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<document id="8BBD102FE5C5D422019FE917971FEDAA" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFAF3C48FC0EF91CFE906B36" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada gardneriana Köhler 2011, n.sp." docType="treatment" docVersion="3" lastPageNumber="189" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="187" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CA9353E4318C04843936BC2953921840">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="8A0880F0497622D759504093A29ABC97" type="personal">
<mods:role id="06BB2BB69E5BBC5ED6F088CB1491AAF9">
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<mods:namePart id="7141DCC12961F9E5B48F50C30A5BE7F2">Köhler, Frank</mods:namePart>
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<mods:title id="9CE82677F4D3C705E5A920BDC482F17A">Records of the Australian Museum</mods:title>
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<mods:part id="0003467C78DD7C3690C9F3EDCE6589D4">
<mods:date id="BDA3FC5EB7B36CF0E08992ADA71FA864">2011</mods:date>
<mods:detail id="C4A9DCB73210CA70671B1E72D28F6361" type="pubDate">
<mods:number id="D33AFFA170B9C603594DA81B110AFF8E">2011-11-30</mods:number>
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<mods:number id="655454E4B39C5FB4F677B487A1D7D608">63</mods:number>
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<mods:url id="8E92CF85F3AA054CA5C1A110159D853D">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
</mods:location>
<mods:classification id="B6F05DA489D7A6D9A9B8A76FBD958AE7">journal article</mods:classification>
<mods:identifier id="996599C70A422FEAED4BC9C228A714C6" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
<mods:identifier id="60336B031B82DBC0B9EBF26E322A8091" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="973C6F79238E19106F1DFD7E5489191B" type="Zenodo-Dep">5239547</mods:identifier>
<mods:identifier id="77DE96E7FB18AFCCBEC7D6BF281AD286" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
</mods:mods>
<treatment id="0384503AFFAF3C48FC0EF91CFE906B36" LSID="urn:lsid:plazi:treatment:0384503AFFAF3C48FC0EF91CFE906B36" httpUri="http://treatment.plazi.org/id/0384503AFFAF3C48FC0EF91CFE906B36" lastPageId="22" lastPageNumber="189" pageId="20" pageNumber="187">
<subSubSection id="C337B2A7FFAF3C4AFC0EF91CFAA168A2" box="[950,1356,1742,1768]" pageId="20" pageNumber="187" type="nomenclature">
<paragraph id="8B92E12CFFAF3C4AFC0EF91CFAA168A2" blockId="20.[950,1356,1742,1768]" box="[950,1356,1742,1768]" pageId="20" pageNumber="187">
<heading id="D0DA5640FFAF3C4AFC0EF91CFAA168A2" box="[950,1356,1742,1768]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="187" reason="2">
<emphasis id="B9593D3EFFAF3C4AFC0EF91CFAA168A2" bold="true" box="[950,1356,1742,1768]" pageId="20" pageNumber="187">
<taxonomicName id="4C2D9AAFFFAF3C4AFC0EF91CFAE568A2" ID-CoL="D7QZ" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[950,1288,1742,1768]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="187" phylum="Mollusca" rank="species" species="gardneriana" status="sp. nov.">
<emphasis id="B9593D3EFFAF3C4AFC0EF91CFAE568A2" bold="true" box="[950,1288,1742,1768]" italics="true" pageId="20" pageNumber="187">Amplirhagada gardneriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFAF3C4AFAA8F91CFAA168A2" box="[1296,1356,1742,1768]" pageId="20" pageNumber="187" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFAF3C4BFCFFF8D0FE3568DC" lastPageId="21" lastPageNumber="188" pageId="20" pageNumber="187" type="materials_examined">
<paragraph id="8B92E12CFFAF3C4AFCFFF8D0FABE691E" blockId="20.[839,1466,1794,1876]" pageId="20" pageNumber="187">
<materialsCitation id="3B45EB71FFAF3C4AFCFFF8D0FAA3691E" collectingDate="1987-06-27" collectionCode="V" collectorName="NW Kimberley" latitude="-15.6" location="Princess May Ranges" longLatPrecision="21" longitude="125.441666" pageId="20" pageNumber="187" specimenCode="RFS 27-3" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B9593D3EFFAF3C4AFCFFF8D0FC386950" bold="true" box="[839,981,1794,1818]" pageId="20" pageNumber="187">
<typeStatus id="54965F8EFFAF3C4AFCFFF8D0FC926950" box="[839,895,1794,1818]" pageId="20" pageNumber="187">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="49E92FCEFFAF3C4AFC67F8D1FB746950" box="[991,1177,1795,1818]" country="Australia" name="Western Australia" pageId="20" pageNumber="187">Western Australia</collectingRegion>
,
<collectorName id="26D884FAFFAF3C4AFB19F8D1FAAF6950" box="[1185,1346,1795,1818]" pageId="20" pageNumber="187">NW Kimberley</collectorName>
,
<locationDeviation id="9A20F80EFFAF3C4AFAF3F8D1FC38697D" location="King Cascade" pageId="20" pageNumber="187">
<quantity id="4CD54CC9FFAF3C4AFAF3F8D1FA666950" box="[1355,1419,1795,1818]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="187" unit="km" value="15.0">15 km</quantity>
E of King Cascade
</locationDeviation>
,
<location id="8EF2B7F7FFAF3C4AFC66F8F2FB55697D" LSID="urn:lsid:plazi:treatment:0384503AFFAF3C48FC0EF91CFE906B36:8EF2B7F7FFAF3C4AFC66F8F2FB55697D" box="[990,1208,1824,1847]" latitude="-15.6" longLatPrecision="21" longitude="125.441666" name="Princess May Ranges" pageId="20" pageNumber="187" stateProvince="Western Australia">Princess May Ranges</location>
;
<geoCoordinate id="EE1987EBFFAF3C4AFB78F8F2FAD9697D" box="[1216,1332,1824,1847]" degrees="15" direction="south" minutes="36" orientation="latitude" pageId="20" pageNumber="187" precision="15" seconds="00" value="-15.6">15°36'00&quot;S</geoCoordinate>
<geoCoordinate id="EE1987EBFFAF3C4AFA80F8F2FA57697D" box="[1336,1466,1824,1847]" degrees="125" direction="east" minutes="26" orientation="longitude" pageId="20" pageNumber="187" precision="15" seconds="30" value="125.441666">125°26'30&quot;E</geoCoordinate>
(
<specimenCode id="DB8B4957FFAF3C4AFCF6F8EFFC55691E" box="[846,952,1853,1876]" collectionCode="RFS" pageId="20" pageNumber="187">RFS 27-3</specimenCode>
; coll.
<collectionCode id="ED3C79E9FFAF3C4AFC4CF8EFFBEA691E" box="[1012,1031,1853,1876]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="187" type="Museum">V</collectionCode>
. Kessner,
<date id="FF93C7ECFFAF3C4AFBC8F8EFFB1D691E" box="[1136,1264,1853,1876]" pageId="20" pageNumber="187" value="1987-06-27">
<collectingDate id="EFD73E04FFAF3C4AFBC8F8EFFB1D691E" box="[1136,1264,1853,1876]" pageId="20" pageNumber="187" value="1987-06-27">27 Jun 1987</collectingDate>
</date>
) (
<figureCitation id="1316FDA9FFAF3C4AFABEF8EFFAAE691E" box="[1286,1347,1853,1876]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="20" pageNumber="187">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B92E12CFFAF3C4AFCFFF8BAFA57699D" blockId="20.[839,1467,1896,2007]" pageId="20" pageNumber="187">
<emphasis id="B9593D3EFFAF3C4AFCFFF8BAFC1669CA" bold="true" box="[839,1019,1896,1920]" pageId="20" pageNumber="187">Type material</emphasis>
.
