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<document id="475338C9DD3822703FF94F536EF4B3D1" ID-CLB-Dataset="27906" ID-DOI="10.11646/zootaxa.4525.1.1" ID-GBIF-Dataset="aab5d525-1d70-4a48-9469-43d4e446685d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610805" ID-ZooBank="9D378750-982F-4061-A419-B28E8DDFF825" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553626294178" checkinUser="plazi" docAuthor="E. B, Neo, Adrain, Jonathan M. &amp; Karim, Talia S." docDate="2018" docId="0384871CFFA5505CFF77FD7EFCBAF9C2" docLanguage="en" docName="zootaxa.4525.1.1.pdf" docOrigin="Zootaxa 4525 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Deltapliomera E.B &amp; Adrain &amp; Karim 2018, n. gen." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFBDFF64FFB3504AFFE0FF82FF8BFFE8" masterDocTitle="The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA" masterLastPageNumber="152" masterPageNumber="1" pageNumber="23" updateTime="1698714183241" updateUser="ExternalLinkService">
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<mods:title id="A1BA6A4D3998ACE1559D526D32A8BBBA">The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA</mods:title>
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<mods:namePart id="91322B7C38524DBF3AE961EAEAB62B9E">E. B, Neo</mods:namePart>
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<mods:namePart id="0E70FEB7851F8AD570667A0A43921882">Adrain, Jonathan M.</mods:namePart>
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<mods:namePart id="CBBAF9E23D7AE80839E275775655C139">Karim, Talia S.</mods:namePart>
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<treatment id="0384871CFFA5505CFF77FD7EFCBAF9C2" ID-DOI="http://doi.org/10.5281/zenodo.5969625" ID-GBIF-Taxon="154499792" ID-Zenodo-Dep="5969625" LSID="urn:lsid:plazi:treatment:0384871CFFA5505CFF77FD7EFCBAF9C2" httpUri="http://treatment.plazi.org/id/0384871CFFA5505CFF77FD7EFCBAF9C2" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C3376581FFA5505CFF77FD7EFE17FCFE" box="[151,412,764,790]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8B92360AFFA5505CFF77FD7EFE17FCFE" blockId="22.[151,412,764,790]" box="[151,412,764,790]" pageId="22" pageNumber="23">
<heading id="D0DA8166FFA5505CFF77FD7EFE17FCFE" bold="true" box="[151,412,764,790]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B959EA18FFA5505CFF77FD7EFE17FCFE" bold="true" box="[151,412,764,790]" pageId="22" pageNumber="23">
<taxonomicName id="4C2D4D89FFA5505CFF77FD7EFEC8FCFE" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[151,323,764,790]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EA18FFA5505CFF77FD7EFEC8FCFE" bold="true" box="[151,323,764,790]" italics="true" pageId="22" pageNumber="23">Deltapliomera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5763FFA5505CFEAAFD7EFE17FCFE" box="[330,412,764,790]" pageId="22" pageNumber="23" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3376581FFA5505CFF77FCC0FC46FBE7" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8B92360AFFA5505CFF77FCC0FB38FC97" blockId="22.[151,1436,834,1579]" pageId="22" pageNumber="23">
<emphasis id="B959EA18FFA5505CFF77FCC0FEB6FCB3" bold="true" box="[151,317,834,859]" pageId="22" pageNumber="23">
<typeStatus id="549688A8FFA5505CFF77FCC0FF59FCB3" box="[151,210,834,859]" pageId="22" pageNumber="23">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C2D4D89FFA5505CFEAEFCC0FD0AFCB3" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[334,641,834,859]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="humphriesi" status="sp. nov.">
<emphasis id="B959EA18FFA5505CFEAEFCC0FD0AFCB3" box="[334,641,834,859]" italics="true" pageId="22" pageNumber="23">Deltapliomera humphriesi</emphasis>
</taxonomicName>
<emphasis id="B959EA18FFA5505CFD72FCC0FD57FCB3" bold="true" box="[658,732,834,859]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A26A5763FFA5505CFD72FCC0FD57FCB3" box="[658,732,834,859]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, from the Fillmore Formation (Blackhillsian;
<taxonomicName id="4C2D4D89FFA5505CFAFFFCC0FEDCFC97" authority="Zone" authorityName="Zone" class="Trilobita" family="Telephinidae" genus="Carolinites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proetida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B959EA18FFA5505CFAFFFCC0FE99FC97" italics="true" pageId="22" pageNumber="23">Carolinites nevadensis</emphasis>
Zone
</taxonomicName>
), southern Confusion Range, Ibex area, Millard County, western
<collectingRegion id="49E9F8E8FFA5505CFBCFFCE4FBE3FC97" box="[1071,1128,870,895]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
,
<collectingCountry id="F33A769AFFA5505CFB94FCE5FB3BFC97" box="[1140,1200,871,895]" name="United States of America" pageId="22" pageNumber="23">USA</collectingCountry>
.
