260 lines
36 KiB
XML
260 lines
36 KiB
XML
<document id="1F616DA0FAC04B5198F090A8234ACDB8" ID-DOI="10.11646/zootaxa.5447.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150353" ID-ZooBank="F9FB91ED-5536-4CBD-8D4D-61439605BD94" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715181655124" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2024" docId="0381F17A9C4EFFE9F1C8F76130B5C570" docLanguage="en" docName="zootaxa.5447.4.3.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus davidiani Mikhaljova 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="498" masterDocId="FFB889029C4FFFEAF15FF47F3320C415" masterDocTitle="New species and new records of millipedes of the genera Nepalmatoiulus Mauriès, 1983 and Anaulaciulus Pocock, 1895 (Diplopoda, Julida, Julidae) from mainland China" masterLastPageNumber="512" masterPageNumber="496" pageNumber="497" updateTime="1715182081406" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="99A46DC91575071C242005828ABCC66F">New species and new records of millipedes of the genera Nepalmatoiulus Mauriès, 1983 and Anaulaciulus Pocock, 1895 (Diplopoda, Julida, Julidae) from mainland China</mods:title>
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<mods:namePart id="DFD2787239CAC99EF00A5A2A6409C989">Mikhaljova, Elena V.</mods:namePart>
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<mods:typeOfResource id="BF6275D192A9DA845B689EC0D738A960">text</mods:typeOfResource>
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<mods:title id="8CAC7A60A3F19A8D6F9C3F74D9C83C9A">Zootaxa</mods:title>
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<mods:part id="3ED3538A0ED59D4B82D20D02444E1896">
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<mods:date id="918845E4B03EEC274619CF756D833094">2024</mods:date>
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<mods:number id="C694E914F9D98EB8837A973CFBE8DF66">2024-05-08</mods:number>
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<mods:number id="519A0DE334AE849126F50AFDA7D0C283">5447</mods:number>
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<mods:classification id="E406B2F327AAE7C3C9017DF57510D47C">journal article</mods:classification>
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<mods:identifier id="DD81719555482E8AA4F2320C88CCCE6C" type="DOI">10.11646/zootaxa.5447.4.3</mods:identifier>
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<treatment id="0381F17A9C4EFFE9F1C8F76130B5C570" ID-DOI="http://doi.org/10.5281/zenodo.11150365" ID-Zenodo-Dep="11150365" LSID="urn:lsid:plazi:treatment:0381F17A9C4EFFE9F1C8F76130B5C570" httpUri="http://treatment.plazi.org/id/0381F17A9C4EFFE9F1C8F76130B5C570" lastPageId="3" lastPageNumber="498" pageId="1" pageNumber="497">
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<subSubSection id="C33213E79C4EFFEBF1C8F7613114C72C" box="[151,564,798,825]" pageId="1" pageNumber="497" type="nomenclature">
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<paragraph id="8B97406C9C4EFFEBF1C8F7613114C72C" blockId="1.[151,564,798,861]" box="[151,564,798,825]" pageId="1" pageNumber="497">
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<heading id="D0DFF7009C4EFFEBF1C8F7613114C72C" bold="true" box="[151,564,798,825]" fontSize="11" level="1" pageId="1" pageNumber="497" reason="1">
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<emphasis id="B95C9C7E9C4EFFEBF1C8F7613114C72C" bold="true" box="[151,564,798,825]" pageId="1" pageNumber="497">
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<taxonomicName id="4C283BEF9C4EFFEBF1C8F76132EFC72C" authority="Mikhaljova, 2024" authorityName="Mikhaljova" authorityYear="2024" box="[151,463,798,825]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="davidiani" status="sp. nov.">
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<emphasis id="B95C9C7E9C4EFFEBF1C8F76132EFC72C" bold="true" box="[151,463,798,825]" italics="true" pageId="1" pageNumber="497">Nepalmatoiulus davidiani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26F21059C4EFFEBF088F7603114C72C" box="[471,564,799,825]" pageId="1" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33213E79C4EFFEBF1C8F73C33D9C748" box="[151,249,835,861]" pageId="1" pageNumber="497" type="description">