<materialsCitation id="3B45EB71FFAF3C4AFBA8F8BAFB7B69D6" accessionNumber="S34734" collectionCode="WAM" pageId="20" pageNumber="187" specimenCount="1" typeStatus="holotype">
<typeStatus id="54965F8EFFAF3C4AFBA8F8BAFB936935" box="[1040,1150,1896,1919]" pageId="20" pageNumber="187" type="holotype">Holotype</typeStatus>
<collectionCode id="ED3C79E9FFAF3C4AFB30F8BAFB206935" box="[1160,1229,1896,1919]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="187" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFAF3C4AFB61F8BAFADE6935" box="[1241,1331,1896,1919]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34734" pageId="20" pageNumber="187" type="EnaNcbi">S34734</accessionNumber>
(preserved specimen) (Pl. 1.11;
<tableCitation id="C6AFD497FFAF3C4AFB8EF857FB6769D6" box="[1078,1162,1925,1948]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="20" pageNumber="187">Table 1</tableCitation>
)
</materialsCitation>
.
<materialsCitation id="3B45EB71FFAF3C4AFB1DF857FC2C699D" accessionNumber="S34735" collectionCode="AM, WAM" pageId="20" pageNumber="187" specimenCount="1" typeStatus="paratype">
<typeStatus id="54965F8EFFAF3C4AFB1DF857FAFD69D6" box="[1189,1296,1925,1948]" pageId="20" pageNumber="187" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED3C79E9FFAF3C4AFAA1F857FAA869D6" box="[1305,1349,1925,1948]" country="Australia" name="Australian Museum" pageId="20" pageNumber="187" type="Museum">AM</collectionCode>
C.472934 (8 preserved specimens),
<collectionCode id="ED3C79E9FFAF3C4AFBC8F871FB5E69F0" box="[1136,1203,1955,1978]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="187" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFAF3C4AFB06F871FAFE69F0" box="[1214,1299,1955,1978]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34735" pageId="20" pageNumber="187" type="EnaNcbi">S34735</accessionNumber>
(13 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFAF3C4AFC74F812FA6A699D" accessionNumber="S34735" box="[972,1415,1984,2007]" collectionCode="FMNH" isTailEnumeration="true" pageId="20" pageNumber="187" specimenCode="FMNH 221084" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFAF3C4AFC74F812FB82699D" box="[972,1135,1984,2007]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="187" type="Museum">FMNH 221084</specimenCode>
(22 preserved specimens)
</materialsCitation>
,
<collectionCode id="ED3C79E9FFAF3C4AFA37F812FA57699D" box="[1423,1466,1984,2007]" country="Australia" name="Australian Museum" pageId="20" pageNumber="187" type="Museum">AM</collectionCode>
</paragraph>
<caption id="DF52B1A4FFAE3C4BFFC1FA07FDE26870" ID-DOI="http://doi.org/10.5281/zenodo.5239610" ID-Zenodo-Dep="5239610" httpUri="https://zenodo.org/record/5239610/files/figure.png" pageId="21" pageNumber="188" startId="21.[121,184,1493,1514]" targetBox="[121,1421,151,1476]" targetPageId="21">
<paragraph id="8B92E12CFFAE3C4BFFC1FA07FDE26870" blockId="21.[121,1422,1493,1594]" pageId="21" pageNumber="188">
Figure 31.
<taxonomicName id="4C2D9AAFFFAE3C4BFF5DFA07FE036BA0" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[229,494,1493,1514]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="188" phylum="Mollusca" rank="species" species="gardneriana" status="sp. nov.">
<emphasis id="B9593D3EFFAE3C4BFF5DFA07FE036BA0" box="[229,494,1493,1514]" italics="true" pageId="21" pageNumber="188">Amplirhagada gardneriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFAE3C4BFE4EFA07FDC96BA0" box="[502,548,1493,1514]" pageId="21" pageNumber="188" rank="species">n.sp.</taxonomicNameLabel>
SEM of shell (AM C.472935), radula and jaw (AM C.472934).
<emphasis id="B9593D3EFFAE3C4BFB1AFA07FB2D6BA0" box="[1186,1216,1493,1514]" italics="true" pageId="21" pageNumber="188">(A)</emphasis>
Protoconch and first teleoconch whorls viewed from above.
<emphasis id="B9593D3EFFAE3C4BFE49FA3DFDE2684E" box="[497,527,1519,1540]" italics="true" pageId="21" pageNumber="188">(B)</emphasis>
Detail of first teleoconch whorls viewed from above.
<emphasis id="B9593D3EFFAE3C4BFBA9FA3DFBDC684E" box="[1041,1073,1519,1540]" italics="true" pageId="21" pageNumber="188">(C)</emphasis>
Close-up of sculpture on last whorl, lateral view.
<emphasis id="B9593D3EFFAE3C4BFF4BF9D8FEF96855" box="[243,276,1546,1567]" italics="true" pageId="21" pageNumber="188">(D)</emphasis>
Jaw.
<emphasis id="B9593D3EFFAE3C4BFEF1F9D8FE8A6855" box="[329,359,1546,1567]" italics="true" pageId="21" pageNumber="188">(E)</emphasis>
Central and inner lateral teeth from above.
<emphasis id="B9593D3EFFAE3C4BFCBFF9D8FCCB6855" box="[775,806,1546,1567]" italics="true" pageId="21" pageNumber="188">(F)</emphasis>
Marginal teeth viewed from above. Scale bars:
<emphasis id="B9593D3EFFAE3C4BFB56F9D8FAFA6855" box="[1262,1303,1546,1567]" italics="true" pageId="21" pageNumber="188">AB</emphasis>
, 200 µm;
<emphasis id="B9593D3EFFAE3C4BFACFF9D8FA6A6855" box="[1399,1415,1546,1567]" italics="true" pageId="21" pageNumber="188">C</emphasis>
, 1 mm;
<emphasis id="B9593D3EFFAE3C4BFF05F9F7FF236870" box="[189,206,1573,1594]" italics="true" pageId="21" pageNumber="188">D</emphasis>
, 100 µm;
<emphasis id="B9593D3EFFAE3C4BFE88F9F7FED26870" box="[304,319,1573,1594]" italics="true" pageId="21" pageNumber="188">E</emphasis>
, 10 µm;
<emphasis id="B9593D3EFFAE3C4BFE2CF9F7FE526870" box="[404,447,1573,1594]" italics="true" pageId="21" pageNumber="188">FG</emphasis>
, 20 µm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFAE3C4BFFC1F9B0FE3568DC" blockId="21.[121,748,1634,1686]" pageId="21" pageNumber="188">
C.472935 (14 dried shells),
<collectionCode id="ED3C79E9FFAE3C4BFE25F9B0FE336833" box="[413,478,1634,1657]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="21" pageNumber="188" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFAE3C4BFE5CF9B0FDDB6833" box="[484,566,1634,1657]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34736" pageId="21" pageNumber="188" type="EnaNcbi">S34736</accessionNumber>
(25 dried shells),
<collectionCode id="ED3C79E9FFAE3C4BFFC1F9ADFF2B68DC" box="[121,198,1663,1686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="188" type="Museum">FMNH</collectionCode>
221083 (34 dried shells).
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFAE3C4BFFC1F978FEF76894" pageId="21" pageNumber="188" type="etymology">
<paragraph id="8B92E12CFFAE3C4BFFC1F978FEF76894" blockId="21.[121,749,1706,1758]" pageId="21" pageNumber="188">
<emphasis id="B9593D3EFFAE3C4BFFC1F978FF196888" bold="true" box="[121,244,1706,1730]" pageId="21" pageNumber="188">Etymology</emphasis>
. In reference to the Gardner Plateau, where this species occurs.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFAE3C48FED1F92AFDF26AE8" lastPageId="22" lastPageNumber="189" pageId="21" pageNumber="188" type="description">
<paragraph id="8B92E12CFFAE3C4BFED1F92AFE106958" blockId="21.[361,509,1784,1810]" box="[361,509,1784,1810]" pageId="21" pageNumber="188">
<emphasis id="B9593D3EFFAE3C4BFED1F92AFE106958" bold="true" box="[361,509,1784,1810]" pageId="21" pageNumber="188">Description</emphasis>
</paragraph>
<paragraph id="8B92E12CFFAE3C4BFFC1F8E2FC5C69CB" blockId="21.[121,749,1839,2009]" lastBlockId="21.[794,1422,1634,2009]" pageId="21" pageNumber="188">
<emphasis id="B9593D3EFFAE3C4BFFC1F8E2FF43690D" box="[121,174,1840,1863]" italics="true" pageId="21" pageNumber="188">Shell</emphasis>
(Pl. 1.11;
<figureCitation id="1316FDA9FFAE3C4BFE9BF8FDFE43690C" box="[291,430,1839,1862]" captionStart="Figure 31" captionStartId="21.[121,184,1493,1514]" captionTargetBox="[121,1421,151,1476]" captionTargetId="figure-89@21.[121,1421,151,1476]" captionTargetPageId="21" captionText="Figure 31. Amplirhagada gardneriana n.sp. SEM of shell (AM C.472935), radula and jaw (AM C.472934). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth from above. (F) Marginal teeth viewed from above. Scale bars: AB, 200 µm; C, 1 mm; D, 100 µm; E, 10 µm; FG, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5239610" httpUri="https://zenodo.org/record/5239610/files/figure.png" pageId="21" pageNumber="188">Fig. 31 AC</figureCitation>
). Semi-globose with elevated spire. Solid, not translucent. Periphery well rounded to angulate; upper and basal sectors of whorls well rounded. Umbilicus narrowly winding, 5080% concealed by columellar reflection. Background colour horn to pale brownish ochre; peripheral band rarely present, if present rather distinct, thin, brown, visible on last whorl only; sub-sutural band absent or diffuse, brown, thin; outer and inner lighter than shell. Protoconch
<quantity id="4CD54CC9FFAE3C4BFB14F94FFAEC68FE" box="[1196,1281,1693,1716]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="21" pageNumber="188" unit="mm" value="2.6">2.6 mm</quantity>
in diameter, comprising about one whorl, with very indistinct radially elongated pustulations. Teleoconch with faint, regular axial growth lines; evenly distributed across shell surface. Angle of aperture 4560 degrees; outer lip thin, well rounded, expanded, not reflected; basal node absent or weak. Parietal wall of inner lip thin.Average shell size 14.2±1.2 × 18.0±
<quantity id="4CD54CC9FFAE3C4BFAD3F89FFCAE69CB" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="188" unit="mm" value="0.7">0.7 mm</quantity>
(
<tableCitation id="C6AFD497FFAE3C4BFCE9F8B8FC4C69CB" box="[849,929,1898,1921]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="21" pageNumber="188">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B92E12CFFAE3C4BFC80F85AFA6369F6" blockId="21.[794,1422,1634,2009]" pageId="21" pageNumber="188">
<emphasis id="B9593D3EFFAE3C4BFC80F85AFB8C69D5" box="[824,1121,1928,1951]" italics="true" pageId="21" pageNumber="188">Radular and jaw morphology</emphasis>
(
<figureCitation id="1316FDA9FFAE3C4BFBD4F855FB0369D4" box="[1132,1262,1927,1950]" captionStart="Figure 31" captionStartId="21.[121,184,1493,1514]" captionTargetBox="[121,1421,151,1476]" captionTargetId="figure-89@21.[121,1421,151,1476]" captionTargetPageId="21" captionText="Figure 31. Amplirhagada gardneriana n.sp. SEM of shell (AM C.472935), radula and jaw (AM C.472934). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth from above. (F) Marginal teeth viewed from above. Scale bars: AB, 200 µm; C, 1 mm; D, 100 µm; E, 10 µm; FG, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5239610" httpUri="https://zenodo.org/record/5239610/files/figure.png" pageId="21" pageNumber="188">Fig. 31 DG</figureCitation>
). Tooth formula C + 11 + 4 + 16; 125 rows of teeth (n = 1). Jaw with 10 plates.
</paragraph>
<paragraph id="8B92E12CFFAE3C48FC80F810FCE06ACE" blockId="21.[794,1422,1634,2009]" lastBlockId="22.[166,794,781,1186]" lastPageId="22" lastPageNumber="189" pageId="21" pageNumber="188">
<emphasis id="B9593D3EFFAE3C4BFC80F810FBE66993" box="[824,1035,1986,2009]" italics="true" pageId="21" pageNumber="188">Genital morphology</emphasis>
(
<figureCitation id="1316FDA9FFAE3C4BFBA1F810FB7E6993" box="[1049,1171,1986,2009]" captionStart-0="Figure 32" captionStart-1="Figure 33" captionStartId-0="22.[166,230,676,697]" captionStartId-1="22.[839,902,154,175]" captionTargetBox-0="[172,781,125,660]" captionTargetBox-1="[839,1464,150,1527]" captionTargetId-0="figure-323@22.[166,793,119,670]" captionTargetId-1="figure-346@22.[877,1479,145,1538]" captionTargetPageId-0="22" captionTargetPageId-1="22" captionText-0="Figure 32. Genitalia of Amplirhagada gardneriana n.sp. (AM C.472934).For labelling of structures see Fig.3. Scale bar = 5 mm." captionText-1="Figure 33. Penial anatomy of Amplirhagada" figureDoi-0="http://doi.org/10.5281/zenodo.5239614" figureDoi-1="http://doi.org/10.5281/zenodo.5239612" httpUri-0="https://zenodo.org/record/5239614/files/figure.png" httpUri-1="https://zenodo.org/record/5239612/files/figure.png" pageId="21" pageNumber="188">Figs 3233</figureCitation>
). Penis straight, tubular, thin, as long as anterior part of oviduct. Penial sheath distally thick, proximally delicate. Penial retractor about half as long as penis. Penial verge conical, with pointed tip, extending about ¼ of length of penial chamber. Inner penial wall entirely covered by very fine, dense pustulation. Main stimulatory pilaster weakly developed, extending distal half of inner penial wall, corrugated. Vas deferens rather thin, entering penial sheath in distal third of penial complex. Vagina moderately long, tubular to slightly inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct. Free oviduct comprising about half of length of anterior part of oviduct. Spermoviduct longer than anterior part of oviduct.
</paragraph>
<caption id="DF52B1A4FFAD3C48FCFFFF48FB036EE5" ID-DOI="http://doi.org/10.5281/zenodo.5239612" ID-Zenodo-Dep="5239612" box="[839,1262,154,175]" httpUri="https://zenodo.org/record/5239612/files/figure.png" pageId="22" pageNumber="189" startId="22.[839,902,154,175]" targetBox="[839,1464,150,1527]" targetPageId="22">
<paragraph id="8B92E12CFFAD3C48FCFFFF48FB036EE5" blockId="22.[839,1262,154,255]" box="[839,1262,154,175]" pageId="22" pageNumber="189">
Figure 33. Penial anatomy of
<taxonomicName id="4C2D9AAFFFAD3C48FBDCFF48FB036EE5" box="[1124,1262,154,175]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="189" phylum="Mollusca" rank="genus">Amplirhagada</taxonomicName>
</paragraph>
</caption>
<caption id="DF52B1A4FFAD3C48FF1EFD76FCF66C9E" ID-DOI="http://doi.org/10.5281/zenodo.5239614" ID-Zenodo-Dep="5239614" httpUri="https://zenodo.org/record/5239614/files/figure.png" pageId="22" pageNumber="189" startId="22.[166,230,676,697]" targetBox="[172,781,125,660]" targetPageId="22">
<paragraph id="8B92E12CFFAD3C48FF1EFD76FCF66C9E" blockId="22.[166,795,676,724]" pageId="22" pageNumber="189">
Figure 32. Genitalia of
<taxonomicName id="4C2D9AAFFFAD3C48FE21FD76FD446CF3" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[409,681,676,697]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="189" phylum="Mollusca" rank="species" species="gardneriana" status="sp. nov.">
<emphasis id="B9593D3EFFAD3C48FE21FD76FD446CF3" box="[409,681,676,697]" italics="true" pageId="22" pageNumber="189">Amplirhagada gardneriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFAD3C48FD0BFD76FD0F6CF3" box="[691,738,676,697]" pageId="22" pageNumber="189" rank="species">n.sp.</taxonomicNameLabel>
(AM C.472934). For labelling of structures see Fig. 3. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFAD3C48FF7CFB59FDF26AE8" blockId="22.[166,794,781,1186]" box="[196,543,1163,1186]" pageId="22" pageNumber="189">
<emphasis id="B9593D3EFFAD3C48FF7CFB59FE7A6AE8" box="[196,407,1163,1186]" italics="true" pageId="22" pageNumber="189">Aestivation strategy</emphasis>
. Free sealer.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFAD3C48FF1EFB67FE906B36" pageId="22" pageNumber="189" type="discussion">
<paragraph id="8B92E12CFFAD3C48FF1EFB67FE906B36" blockId="22.[166,794,1205,1404]" pageId="22" pageNumber="189">
<emphasis id="B9593D3EFFAD3C48FF1EFB67FEE36A87" bold="true" box="[166,270,1205,1229]" pageId="22" pageNumber="189">Remarks</emphasis>
. Anatomical description based on dissections of
<specimenCount id="9D2B2AA5FFAD3C48FF1EFB01FEB96AA0" box="[166,340,1235,1258]" pageId="22" pageNumber="189" type="generic">three specimens</specimenCount>
. Listed by Solem (1991) as “
<taxonomicName id="4C2D9AAFFFAD3C48FD38FB01FF316B4D" authority="NSP" authorityName="NSP" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="189" phylum="Mollusca" rank="genus">
<emphasis id="B9593D3EFFAD3C48FD38FB01FCF76AA0" box="[640,794,1235,1258]" italics="true" pageId="22" pageNumber="189">Amplirhagada</emphasis>
NSP
</taxonomicName>
28”. According to Solem (1991) the species differs from
<taxonomicName id="4C2D9AAFFFAD3C48FF59FADCFE396B6E" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[225,468,1293,1317]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="189" phylum="Mollusca" rank="species" species="pusilla">
<emphasis id="B9593D3EFFAD3C48FF59FADCFEA56B6F" box="[225,328,1294,1317]" italics="true" pageId="22" pageNumber="189">A. pusilla</emphasis>
<bibRefCitation id="EFBC9CDDFFAD3C48FEF6FADFFE396B6E" author="Solem, A" box="[334,468,1293,1316]" pageId="22" pageNumber="189" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
</taxonomicName>
by its narrower umbilicus and more elevated spire and from
<taxonomicName id="4C2D9AAFFFAD3C48FE50FAF9FCF46B08" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[488,793,1323,1346]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="189" phylum="Mollusca" rank="species" species="drysdaleana">
<emphasis id="B9593D3EFFAD3C48FE50FAF9FD666B08" box="[488,651,1323,1346]" italics="true" pageId="22" pageNumber="189">A. drysdaleana</emphasis>
<bibRefCitation id="EFBC9CDDFFAD3C48FD2AFAF9FCF46B08" author="Solem, A" box="[658,793,1323,1346]" pageId="22" pageNumber="189" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
</taxonomicName>
by a shorter vagina. It also differs by a distinct pustulation of inner penial wall.
</paragraph>
</subSubSection>
</treatment>
</document>