</paragraph>
<paragraph id="8B92360AFFA5505CFF27FC08FC46FBE7" blockId="22.[151,1436,834,1579]" pageId="22" pageNumber="23">
<emphasis id="B959EA18FFA5505CFF27FC08FEE4FC4B" bold="true" box="[199,367,906,931]" pageId="22" pageNumber="23">Other species.</emphasis>
<taxonomicName id="4C2D4D89FFA5505CFE97FC08FDE9FC4B" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[375,610,906,931]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inglei" status="sp. nov.">
<emphasis id="B959EA18FFA5505CFE97FC08FDE9FC4B" box="[375,610,906,931]" italics="true" pageId="22" pageNumber="23">Deltapliomera inglei</emphasis>
</taxonomicName>
<emphasis id="B959EA18FFA5505CFD89FC08FD23FC4B" bold="true" box="[617,680,906,931]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A26A5763FFA5505CFD89FC08FD23FC4B" box="[617,680,906,931]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Fillmore Formation (Tulean;
<taxonomicName id="4C2D4D89FFA5505CFBE1FC08FABBFC4B" authority="Zone" authorityName="Zone" box="[1025,1328,906,931]" class="Trilobita" family="Dimeropygidae" genus="Heckethornia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proetida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bowiei">
<emphasis id="B959EA18FFA5505CFBE1FC08FB67FC4B" box="[1025,1260,906,931]" italics="true" pageId="22" pageNumber="23">Heckethornia bowiei</emphasis>
Zone
</taxonomicName>
), western
<collectingRegion id="49E9F8E8FFA5505CFF77FC2CFF5AFC2F" box="[151,209,942,967]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
;
<taxonomicName id="4C2D4D89FFA5505CFF3DFC2DFEFAFC2F" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[221,369,942,967]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="heimbergi" status="sp. nov.">
<emphasis id="B959EA18FFA5505CFF3DFC2DFEFAFC2F" box="[221,369,942,967]" italics="true" pageId="22" pageNumber="23">D. heimbergi</emphasis>
</taxonomicName>
<emphasis id="B959EA18FFA5505CFE98FC2CFE33FC2F" bold="true" box="[376,440,942,967]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A26A5763FFA5505CFE98FC2CFE33FC2F" box="[376,440,942,967]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Fillmore Formation (Tulean;
<emphasis id="B959EA18FFA5505CFCF1FC2DFB87FC2E" box="[785,1036,943,966]" italics="true" pageId="22" pageNumber="23">Panisaspis sevierensis</emphasis>
Zone), western
<collectingRegion id="49E9F8E8FFA5505CFB25FC2CFB74FC2F" box="[1221,1279,942,967]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
;
<taxonomicName id="4C2D4D89FFA5505CFAEAFC2DFA17FC2E" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[1290,1436,943,966]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="eppersoni" status="sp. nov.">
<emphasis id="B959EA18FFA5505CFAEAFC2DFA17FC2E" box="[1290,1436,943,966]" italics="true" pageId="22" pageNumber="23">D. eppersoni</emphasis>
</taxonomicName>
<emphasis id="B959EA18FFA5505CFF77FC50FF5DFC03" bold="true" box="[151,214,978,1003]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A26A5763FFA5505CFF77FC50FF5DFC03" box="[151,214,978,1003]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Fillmore Formation (Blackhillsian;
<taxonomicName id="4C2D4D89FFA5505CFD98FC50FC5CFC03" authority="Zone" authorityName="Zone" box="[632,983,978,1003]" class="Trilobita" family="Bathyuridae" genus="Bathyurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proetida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="plicolabeona">
<emphasis id="B959EA18FFA5505CFD98FC50FC1BFC03" box="[632,912,978,1003]" italics="true" pageId="22" pageNumber="23">Bathyurina plicolabeona</emphasis>
Zone
</taxonomicName>
), western
<collectingRegion id="49E9F8E8FFA5505CFBAAFC50FB0FFC03" box="[1098,1156,978,1003]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
;
<taxonomicName id="4C2D4D89FFA5505CFB70FC50FABFFC03" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[1168,1332,978,1003]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EA18FFA5505CFB70FC50FABFFC03" box="[1168,1332,978,1003]" italics="true" pageId="22" pageNumber="23">Deltapliomera</emphasis>
</taxonomicName>
n. sp. A, Fillmore Formation (Tulean;
<emphasis id="B959EA18FFA5505CFE3FFC75FD52FBE6" box="[479,729,1015,1038]" italics="true" pageId="22" pageNumber="23">Panisaspis sevierensis</emphasis>
Zone), western
<collectingRegion id="49E9F8E8FFA5505CFC6FFC74FC42FBE7" box="[911,969,1014,1039]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376581FFA5505CFF27FB98FB27FBDB" box="[199,1196,1050,1075]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="8B92360AFFA5505CFF27FB98FB27FBDB" blockId="22.[151,1436,834,1579]" box="[199,1196,1050,1075]" pageId="22" pageNumber="23">
<emphasis id="B959EA18FFA5505CFF27FB98FEC5FBDB" bold="true" box="[199,334,1050,1075]" pageId="22" pageNumber="23">Etymology.</emphasis>
The town of Delta,
<collectingRegion id="49E9F8E8FFA5505CFDCEFB98FDECFBDB" box="[558,615,1050,1075]" country="United States of America" name="Utah" pageId="22" pageNumber="23">Utah</collectingRegion>
, and the genus name
<taxonomicName id="4C2D4D89FFA5505CFCB8FB98FC4BFBDB" box="[856,960,1050,1075]" class="Trilobita" family="Pliomeridae" genus="Pliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B959EA18FFA5505CFCB8FB98FC4BFBDB" box="[856,960,1050,1075]" italics="true" pageId="22" pageNumber="23">Pliomera</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3376581FFA5505CFF27FBBCFEADFB0F" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8B92360AFFA5505CFF27FBBCFEADFB0F" blockId="22.[151,1436,834,1579]" pageId="22" pageNumber="23">
<emphasis id="B959EA18FFA5505CFF27FBBCFECAFBBF" bold="true" box="[199,321,1086,1111]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Sagittal glabellar height in anterior view 29.935.0% width across posterior projections (excluding genal spines); anterior course of axial furrows strongly anteriorly divergent; size of librigenal field reduced, trapezoidal in shape; posterior branch of facial suture along lateral border very short; tuberculate sculpture on first pygidial axial ring usually effaced at least medially; distal tips of pygidial pleural spines usually expanded and club-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3376581FFA5505CFF27FB70FCBAF9C2" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8B92360AFFA5505CFF27FB70FF7EFA9F" blockId="22.[151,1436,834,1579]" pageId="22" pageNumber="23">
<emphasis id="B959EA18FFA5505CFF27FB70FEC7FAE3" bold="true" box="[199,332,1266,1291]" pageId="22" pageNumber="23">Discussion.</emphasis>
<taxonomicName id="4C2D4D89FFA5505CFEB4FB70FE7CFAE3" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[340,503,1266,1291]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EA18FFA5505CFEB4FB70FE7CFAE3" box="[340,503,1266,1291]" italics="true" pageId="22" pageNumber="23">Deltapliomera</emphasis>
</taxonomicName>
was recovered as sister to
<taxonomicName id="4C2D4D89FFA5505CFCCEFB70FC37FAE3" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[814,956,1266,1291]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EA18FFA5505CFCCEFB70FC37FAE3" box="[814,956,1266,1291]" italics="true" pageId="22" pageNumber="23">Millardaspis</emphasis>
</taxonomicName>
in the phylogenetic analysis, and the taxa share a reduction in later ontogeny of clusters of tubercles on the glabellar lateral lobes that are prominent early; librigenal fields with a swollen rim adjacent to the lateral border furrow; and very broad librigenal lateral border furrows.
</paragraph>
<paragraph id="8B92360AFFA5505CFF27FA00FCBAF9C2" blockId="22.[151,1436,834,1579]" pageId="22" pageNumber="23">
Species of
<taxonomicName id="4C2D4D89FFA5505CFEA6FA00FE61FA73" authority="E.B &amp; Adrain &amp; Karim, 2018" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[326,490,1410,1435]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EA18FFA5505CFEA6FA00FE61FA73" box="[326,490,1410,1435]" italics="true" pageId="22" pageNumber="23">Deltapliomera</emphasis>
</taxonomicName>
occur at several horizons with more or less equally common species of
<taxonomicName id="4C2D4D89FFA5505CFACBFA00FA1EFA73" box="[1323,1429,1410,1435]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B959EA18FFA5505CFACBFA00FA1EFA73" box="[1323,1429,1410,1435]" italics="true" pageId="22" pageNumber="23">Ibexaspis</emphasis>
</taxonomicName>
. There is little question of correct association of sclerites, however, as species belonging to either genus also occur at some horizons in the absence of any member of the other genus.
<taxonomicName id="4C2D4D89FFA5505CFC44FA48FBE2FA0B" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[932,1129,1482,1507]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rupauli">
<emphasis id="B959EA18FFA5505CFC44FA48FBE2FA0B" box="[932,1129,1482,1507]" italics="true" pageId="22" pageNumber="23">Ibexaspis rupauli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D89FFA5505CFB43FA4EFA89FA0B" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[1187,1282,1482,1507]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="leuppi">
<emphasis id="B959EA18FFA5505CFB43FA4EFA89FA0B" box="[1187,1282,1482,1507]" italics="true" pageId="22" pageNumber="23">I. leuppi</emphasis>
</taxonomicName>
do not occur with any other member of the overall clade, nor does
<taxonomicName id="4C2D4D89FFA5505CFCF2FA6DFCFCF9EF" authorityName="E.B &amp; Adrain &amp; Karim" authorityYear="2018" box="[786,887,1518,1543]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inglei">
<emphasis id="B959EA18FFA5505CFCF2FA6DFCFCF9EF" box="[786,887,1518,1543]" italics="true" pageId="22" pageNumber="23">D. inglei</emphasis>
</taxonomicName>
. Reference to the morphology of these species removes any potential confusion when species do co-occur.
</paragraph>
</subSubSection>
</treatment>
</document>