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<paragraph id="8B97406C9C4EFFEBF1C8F73C33D9C748" blockId="1.[151,564,798,861]" box="[151,249,835,861]" pageId="1" pageNumber="497">
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<figureCitation id="13135CE99C4EFFEBF1C8F73C33D9C748" box="[151,249,835,861]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–9" captionStartId-0="2.[152,270,1487,1511]" captionStartId-1="3.[152,270,1410,1434]" captionTargetBox-0="[151,1436,181,1463]" captionTargetBox-1="[418,1170,381,1387]" captionTargetId-0="figure-15@2.[151,1436,181,1463]" captionTargetId-1="figure-155@3.[418,1170,381,1387]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." captionText-1="FIGURES 8–9. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 8. Opisthomere, mesal view. 9. Solenomere, mesal view. Abbreviation: bb, basal blade. Scales: 10 μm (Fig. 9), 100 μm (Fig. 8)." figureDoi-0="http://doi.org/10.5281/zenodo.11150355" figureDoi-1="http://doi.org/10.5281/zenodo.11150359" httpUri-0="https://zenodo.org/record/11150355/files/figure.png" httpUri-1="https://zenodo.org/record/11150359/files/figure.png" pageId="1" pageNumber="497">Figs 1–9</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33213E79C4EFFEBF1C8F7FB37E2C7D7" pageId="1" pageNumber="497" type="materials_examined">
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<paragraph id="8B97406C9C4EFFEBF1C8F7FB37E2C7D7" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
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<emphasis id="B95C9C7E9C4EFFEBF1C8F7FB32A5C78B" bold="true" box="[151,389,900,926]" pageId="1" pageNumber="497">Material examined.</emphasis>
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<materialsCitation id="3B404A319C4EFFEBF0CFF7FB37E2C7D7" collectingDate="2016-05-16" collectionCode="ZMUM" collectorName="I. Kabak & Kabak, G." country="China" elevation="2870" latitude="26.616388" location="Jianchuan" longLatPrecision="20" longitude="99.776115" pageId="1" pageNumber="497" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="5493FECE9C4EFFEBF0CFF7FB32DDC78B" box="[400,509,900,926]" pageId="1" pageNumber="497" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D2E8BE59C4EFFEBF354F7FB317AC78B" box="[523,602,900,926]" count="1" pageId="1" pageNumber="497" type="male">1 male</specimenCount>
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(
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<collectionCode id="ED39D8A99C4EFFEBF332F7FB31EAC78B" box="[621,714,900,926]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="497">ZMUM</collectionCode>
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),
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<collectingCountry id="F33F00FC9C4EFFEBF3BEF7FB3007C78B" box="[737,807,900,926]" name="China" pageId="1" pageNumber="497">China</collectingCountry>
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,
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<collectingRegion id="49EC8E8E9C4EFFEBF26AF7FA30DCC78B" box="[821,1020,900,927]" country="China" name="Yunnan" pageId="1" pageNumber="497">Yunnan Province</collectingRegion>
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, NW of
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<location id="8EF716B79C4EFFEBF538F7FB37FAC78B" LSID="urn:lsid:plazi:treatment:0381F17A9C4EFFE9F1C8F76130B5C570:8EF716B79C4EFFEBF538F7FB37FAC78B" box="[1127,1242,900,926]" country="China" latitude="26.616388" longLatPrecision="20" longitude="99.776115" name="Jianchuan" pageId="1" pageNumber="497" stateProvince="Yunnan">Jianchuan</location>
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,
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<locationDeviation id="9A25594E9C4EFFEBF5B6F7FB3211C7D7" location="Shuanghecun" pageId="1" pageNumber="497">
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<quantity id="4CD0ED899C4EFFEBF5B6F7FB3669C78B" box="[1257,1353,900,927]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="497" unit="km" value="10.5">10.5 km</quantity>
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WNW Shuanghecun
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</locationDeviation>
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,
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<geoCoordinate id="EE1C26AB9C4EFFEBF063F7D732EAC7D6" box="[316,458,936,963]" degrees="26" direction="north" minutes="36" orientation="latitude" pageId="1" pageNumber="497" precision="15" seconds="59" value="26.616388">26°36'59" N</geoCoordinate>
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,
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<geoCoordinate id="EE1C26AB9C4EFFEBF08AF7D7317FC7D6" box="[469,607,936,963]" degrees="99" direction="east" minutes="46" orientation="longitude" pageId="1" pageNumber="497" precision="15" seconds="34" value="99.776115">99°46'34" E</geoCoordinate>
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, H =
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<quantity id="4CD0ED899C4EFFEBF3C2F7D731D4C7D6" box="[669,756,936,963]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="1" pageNumber="497" unit="m" value="2870.0">
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<elevation id="0005A75F9C4EFFEBF3C2F7D731D4C7D6" box="[669,756,936,963]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="1" pageNumber="497" unit="m" value="2870.0">2870 m</elevation>
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</quantity>
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||
,
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<date id="FF9666AC9C4EFFEBF3A0F7D730A7C7D7" box="[767,903,936,963]" pageId="1" pageNumber="497" value="2016-05-16">
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<collectingDate id="EFD29F449C4EFFEBF3A0F7D730A7C7D7" box="[767,903,936,963]" pageId="1" pageNumber="497" value="2016-05-16">16.05. 2016</collectingDate>
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</date>
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, leg.
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<collectorName id="26DD25BA9C4EFFEBF29CF7D63706C7D7" box="[963,1062,936,963]" pageId="1" pageNumber="497">I. Kabak</collectorName>
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, G. Davidian.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33213E79C4EFFEBF198F7B33673C063" pageId="1" pageNumber="497" type="diagnosis">
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<paragraph id="8B97406C9C4EFFEBF198F7B33673C063" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
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<emphasis id="B95C9C7E9C4EFFEBF198F7B33262C7F3" bold="true" box="[199,322,972,998]" pageId="1" pageNumber="497">Diagnosis.</emphasis>
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Differs from Chinese congeners mainly by the combination of: male coxa 2 with one and two mesapical oral setae anteriorly and posteriorly, respectively; a strongly obliquely rounded apex of gonopod promere; relatively small body size; colour pattern of the usual julid
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<typeStatus id="5493FECE9C4EFFEBF271F06A307EC03A" box="[814,862,1045,1071]" pageId="1" pageNumber="497">type</typeStatus>
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(after
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<bibRefCitation id="EFB93D9D9C4EFFEBF2F6F06B3765C03B" author="Enghoff, H." box="[937,1093,1044,1070]" pageId="1" pageNumber="497" pagination="1 - 142" refId="ref7831" refString="Enghoff, H. (1982) The millipede genus Cylindroiulus on Madeira - an insular species swarm (Diplopoda, Julida: Julidae). Entomologica Scandinavica Supplement, 18, 1 - 142." type="journal article" year="1982">Enghoff 1982</bibRefCitation>
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) of rings below ozopore level. Configuration of gonopod opisthomere in
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<taxonomicName id="4C283BEF9C4EFFEBF323F0463029C047" authority="Mikhaljova, 2024" authorityName="Mikhaljova" authorityYear="2024" box="[636,777,1080,1106]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="davidiani" status="sp. nov.">
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<emphasis id="B95C9C7E9C4EFFEBF323F0463029C047" box="[636,777,1080,1106]" italics="true" pageId="1" pageNumber="497">N. davidiani</emphasis>
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</taxonomicName>
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<emphasis id="B95C9C7E9C4EFFEBF24DF047304EC047" bold="true" box="[786,878,1080,1106]" pageId="1" pageNumber="497">
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<taxonomicNameLabel id="A26F21059C4EFFEBF24DF047304EC047" box="[786,878,1080,1106]" pageId="1" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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similar to
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<taxonomicName id="4C283BEF9C4EFFEBF2B2F04733FDC063" authority="Mikhaljova, 2023 b" authorityName="Mikhaljova" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="angustus">
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<emphasis id="B95C9C7E9C4EFFEBF2B2F047362BC047" box="[1005,1291,1080,1106]" italics="true" pageId="1" pageNumber="497">Nepalmatoiulus angustus</emphasis>
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<bibRefCitation id="EFB93D9D9C4EFFEBF44BF04733FDC063" author="Mikhaljova, E. V." pageId="1" pageNumber="497" pagination="285 - 300" refId="ref8433" refString="Mikhaljova, E. V. (2023 b) A contribution to the millipede fauna of China: descriptions of four new species of the genus Nepalmatoiulus Mauries, 1983 from high-altitude areas of Yunnan Province (Diplopoda, Julida, Julidae). Zootaxa, 5357 (2), 285 - 300. https: // doi. org / 10.11646 / zootaxa. 5357.2.6" type="journal article" year="2023">Mikhaljova, 2023b</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C283BEF9C4EFFEBF04BF0223289C063" authority="Mikhaljova, 2024" authorityName="Mikhaljova" authorityYear="2024" box="[276,425,1116,1142]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="shiguensis" status="sp. nov.">
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<emphasis id="B95C9C7E9C4EFFEBF04BF0223289C063" box="[276,425,1116,1142]" italics="true" pageId="1" pageNumber="497">N. shiguensis</emphasis>
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</taxonomicName>
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<emphasis id="B95C9C7E9C4EFFEBF0EFF0233131C063" bold="true" box="[432,529,1116,1142]" pageId="1" pageNumber="497">
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<taxonomicNameLabel id="A26F21059C4EFFEBF0EFF023312BC063" box="[432,523,1116,1142]" pageId="1" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
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,
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</emphasis>
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but differs from both of them by specific characters (see Remarks below).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33213E79C4EFFE8F198F0FF374FC35A" lastPageId="2" lastPageNumber="498" pageId="1" pageNumber="497" type="description">
|
||
<paragraph id="8B97406C9C4EFFEBF198F0FF37D8C113" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
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||
<emphasis id="B95C9C7E9C4EFFEBF198F0FF3279C08F" bold="true" box="[199,345,1152,1178]" pageId="1" pageNumber="497">Description.</emphasis>
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||
<emphasis id="B95C9C7E9C4EFFEBF03EF0FF32BBC08F" box="[353,411,1152,1178]" italics="true" pageId="1" pageNumber="497">Male</emphasis>
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||
. Length in alcohol about 18.0 mm, midbody vertical diameter about
|
||
<quantity id="4CD0ED899C4EFFEBF5EBF0FF362EC08E" box="[1204,1294,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="497" unit="mm" value="1.4">1.4 mm</quantity>
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||
, with 53(-3) rings, excluding telson. Coloration in alcohol dark brown with broad longitudinal slightly lighter dorsal stripe; rings below ozopore level, head, collum and pregonopodal rings of the usual julid
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<typeStatus id="5493FECE9C4EFFEBF575F0B6377AC0F6" box="[1066,1114,1225,1251]" pageId="1" pageNumber="497">type</typeStatus>
|
||
(after
|
||
<bibRefCitation id="EFB93D9D9C4EFFEBF5FCF0B73661C0F7" author="Enghoff, H." box="[1187,1345,1224,1250]" pageId="1" pageNumber="497" pagination="1 - 142" refId="ref7831" refString="Enghoff, H. (1982) The millipede genus Cylindroiulus on Madeira - an insular species swarm (Diplopoda, Julida: Julidae). Entomologica Scandinavica Supplement, 18, 1 - 142." type="journal article" year="1982">Enghoff 1982</bibRefCitation>
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||
). Venter including basal parts of legs beige. Distal parts of legs marbled brown. Antennae brown, eyes black.
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||
</paragraph>
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||
<paragraph id="8B97406C9C4EFFEBF198F16F312CC217" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
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||
Head smooth, 2 epicranial setae, 4 supralabral setae; at least 24 labral setae. Eye patch almost oval, composed of at least 40 ommatidia in
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<typeStatus id="5493FECE9C4EFFEBF08AF14B311BC15B" box="[469,571,1332,1358]" pageId="1" pageNumber="497" type="holotype">holotype</typeStatus>
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. Antennae medium-sized, rather slender and clavate. Antennomeres 5 and 6 with incomplete distodorsal corolla of sensilla basiconica. Mandibular stipites with subtriangular smooth lobes. Gnathochilarium with about 10 nonapical stipital setae; lamellae linguales each with 5 setae arranged longitudinally. Collum laterally with distinct striae of different lengths at posterior margin, the first of which (the longest striae) reaching to anterior margin, dorsally with distinct short striae at posterior margin. A transverse row of very sparse thin setae at hind edge of collum.
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</paragraph>
|
||
<paragraph id="8B97406C9C4EFFEBF198F27331F1C2EA" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
|
||
Body rings circular. Metazona with dense, regular, longitudinal striae reaching hind margin (15–16 striae in an approximate square with sides equal to metazonital length of a dorsal side of a midbody ring). Limbus straight, smooth (of
|
||
<typeStatus id="5493FECE9C4EFFEBF04CF22A326BC27A" box="[275,331,1621,1647]" pageId="1" pageNumber="497">Type</typeStatus>
|
||
<quantity id="4CD0ED899C4EFFEBF010F22B3258C27B" box="[335,376,1620,1646]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="497" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EFB93D9D9C4EFFEBF023F22B3138C27B" author="Enghoff, H." box="[380,536,1620,1646]" pageId="1" pageNumber="497" pagination="241 - 331" refId="ref7912" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
).A transverse row of sparse, thin setae at hind edge of metazonites, setae gradually growing denser and longer toward telson. Ozopores small, lying behind suture dividing pro- and metazona, set off from the suture on almost all rings, in touch with suture only in the most posterior ones. Telson with caudal dorsal projection straight and long, covered with setae and carrying at tip a claw-shaped process curved dorsad. Preanal ring, anal valves, and subanal scale densely setose.
|
||
</paragraph>
|
||
<paragraph id="8B97406C9C4EFFEBF198F376379EC3EF" blockId="1.[151,1437,900,2042]" pageId="1" pageNumber="497">
|
||
Legs relatively short and slender. Very delicately serrate ventral pads present on postfemur and tibia, starting from legs 2. Claw of all legs at base with a long (longer than claw) setiform accessory claw ventrally. Leg pair 1 forming hook, the distal podomere not coming into close contact with the basal podomeres (“open hook”
|
||
<typeStatus id="5493FECE9C4EFFEBF411F32E365EC37E" box="[1358,1406,1873,1899]" pageId="1" pageNumber="497">type</typeStatus>
|
||
in
|
||
<bibRefCitation id="EFB93D9D9C4EFFEBF1C8F30B3216C39B" author="Enghoff, H." box="[151,310,1908,1934]" pageId="1" pageNumber="497" pagination="241 - 331" refId="ref7912" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
); postfemur with inflated scaly-rugose ventral surface, tip indistinctly wrinkled, coxa with one seta, distal segment with one seta laterally and minute seta tarsal remnant (
|
||
<figureCitation id="13135CE99C4EFFEBF29AF3E73729C3A7" box="[965,1033,1944,1970]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="1" pageNumber="497">Fig. 1</figureCitation>
|
||
). Coxa 2 with one long mesapical oral seta anteriorly (
|
||
<figureCitation id="13135CE99C4EFFEBF024F3C3329FC3C3" box="[379,447,1980,2006]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="1" pageNumber="497">Fig. 2</figureCitation>
|
||
) and shorter two ones posteriorly (
|
||
<figureCitation id="13135CE99C4EFFEBF213F3C330B2C3C3" box="[844,914,1980,2006]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="1" pageNumber="497">Fig. 4</figureCitation>
|
||
), gland opening positioned in apical and axial position
|
||
<emphasis id="B95C9C7E9C4EFFEBF1A7F39E3216C3EF" box="[248,310,2017,2042]" italics="true" pageId="1" pageNumber="497">sensu</emphasis>
|
||
<bibRefCitation id="EFB93D9D9C4EFFEBF061F39F32CDC3EF" author="Enghoff, H." box="[318,493,2016,2042]" pageId="1" pageNumber="497" pagination="241 - 331" refId="ref7912" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff (1987)</bibRefCitation>
|
||
. Penis subtrapezoidal, about 1.6 times longer than wide (
|
||
<figureCitation id="13135CE99C4EFFEBF533F39F3790C3EF" box="[1132,1200,2016,2042]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="1" pageNumber="497">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5710E49C4DFFE8F1C7F1B03054C23A" ID-DOI="http://doi.org/10.5281/zenodo.11150355" ID-Zenodo-Dep="11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="2" pageNumber="498" startId="2.[152,270,1487,1511]" targetBox="[151,1436,181,1463]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B97406C9C4DFFE8F1C7F1B03054C23A" blockId="2.[151,1437,1487,1583]" pageId="2" pageNumber="498">
|
||
<emphasis id="B95C9C7E9C4DFFE8F1C7F1B03260C1F2" bold="true" box="[152,320,1487,1511]" pageId="2" pageNumber="498">FIGURES 1–7.</emphasis>
|
||
<taxonomicName id="4C283BEF9C4DFFE8F01AF1B0316BC1F2" authority="Mikhaljova, 2024" authorityName="Mikhaljova" authorityYear="2024" box="[325,587,1487,1511]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="davidiani" status="sp. nov.">
|
||
<emphasis id="B95C9C7E9C4DFFE8F01AF1B0316BC1F2" box="[325,587,1487,1511]" italics="true" pageId="2" pageNumber="498">Nepalmatoiulus davidiani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C9C7E9C4DFFE8F30EF1B03180C1F2" bold="true" box="[593,672,1487,1511]" pageId="2" pageNumber="498">
|
||
<taxonomicNameLabel id="A26F21059C4DFFE8F30EF1B03180C1F2" box="[593,672,1487,1511]" pageId="2" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype (ZMUM).
|
||
<emphasis id="B95C9C7E9C4DFFE8F2F2F1B0309AC1F2" bold="true" box="[941,954,1487,1511]" pageId="2" pageNumber="498">1</emphasis>
|
||
. Leg 1, anterior view.
|
||
<emphasis id="B95C9C7E9C4DFFE8F5C4F1B03788C1F2" bold="true" box="[1179,1192,1487,1511]" pageId="2" pageNumber="498">2</emphasis>
|
||
. Coxae 2, anterior view.
|
||
<emphasis id="B95C9C7E9C4DFFE8F1C8F18C3384C21E" bold="true" box="[151,164,1523,1547]" pageId="2" pageNumber="498">3</emphasis>
|
||
. Penis, caudal view.
|
||
<emphasis id="B95C9C7E9C4DFFE8F02AF18C32A2C21E" bold="true" box="[373,386,1523,1547]" pageId="2" pageNumber="498">4</emphasis>
|
||
. Coxa 2, posterior view.
|
||
<emphasis id="B95C9C7E9C4DFFE8F324F18C31AEC21E" bold="true" box="[635,654,1523,1547]" pageId="2" pageNumber="498">5.</emphasis>
|
||
Apex of flagellum.
|
||
<emphasis id="B95C9C7E9C4DFFE8F20AF18C3048C21E" bold="true" box="[853,872,1523,1547]" pageId="2" pageNumber="498">6.</emphasis>
|
||
Distal part of right promere.
|
||
<emphasis id="B95C9C7E9C4DFFE8F5D3F18C37BFC21E" bold="true" box="[1164,1183,1523,1547]" pageId="2" pageNumber="498">7.</emphasis>
|
||
Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B97406C9C4DFFE8F198F2FE374FC35A" blockId="2.[151,1437,1665,2016]" pageId="2" pageNumber="498">
|
||
Gonopods slightly protruding. Anterior gonopod flattened, with parallel margins, apically strongly obliquely rounded, in posterior view apically excavated for accommodation of mesomeral process, distal margins of the excavation papillate (
|
||
<figureCitation id="13135CE99C4DFFE8F0D0F2B632DAC2F6" box="[399,506,1737,1763]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="2" pageNumber="498">Figs 6–7</figureCitation>
|
||
). Flagellum slender, of medium length, caudally covered with cuticular conical spikes (
|
||
<figureCitation id="13135CE99C4DFFE8F1B0F2923215C31D" box="[239,309,1773,1800]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1487,1511]" captionTargetBox="[151,1436,181,1463]" captionTargetId="figure-15@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURES 1–7. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 1. Leg 1, anterior view. 2. Coxae 2, anterior view. 3. Penis, caudal view. 4. Coxa 2, posterior view. 5. Apex of flagellum. 6. Distal part of right promere. 7. Right promere, posterior view. Scales: 10 μm (Figs 5, 6), 20 μm (Figs 1, 2, 3, 4), 100 μm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.11150355" httpUri="https://zenodo.org/record/11150355/files/figure.png" pageId="2" pageNumber="498">Fig. 5</figureCitation>
|
||
). Opisthomere slender (
|
||
<figureCitation id="13135CE99C4DFFE8F315F29231B0C312" box="[586,656,1773,1799]" captionStart="FIGURES 8–9" captionStartId="3.[152,270,1410,1434]" captionTargetBox="[418,1170,381,1387]" captionTargetId="figure-155@3.[418,1170,381,1387]" captionTargetPageId="3" captionText="FIGURES 8–9. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 8. Opisthomere, mesal view. 9. Solenomere, mesal view. Abbreviation: bb, basal blade. Scales: 10 μm (Fig. 9), 100 μm (Fig. 8)." figureDoi="http://doi.org/10.5281/zenodo.11150359" httpUri="https://zenodo.org/record/11150359/files/figure.png" pageId="2" pageNumber="498">Fig. 8</figureCitation>
|
||
). Mesomeral process slightly arched forward, its apex covered with papillae. Margin of velum smooth, arcuate, with a deep notch near the mesomeral process. Additional membrane with a serrate edge. Solenomere spinose throughout, basally with blade (
|
||
<emphasis id="B95C9C7E9C4DFFE8F29AF34A30C4C35A" bold="true" box="[965,996,1845,1871]" pageId="2" pageNumber="498">bb</emphasis>
|
||
) (
|
||
<figureCitation id="13135CE99C4DFFE8F2A3F34A3742C35A" box="[1020,1122,1845,1871]" captionStart="FIGURES 8–9" captionStartId="3.[152,270,1410,1434]" captionTargetBox="[418,1170,381,1387]" captionTargetId="figure-155@3.[418,1170,381,1387]" captionTargetPageId="3" captionText="FIGURES 8–9. Nepalmatoiulus davidiani sp. nov., male holotype (ZMUM). 8. Opisthomere, mesal view. 9. Solenomere, mesal view. Abbreviation: bb, basal blade. Scales: 10 μm (Fig. 9), 100 μm (Fig. 8)." figureDoi="http://doi.org/10.5281/zenodo.11150359" httpUri="https://zenodo.org/record/11150359/files/figure.png" pageId="2" pageNumber="498">Figs 8–9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33213E79C4DFFE8F198F326362CC366" box="[199,1292,1881,1908]" pageId="2" pageNumber="498" type="etymology">
|
||
<paragraph id="8B97406C9C4DFFE8F198F326362CC366" blockId="2.[151,1437,1665,2016]" box="[199,1292,1881,1908]" pageId="2" pageNumber="498">
|
||
<emphasis id="B95C9C7E9C4DFFE8F198F326326EC366" bold="true" box="[199,334,1881,1907]" pageId="2" pageNumber="498">Etymology.</emphasis>
|
||
Honours Genrikh Davidian, one of the collectors of this material. A noun in genitive.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33213E79C4DFFE9F198F30230B5C570" lastPageId="3" lastPageNumber="499" pageId="2" pageNumber="498" type="discussion">
|
||
<paragraph id="8B97406C9C4DFFE9F198F30230B5C570" blockId="2.[151,1437,1665,2016]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="499" pageId="2" pageNumber="498">
|
||
<emphasis id="B95C9C7E9C4DFFE8F198F302321BC382" bold="true" box="[199,315,1917,1943]" pageId="2" pageNumber="498">Remarks.</emphasis>
|
||
<taxonomicName id="4C283BEF9C4DFFE8F019F302314DC382" authority="Mikhaljova, 2024" authorityName="Mikhaljova" authorityYear="2024" box="[326,621,1917,1943]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="davidiani" status="sp. nov.">
|
||
<emphasis id="B95C9C7E9C4DFFE8F019F302314DC382" box="[326,621,1917,1943]" italics="true" pageId="2" pageNumber="498">Nepalmatoiulus davidiani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C9C7E9C4DFFE8F328F30231F5C382" bold="true" box="[631,725,1917,1943]" pageId="2" pageNumber="498">
|
||
<taxonomicNameLabel id="A26F21059C4DFFE8F328F30231F5C382" box="[631,725,1917,1943]" pageId="2" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C283BEF9C4DFFE8F22AF30130DEC382" authorityName="Mikhaljova" authorityYear="2023" box="[885,1022,1918,1943]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4DFFE8F22AF30130DEC382" box="[885,1022,1918,1943]" italics="true" pageId="2" pageNumber="498">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
mainly by the smaller body length (about 18.0 mm) and larger midbody vertical diameter (about
|
||
<quantity id="4CD0ED899C4DFFE8F227F3DE30F4C3AE" box="[888,980,1953,1979]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="498" unit="mm" value="1.4">1.4 mm</quantity>
|
||
) (vs. length—32.0–34.0 mm, midbody vertical diameter—about
|
||
<quantity id="4CD0ED899C4DFFE8F0E8F3BA312EC3F5" box="[439,526,1989,2016]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="498" unit="mm" value="2.5">2.5 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C283BEF9C4DFFE8F36CF3B93198C3CA" authorityName="Mikhaljova" authorityYear="2023" box="[563,696,1990,2015]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4DFFE8F36CF3B93198C3CA" box="[563,696,1990,2015]" italics="true" pageId="2" pageNumber="498">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
), by the coloration of the usual julid
|
||
<typeStatus id="5493FECE9C4DFFE8F500F3B937AFC3F5" box="[1119,1167,1990,2016]" pageId="2" pageNumber="498">type</typeStatus>
|
||
of rings below ozopore level, (vs. from dark gray with a pale swoop to gray-dark brown in
|
||
<taxonomicName id="4C283BEF9C4CFFE9F2D9F4E8372BC4A5" authorityName="Mikhaljova" authorityYear="2023" box="[902,1035,151,176]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4CFFE9F2D9F4E8372BC4A5" box="[902,1035,151,176]" italics="true" pageId="3" pageNumber="499">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
), by the strongly obliquely rounded apex of gonopod promere, by the male coxa 2 with one mesapical oral seta anteriorly (vs. male coxa 2 with two ones anteriorly in
|
||
<taxonomicName id="4C283BEF9C4CFFE9F03EF4A032C6C4ED" authorityName="Mikhaljova" authorityYear="2023" box="[353,486,223,248]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4CFFE9F03EF4A032C6C4ED" box="[353,486,223,248]" italics="true" pageId="3" pageNumber="499">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
), by the distal podomere of the male legpair 1 with one seta laterally and minute seta tarsal remnant (vs. distal podomere of the male legpair 1 without seta laterally and tarsal remnant in
|
||
<taxonomicName id="4C283BEF9C4CFFE9F458F57C36ACC509" authorityName="Mikhaljova" authorityYear="2023" box="[1287,1420,259,284]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4CFFE9F458F57C36ACC509" box="[1287,1420,259,284]" italics="true" pageId="3" pageNumber="499">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
), by the long (longer than claw) setiform accessory claw ventrally in claw of all legs (vs. short, (equal to claw length) setiform accessory claw ventrally in claw of all legs in
|
||
<taxonomicName id="4C283BEF9C4CFFE9F25EF53430A5C571" authorityName="Mikhaljova" authorityYear="2023" box="[769,901,331,356]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="B95C9C7E9C4CFFE9F25EF53430A5C571" box="[769,901,331,356]" italics="true" pageId="3" pageNumber="499">N. angustus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |