445" refString="Norman, J. R. (1939) Fishes. The John Murray Expedition 1933 - 34. Scientific Reports, John Murray Expedition, 7 (1), 1 - 116." type="journal article" year="1939">Norman, 1939</bibRefCitation>
</taxonomicName>
). 
<taxonomicName id="4C2D4D09F547FF9E64B7FE85FB289210" box="[965,1187,332,356]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B959EA98F547FF9E64B7FE85FB289210" box="[965,1187,332,356]" italics="true" pageId="5" pageNumber="445">Saurida macrolepis</emphasis>
</taxonomicName>
(type-locality 
<collectingCountry id="F33A761AF547FF9E623EFE85FA1B9210" box="[1356,1424,332,356]" name="Japan" pageId="5" pageNumber="445">Japan</collectingCountry>
), formerly synonymised with 
<taxonomicName id="4C2D4D09F547FF9E66AEFEB9FD0492FC" box="[476,655,368,392]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E66AEFEB9FD0492FC" box="[476,655,368,392]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFBC4B7BF547FF9E65CCFEB9FCFB92FC" author="Norman" box="[702,880,367,392]" pageId="5" pageNumber="445" refString="Norman, J. R. (1935) A revision of the Lizard-fishes of the genera Synodus, Trachinocephalus, and Saurida. Proceedings of the Zoological Society of London, 1935 (1), 99 - 135. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1935. tb 06233. x" type="journal article" year="1935">Norman (1935)</bibRefCitation>
, was recognised as distinct by 
<bibRefCitation id="EFBC4B7BF547FF9E63A9FEB9FF6F92D9" author="Inoue" pageId="5" pageNumber="445" refString="Inoue, T. &amp; Nakabo, T. (2006) The Saurida undosquamis group (Aulopiformes: Synodontidae), with description of a new species from southern Japan. Ichthyological research, 53 (4), 379 - 397. http: // dx. doi. org / 10.1007 / s 10228 - 006 - 0358 - y" type="journal article" year="2006">Inoue &amp; Nakabo (2006)</bibRefCitation>
, who applied the name to the species previously recorded as 
<taxonomicName id="4C2D4D09F547FF9E64E8FE5DFBC092D8" box="[922,1099,404,428]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E64E8FE5DFBC092D8" box="[922,1099,404,428]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
in the Western Indian Ocean. 
<bibRefCitation id="EFBC4B7BF547FF9E67E5FE71FE2F92A4" author="Inoue" box="[151,420,439,464]" pageId="5" pageNumber="445" refString="Inoue, T. &amp; Nakabo, T. (2006) The Saurida undosquamis group (Aulopiformes: Synodontidae), with description of a new species from southern Japan. Ichthyological research, 53 (4), 379 - 397. http: // dx. doi. org / 10.1007 / s 10228 - 006 - 0358 - y" type="journal article" year="2006">Inoue &amp; Nakabo (2006)</bibRefCitation>
considered 
<taxonomicName id="4C2D4D09F547FF9E655EFE71FD4E92A4" box="[556,709,440,464]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B959EA98F547FF9E655EFE71FD4E92A4" box="[556,709,440,464]" italics="true" pageId="5" pageNumber="445">S. macrolepis</emphasis>
</taxonomicName>
to be a wide-ranging, geographically variable species, occurring in the Western Pacific from southern 
<collectingCountry id="F33A761AF547FF9E6545FE14FDFC9280" box="[567,631,477,500]" name="Japan" pageId="5" pageNumber="445">Japan</collectingCountry>
to 
<collectingCountry id="F33A761AF547FF9E65E8FE15FC889281" box="[666,771,476,501]" name="Australia" pageId="5" pageNumber="445">Australia</collectingCountry>
, and in the Indian Ocean westwards to East Africa, the Red Sea and Persian Gulf. However, examination of specimens from the Red Sea-Mediterranean shows they are morphologically and genetically distinct from 
<taxonomicName id="4C2D4D09F547FF9E65EFFDEDFCBF9148" box="[669,820,548,572]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B959EA98F547FF9E65EFFDEDFCBF9148" box="[669,820,548,572]" italics="true" pageId="5" pageNumber="445">S. macrolepis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F33A761AF547FF9E640AFDECFC309148" box="[888,955,549,572]" name="Japan" pageId="5" pageNumber="445">Japan</collectingCountry>
, and represent a new species (Russell 
<emphasis id="B959EA98F547FF9E6216FDECFA179148" box="[1380,1436,548,572]" italics="true" pageId="5" pageNumber="445">et al.</emphasis>
in press). 
<taxonomicName id="4C2D4D09F547FF9E6676FD81FCC59114" authority="Inoue &amp; Nakabo (2006)" authorityName="Inoue &amp; Nakabo" authorityYear="2006" box="[260,846,583,608]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E6676FD81FDB79114" box="[260,572,584,608]" italics="true" pageId="5" pageNumber="445">Saurida undosquamis sensu</emphasis>
<bibRefCitation id="EFBC4B7BF547FF9E6531FD81FCC59114" author="Inoue" box="[579,846,583,608]" pageId="5" pageNumber="445" refString="Inoue, T. &amp; Nakabo, T. (2006) The Saurida undosquamis group (Aulopiformes: Synodontidae), with description of a new species from southern Japan. Ichthyological research, 53 (4), 379 - 397. http: // dx. doi. org / 10.1007 / s 10228 - 006 - 0358 - y" type="journal article" year="2006">Inoue &amp; Nakabo (2006)</bibRefCitation>
</taxonomicName>
also appears to comprise more than one species, and is the subject of ongoing study by the author.
</paragraph>
<paragraph id="8B92368AF547FF9E67B5FD59FBB690F4" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="445">
<taxonomicName id="4C2D4D09F547FF9E67B5FD59FE0D91DC" box="[199,390,656,680]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F547FF9E67B5FD59FE0D91DC" box="[199,390,656,680]" italics="true" pageId="5" pageNumber="445">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F547FF9E66FFFD46FE4E91DC" box="[397,453,655,680]" pageId="5" pageNumber="445" rank="species">
<emphasis id="B959EA98F547FF9E66FFFD46FE4E91DC" bold="true" box="[397,453,655,680]" pageId="5" pageNumber="445">n.sp.</emphasis>
</taxonomicNameLabel>
differs from the species in the Red Sea-Mediterranean misidentified as 
<taxonomicName id="4C2D4D09F547FF9E639EFD59FA1791DC" box="[1260,1436,656,680]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E639EFD59FA1791DC" box="[1260,1436,656,680]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2D4D09F547FF9E67B4FD7DFED691B8" box="[198,349,692,716]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B959EA98F547FF9E67B4FD7DFED691B8" box="[198,349,692,716]" italics="true" pageId="5" pageNumber="445">S. macrolepis</emphasis>
</taxonomicName>
, most notably in having a greater number of lateral line scales (53–55 versus 
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<emphasis id="B959EA98F547FF9E625CFD7DFAC891BF" box="[1326,1347,692,715]" italics="true" pageId="5" pageNumber="445">S.</emphasis>
sp) and vertebrae (49–50 versus 
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<emphasis id="B959EA98F547FF9E6566FD11FDA9919B" box="[532,546,728,751]" italics="true" pageId="5" pageNumber="445">S</emphasis>
. sp). It also differs from 
<emphasis id="B959EA98F547FF9E644BFD11FBF6919B" box="[825,1149,728,752]" italics="true" pageId="5" pageNumber="445">
<taxonomicName id="4C2D4D09F547FF9E644BFD11FBA59184" box="[825,1070,728,752]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="subSpecies" species="undosquamis" subSpecies="sensu">
S. undosquamis 
<taxonomicNameLabel id="A26A57E3F547FF9E6482FD10FBA59184" box="[1008,1070,729,752]" pageId="5" pageNumber="445" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
in having longer pectoral fins (extending to well beyond P-D line, versus pectoral fins shorter, not reaching beyond P-D line in 
<taxonomicName id="4C2D4D09F547FF9E62F5FD35FEA0904C" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E62F5FD35FEA0904C" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
); lateral line scales 53–55 (versus 
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<taxonomicName id="4C2D4D09F547FF9E645CFCE9FC6B904C" box="[814,992,800,824]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E645CFCE9FC6B904C" box="[814,992,800,824]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
); teeth on tongue moderately large, in about 4–5 rows (versus teeth on tongue small, in about 7–12 rows in 
<taxonomicName id="4C2D4D09F547FF9E64DDFC8DFBEB9028" box="[943,1120,836,860]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E64DDFC8DFBEB9028" box="[943,1120,836,860]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
); and 8–16 (usually 11–14) blackish spots on leading edge of caudal fin (versus 5–12 spots in 
<taxonomicName id="4C2D4D09F547FF9E640EFCA1FBA690F4" box="[892,1069,872,896]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="undosquamis">
<emphasis id="B959EA98F547FF9E640EFCA1FBA690F4" box="[892,1069,872,896]" italics="true" pageId="5" pageNumber="445">S. undosquamis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B92368AF547FF9E67B5FC45FA1F90D1" blockId="5.[151,1437,151,933]" box="[199,1428,908,933]" pageId="5" pageNumber="445">
<taxonomicName id="4C2D4D09F547FF9E67B5FC45FE0E90D0" box="[199,389,908,932]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="tweddlei" status="sp. nov.">
<emphasis id="B959EA98F547FF9E67B5FC45FE0E90D0" box="[199,389,908,932]" italics="true" pageId="5" pageNumber="445">Saurida tweddlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57E3F547FF9E66FEFC45FE4F90D1" box="[396,452,908,933]" pageId="5" pageNumber="445" rank="species">
<emphasis id="B959EA98F547FF9E66FEFC45FE4F90D1" bold="true" box="[396,452,908,933]" pageId="5" pageNumber="445">n.sp.</emphasis>
</taxonomicNameLabel>
is distinguished from other Western Indian Ocean species of 
<taxonomicName id="4C2D4D09F547FF9E6306FC45FB4690D0" box="[1140,1229,908,932]" class="Actinopterygii" family="Synodontidae" genus="Saurida" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="445" phylum="Chordata" rank="genus">
<emphasis id="B959EA98F547FF9E6306FC45FB4690D0" box="[1140,1229,908,932]" italics="true" pageId="5" pageNumber="445">Saurida</emphasis>
</taxonomicName>
in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>