170 lines
21 KiB
XML
170 lines
21 KiB
XML
<document id="D6EEDEE4B6700858B4824E3D9327CAFE" ID-CLB-Dataset="27906" ID-DOI="10.11646/zootaxa.4525.1.1" ID-GBIF-Dataset="aab5d525-1d70-4a48-9469-43d4e446685d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610805" ID-ZooBank="9D378750-982F-4061-A419-B28E8DDFF825" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553626294178" checkinUser="plazi" docAuthor="E. B, Neo, Adrain, Jonathan M. & Karim, Talia S." docDate="2018" docId="0384871CFF915069FF77F926FB35FBF8" docLanguage="en" docName="zootaxa.4525.1.1.pdf" docOrigin="Zootaxa 4525 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tuleaspis E.B & Adrain & Karim 2018" docType="treatment" docVersion="10" lastPageNumber="36" masterDocId="FFBDFF64FFB3504AFFE0FF82FF8BFFE8" masterDocTitle="The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA" masterLastPageNumber="152" masterPageNumber="1" pageNumber="35" updateTime="1698714183241" updateUser="ExternalLinkService">
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<mods:title id="6A1A5ADE237A873935D5D195AAED7C2B">The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA</mods:title>
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<mods:namePart id="5C9C62C4C94F4C45B8D5354F8D50177D">E. B, Neo</mods:namePart>
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<mods:namePart id="D8F5FF977F169FD98361F997D963E23B">Adrain, Jonathan M.</mods:namePart>
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<mods:namePart id="6F304298E3C82108D0CB59A647258D54">Karim, Talia S.</mods:namePart>
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<mods:title id="F95C2AD2BB5B51CE63302F6C16878CD8">Zootaxa</mods:title>
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<mods:date id="247BB2811AE70FA76640B09A7D3FE349">2018</mods:date>
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<treatment id="0384871CFF915069FF77F926FB35FBF8" ID-DOI="http://doi.org/10.5281/zenodo.5969674" ID-GBIF-Taxon="154499799" ID-Zenodo-Dep="5969674" LSID="urn:lsid:plazi:treatment:0384871CFF915069FF77F926FB35FBF8" httpUri="http://treatment.plazi.org/id/0384871CFF915069FF77F926FB35FBF8" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<subSubSection id="C3376581FF915068FF77F926FEE4F956" box="[151,367,1700,1726]" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="8B92360AFF915068FF77F926FEE4F956" blockId="34.[151,375,1700,1759]" box="[151,367,1700,1726]" pageId="34" pageNumber="35">
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<heading id="D0DA8166FF915068FF77F926FEE4F956" bold="true" box="[151,367,1700,1726]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<emphasis id="B959EA18FF915068FF77F926FEE4F956" bold="true" box="[151,367,1700,1726]" pageId="34" pageNumber="35">
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<taxonomicName id="4C2D4D89FF915068FF77F926FE81F956" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[151,266,1700,1726]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="34" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B959EA18FF915068FF77F926FE81F956" bold="true" box="[151,266,1700,1726]" italics="true" pageId="34" pageNumber="35">Tuleaspis</emphasis>
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</taxonomicName>
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n. sp. A
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF915068FF77F944FEFCF937" box="[151,375,1734,1759]" pageId="34" pageNumber="35" type="description">
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<paragraph id="8B92360AFF915068FF77F944FEFCF937" blockId="34.[151,375,1700,1759]" box="[151,375,1734,1759]" pageId="34" pageNumber="35">
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<figureCitation id="13162A8FFF915068FF77F944FF7FF937" box="[151,244,1734,1759]" captionStart="PLATE 42" captionStartId="125.[151,234,1474,1496]" captionText="PLATE 42. Deltapliomera humphriesi n. sp., from Section H 222.1 m, Fillmore Formation (Floian; Blackhillsian; Carolinites nevadensis Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." pageId="34" pageNumber="35">Plate 42</figureCitation>
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, figs 35–41
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF915068FF77F88CFC3EF87B" pageId="34" pageNumber="35" type="materials_examined">
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<paragraph id="8B92360AFF915068FF77F88CFC3EF87B" blockId="34.[151,1436,1806,2011]" pageId="34" pageNumber="35">
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<emphasis id="B959EA18FF915068FF77F88CFE8DF8CF" bold="true" box="[151,262,1806,1831]" pageId="34" pageNumber="35">Material.</emphasis>
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Assigned specimen SUI 129753 from Section G 155.6 m, Fillmore Formation, southern Confusion Range, Ibex area, Millard County, western
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<collectingRegion id="49E9F8E8FF915068FD75F8B0FD45F8A3" box="[661,718,1842,1867]" country="United States of America" name="Utah" pageId="34" pageNumber="35">Utah</collectingRegion>
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,
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<collectingCountry id="F33A769AFF915068FD3EF8B1FC92F8A3" box="[734,793,1843,1867]" name="United States of America" pageId="34" pageNumber="35">USA</collectingCountry>
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, and SUI 129754 from Section HC5 203.7–204.2T m, Garden City Formation, east side of Hillyard Canyon, Bear River Range, Franklin County, southeastern
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<collectingRegion id="49E9F8E8FF915068FAB5F8D4FA13F887" box="[1365,1432,1878,1903]" country="United States of America" name="Idaho" pageId="34" pageNumber="35">Idaho</collectingRegion>
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,
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<collectingCountry id="F33A769AFF915068FF77F8F9FF58F87B" box="[151,211,1915,1939]" name="United States of America" pageId="34" pageNumber="35">USA</collectingCountry>
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. Both are from the Tulean (low
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<taxonomicName id="4C2D4D89FF915068FDD8F8F8FC22F87B" authority="Zone" authorityName="Zone" box="[568,937,1914,1939]" class="Trilobita" family="Bathyuridae" genus="Psalikilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proetida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cuspidicauda">
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<emphasis id="B959EA18FF915068FDD8F8F8FCE8F87B" box="[568,867,1914,1939]" italics="true" pageId="34" pageNumber="35">Psalikilopsis cuspidicauda</emphasis>
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Zone
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF915069FF27F81CFB35FBF8" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
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<paragraph id="8B92360AFF915069FF27F81CFE4FFE45" blockId="34.[151,1436,1806,2011]" lastBlockId="35.[151,1437,151,1040]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<emphasis id="B959EA18FF915068FF27F81CFEC7F85F" bold="true" box="[199,332,1950,1975]" pageId="34" pageNumber="35">Discussion.</emphasis>
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One cranidium and one pygidium are assigned to
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<taxonomicName id="4C2D4D89FF915068FC76F81CFB8BF85F" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[918,1024,1950,1975]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="34" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B959EA18FF915068FC76F81CFB8BF85F" box="[918,1024,1950,1975]" italics="true" pageId="34" pageNumber="35">Tuleaspis</emphasis>
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</taxonomicName>
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n. sp. A. Although the pygidium is from Section G in
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<collectingRegion id="49E9F8E8FF915068FE91F840FE23F833" box="[369,424,1986,2011]" country="United States of America" name="Utah" pageId="34" pageNumber="35">Utah</collectingRegion>
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and the cranidium is from Section HC
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<quantity id="4CD59BEFFF915068FC8BF840FC1CF833" box="[875,919,1986,2011]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">5 in</quantity>
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<collectingRegion id="49E9F8E8FF915068FC40F840FC68F833" box="[928,995,1986,2011]" country="United States of America" name="Idaho" pageId="34" pageNumber="35">Idaho</collectingRegion>
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, we feel justified that they represent a single species because each sclerite clearly represents an
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<taxonomicName id="4C2D4D89FF905069FCDEFF15FC23FF58" box="[830,936,151,176]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA18FF905069FCDEFF15FC23FF58" box="[830,936,151,176]" italics="true" pageId="35" pageNumber="36">Ibexaspis</emphasis>
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</taxonomicName>
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group pliomerid, and the only known cooccurring pliomerids are well-known species of
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<emphasis id="B959EA18FF905069FD24FF3FFCB6FF3C" box="[708,829,189,212]" italics="true" pageId="35" pageNumber="36">Panisaspis</emphasis>
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<bibRefCitation id="EFBC4BFBFF905069FCA7FF3EFB16FF3D" author="McAdams, N. E. B. & Adrain, J. M." box="[839,1181,188,213]" pageId="35" pageNumber="36" pagination="1 - 68" refId="ref37597" refString="McAdams, N. E. B. & Adrain, J. M. (2011 b) Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Zootaxa, 2969, 1 - 68." type="journal article" year="2011">McAdams and Adrain, 2011b</bibRefCitation>
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, and
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<taxonomicName id="4C2D4D89FF905069FB3DFF3EFE96FF10" authority="Ross, 1951" authorityName="Ross" authorityYear="1951" class="Trilobita" family="Pliomeridae" genus="Protopliomerella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA18FF905069FB3DFF3EFA17FF3D" box="[1245,1436,188,213]" italics="true" pageId="35" pageNumber="36">Protopliomerella</emphasis>
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<bibRefCitation id="EFBC4BFBFF905069FF77FF62FE96FF10" author="Ross, R. J., Jr." box="[151,285,223,248]" pageId="35" pageNumber="36" pagination="1 - 161" refId="ref37889" refString="Ross, R. J., Jr. (1951) Stratigraphy of the Garden City Formation in northeastern Utah, and its trilobite faunas. Peabody Museum of Natural History, Yale University Bulletin, 6, 1 - 161." type="journal article" year="1951">Ross, 1951</bibRefCitation>
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</taxonomicName>
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(revised by
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<bibRefCitation id="EFBC4BFBFF905069FE27FF5DFCB4FF10" author="McAdams, N. E. B. & Adrain, J. M." box="[455,831,223,248]" pageId="35" pageNumber="36" pagination="1 - 113" refId="ref37651" refString="McAdams, N. E. B. & Adrain, J. M. (2011 c) Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA. Zootaxa, 3144, 1 - 113." type="journal article" year="2011">McAdams and Adrain [2011c]</bibRefCitation>
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). Additionally, both horizons are from the same biostratigraphic zone, and share many species in common. The pygidium overall is rather different to that of
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<taxonomicName id="4C2D4D89FF905069FA67FE87FF51FEA8" authorityName="E.B & Adrain & Karim" authorityYear="2018" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="jeneki">
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<emphasis id="B959EA18FF905069FA67FE87FF51FEA8" italics="true" pageId="35" pageNumber="36">T. jeneki</emphasis>
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</taxonomicName>
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, but
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<taxonomicName id="4C2D4D89FF905069FEF2FEA5FEF7FEA8" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[274,380,295,320]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B959EA18FF905069FEF2FEA5FEF7FEA8" box="[274,380,295,320]" italics="true" pageId="35" pageNumber="36">Tuleaspis</emphasis>
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</taxonomicName>
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n. sp. A is also a considerably older taxon, and the additional segment and terminal piece of
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<taxonomicName id="4C2D4D89FF905069FA67FEABFF51FE8D" authorityName="E.B & Adrain & Karim" authorityYear="2018" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="jeneki">
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<emphasis id="B959EA18FF905069FA67FEABFF51FE8D" italics="true" pageId="35" pageNumber="36">T. jeneki</emphasis>
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</taxonomicName>
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likely represent a derived condition. The sculpture of the pygidium is coarser and more widely spaced than that of the cranidium, but this is also true of
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<taxonomicName id="4C2D4D89FF905069FD4CFEF3FC85FE60" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[684,782,367,392]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="jeneki">
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<emphasis id="B959EA18FF905069FD4CFEF3FC85FE60" box="[684,782,367,392]" italics="true" pageId="35" pageNumber="36">T. jeneki</emphasis>
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</taxonomicName>
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, and therefore does not indicate that the sclerites might represent different species.
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</paragraph>
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<paragraph id="8B92360AFF905069FF27FE35FE2FFD18" blockId="35.[151,1437,151,1040]" pageId="35" pageNumber="36">
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The cranidium of
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<taxonomicName id="4C2D4D89FF905069FE74FE35FE75FE38" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[404,510,439,464]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B959EA18FF905069FE74FE35FE75FE38" box="[404,510,439,464]" italics="true" pageId="35" pageNumber="36">Tuleaspis</emphasis>
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</taxonomicName>
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n. sp. A differs from those of
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<taxonomicName id="4C2D4D89FF905069FCBCFE3BFC37FE38" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[860,956,439,464]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="jeneki">
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<emphasis id="B959EA18FF905069FCBCFE3BFC37FE38" box="[860,956,439,464]" italics="true" pageId="35" pageNumber="36">T. jeneki</emphasis>
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</taxonomicName>
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(cf. Pl. 42, fig. 40, Pl. 44, fig. 3) in being much more strongly vaulted (sag., particularly tr.) and narrower relative to sagittal length; with sculpture of slightly larger tubercles, evenly distributed on the glabella rather than concentrated on the lateral lobes; a slightly longer, more inflated, and more strongly anteriorly bowed anterior border; slightly shorter, wider, and considerably more inflated glabella with a smaller L1 relative to the other lobes; longer SO; LO with a more prominent median tubercle and fewer tubercles at the lateral corners; wider axial furrows; shorter, slightly wider, more anterolaterally positioned palpebral lobes; posterior portion of the palpebro-ocular suture describing a much less acute angle opposite the posterior fixigenae; longer, narrower posterior fixigenae; and a posterior border with more tubercles on the posterior margin.
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</paragraph>
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<paragraph id="8B92360AFF905069FF27FD7EFBA9FC05" blockId="35.[151,1437,151,1040]" pageId="35" pageNumber="36">
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The pygidium of
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<taxonomicName id="4C2D4D89FF905069FE6DFD7EFE7CFCFD" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[397,503,764,789]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B959EA18FF905069FE6DFD7EFE7CFCFD" box="[397,503,764,789]" italics="true" pageId="35" pageNumber="36">Tuleaspis</emphasis>
|
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</taxonomicName>
|
||
n. sp. A is quite different from that of
|
||
<taxonomicName id="4C2D4D89FF905069FC54FD7FFB9FFCFD" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[948,1044,764,789]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="jeneki">
|
||
<emphasis id="B959EA18FF905069FC54FD7FFB9FFCFD" box="[948,1044,764,789]" italics="true" pageId="35" pageNumber="36">T. jeneki</emphasis>
|
||
</taxonomicName>
|
||
(cf. Pl. 42, fig. 41, Pl. 46, figs 24, 29) because it possesses four segments and lacks a terminal piece, compared to five segments and a large terminal piece; it is relatively shorter, wider, and less posteriorly tapered in outline, as well as slightly more vaulted (sag., particularly tr.); the axis is shorter and broader, with more strongly inflated and medially shortened axial rings, and longer articulating and inter-ring furrows; the axial furrows are wider and shallower; the pleural spines are narrower and more closely spaced; and the sculpture is generally smaller, more sparsely distributed tubercles, with effaced areas along the midlength of the first axial ring and on the pleural spines.
|
||
</paragraph>
|
||
<paragraph id="8B92360AFF905069FF27FC75FB35FBF8" blockId="35.[151,1437,151,1040]" box="[199,1214,1015,1040]" pageId="35" pageNumber="36">
|
||
<taxonomicName id="4C2D4D89FF905069FF27FC75FEBAFBF8" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[199,305,1015,1040]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B959EA18FF905069FF27FC75FEBAFBF8" box="[199,305,1015,1040]" italics="true" pageId="35" pageNumber="36">Tuleaspis</emphasis>
|
||
</taxonomicName>
|
||
n. sp. A is compared to
|
||
<taxonomicName id="4C2D4D89FF905069FDA2FC75FD27FBF8" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[578,684,1015,1040]" class="Trilobita" family="Pliomeridae" genus="Tuleaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B959EA18FF905069FDA2FC75FD27FBF8" box="[578,684,1015,1040]" italics="true" pageId="35" pageNumber="36">Tuleaspis</emphasis>
|
||
</taxonomicName>
|
||
? n. sp. B in the discussion of the latter species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |