treatments-xml/data/03/81/E1/0381E12F8430FF82FF7AFDED3833FDCA.xml

<document id="FF973C4CEB96D010E1E58FF15FDAFB35" ID-CLB-Dataset="55760" ID-DOI="10.11646/zootaxa.5129.4.4" ID-GBIF-Dataset="2e2fb97f-7d86-4289-8e27-bf81bfacc4db" ID-ISSN="1175-5326" ID-Zenodo-Dep="6504104" ID-ZooBank="B0368310-A732-4191-8F41-B213BC89BDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651218560238" checkinUser="plazi" docAuthor="Ponce, Marcos, Navarro, Deivy, Morales, Roger &amp; Batista, Abel" docDate="2022" docId="0381E12F8430FF82FF7AFDED3833FDCA" docLanguage="en" docName="zootaxa.5129.4.4.pdf" docOrigin="Zootaxa 5129 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolitoglossa cathyledecae Ponce &amp; Navarro &amp; Morales &amp; Batista 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="553" masterDocId="FFB899578433FF88FFEDFFAB3D11FFAB" masterDocTitle="A new salamander of the genus Bolitoglossa (Caudata: Plethodontidae) from the highlands of western Panama" masterLastPageNumber="556" masterPageNumber="543" pageNumber="546" updateTime="1699329450438" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8A542878293EEB8A8788A99D5A4850A9">A new salamander of the genus Bolitoglossa (Caudata: Plethodontidae) from the highlands of western Panama</mods:title>
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<mods:namePart id="4F2C8F926AC7A49DB7A949828A5BB9B7">Ponce, Marcos</mods:namePart>
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<mods:affiliation id="A494D559E76E9A4FA381E494548FD611">Bioconsultant, David, Provincia de Chiriquí, Panamá &amp; https: // orcid. org / 0000 - 0002 - 6850 - 178 X</mods:affiliation>
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<mods:namePart id="0534A4E562EE33B2B745E9F719DCC443">Navarro, Deivy</mods:namePart>
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<mods:affiliation id="6245C42F582C48AA2AEC4CA9F12DC733">Protecma Panamá S. A. Santiago, Veraguas, Panamá &amp; https: // orcid. org / 0000 - 0002 - 7179 - 283 X</mods:affiliation>
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<mods:namePart id="889CCE599D80C14DA906394D86BCA705">Morales, Roger</mods:namePart>
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<mods:affiliation id="57203C1471830FC304C6101D23920D98">Bioconsultant, David, Provincia de Chiriquí, Panamá &amp; Fundación Los Naturalistas, P. O. Box 0426 - 01459 David, Provincia de Chiriquí, Panamá &amp; https: // orcid. org / 0000 - 0001 - 5511 - 2964</mods:affiliation>
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<mods:namePart id="4AB2F4D557F4128C161374F97052E437">Batista, Abel</mods:namePart>
<mods:affiliation id="E9DC64B1C2B2D2228B5F4295260ABDEF">Fundación Los Naturalistas, P. O. Box 0426 - 01459 David, Provincia de Chiriquí, Panamá &amp; Vicerrectoría de Investigación y Postgrado, Universidad Autónoma de Chiriquí, Ciudad Universitaria El Cabrero David, Chiriquí, 427, Panamá &amp; Asociación ADOPTA el Bosque Panamá, 119 x Gamboa, Panamá</mods:affiliation>
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<treatment id="0381E12F8430FF82FF7AFDED3833FDCA" ID-DOI="http://doi.org/10.5281/zenodo.6506304" ID-GBIF-Taxon="194951829" ID-Zenodo-Dep="6506304" LSID="urn:lsid:plazi:treatment:0381E12F8430FF82FF7AFDED3833FDCA" httpUri="http://treatment.plazi.org/id/0381E12F8430FF82FF7AFDED3833FDCA" lastPageId="10" lastPageNumber="553" pageId="3" pageNumber="546">
<subSubSection id="C33203B28430FF8BFF7AFDED3F25FDCA" box="[151,564,582,609]" pageId="3" pageNumber="546" type="nomenclature">
<paragraph id="8B9750398430FF8BFF7AFDED3F25FDCA" blockId="3.[151,784,582,682]" box="[151,564,582,609]" pageId="3" pageNumber="546">
<heading id="D0DFE7558430FF8BFF7AFDED3F25FDCA" bold="true" box="[151,564,582,609]" fontSize="11" level="1" pageId="3" pageNumber="546" reason="1">
<emphasis id="B95C8C2B8430FF8BFF7AFDED3F25FDCA" bold="true" box="[151,564,582,609]" italics="true" pageId="3" pageNumber="546">
<taxonomicName id="4C282BBA8430FF8BFF7AFDED3CDFFDCA" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[151,462,582,609]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">Bolitoglossa cathyledecae</taxonomicName>
<taxonomicNameLabel id="A26F31508430FF8BFE38FDEC3F25FDCA" box="[469,564,583,609]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33203B28430FF8BFF7AFDC03E01FD2E" box="[151,784,619,645]" pageId="3" pageNumber="546" type="vernacular_names">
<paragraph id="8B9750398430FF8BFF7AFDC03E01FD2E" blockId="3.[151,784,582,682]" box="[151,784,619,645]" pageId="3" pageNumber="546">Chiriqui fire salamander, Salamandra de fuego chiricana</paragraph>
</subSubSection>
<subSubSection id="C33203B28430FF8BFF7AFD243C3EFD01" box="[151,303,655,682]" pageId="3" pageNumber="546" type="description">
<paragraph id="8B9750398430FF8BFF7AFD243C3EFD01" blockId="3.[151,784,582,682]" box="[151,303,655,682]" pageId="3" pageNumber="546">
(
<figureCitation id="13134CBC8430FF8BFF72FD243C37FD01" box="[159,294,655,682]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="6.[151,250,1936,1960]" captionStartId-1="7.[151,250,1962,1986]" captionStartId-2="8.[151,250,1296,1320]" captionStartId-3="9.[151,250,1234,1258]" captionTargetBox-0="[172,1414,669,1913]" captionTargetBox-1="[156,1431,1040,1939]" captionTargetBox-2="[151,1436,181,1272]" captionTargetBox-3="[198,1430,309,1211]" captionTargetId-0="figure-274@6.[172,1414,669,1913]" captionTargetId-1="figure-277@7.[156,1431,1040,1939]" captionTargetId-2="figure-21@8.[151,1436,181,1272]" captionTargetId-3="figure-73@9.[157,1430,309,1211]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 3. Details of hand and foot of preserved holotype of Bolitoglossa cathyledecae sp. nov. (MHCH 3240); A) Right hand, left: dorsal view, right: ventral view; B) Right foot, left: dorsal view, right: ventral view. Scale bars: 5 mm. Photographs taken by MP." captionText-1="FIGURE 4. Holotype of Bolitoglossa cathyledecae sp. nov. (MHCH 3240) in life; A–B) Details of head; C–D) Body coloration in life. Photograph by MP." captionText-2="FIGURE 5. Holotype after three years in 70 % ethanol; A) Dorsal view; B) ventral view. Scale bars= 2 cm. Photographs taken by MP." captionText-3="FIGURE 6. Color variation in the paratypes.A) Dorsum in daytime coloration (MHCH 3242); B-C) Details of head and dorsum in daytime coloration (MHCH 3241); D) Paratype at the moment of encounter at night (MHCH 3242). Photographs taken by MP." figureDoi-0="http://doi.org/10.5281/zenodo.6504112" figureDoi-1="http://doi.org/10.5281/zenodo.6504114" figureDoi-2="http://doi.org/10.5281/zenodo.6504116" figureDoi-3="http://doi.org/10.5281/zenodo.6504118" httpUri-0="https://zenodo.org/record/6504112/files/figure.png" httpUri-1="https://zenodo.org/record/6504114/files/figure.png" httpUri-2="https://zenodo.org/record/6504116/files/figure.png" httpUri-3="https://zenodo.org/record/6504118/files/figure.png" pageId="3" pageNumber="546">Figures 3–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33203B28430FF8BFF7AFD7C3DD0FC0E" pageId="3" pageNumber="546" type="materials_examined">
<paragraph id="8B9750398430FF8BFF7AFD7C396EFC92" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
<materialsCitation id="3B405A648430FF8BFF7AFD7C396EFC92" ID-GBIF-Occurrence="3758761302" collectingDate="2017-05-20" collectionCode="PILA" collectorName="La Amistad International Park &amp; Deivy Navarro &amp; Carlos Castillo &amp; Erick Nunez &amp; Marcos Ponce" country="Panama" elevation="1969" latitude="8.8635" location="La Amistad International Park" longLatPrecision="7" longitude="-82.4864" municipality="Boquete" pageId="3" pageNumber="546" specimenCode="MHCH 3240, MP 12" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<emphasis id="B95C8C2B8430FF8BFF7AFD7C3C15FD5A" bold="true" box="[151,260,727,753]" pageId="3" pageNumber="546">
<typeStatus id="5493EE9B8430FF8BFF7AFD7C3C15FD5A" box="[151,260,727,753]" pageId="3" pageNumber="546" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<specimenCode id="DB8EF8428430FF8BFEF8FD7C3CBDFD5A" box="[277,428,727,753]" collectionCode="MHCH" pageId="3" pageNumber="546">MHCH 3240</specimenCode>
(original field number 
<specimenCode id="DB8EF8428430FF8BFD53FD7C3E1FFD5A" box="[702,782,727,753]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="3" pageNumber="546">MP 12</specimenCode>
), an adult male from 
<collectingCountry id="F33F10A98430FF8BFBFDFD7C397CFD5A" box="[1040,1133,727,753]" name="Panama" pageId="3" pageNumber="546">Panama</collectingCountry>
: 
<collectingRegion id="49EC9EDB8430FF8BFB97FD7C388DFD5A" box="[1146,1436,727,753]" country="Panama" name="Chiriqui" pageId="3" pageNumber="546">Chiriquí province district</collectingRegion>
<collectingMunicipality id="6BF3CA438430FF8BFF7AFD503DE6FCBE" box="[151,247,763,789]" pageId="3" pageNumber="546">Boquete</collectingMunicipality>
, 
<collectorName id="26DD35EF8430FF8BFEEBFD503F72FCBE" box="[262,611,763,789]" pageId="3" pageNumber="546">
<location id="8EF706E28430FF8BFEEBFD503F72FCBE" LSID="urn:lsid:plazi:treatment:0381E12F8430FF82FF7AFDED3833FDCA:8EF706E28430FF8BFEEBFD503F72FCBE" box="[262,611,763,789]" country="Panama" latitude="8.8635" longLatPrecision="7" longitude="-82.4864" municipality="Boquete" name="La Amistad International Park" pageId="3" pageNumber="546" stateProvince="Chiriqui">La Amistad International Park</location>
</collectorName>
(
<collectionCode id="ED39C8FC8430FF8BFD98FD503FA6FCBE" box="[629,695,763,789]" pageId="3" pageNumber="546">PILA</collectionCode>
) (
<geoCoordinate id="EE1C36FE8430FF8BFD3CFD513E53FCBE" box="[721,834,762,789]" degrees="8.8635" direction="north" orientation="latitude" pageId="3" pageNumber="546" precision="5" value="8.8635">8.8635°N</geoCoordinate>
, 
<geoCoordinate id="EE1C36FE8430FF8BFCBDFD503EC0FCBE" box="[848,977,763,789]" degrees="82.4864" direction="west" orientation="longitude" pageId="3" pageNumber="546" precision="5" value="-82.4864">82.4864°W</geoCoordinate>
, 
<quantity id="4CD0FDDC8430FF8BFC0DFD503955FCBE" box="[992,1092,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.969" pageId="3" pageNumber="546" unit="m" value="1969.0">
<elevation id="0005B70A8430FF8BFC0DFD503955FCBE" box="[992,1092,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.969" pageId="3" pageNumber="546" unit="m" value="1969.0">1,969 m</elevation>
</quantity>
); collected by 
<collectorName id="26DD35EF8430FF8BFB1CFD50388DFCBE" box="[1265,1436,763,789]" pageId="3" pageNumber="546">Deivy Navarro</collectorName>
along with 
<collectorName id="26DD35EF8430FF8BFEF8FCB43CAEFC92" box="[277,447,799,825]" pageId="3" pageNumber="546">Carlos Castillo</collectorName>
, 
<collectorName id="26DD35EF8430FF8BFE27FCB43F46FC92" box="[458,599,799,825]" pageId="3" pageNumber="546">Erick Nuñez</collectorName>
and 
<collectorName id="26DD35EF8430FF8BFD63FCB43E3CFC92" box="[654,813,799,825]" pageId="3" pageNumber="546">Marcos Ponce</collectorName>
on 
<date id="FF9676F98430FF8BFCBAFCB43EFDFC92" box="[855,1004,799,825]" pageId="3" pageNumber="546" value="2017-05-20">
<collectingDate id="EFD28F118430FF8BFCBAFCB43EFDFC92" box="[855,1004,799,825]" pageId="3" pageNumber="546" value="2017-05-20">20 May 2017</collectingDate>
</date>
at 19:30 hrs.
</materialsCitation>
</paragraph>
<paragraph id="8B9750398430FF8BFF2AFCE83DD0FC0E" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
<materialsCitation id="3B405A648430FF8BFF2AFCE83E8BFCF6" ID-GBIF-Occurrence="3758761301" box="[199,922,834,861]" collectionCode="MHCH" pageId="3" pageNumber="546" specimenCode="MHCH 3241-42, MP 175, 185" specimenCount="1" typeStatus="paratype">
<emphasis id="B95C8C2B8430FF8BFF2AFCE83C2EFCF6" bold="true" box="[199,319,835,861]" pageId="3" pageNumber="546">
<typeStatus id="5493EE9B8430FF8BFF2AFCE83C2EFCF6" box="[199,319,835,861]" pageId="3" pageNumber="546" type="paratype">Paratypes</typeStatus>
</emphasis>
. 
<specimenCode id="DB8EF8428430FF8BFEA2FCE83F00FCF6" box="[335,529,835,861]" collectionCode="MHCH" pageId="3" pageNumber="546">MHCH 3241–42</specimenCode>
, both are juveniles (
<specimenCode id="DB8EF8428430FF8BFD16FCE83E48FCF6" box="[763,857,834,861]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="3" pageNumber="546">MP 175</specimenCode>
, 
<specimenCode id="DB8EF8428430FF8BFC8AFCE93E83FCF6" box="[871,914,834,861]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="3" pageNumber="546">185</specimenCode>
)
</materialsCitation>
collected 
<quantity id="4CD0FDDC8430FF8BFBFFFCE83973FCF6" box="[1042,1122,835,861]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="546" unit="km" value="0.9">0.9 km</quantity>
ca. NE of the type locality (
<geoCoordinate id="EE1C36FE8430FF8BFF72FCCC3C1EFC2A" box="[159,271,871,897]" degrees="8.8682" direction="north" orientation="latitude" pageId="3" pageNumber="546" precision="5" value="8.8682">8.8682°N</geoCoordinate>
, 
<geoCoordinate id="EE1C36FE8430FF8BFEF1FCCC3C8CFC2A" box="[284,413,871,897]" degrees="82.4844" direction="west" orientation="longitude" pageId="3" pageNumber="546" precision="5" value="-82.4844">82.4844°W</geoCoordinate>
, 
<quantity id="4CD0FDDC8430FF8BFE47FCCC3F10FC2A" box="[426,513,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.983" pageId="3" pageNumber="546" unit="m" value="1983.0">1983 m</quantity>
) collected by Marcos Ponce and Roger Morales on 
<date id="FF9676F98430FF8BFBB9FCCC3822FC2A" box="[1108,1331,871,897]" pageId="3" pageNumber="546" value="2017-09-31">31 September 2017</date>
, at 23:00 hrs.
</paragraph>
</subSubSection>
<subSubSection id="C33203B28430FF8DFF2AFC0539DEF850" lastPageId="5" lastPageNumber="548" pageId="3" pageNumber="546" type="diagnosis">
<paragraph id="8B9750398430FF8BFF2AFC053C6CFB0A" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
<emphasis id="B95C8C2B8430FF8BFF2AFC053C55FC63" bold="true" box="[199,324,942,968]" pageId="3" pageNumber="546">Diagnosis:</emphasis>
Assigned to genus 
<taxonomicName id="4C282BBA8430FF8BFDF6FC043FB6FC62" baseAuthorityName="AmphibiaWeb" baseAuthorityYear="2022" box="[539,679,943,969]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8430FF8BFDF6FC043FB6FC62" box="[539,679,943,969]" italics="true" pageId="3" pageNumber="546">Bolitoglossa</emphasis>
</taxonomicName>
due to having fewer than 14 costal grooves and lacking a sublingual fold, and to subgenus 
<taxonomicName id="4C282BBA8430FF8BFE79FC783CE8FC46" box="[404,505,979,1005]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8430FF8BFE79FC783CE8FC46" box="[404,505,979,1005]" italics="true" pageId="3" pageNumber="546">Eladinea</emphasis>
</taxonomicName>
based on mtDNA sequence data. 
<taxonomicName id="4C282BBA8430FF8BFC90FC7839B2FC46" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[893,1187,979,1005]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B8430FF8BFC90FC7839B2FC46" box="[893,1187,979,1005]" italics="true" pageId="3" pageNumber="546">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
is differentiated from all known species of the subgenus 
<taxonomicName id="4C282BBA8430FF8BFDF2FC5C3F95FBBA" box="[543,644,1015,1041]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8430FF8BFDF2FC5C3F95FBBA" box="[543,644,1015,1041]" italics="true" pageId="3" pageNumber="546">Eladinea</emphasis>
</taxonomicName>
by the combination of the following characters: (1) Unique coloration consisting of pinkish-flesh with flame-scarlet speckles, ventral salmon color; (2) Moderately small size with wide head HW/SVL=0.21; (3) moderate -sized extremities; (4) digits with moderate to extensive webbing on the hands and feet; (5) prehensile and considerably longer tail than body TL/SVL=1.5; (6) high tooth counts: premaxillary (7), maxillary teeth (78).
</paragraph>
<paragraph id="8B9750398430FF8BFF2AFB013F04FA69" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
<emphasis id="B95C8C2B8430FF8BFF2AFB013C99FB6F" bold="true" box="[199,392,1194,1220]" pageId="3" pageNumber="546">Similar species:</emphasis>
<taxonomicName id="4C282BBA8430FF8BFE78FB003FD1FB6E" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[405,704,1195,1221]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B8430FF8BFE78FB003FD1FB6E" box="[405,704,1195,1221]" italics="true" pageId="3" pageNumber="546">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
is well differentiated from all other described species of the subgenus 
<taxonomicName id="4C282BBA8430FF8BFEE5FB643C7CFB42" box="[264,365,1231,1257]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8430FF8BFEE5FB643C7CFB42" box="[264,365,1231,1257]" italics="true" pageId="3" pageNumber="546">Eladinea</emphasis>
</taxonomicName>
by 16S mtDNA distances (
<figureCitation id="13134CBC8430FF8BFD44FB643FE1FB42" box="[681,752,1231,1257]" captionStart="FIGURE 2" captionStartId="5.[151,250,901,925]" captionTargetBox="[170,1419,185,875]" captionTargetId="figure-19@5.[168,1419,181,878]" captionTargetPageId="5" captionText="FIGURE 2. Maximum likelihood trees based on 16S mtDNA sequence of Bolitoglossa spp. of the Eladinea subgenus. SHaLRT (Shimodaira and Hasegawa approximate likelihood ratio test) support (%) / ultrafast bootstrap support (%) are shown on the internodes. Outgroups not shown." figureDoi="http://doi.org/10.5281/zenodo.6504108" httpUri="https://zenodo.org/record/6504108/files/figure.png" pageId="3" pageNumber="546">Fig. 2</figureCitation>
; 
<tableCitation id="C6AA65828430FF8BFD10FB643E40FB42" box="[765,849,1231,1257]" captionStart="TABLE 1" captionStartId="4.[151,244,680,707]" captionTargetPageId="4" captionText="TABLE 1. Estimates of uncorrected p-distances between sequences of 16S mtDNA (in %), between Bolitoglossa subpalmata, B. epimela and B. robinsoni species group and B. cathyledecae sp. nov." httpUri="http://table.plazi.org/id/DF5700B18437FF8CFF7AFD033CC9FD4C" pageId="3" pageNumber="546" tableUuid="DF5700B18437FF8CFF7AFD033CC9FD4C">Table 1</tableCitation>
). It is further distinguished morphologically by the following characteristics (condition for 
<taxonomicName id="4C282BBA8430FF8BFD8FFB583E06FAA6" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[610,791,1267,1293]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B8430FF8BFD8FFB583E06FAA6" box="[610,791,1267,1293]" italics="true" pageId="3" pageNumber="546">B. cathyledecae</emphasis>
</taxonomicName>
in parentheses): It differs from 
<taxonomicName id="4C282BBA8430FF8BFB7CFB583867FAA6" authority="Taylor" authorityName="Taylor" authorityYear="1954" box="[1169,1398,1267,1293]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="alvaradoi">
<emphasis id="B95C8C2B8430FF8BFB7CFB583831FAA6" box="[1169,1312,1267,1293]" italics="true" pageId="3" pageNumber="546">B. alvaradoi</emphasis>
Taylor
</taxonomicName>
, 
<taxonomicName id="4C282BBA8430FF8BFA69FB583C43FA9A" authority="Tanner" authorityName="Tanner" authorityYear="1962" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="biseriata">
<emphasis id="B95C8C2B8430FF8BFA69FB583DEAFA9A" italics="true" pageId="3" pageNumber="546">B. biseriata</emphasis>
Tanner
</taxonomicName>
, 
<taxonomicName id="4C282BBA8430FF8BFEB3FABC3EBFFA9A" authority="Batista, Kohler, Mebert and Vesely" authorityName="Batista, Kohler, Mebert and Vesely" box="[350,942,1303,1329]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="chucantiensis">
<emphasis id="B95C8C2B8430FF8BFEB3FABC3F06FA9A" box="[350,535,1303,1329]" italics="true" pageId="3" pageNumber="546">B. chucantiensis</emphasis>
Batista, Köhler, Mebert and Vesely
</taxonomicName>
, 
<taxonomicName id="4C282BBA8430FF8BFC57FABC399BFA9A" authority="Dunn" authorityName="Dunn" box="[954,1162,1303,1329]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="colonnea">
<emphasis id="B95C8C2B8430FF8BFC57FABC3950FA9A" box="[954,1089,1303,1329]" italics="true" pageId="3" pageNumber="546">B. colonnea</emphasis>
Dunn
</taxonomicName>
, 
<taxonomicName id="4C282BBA8430FF8BFB7AFABC39FCFA9B" authorityName="Wake, Brame &amp; Duellman" authorityYear="1973" box="[1175,1261,1303,1328]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cuna">
<emphasis id="B95C8C2B8430FF8BFB7AFABC39FCFA9B" box="[1175,1261,1303,1328]" italics="true" pageId="3" pageNumber="546">B. cuna</emphasis>
</taxonomicName>
, 
<collectingCountry id="F33F10A98430FF8BFB11FABC382FFA9A" box="[1276,1342,1303,1329]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, Brame, and Duellman 
<taxonomicName id="4C282BBA8430FF8BFED6FA903CD6FAFE" box="[315,455,1339,1365]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="lignicolor">
<emphasis id="B95C8C2B8430FF8BFED6FA903CD6FAFE" box="[315,455,1339,1365]" italics="true" pageId="3" pageNumber="546">B. lignicolor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8430FF8BFE38FA903F84FAFE" authority="Brame" authorityName="Brame" box="[469,661,1339,1365]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="medemi">
<emphasis id="B95C8C2B8430FF8BFE38FA903F55FAFE" box="[469,580,1339,1365]" italics="true" pageId="3" pageNumber="546">B.medemi</emphasis>
Brame
</taxonomicName>
and 
<collectingCountry id="F33F10A98430FF8BFD24FA903E1AFAFE" box="[713,779,1339,1365]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, 
<taxonomicName id="4C282BBA8430FF8BFCFBFA903E89FAFE" box="[790,920,1339,1365]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="minutula">
<emphasis id="B95C8C2B8430FF8BFCFBFA903E89FAFE" box="[790,920,1339,1365]" italics="true" pageId="3" pageNumber="546">B. minutula</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98430FF8BFC73FA903EF1FAFE" box="[926,992,1339,1365]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, Brame and Duellman, 
<taxonomicName id="4C282BBA8430FF8BFB08FA90388DFAFE" authorityName="KRL" authorityYear="1179" box="[1253,1436,1339,1365]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="schizodactyla">
<emphasis id="B95C8C2B8430FF8BFB08FA90388DFAFE" box="[1253,1436,1339,1365]" italics="true" pageId="3" pageNumber="546">B. schizodactyla</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98430FF8BFF7AFAF43DC7FAD2" box="[151,214,1375,1401]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
and Brame, 
<taxonomicName id="4C282BBA8430FF8BFE83FAF43CCDFAD2" authorityName="MHCH" authorityYear="2668" box="[366,476,1375,1401]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B95C8C2B8430FF8BFE83FAF43CCDFAD2" box="[366,476,1375,1401]" italics="true" pageId="3" pageNumber="546">B. taylori</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98430FF8BFE08FAF43F36FAD2" box="[485,551,1375,1401]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, Brame and Myers, by its less extensively webbed hands and feet. All these species have fully webbed hands and feet (well-developed digital webbing; only portions of distal-most phalanges of longest digits free of webbing).
</paragraph>
<paragraph id="8B9750398430FF8BFF2AFA603E9EF9DE" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
<taxonomicName id="4C282BBA8430FF8BFF2AFA603CFFFA4E" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[199,494,1483,1509]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B8430FF8BFF2AFA603CFFFA4E" box="[199,494,1483,1509]" italics="true" pageId="3" pageNumber="546">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
is easily distinguishable by its unique coloration (pinkish flesh with flame scarlet speckles and ventral salmon color) from the large black salamanders found between 
<collectingCountry id="F33F10A98430FF8BFB92FA4439ECF9A2" box="[1151,1277,1519,1545]" name="Costa Rica" pageId="3" pageNumber="546">Costa Rica</collectingCountry>
and 
<collectingCountry id="F33F10A98430FF8BFAD0FA443888F9A2" box="[1341,1433,1519,1545]" name="Panama" pageId="3" pageNumber="546">Panamá</collectingCountry>
, including the almost entirely black species 
<taxonomicName id="4C282BBA8430FF8BFD6EF9B83EC4F986" authority="Brame, Savage" authorityName="Brame, Savage" box="[643,981,1555,1581]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="anthracina">
<emphasis id="B95C8C2B8430FF8BFD6EF9B83E0EF986" box="[643,799,1555,1581]" italics="true" pageId="3" pageNumber="546">B. anthracina</emphasis>
Brame, Savage
</taxonomicName>
, 
<collectingCountry id="F33F10A98430FF8BFC0CF9B83931F986" box="[993,1056,1555,1581]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
and Hanken; 
<taxonomicName id="4C282BBA8430FF8BFB2CF9B8385FF987" box="[1217,1358,1555,1580]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="compacta">
<emphasis id="B95C8C2B8430FF8BFB2CF9B8385FF987" box="[1217,1358,1555,1580]" italics="true" pageId="3" pageNumber="546">B. compacta</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98430FF8BFABBF9B83889F986" box="[1366,1432,1555,1581]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, Brame and Duellman; 
<taxonomicName id="4C282BBA8430FF8BFE7AF99C3CE4F9FB" authorityName="Wake, Hanken &amp; Ibanez" authorityYear="2005" box="[407,501,1591,1616]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="copia">
<emphasis id="B95C8C2B8430FF8BFE7AF99C3CE4F9FB" box="[407,501,1591,1616]" italics="true" pageId="3" pageNumber="546">B. copia</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98430FF8BFE10F99C3F2EF9FA" box="[509,575,1591,1617]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
, Hanken and Ibañez; 
<taxonomicName id="4C282BBA8430FF8BFCDEF99C3935F9FA" authority="Hanken" authorityName="Hanken" box="[819,1060,1591,1617]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="magnifica">
<emphasis id="B95C8C2B8430FF8BFCDEF99C3ED3F9FA" box="[819,962,1591,1617]" italics="true" pageId="3" pageNumber="546">B. magnifica</emphasis>
Hanken
</taxonomicName>
, 
<collectingCountry id="F33F10A98430FF8BFBC2F99C397FF9FA" box="[1071,1134,1591,1617]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
and Savage; 
<taxonomicName id="4C282BBA8430FF8BFAEBF99C3DF2F9DE" authority="Taylor" authorityName="Taylor" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="nigrescens">
<emphasis id="B95C8C2B8430FF8BFAEBF99C388CF9FB" box="[1286,1437,1591,1616]" italics="true" pageId="3" pageNumber="546">B. nigrescens</emphasis>
Taylor
</taxonomicName>
; 
<taxonomicName id="4C282BBA8430FF8BFF02F9F03CB9F9DE" authority="Cope" authorityName="Cope" box="[239,424,1627,1653]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="robusta">
<emphasis id="B95C8C2B8430FF8BFF02F9F03C72F9DE" box="[239,355,1627,1653]" italics="true" pageId="3" pageNumber="546">B. robusta</emphasis>
Cope
</taxonomicName>
; and 
<taxonomicName id="4C282BBA8430FF8BFE0EF9F03FA6F9DE" authority="Hanken" authorityName="Hanken" box="[483,695,1627,1653]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="sombra">
<emphasis id="B95C8C2B8430FF8BFE0EF9F03F47F9DE" box="[483,598,1627,1653]" italics="true" pageId="3" pageNumber="546">B. sombra</emphasis>
Hanken
</taxonomicName>
, 
<collectingCountry id="F33F10A98430FF8BFD2CF9F03E11F9DE" box="[705,768,1627,1653]" name="Wake Island" pageId="3" pageNumber="546">Wake</collectingCountry>
and Savage.
</paragraph>
<paragraph id="8B9750398430FF8BFF2AF9D43C06F811" blockId="3.[151,1437,727,1978]" pageId="3" pageNumber="546">
This new species is differentiated from other salamanders found in the Talamanca Mountain range between 
<collectingCountry id="F33F10A98430FF8BFF7AF9083C03F916" box="[151,274,1699,1725]" name="Costa Rica" pageId="3" pageNumber="546">Costa Rica</collectingCountry>
and 
<collectingCountry id="F33F10A98430FF8BFEA6F9083CB5F916" box="[331,420,1699,1725]" name="Panama" pageId="3" pageNumber="546">Panama</collectingCountry>
as follows: 
<taxonomicName id="4C282BBA8430FF8BFDC2F9083ED2F916" authority="Boulenger" authorityName="Boulenger" box="[559,963,1699,1725]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8430FF8BFDC2F9083E57F916" box="[559,838,1699,1725]" italics="true" pageId="3" pageNumber="546">Bolitoglossa subpalmata</emphasis>
Boulenger
</taxonomicName>
a moderate-sized species with SVL of 
<quantity id="4CD0FDDC8430FF8BFA6DF9083DD4F94A" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="546" unit="mm" value="60.0">60 mm</quantity>
, (
<quantity id="4CD0FDDC8430FF8BFF35F96C3C36F94A" box="[216,295,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="546" unit="mm" value="46.0">46 mm</quantity>
in adult male), slate-gray color with brownish flecking evident on the back; grayish venter, the median area darker than outer portions (pinkish flesh with flame scarlet speckles and ventral salmon color). 
<taxonomicName id="4C282BBA8430FF8BFAFDF9403C5EF882" authority="Taylor" authorityName="Taylor" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B95C8C2B8430FF8BFAFDF9403DEDF882" italics="true" pageId="3" pageNumber="546">Bolitoglossa pesrubra</emphasis>
Taylor
</taxonomicName>
, is a moderate-sized salamander with an SVL = 
<quantity id="4CD0FDDC8430FF8BFC9CF8A43ED2F882" box="[881,963,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="546" unit="mm" value="67.0">67 mm</quantity>
, (
<quantity id="4CD0FDDC8430FF8BFC3AF8A43938F882" box="[983,1065,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="546" unit="mm" value="46.0">46 mm</quantity>
) and has fewer premaxillary and maxillary teeth, with 3 PMT (7), MT and 66 MT (78), has a shorter tail in proportion to body size, TL/SVL = 1, (1.5). 
<taxonomicName id="4C282BBA8430FF8BFF36F8FC3CF2F8DB" box="[219,483,1879,1905]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="marmorea">
<emphasis id="B95C8C2B8430FF8BFF36F8FC3CF2F8DB" box="[219,483,1879,1905]" italics="true" pageId="3" pageNumber="546">Bolitoglossa marmorea</emphasis>
</taxonomicName>
a moderated-sized species with an SVL of 
<quantity id="4CD0FDDC8430FF8BFC21F8FC3922F8DA" box="[972,1075,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="6.01" pageId="3" pageNumber="546" unit="mm" value="60.1">60.1 mm</quantity>
, (
<quantity id="4CD0FDDC8430FF8BFBAAF8FC3988F8DA" box="[1095,1177,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="546" unit="mm" value="46.0">46 mm</quantity>
) having longer legs, in which the toes of the adpressed legs are usually in contact (limb interval between toes of: 3, not in contact when adpressed).
</paragraph>
<caption id="DF5700B18437FF8CFF7AFD033CC9FD4C" ID-Table-UUID="DF5700B18437FF8CFF7AFD033CC9FD4C" httpUri="http://table.plazi.org/id/DF5700B18437FF8CFF7AFD033CC9FD4C" pageId="4" pageNumber="547" startId="4.[151,244,680,707]" targetBox="[165,1996,758,1508]" targetIsTable="true" targetPageId="4">
<paragraph id="8B9750398437FF8CFF7AFD033CC9FD4C" blockId="4.[151,2038,680,743]" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFF7AFD033C01FD68" bold="true" box="[151,272,680,707]" pageId="4" pageNumber="547">TABLE 1.</emphasis>
Estimates of uncorrected p-distances between sequences of 16S mtDNA (in %), between 
<taxonomicName id="4C282BBA8437FF8CFB11FD033B03FD68" box="[1276,1554,680,707]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8437FF8CFB11FD033B03FD68" box="[1276,1554,680,707]" italics="true" pageId="4" pageNumber="547">Bolitoglossa subpalmata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8437FF8CF9CDFD033B86FD68" box="[1568,1687,680,707]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B8437FF8CF9CDFD033B86FD68" box="[1568,1687,680,707]" italics="true" pageId="4" pageNumber="547">B. epimela</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C282BBA8437FF8CF920FD033A47FD68" box="[1741,1878,680,707]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="B95C8C2B8437FF8CF920FD033A47FD68" box="[1741,1878,680,707]" italics="true" pageId="4" pageNumber="547">B. robinsoni</emphasis>
</taxonomicName>
species group and 
<taxonomicName id="4C282BBA8437FF8CFF2BFD673C66FD4C" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[198,375,716,743]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B8437FF8CFF2BFD673C66FD4C" box="[198,375,716,743]" italics="true" pageId="4" pageNumber="547">B. cathyledecae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F31508437FF8CFE92FD673CC9FD4C" box="[383,472,716,743]" pageId="4" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B9750398437FF8CFF48FD5D3ADDFA48" pageId="4" pageNumber="547">
<table id="F928A29984370077FF48FD5D3ADDFA4F" box="[165,1996,758,1508]" gridcols="19" gridrows="19" pageId="4" pageNumber="547">
<tr id="3518527B84370077FF48FD5D3ADDFCA4" box="[165,1996,758,783]" gridrow="0" pageId="4" pageNumber="547">
<th id="76C93B0784370077FF48FD5D3CE8FCA4" box="[165,505,758,783]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="4" pageNumber="547">Species</th>
<th id="76C93B0784370077FDBFFD5D3F63FCA4" box="[594,626,758,783]" gridcol="2" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFDBFFD5D3F4EFCA4" bold="true" box="[594,607,758,783]" pageId="4" pageNumber="547">1</emphasis>
</th>
<th id="76C93B0784370077FD45FD5D3FD9FCA4" box="[680,712,758,783]" gridcol="3" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFD45FD5D3FA4FCA4" bold="true" box="[680,693,758,783]" pageId="4" pageNumber="547">2</emphasis>
</th>
<th id="76C93B0784370077FD10FD5D3E0CFCA4" box="[765,797,758,783]" gridcol="4" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFD10FD5D3E1BFCA4" bold="true" box="[765,778,758,783]" pageId="4" pageNumber="547">3</emphasis>
</th>
<th id="76C93B0784370077FCBEFD5D3E62FCA4" box="[851,883,758,783]" gridcol="5" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFCBEFD5D3E71FCA4" bold="true" box="[851,864,758,783]" pageId="4" pageNumber="547">4</emphasis>
</th>
<th id="76C93B0784370077FC44FD5D3ED8FCA4" box="[937,969,758,783]" gridcol="6" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFC44FD5D3EA7FCA4" bold="true" box="[937,950,758,783]" pageId="4" pageNumber="547">5</emphasis>
</th>
<th id="76C93B0784370077FC13FD5D390FFCA4" box="[1022,1054,758,783]" gridcol="7" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFC13FD5D391AFCA4" bold="true" box="[1022,1035,758,783]" pageId="4" pageNumber="547">6</emphasis>
</th>
<th id="76C93B0784370077FBB9FD5D3965FCA4" box="[1108,1140,758,783]" gridcol="8" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFBB9FD5D3970FCA4" bold="true" box="[1108,1121,758,783]" pageId="4" pageNumber="547">7</emphasis>
</th>
<th id="76C93B0784370077FB47FD5D39DBFCA4" box="[1194,1226,758,783]" gridcol="9" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFB47FD5D39A6FCA4" bold="true" box="[1194,1207,758,783]" pageId="4" pageNumber="547">8</emphasis>
</th>
<th id="76C93B0784370077FB12FD5D380EFCA4" box="[1279,1311,758,783]" gridcol="10" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFB12FD5D381DFCA4" bold="true" box="[1279,1292,758,783]" pageId="4" pageNumber="547">9</emphasis>
</th>
<th id="76C93B0784370077FAB8FD5D3864FCA4" box="[1365,1397,758,783]" gridcol="11" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFAB8FD5D387FFCA4" bold="true" box="[1365,1390,758,783]" pageId="4" pageNumber="547">10</emphasis>
</th>
<th id="76C93B0784370077FA47FD5D38DBFCA4" box="[1450,1482,758,783]" gridcol="12" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CFA47FD5D38D3FCA4" bold="true" box="[1450,1474,758,783]" pageId="4" pageNumber="547">11</emphasis>
</th>
<th id="76C93B0784370077F9EDFD5D3B31FCA4" box="[1536,1568,758,783]" gridcol="13" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF9EDFD5D3B08FCA4" bold="true" box="[1536,1561,758,783]" pageId="4" pageNumber="547">12</emphasis>
</th>
<th id="76C93B0784370077F9B8FD5D3B64FCA4" box="[1621,1653,758,783]" gridcol="14" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF9B8FD5D3B7FFCA4" bold="true" box="[1621,1646,758,783]" pageId="4" pageNumber="547">13</emphasis>
</th>
<th id="76C93B0784370077F946FD5D3BDAFCA4" box="[1707,1739,758,783]" gridcol="15" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF946FD5D3BD5FCA4" bold="true" box="[1707,1732,758,783]" pageId="4" pageNumber="547">14</emphasis>
</th>
<th id="76C93B0784370077F8ECFD5D3A30FCA4" box="[1793,1825,758,783]" gridcol="16" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF8ECFD5D3A0BFCA4" bold="true" box="[1793,1818,758,783]" pageId="4" pageNumber="547">15</emphasis>
</th>
<th id="76C93B0784370077F8BBFD5D3A67FCA4" box="[1878,1910,758,783]" gridcol="17" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF8BBFD5D3A7EFCA4" bold="true" box="[1878,1903,758,783]" pageId="4" pageNumber="547">16</emphasis>
</th>
<th id="76C93B0784370077F841FD5D3ADDFCA4" box="[1964,1996,758,783]" gridcol="18" gridrow="0" pageId="4" pageNumber="547">
<emphasis id="B95C8C2B8437FF8CF841FD5D3AD4FCA4" bold="true" box="[1964,1989,758,783]" pageId="4" pageNumber="547">17</emphasis>
</th>
</tr>
<tr id="3518527B84370077FF48FCB53ADDFC9C" box="[165,1996,798,823]" gridrow="1" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FCB53DD4FC9C" box="[165,197,798,823]" gridcol="0" gridrow="1" pageId="4" pageNumber="547">1.</th>
<td id="76C93B0784370077FEECFCB53CE8FC9C" box="[257,505,798,823]" gridcol="1" gridrow="1" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFCB53C68FC9C" box="[257,377,798,823]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B95C8C2B8437FF8CFEECFCB53C68FC9C" box="[257,377,798,823]" italics="true" pageId="4" pageNumber="547">B. pesrubra</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3518527B84370077FF48FCED3ADDFCF4" box="[165,1996,838,863]" gridrow="2" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FCED3DD4FCF4" box="[165,197,838,863]" gridcol="0" gridrow="2" pageId="4" pageNumber="547">2.</th>
<td id="76C93B0784370077FEECFCED3CE8FCF4" box="[257,505,838,863]" gridcol="1" gridrow="2" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFCED3C71FCF4" box="[257,352,838,863]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B95C8C2B8437FF8CFEECFCED3C71FCF4" box="[257,352,838,863]" italics="true" pageId="4" pageNumber="547">B. kamuk</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFCED3F63FCF4" box="[594,626,838,863]" gridcol="2" gridrow="2" pageId="4" pageNumber="547">1.3</td>
</tr>
<tr id="3518527B84370077FF48FCC43ADDFC23" box="[165,1996,879,904]" gridrow="3" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FCC43DD4FC23" box="[165,197,879,904]" gridcol="0" gridrow="3" pageId="4" pageNumber="547">3.</th>
<td id="76C93B0784370077FEECFCC43CE8FC23" box="[257,505,879,904]" gridcol="1" gridrow="3" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFCC43C9CFC23" box="[257,397,879,904]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="anthracina">
<emphasis id="B95C8C2B8437FF8CFEECFCC43C9CFC23" box="[257,397,879,904]" italics="true" pageId="4" pageNumber="547">B. anthracina</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFCC43F63FC23" box="[594,626,879,904]" gridcol="2" gridrow="3" pageId="4" pageNumber="547">3.5</td>
<td id="76C93B0784370077FD45FCC43FD9FC23" box="[680,712,879,904]" gridcol="3" gridrow="3" pageId="4" pageNumber="547">3.0</td>
</tr>
<tr id="3518527B84370077FF48FC3C3ADDFC1B" box="[165,1996,919,944]" gridrow="4" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FC3C3DD4FC1B" box="[165,197,919,944]" gridcol="0" gridrow="4" pageId="4" pageNumber="547">4.</th>
<td id="76C93B0784370077FEECFC3C3CE8FC1B" box="[257,505,919,944]" gridcol="1" gridrow="4" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFC3C3C90FC1B" box="[257,385,919,944]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="splendida">
<emphasis id="B95C8C2B8437FF8CFEECFC3C3C90FC1B" box="[257,385,919,944]" italics="true" pageId="4" pageNumber="547">B. splendida</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFC3C3F63FC1B" box="[594,626,919,944]" gridcol="2" gridrow="4" pageId="4" pageNumber="547">3.2</td>
<td id="76C93B0784370077FD45FC3C3FD9FC1B" box="[680,712,919,944]" gridcol="3" gridrow="4" pageId="4" pageNumber="547">2.6</td>
<td id="76C93B0784370077FD10FC3C3E0CFC1B" box="[765,797,919,944]" gridcol="4" gridrow="4" pageId="4" pageNumber="547">3.7</td>
</tr>
<tr id="3518527B84370077FF48FC143ADDFC73" box="[165,1996,959,984]" gridrow="5" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FC143DD4FC73" box="[165,197,959,984]" gridcol="0" gridrow="5" pageId="4" pageNumber="547">5.</th>
<td id="76C93B0784370077FEECFC143CE8FC73" box="[257,505,959,984]" gridcol="1" gridrow="5" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFC143C85FC73" box="[257,404,959,984]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8437FF8CFEECFC143C85FC73" box="[257,404,959,984]" italics="true" pageId="4" pageNumber="547">B. subpalmata</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFC143F63FC73" box="[594,626,959,984]" gridcol="2" gridrow="5" pageId="4" pageNumber="547">2.4</td>
<td id="76C93B0784370077FD45FC143FD9FC73" box="[680,712,959,984]" gridcol="3" gridrow="5" pageId="4" pageNumber="547">1.3</td>
<td id="76C93B0784370077FD10FC143E0CFC73" box="[765,797,959,984]" gridcol="4" gridrow="5" pageId="4" pageNumber="547">3.4</td>
<td id="76C93B0784370077FCBEFC143E62FC73" box="[851,883,959,984]" gridcol="5" gridrow="5" pageId="4" pageNumber="547">2.6</td>
</tr>
<tr id="3518527B84370077FF48FC433ADDFBAA" box="[165,1996,1000,1025]" gridrow="6" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FC433DD4FBAA" box="[165,197,1000,1025]" gridcol="0" gridrow="6" pageId="4" pageNumber="547">6.</th>
<td id="76C93B0784370077FEECFC433CE8FBAA" box="[257,505,1000,1025]" gridcol="1" gridrow="6" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFC433C55FBAA" box="[257,324,1000,1025]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="tica">
<emphasis id="B95C8C2B8437FF8CFEECFC433C55FBAA" box="[257,324,1000,1025]" italics="true" pageId="4" pageNumber="547">B. tica</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFC433F63FBAA" box="[594,626,1000,1025]" gridcol="2" gridrow="6" pageId="4" pageNumber="547">3.4</td>
<td id="76C93B0784370077FD45FC433FD9FBAA" box="[680,712,1000,1025]" gridcol="3" gridrow="6" pageId="4" pageNumber="547">3.0</td>
<td id="76C93B0784370077FD10FC433E0CFBAA" box="[765,797,1000,1025]" gridcol="4" gridrow="6" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FCBEFC433E62FBAA" box="[851,883,1000,1025]" gridcol="5" gridrow="6" pageId="4" pageNumber="547">4.5</td>
<td id="76C93B0784370077FC44FC433ED8FBAA" box="[937,969,1000,1025]" gridcol="6" gridrow="6" pageId="4" pageNumber="547">1.9</td>
</tr>
<tr id="3518527B84370077FF48FBBB3ADDFB82" box="[165,1996,1040,1065]" gridrow="7" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-8="1" rowspan-9="1">
<th id="76C93B0784370077FF48FBBB3DD4FB82" box="[165,197,1040,1065]" gridcol="0" gridrow="7" pageId="4" pageNumber="547">7.</th>
<td id="76C93B0784370077FEECFBBB3CE8FB82" box="[257,505,1040,1065]" gridcol="1" gridrow="7" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFBBB3C7AFB82" box="[257,363,1040,1065]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B95C8C2B8437FF8CFEECFBBB3C7AFB82" box="[257,363,1040,1065]" italics="true" pageId="4" pageNumber="547">B. gracilis</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFBBB3F63FB82" box="[594,626,1040,1065]" gridcol="2" gridrow="7" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FD45FBBB3FD9FB82" box="[680,712,1040,1065]" gridcol="3" gridrow="7" pageId="4" pageNumber="547">3.6</td>
<td id="76C93B0784370077FD10FBBB3E0CFB82" box="[765,797,1040,1065]" gridcol="4" gridrow="7" pageId="4" pageNumber="547">4.5</td>
<td id="76C93B0784370077FCBEFBBB3E62FB82" box="[851,883,1040,1065]" gridcol="5" gridrow="7" pageId="4" pageNumber="547">4.7</td>
<td id="76C93B0784370077FC44FBBB3ED8FB82" box="[937,969,1040,1065]" gridcol="6" gridrow="7" pageId="4" pageNumber="547">3.0</td>
<td id="76C93B0784370077FC13FBBB390FFB82" box="[1022,1054,1040,1065]" gridcol="7" gridrow="7" pageId="4" pageNumber="547">4.1</td>
</tr>
<tr id="3518527B84370077FF48FB933ADDFBFA" box="[165,1996,1080,1105]" gridrow="8" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-9="1">
<th id="76C93B0784370077FF48FB933DD4FBFA" box="[165,197,1080,1105]" gridcol="0" gridrow="8" pageId="4" pageNumber="547">8.</th>
<td id="76C93B0784370077FEECFB933CE8FBFA" box="[257,505,1080,1105]" gridcol="1" gridrow="8" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFB933C77FBFA" box="[257,358,1080,1105]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="bramei">
<emphasis id="B95C8C2B8437FF8CFEECFB933C77FBFA" box="[257,358,1080,1105]" italics="true" pageId="4" pageNumber="547">B. bramei</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFB933F63FBFA" box="[594,626,1080,1105]" gridcol="2" gridrow="8" pageId="4" pageNumber="547">3.2</td>
<td id="76C93B0784370077FD45FB933FD9FBFA" box="[680,712,1080,1105]" gridcol="3" gridrow="8" pageId="4" pageNumber="547">2.4</td>
<td id="76C93B0784370077FD10FB933E0CFBFA" box="[765,797,1080,1105]" gridcol="4" gridrow="8" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FCBEFB933E62FBFA" box="[851,883,1080,1105]" gridcol="5" gridrow="8" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FC44FB933ED8FBFA" box="[937,969,1080,1105]" gridcol="6" gridrow="8" pageId="4" pageNumber="547">2.6</td>
<td id="76C93B0784370077FC13FB93390FFBFA" box="[1022,1054,1080,1105]" gridcol="7" gridrow="8" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FBB9FB933965FBFA" box="[1108,1140,1080,1105]" gridcol="8" gridrow="8" pageId="4" pageNumber="547">3.0</td>
</tr>
<tr id="3518527B84370077FF48FBCB3ADDFBD2" box="[165,1996,1120,1145]" gridrow="9" pageId="4" pageNumber="547" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FBCB3DD4FBD2" box="[165,197,1120,1145]" gridcol="0" gridrow="9" pageId="4" pageNumber="547">9.</th>
<td id="76C93B0784370077FEECFBCB3CE8FBD2" box="[257,505,1120,1145]" gridcol="1" gridrow="9" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFBCB3C77FBD2" box="[257,358,1120,1145]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="gomezi">
<emphasis id="B95C8C2B8437FF8CFEECFBCB3C77FBD2" box="[257,358,1120,1145]" italics="true" pageId="4" pageNumber="547">B. gomezi</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFBCB3F63FBD2" box="[594,626,1120,1145]" gridcol="2" gridrow="9" pageId="4" pageNumber="547">3.6</td>
<td id="76C93B0784370077FD45FBCB3FD9FBD2" box="[680,712,1120,1145]" gridcol="3" gridrow="9" pageId="4" pageNumber="547">2.6</td>
<td id="76C93B0784370077FD10FBCB3E0CFBD2" box="[765,797,1120,1145]" gridcol="4" gridrow="9" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077FCBEFBCB3E62FBD2" box="[851,883,1120,1145]" gridcol="5" gridrow="9" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FC44FBCB3ED8FBD2" box="[937,969,1120,1145]" gridcol="6" gridrow="9" pageId="4" pageNumber="547">3.0</td>
<td id="76C93B0784370077FC13FBCB390FFBD2" box="[1022,1054,1120,1145]" gridcol="7" gridrow="9" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077FBB9FBCB3965FBD2" box="[1108,1140,1120,1145]" gridcol="8" gridrow="9" pageId="4" pageNumber="547">4.7</td>
<td id="76C93B0784370077FB47FBCB39DBFBD2" box="[1194,1226,1120,1145]" gridcol="9" gridrow="9" pageId="4" pageNumber="547">3.0</td>
</tr>
<tr id="3518527B84370077FF48FB223ADDFB09" box="[165,1996,1161,1186]" gridrow="10" pageId="4" pageNumber="547" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FB223DD4FB09" box="[165,197,1161,1186]" gridcol="0" gridrow="10" pageId="4" pageNumber="547">10.</th>
<td id="76C93B0784370077FEECFB223CE8FB09" box="[257,505,1161,1186]" gridcol="1" gridrow="10" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFB223C93FB09" box="[257,386,1161,1186]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="sooyorum">
<emphasis id="B95C8C2B8437FF8CFEECFB223C93FB09" box="[257,386,1161,1186]" italics="true" pageId="4" pageNumber="547">B. sooyorum</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFB223F63FB09" box="[594,626,1161,1186]" gridcol="2" gridrow="10" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077FD45FB223FD9FB09" box="[680,712,1161,1186]" gridcol="3" gridrow="10" pageId="4" pageNumber="547">3.3</td>
<td id="76C93B0784370077FD10FB223E0CFB09" box="[765,797,1161,1186]" gridcol="4" gridrow="10" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FCBEFB223E62FB09" box="[851,883,1161,1186]" gridcol="5" gridrow="10" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FC44FB223ED8FB09" box="[937,969,1161,1186]" gridcol="6" gridrow="10" pageId="4" pageNumber="547">3.5</td>
<td id="76C93B0784370077FC13FB22390FFB09" box="[1022,1054,1161,1186]" gridcol="7" gridrow="10" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FBB9FB223965FB09" box="[1108,1140,1161,1186]" gridcol="8" gridrow="10" pageId="4" pageNumber="547">5.7</td>
<td id="76C93B0784370077FB47FB2239DBFB09" box="[1194,1226,1161,1186]" gridcol="9" gridrow="10" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077FB12FB22380EFB09" box="[1279,1311,1161,1186]" gridcol="10" gridrow="10" pageId="4" pageNumber="547">3.9</td>
</tr>
<tr id="3518527B84370077FF48FB1A3ADDFB61" box="[165,1996,1201,1226]" gridrow="11" pageId="4" pageNumber="547" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FB1A3DD4FB61" box="[165,197,1201,1226]" gridcol="0" gridrow="11" pageId="4" pageNumber="547">11.</th>
<td id="76C93B0784370077FEECFB1A3CE8FB61" box="[257,505,1201,1226]" gridcol="1" gridrow="11" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFB1A3C69FB61" box="[257,376,1201,1226]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="minutula">
<emphasis id="B95C8C2B8437FF8CFEECFB1A3C69FB61" box="[257,376,1201,1226]" italics="true" pageId="4" pageNumber="547">B. minutula</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFB1A3F63FB61" box="[594,626,1201,1226]" gridcol="2" gridrow="11" pageId="4" pageNumber="547">3.5</td>
<td id="76C93B0784370077FD45FB1A3FD9FB61" box="[680,712,1201,1226]" gridcol="3" gridrow="11" pageId="4" pageNumber="547">2.6</td>
<td id="76C93B0784370077FD10FB1A3E0CFB61" box="[765,797,1201,1226]" gridcol="4" gridrow="11" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FCBEFB1A3E62FB61" box="[851,883,1201,1226]" gridcol="5" gridrow="11" pageId="4" pageNumber="547">4.6</td>
<td id="76C93B0784370077FC44FB1A3ED8FB61" box="[937,969,1201,1226]" gridcol="6" gridrow="11" pageId="4" pageNumber="547">3.3</td>
<td id="76C93B0784370077FC13FB1A390FFB61" box="[1022,1054,1201,1226]" gridcol="7" gridrow="11" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077FBB9FB1A3965FB61" box="[1108,1140,1201,1226]" gridcol="8" gridrow="11" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FB47FB1A39DBFB61" box="[1194,1226,1201,1226]" gridcol="9" gridrow="11" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FB12FB1A380EFB61" box="[1279,1311,1201,1226]" gridcol="10" gridrow="11" pageId="4" pageNumber="547">3.7</td>
<td id="76C93B0784370077FAB8FB1A3864FB61" box="[1365,1397,1201,1226]" gridcol="11" gridrow="11" pageId="4" pageNumber="547">1.7</td>
</tr>
<tr id="3518527B84370077FF48FB723ADDFB59" box="[165,1996,1241,1266]" gridrow="12" pageId="4" pageNumber="547" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FB723DD4FB59" box="[165,197,1241,1266]" gridcol="0" gridrow="12" pageId="4" pageNumber="547">12.</th>
<td id="76C93B0784370077FEECFB723CE8FB59" box="[257,505,1241,1266]" gridcol="1" gridrow="12" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFB723C96FB59" box="[257,391,1241,1266]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="marmorea">
<emphasis id="B95C8C2B8437FF8CFEECFB723C96FB59" box="[257,391,1241,1266]" italics="true" pageId="4" pageNumber="547">B. marmorea</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFB723F63FB59" box="[594,626,1241,1266]" gridcol="2" gridrow="12" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FD45FB723FD9FB59" box="[680,712,1241,1266]" gridcol="3" gridrow="12" pageId="4" pageNumber="547">3.0</td>
<td id="76C93B0784370077FD10FB723E0CFB59" box="[765,797,1241,1266]" gridcol="4" gridrow="12" pageId="4" pageNumber="547">5.3</td>
<td id="76C93B0784370077FCBEFB723E62FB59" box="[851,883,1241,1266]" gridcol="5" gridrow="12" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FC44FB723ED8FB59" box="[937,969,1241,1266]" gridcol="6" gridrow="12" pageId="4" pageNumber="547">3.3</td>
<td id="76C93B0784370077FC13FB72390FFB59" box="[1022,1054,1241,1266]" gridcol="7" gridrow="12" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FBB9FB723965FB59" box="[1108,1140,1241,1266]" gridcol="8" gridrow="12" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FB47FB7239DBFB59" box="[1194,1226,1241,1266]" gridcol="9" gridrow="12" pageId="4" pageNumber="547">3.9</td>
<td id="76C93B0784370077FB12FB72380EFB59" box="[1279,1311,1241,1266]" gridcol="10" gridrow="12" pageId="4" pageNumber="547">3.7</td>
<td id="76C93B0784370077FAB8FB723864FB59" box="[1365,1397,1241,1266]" gridcol="11" gridrow="12" pageId="4" pageNumber="547">2.0</td>
<td id="76C93B0784370077FA47FB7238DBFB59" box="[1450,1482,1241,1266]" gridcol="12" gridrow="12" pageId="4" pageNumber="547">2.0</td>
</tr>
<tr id="3518527B84370077FF48FAAA3ADDFAB1" box="[165,1996,1281,1306]" gridrow="13" pageId="4" pageNumber="547" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FAAA3DD4FAB1" box="[165,197,1281,1306]" gridcol="0" gridrow="13" pageId="4" pageNumber="547">13.</th>
<td id="76C93B0784370077FEECFAAA3CE8FAB1" box="[257,505,1281,1306]" gridcol="1" gridrow="13" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFAAA3C96FAB1" box="[257,391,1281,1306]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="cerroensis">
<emphasis id="B95C8C2B8437FF8CFEECFAAA3C96FAB1" box="[257,391,1281,1306]" italics="true" pageId="4" pageNumber="547">B. cerroensis</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFAAA3F63FAB1" box="[594,626,1281,1306]" gridcol="2" gridrow="13" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FD45FAAA3FD9FAB1" box="[680,712,1281,1306]" gridcol="3" gridrow="13" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FD10FAAA3E0CFAB1" box="[765,797,1281,1306]" gridcol="4" gridrow="13" pageId="4" pageNumber="547">6.3</td>
<td id="76C93B0784370077FCBEFAAA3E62FAB1" box="[851,883,1281,1306]" gridcol="5" gridrow="13" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FC44FAAA3ED8FAB1" box="[937,969,1281,1306]" gridcol="6" gridrow="13" pageId="4" pageNumber="547">4.6</td>
<td id="76C93B0784370077FC13FAAA390FFAB1" box="[1022,1054,1281,1306]" gridcol="7" gridrow="13" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FBB9FAAA3965FAB1" box="[1108,1140,1281,1306]" gridcol="8" gridrow="13" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077FB47FAAA39DBFAB1" box="[1194,1226,1281,1306]" gridcol="9" gridrow="13" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077FB12FAAA380EFAB1" box="[1279,1311,1281,1306]" gridcol="10" gridrow="13" pageId="4" pageNumber="547">4.6</td>
<td id="76C93B0784370077FAB8FAAA3864FAB1" box="[1365,1397,1281,1306]" gridcol="11" gridrow="13" pageId="4" pageNumber="547">3.5</td>
<td id="76C93B0784370077FA47FAAA38DBFAB1" box="[1450,1482,1281,1306]" gridcol="12" gridrow="13" pageId="4" pageNumber="547">3.1</td>
<td id="76C93B0784370077F9EDFAAA3B31FAB1" box="[1536,1568,1281,1306]" gridcol="13" gridrow="13" pageId="4" pageNumber="547">2.8</td>
</tr>
<tr id="3518527B84370077FF48FA813ADDFAE8" box="[165,1996,1322,1347]" gridrow="14" pageId="4" pageNumber="547" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FA813DD4FAE8" box="[165,197,1322,1347]" gridcol="0" gridrow="14" pageId="4" pageNumber="547">14.</th>
<td id="76C93B0784370077FEECFA813CE8FAE8" box="[257,505,1322,1347]" gridcol="1" gridrow="14" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFA813C7FFAE8" box="[257,366,1322,1347]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B8437FF8CFEECFA813C7FFAE8" box="[257,366,1322,1347]" italics="true" pageId="4" pageNumber="547">B. epimela</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFA813F63FAE8" box="[594,626,1322,1347]" gridcol="2" gridrow="14" pageId="4" pageNumber="547">6.3</td>
<td id="76C93B0784370077FD45FA813FD9FAE8" box="[680,712,1322,1347]" gridcol="3" gridrow="14" pageId="4" pageNumber="547">5.5</td>
<td id="76C93B0784370077FD10FA813E0CFAE8" box="[765,797,1322,1347]" gridcol="4" gridrow="14" pageId="4" pageNumber="547">5.9</td>
<td id="76C93B0784370077FCBEFA813E62FAE8" box="[851,883,1322,1347]" gridcol="5" gridrow="14" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077FC44FA813ED8FAE8" box="[937,969,1322,1347]" gridcol="6" gridrow="14" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FC13FA81390FFAE8" box="[1022,1054,1322,1347]" gridcol="7" gridrow="14" pageId="4" pageNumber="547">5.9</td>
<td id="76C93B0784370077FBB9FA813965FAE8" box="[1108,1140,1322,1347]" gridcol="8" gridrow="14" pageId="4" pageNumber="547">6.8</td>
<td id="76C93B0784370077FB47FA8139DBFAE8" box="[1194,1226,1322,1347]" gridcol="9" gridrow="14" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FB12FA81380EFAE8" box="[1279,1311,1322,1347]" gridcol="10" gridrow="14" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FAB8FA813864FAE8" box="[1365,1397,1322,1347]" gridcol="11" gridrow="14" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077FA47FA8138DBFAE8" box="[1450,1482,1322,1347]" gridcol="12" gridrow="14" pageId="4" pageNumber="547">5.3</td>
<td id="76C93B0784370077F9EDFA813B31FAE8" box="[1536,1568,1322,1347]" gridcol="13" gridrow="14" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077F9B8FA813B64FAE8" box="[1621,1653,1322,1347]" gridcol="14" gridrow="14" pageId="4" pageNumber="547">4.2</td>
</tr>
<tr id="3518527B84370077FF48FAF93ADDFAC0" box="[165,1996,1362,1387]" gridrow="15" pageId="4" pageNumber="547" rowspan-16="1" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FAF93DD4FAC0" box="[165,197,1362,1387]" gridcol="0" gridrow="15" pageId="4" pageNumber="547">15.</th>
<td id="76C93B0784370077FEECFAF93CE8FAC0" box="[257,505,1362,1387]" gridcol="1" gridrow="15" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFAF93C9CFAC0" box="[257,397,1362,1387]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="jugivagans">
<emphasis id="B95C8C2B8437FF8CFEECFAF93C9CFAC0" box="[257,397,1362,1387]" italics="true" pageId="4" pageNumber="547">B. jugivagans</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFAF93F63FAC0" box="[594,626,1362,1387]" gridcol="2" gridrow="15" pageId="4" pageNumber="547">6.6</td>
<td id="76C93B0784370077FD45FAF93FD9FAC0" box="[680,712,1362,1387]" gridcol="3" gridrow="15" pageId="4" pageNumber="547">6.3</td>
<td id="76C93B0784370077FD10FAF93E0CFAC0" box="[765,797,1362,1387]" gridcol="4" gridrow="15" pageId="4" pageNumber="547">8.1</td>
<td id="76C93B0784370077FCBEFAF93E62FAC0" box="[851,883,1362,1387]" gridcol="5" gridrow="15" pageId="4" pageNumber="547">6.6</td>
<td id="76C93B0784370077FC44FAF93ED8FAC0" box="[937,969,1362,1387]" gridcol="6" gridrow="15" pageId="4" pageNumber="547">6.6</td>
<td id="76C93B0784370077FC13FAF9390FFAC0" box="[1022,1054,1362,1387]" gridcol="7" gridrow="15" pageId="4" pageNumber="547">6.9</td>
<td id="76C93B0784370077FBB9FAF93965FAC0" box="[1108,1140,1362,1387]" gridcol="8" gridrow="15" pageId="4" pageNumber="547">8.6</td>
<td id="76C93B0784370077FB47FAF939DBFAC0" box="[1194,1226,1362,1387]" gridcol="9" gridrow="15" pageId="4" pageNumber="547">6.3</td>
<td id="76C93B0784370077FB12FAF9380EFAC0" box="[1279,1311,1362,1387]" gridcol="10" gridrow="15" pageId="4" pageNumber="547">7.2</td>
<td id="76C93B0784370077FAB8FAF93864FAC0" box="[1365,1397,1362,1387]" gridcol="11" gridrow="15" pageId="4" pageNumber="547">5.8</td>
<td id="76C93B0784370077FA47FAF938DBFAC0" box="[1450,1482,1362,1387]" gridcol="12" gridrow="15" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077F9EDFAF93B31FAC0" box="[1536,1568,1362,1387]" gridcol="13" gridrow="15" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077F9B8FAF93B64FAC0" box="[1621,1653,1362,1387]" gridcol="14" gridrow="15" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077F946FAF93BDAFAC0" box="[1707,1739,1362,1387]" gridcol="15" gridrow="15" pageId="4" pageNumber="547">5.3</td>
</tr>
<tr id="3518527B84370077FF48FAD13ADDFA38" box="[165,1996,1402,1427]" gridrow="16" pageId="4" pageNumber="547" rowspan-17="1" rowspan-18="1">
<th id="76C93B0784370077FF48FAD13DD4FA38" box="[165,197,1402,1427]" gridcol="0" gridrow="16" pageId="4" pageNumber="547">16.</th>
<td id="76C93B0784370077FEECFAD13CE8FA38" box="[257,505,1402,1427]" gridcol="1" gridrow="16" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFAD13C6FFA38" box="[257,382,1402,1427]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="B95C8C2B8437FF8CFEECFAD13C6FFA38" box="[257,382,1402,1427]" italics="true" pageId="4" pageNumber="547">B. robinsoni</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFAD13F63FA38" box="[594,626,1402,1427]" gridcol="2" gridrow="16" pageId="4" pageNumber="547">5.6</td>
<td id="76C93B0784370077FD45FAD13FD9FA38" box="[680,712,1402,1427]" gridcol="3" gridrow="16" pageId="4" pageNumber="547">4.9</td>
<td id="76C93B0784370077FD10FAD13E0CFA38" box="[765,797,1402,1427]" gridcol="4" gridrow="16" pageId="4" pageNumber="547">6.5</td>
<td id="76C93B0784370077FCBEFAD13E62FA38" box="[851,883,1402,1427]" gridcol="5" gridrow="16" pageId="4" pageNumber="547">6.0</td>
<td id="76C93B0784370077FC44FAD13ED8FA38" box="[937,969,1402,1427]" gridcol="6" gridrow="16" pageId="4" pageNumber="547">5.2</td>
<td id="76C93B0784370077FC13FAD1390FFA38" box="[1022,1054,1402,1427]" gridcol="7" gridrow="16" pageId="4" pageNumber="547">6.0</td>
<td id="76C93B0784370077FBB9FAD13965FA38" box="[1108,1140,1402,1427]" gridcol="8" gridrow="16" pageId="4" pageNumber="547">6.9</td>
<td id="76C93B0784370077FB47FAD139DBFA38" box="[1194,1226,1402,1427]" gridcol="9" gridrow="16" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FB12FAD1380EFA38" box="[1279,1311,1402,1427]" gridcol="10" gridrow="16" pageId="4" pageNumber="547">5.6</td>
<td id="76C93B0784370077FAB8FAD13864FA38" box="[1365,1397,1402,1427]" gridcol="11" gridrow="16" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077FA47FAD138DBFA38" box="[1450,1482,1402,1427]" gridcol="12" gridrow="16" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077F9EDFAD13B31FA38" box="[1536,1568,1402,1427]" gridcol="13" gridrow="16" pageId="4" pageNumber="547">5.0</td>
<td id="76C93B0784370077F9B8FAD13B64FA38" box="[1621,1653,1402,1427]" gridcol="14" gridrow="16" pageId="4" pageNumber="547">4.8</td>
<td id="76C93B0784370077F946FAD13BDAFA38" box="[1707,1739,1402,1427]" gridcol="15" gridrow="16" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077F8ECFAD13A30FA38" box="[1793,1825,1402,1427]" gridcol="16" gridrow="16" pageId="4" pageNumber="547">4.0</td>
</tr>
<tr id="3518527B84370077FF48FA093ADDFA10" box="[165,1996,1442,1467]" gridrow="17" pageId="4" pageNumber="547" rowspan-18="1">
<th id="76C93B0784370077FF48FA093DD4FA10" box="[165,197,1442,1467]" gridcol="0" gridrow="17" pageId="4" pageNumber="547">17.</th>
<td id="76C93B0784370077FEECFA093CE8FA10" box="[257,505,1442,1467]" gridcol="1" gridrow="17" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFA093CB1FA10" box="[257,416,1442,1467]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="aureogularis">
<emphasis id="B95C8C2B8437FF8CFEECFA093CB1FA10" box="[257,416,1442,1467]" italics="true" pageId="4" pageNumber="547">B. aureogularis</emphasis>
</taxonomicName>
</td>
<td id="76C93B0784370077FDBFFA093F63FA10" box="[594,626,1442,1467]" gridcol="2" gridrow="17" pageId="4" pageNumber="547">4.9</td>
<td id="76C93B0784370077FD45FA093FD9FA10" box="[680,712,1442,1467]" gridcol="3" gridrow="17" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077FD10FA093E0CFA10" box="[765,797,1442,1467]" gridcol="4" gridrow="17" pageId="4" pageNumber="547">5.8</td>
<td id="76C93B0784370077FCBEFA093E62FA10" box="[851,883,1442,1467]" gridcol="5" gridrow="17" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FC44FA093ED8FA10" box="[937,969,1442,1467]" gridcol="6" gridrow="17" pageId="4" pageNumber="547">4.5</td>
<td id="76C93B0784370077FC13FA09390FFA10" box="[1022,1054,1442,1467]" gridcol="7" gridrow="17" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FBB9FA093965FA10" box="[1108,1140,1442,1467]" gridcol="8" gridrow="17" pageId="4" pageNumber="547">6.2</td>
<td id="76C93B0784370077FB47FA0939DBFA10" box="[1194,1226,1442,1467]" gridcol="9" gridrow="17" pageId="4" pageNumber="547">4.7</td>
<td id="76C93B0784370077FB12FA09380EFA10" box="[1279,1311,1442,1467]" gridcol="10" gridrow="17" pageId="4" pageNumber="547">4.9</td>
<td id="76C93B0784370077FAB8FA093864FA10" box="[1365,1397,1442,1467]" gridcol="11" gridrow="17" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077FA47FA0938DBFA10" box="[1450,1482,1442,1467]" gridcol="12" gridrow="17" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077F9EDFA093B31FA10" box="[1536,1568,1442,1467]" gridcol="13" gridrow="17" pageId="4" pageNumber="547">4.3</td>
<td id="76C93B0784370077F9B8FA093B64FA10" box="[1621,1653,1442,1467]" gridcol="14" gridrow="17" pageId="4" pageNumber="547">4.1</td>
<td id="76C93B0784370077F946FA093BDAFA10" box="[1707,1739,1442,1467]" gridcol="15" gridrow="17" pageId="4" pageNumber="547">5.5</td>
<td id="76C93B0784370077F8ECFA093A30FA10" box="[1793,1825,1442,1467]" gridcol="16" gridrow="17" pageId="4" pageNumber="547">3.4</td>
<td id="76C93B0784370077F8BBFA093A67FA10" box="[1878,1910,1442,1467]" gridcol="17" gridrow="17" pageId="4" pageNumber="547">0.6</td>
</tr>
<tr id="3518527B84370077FF48FA613ADDFA4F" box="[165,1996,1482,1508]" gridrow="18" pageId="4" pageNumber="547">
<th id="76C93B0784370077FF48FA613DD4FA4F" box="[165,197,1482,1508]" gridcol="0" gridrow="18" pageId="4" pageNumber="547">18.</th>
<td id="76C93B0784370077FEECFA613CE8FA4F" box="[257,505,1482,1508]" gridcol="1" gridrow="18" pageId="4" pageNumber="547">
<taxonomicName id="4C282BBA8437FF8CFEECFA603CB3FA4F" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[257,418,1483,1508]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B8437FF8CFEECFA603CB3FA4F" box="[257,418,1483,1508]" italics="true" pageId="4" pageNumber="547">B. cathyledecae</emphasis>
</taxonomicName>
<emphasis id="B95C8C2B8437FF8CFE45FA603CE8FA4F" bold="true" box="[424,505,1483,1508]" pageId="4" pageNumber="547">
<taxonomicNameLabel id="A26F31508437FF8CFE45FA603CE8FA4F" box="[424,505,1483,1508]" pageId="4" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
<td id="76C93B0784370077FDBFFA613F63FA4F" box="[594,626,1482,1508]" gridcol="2" gridrow="18" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077FD45FA613FD9FA4F" box="[680,712,1482,1508]" gridcol="3" gridrow="18" pageId="4" pageNumber="547">5.7</td>
<td id="76C93B0784370077FD10FA613E0CFA4F" box="[765,797,1482,1508]" gridcol="4" gridrow="18" pageId="4" pageNumber="547">7.2</td>
<td id="76C93B0784370077FCBEFA613E62FA4F" box="[851,883,1482,1508]" gridcol="5" gridrow="18" pageId="4" pageNumber="547">6.8</td>
<td id="76C93B0784370077FC44FA613ED8FA4F" box="[937,969,1482,1508]" gridcol="6" gridrow="18" pageId="4" pageNumber="547">6.5</td>
<td id="76C93B0784370077FC13FA61390FFA4F" box="[1022,1054,1482,1508]" gridcol="7" gridrow="18" pageId="4" pageNumber="547">7.2</td>
<td id="76C93B0784370077FBB9FA613965FA4F" box="[1108,1140,1482,1508]" gridcol="8" gridrow="18" pageId="4" pageNumber="547">7.4</td>
<td id="76C93B0784370077FB47FA6139DBFA4F" box="[1194,1226,1482,1508]" gridcol="9" gridrow="18" pageId="4" pageNumber="547">5.4</td>
<td id="76C93B0784370077FB12FA61380EFA4F" box="[1279,1311,1482,1508]" gridcol="10" gridrow="18" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077FAB8FA613864FA4F" box="[1365,1397,1482,1508]" gridcol="11" gridrow="18" pageId="4" pageNumber="547">6.4</td>
<td id="76C93B0784370077FA47FA6138DBFA4F" box="[1450,1482,1482,1508]" gridcol="12" gridrow="18" pageId="4" pageNumber="547">6.2</td>
<td id="76C93B0784370077F9EDFA613B31FA4F" box="[1536,1568,1482,1508]" gridcol="13" gridrow="18" pageId="4" pageNumber="547">6.2</td>
<td id="76C93B0784370077F9B8FA613B64FA4F" box="[1621,1653,1482,1508]" gridcol="14" gridrow="18" pageId="4" pageNumber="547">5.9</td>
<td id="76C93B0784370077F946FA613BDAFA4F" box="[1707,1739,1482,1508]" gridcol="15" gridrow="18" pageId="4" pageNumber="547">7.3</td>
<td id="76C93B0784370077F8ECFA613A30FA4F" box="[1793,1825,1482,1508]" gridcol="16" gridrow="18" pageId="4" pageNumber="547">6.7</td>
<td id="76C93B0784370077F8BBFA613A67FA4F" box="[1878,1910,1482,1508]" gridcol="17" gridrow="18" pageId="4" pageNumber="547">6.1</td>
<td id="76C93B0784370077F841FA613ADDFA4F" box="[1964,1996,1482,1508]" gridcol="18" gridrow="18" pageId="4" pageNumber="547">5.9</td>
</tr>
</table>
</paragraph>
<caption id="DF5700B18436FF8DFF7AFC2E3F05FC4D" ID-DOI="http://doi.org/10.5281/zenodo.6504108" ID-Zenodo-Dep="6504108" httpUri="https://zenodo.org/record/6504108/files/figure.png" pageId="5" pageNumber="548" startId="5.[151,250,901,925]" targetBox="[170,1419,185,875]" targetPageId="5">
<paragraph id="8B9750398436FF8DFF7AFC2E3F05FC4D" blockId="5.[151,1436,901,998]" pageId="5" pageNumber="548">
<emphasis id="B95C8C2B8436FF8DFF7AFC2E3C04FC35" bold="true" box="[151,277,901,926]" pageId="5" pageNumber="548">FIGURE 2.</emphasis>
Maximum likelihood trees based on 16S mtDNA sequence of 
<taxonomicName id="4C282BBA8436FF8DFC48FC2D3947FC36" box="[933,1110,902,926]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95C8C2B8436FF8DFC48FC2D3935FC35" box="[933,1060,902,926]" italics="true" pageId="5" pageNumber="548">Bolitoglossa</emphasis>
spp.
</taxonomicName>
of the 
<taxonomicName id="4C282BBA8436FF8DFB4EFC2D39EFFC35" box="[1187,1278,902,926]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8436FF8DFB4EFC2D39EFFC35" box="[1187,1278,902,926]" italics="true" pageId="5" pageNumber="548">Eladinea</emphasis>
</taxonomicName>
subgenus. SHaLRT (Shimodaira and Hasegawa approximate likelihood ratio test) support (%) / ultrafast bootstrap support (%) are shown on the internodes. Outgroups not shown.
</paragraph>
</caption>
<paragraph id="8B9750398436FF8DFF2AFBBF3E82FA10" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="548">
<taxonomicName id="4C282BBA8436FF8DFF2AFBBF3F32FB85" authority="Bolanos" authorityName="Bolanos" box="[199,547,1044,1070]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="pygmaea">
<emphasis id="B95C8C2B8436FF8DFF2AFBBF3CD1FB85" box="[199,448,1044,1070]" italics="true" pageId="5" pageNumber="548">Bolitoglossa pygmaea</emphasis>
Bolaños
</taxonomicName>
and 
<collectingCountry id="F33F10A98436FF8DFDB7FBBF3F8DFB85" box="[602,668,1044,1070]" name="Wake Island" pageId="5" pageNumber="548">Wake</collectingCountry>
, 
<taxonomicName id="4C282BBA8436FF8DFD4AFBBE3EE8FB85" authority="Bolanos, Robinson" authorityName="Bolanos, Robinson" box="[679,1017,1044,1070]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B95C8C2B8436FF8DFD4AFBBE3E0CFB85" box="[679,797,1044,1070]" italics="true" pageId="5" pageNumber="548">B. gracilis</emphasis>
Bolaños, Robinson
</taxonomicName>
and 
<collectingCountry id="F33F10A98436FF8DFBDDFBBF397EFB85" box="[1072,1135,1044,1070]" name="Wake Island" pageId="5" pageNumber="548">Wake</collectingCountry>
and 
<taxonomicName id="4C282BBA8436FF8DFB4BFBBE388DFB85" authority="Robinson" authorityName="Robinson" authorityYear="1976" box="[1190,1436,1044,1070]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="diminuta">
<emphasis id="B95C8C2B8436FF8DFB4BFBBE383BFB85" box="[1190,1322,1044,1070]" italics="true" pageId="5" pageNumber="548">B. diminuta</emphasis>
Robinson
</taxonomicName>
are small species with SVL ≤ 
<quantity id="4CD0FDDC8436FF8DFE1BFB933F59FBF9" box="[502,584,1080,1106]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="548" unit="mm" value="43.0">43 mm</quantity>
(46.0 mm), also, their hands and feet have less webbing, digital webbing well developed). 
<taxonomicName id="4C282BBA8436FF8DFEB4FBF73F47FBDD" box="[345,598,1116,1142]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="B95C8C2B8436FF8DFEB4FBF73F47FBDD" box="[345,598,1116,1142]" italics="true" pageId="5" pageNumber="548">Bolitoglossa robinsoni</emphasis>
</taxonomicName>
is a relatively large SVL in adult males with 
<quantity id="4CD0FDDC8436FF8DFBB9FBF739E6FBDD" box="[1108,1271,1116,1143]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" metricValueMax="6.35" metricValueMin="4.59" pageId="5" pageNumber="548" unit="mm" value="54.7" valueMax="63.5" valueMin="45.9">45.9‒63.5 mm</quantity>
(
<quantity id="4CD0FDDC8436FF8DFAEBFBF7384AFBDD" box="[1286,1371,1116,1142]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="548" unit="mm" value="46.0">46 mm</quantity>
), tails are moderate, shorter in proportion to body size TL/SVL = 1.06‒1.34 (1.5); hands and feet with little webbing; fewer than two distal-most phalanges of the longest digits are free of webbing (digital webbing well developed; only portions of distal-most phalanges of longest digits free). 
<taxonomicName id="4C282BBA8436FF8DFCB9FB633926FB49" box="[852,1079,1224,1250]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="gomezi">
<emphasis id="B95C8C2B8436FF8DFCB9FB633926FB49" box="[852,1079,1224,1250]" italics="true" pageId="5" pageNumber="548">Bolitoglossa gomezi</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98436FF8DFBADFB633993FB49" box="[1088,1154,1224,1250]" name="Wake Island" pageId="5" pageNumber="548">Wake</collectingCountry>
, Savage and Hanken has a shorter tail in proportion to body size TL/SVL: 1.15–1.19 (TL/SVL = 1.5) and a lower tooth count in PMT: 2 or 3 (7), MT: 
<quantity id="4CD0FDDC8436FF8DFEFBFABB3C6AFA81" box="[278,379,1296,1323]" metricMagnitude="0" metricUnit="m" metricValue="1.0287000000000002" metricValueMax="1.3208000000000002" metricValueMin="0.7365999999999999" pageId="5" pageNumber="548" unit="in" value="40.5" valueMax="52.0" valueMin="29.0">29–52 in</quantity>
males (78), and VT: 17–30 (33). 
<taxonomicName id="4C282BBA8436FF8DFD16FABB3ECFFA81" box="[763,990,1296,1322]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="bramei">
<emphasis id="B95C8C2B8436FF8DFD16FABB3ECFFA81" box="[763,990,1296,1322]" italics="true" pageId="5" pageNumber="548">Bolitoglossa bramei</emphasis>
</taxonomicName>
<collectingCountry id="F33F10A98436FF8DFC0AFABB3938FA81" box="[999,1065,1296,1322]" name="Wake Island" pageId="5" pageNumber="548">Wake</collectingCountry>
, Savage and Hanken as well has fewer premaxillary, maxillary and vomerine teeth PMT: 3‒4 (7), MT: 49–66 (78), VT: 24–28 (33), limbs long; limb interval 0–0.5 (3). 
<taxonomicName id="4C282BBA8436FF8DFE4EFAF3388DFAD9" authority="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" authorityName="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" box="[419,1436,1368,1394]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="splendida">
<emphasis id="B95C8C2B8436FF8DFE4EFAF33FB5FAD9" box="[419,676,1368,1394]" italics="true" pageId="5" pageNumber="548">Bolitoglossa splendida</emphasis>
Boza-Oviedo, Rovito, Chaves, García-Rodríguez, Artavia, Bolaños
</taxonomicName>
and Wake presents a coloration with bright-red dorsal band and bright-yellow spots laterally and ventro-laterally (pinkish flesh with flame scarlet speckles and ventral salmon color).
</paragraph>
<paragraph id="8B9750398436FF8DFF2AFA6F381EF971" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="548">
<taxonomicName id="4C282BBA8436FF8DFF2AFA6F39B6FA75" authority="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" authorityName="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" box="[199,1191,1476,1502]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B95C8C2B8436FF8DFF2AFA6F3CB4FA75" box="[199,421,1476,1502]" italics="true" pageId="5" pageNumber="548">Bolitoglossa kamuk</emphasis>
Boza-Oviedo, Rovito, Chaves, García-Rodríguez, Artavia, Bolaños
</taxonomicName>
and Wake has a dark ground coloration with golden flecking on the posterior part of the body and the tail (pinkish -flesh with flame scarlet speckles, ventral salmon color), as well as fewer maxillary and vomerine teeth MT: 34 (78), VT: 16 (33), and a shorter tail in proportion to body size TL/SVL ≈ 1 (TL/SVL = 1.5). 
<taxonomicName id="4C282BBA8436FF8DFC3BF99B3FF7F9C5" authority="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" authorityName="Boza-Oviedo, Rovito, Chaves, Garcia-Rodriguez, Artavia, Bolanos" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="aureogularis">
<emphasis id="B95C8C2B8436FF8DFC3BF99B39E8F9E1" box="[982,1273,1584,1610]" italics="true" pageId="5" pageNumber="548">Bolitoglossa aureogularis</emphasis>
Boza-Oviedo, Rovito, Chaves, García-Rodríguez, Artavia, Bolaños
</taxonomicName>
and Wake with dorsal coloration golden-tan with some bright highlights on dorsolateral regions and with some narrow streaks of dark brown, gular area bright yellow, which becomes golden on chest before fading into darker color in the midtrunk region (pinkish flesh with flame scarlet speckles and ventral salmon color), Head is narrow in proportion to body size HW/SVL = 0.13 (0.21).
</paragraph>
<paragraph id="8B9750398436FF8DFF2AF94F39DEF850" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="548">
<taxonomicName id="4C282BBA8436FF8DFF2AF94F3E1EF955" authority=", Hertz, Lotzkat and Kohler" authorityName="Hertz, Lotzkat and Kohler" box="[199,783,1764,1790]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="jugivagans">
<emphasis id="B95C8C2B8436FF8DFF2AF94F3CC7F955" box="[199,470,1764,1790]" italics="true" pageId="5" pageNumber="548">Bolitoglossa jugivagans</emphasis>
, Hertz, Lotzkat and Köhler
</taxonomicName>
is a small thin 
<taxonomicName id="4C282BBA8436FF8DFC53F94F395BF955" baseAuthorityName="AmphibiaWeb" baseAuthorityYear="2022" box="[958,1098,1764,1790]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8436FF8DFC53F94F395BF955" box="[958,1098,1764,1790]" italics="true" pageId="5" pageNumber="548">Bolitoglossa</emphasis>
</taxonomicName>
with an SVL of 
<quantity id="4CD0FDDC8436FF8DFAE7F94F387EF955" box="[1290,1391,1764,1790]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="5" pageNumber="548" unit="mm" value="31.2">31.2 mm</quantity>
(
<quantity id="4CD0FDDC8436FF8DFA92F94F3DD5F889" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="548" unit="mm" value="46.0">46 mm</quantity>
) with a narrow head in proportion to body size HW/SVL = 0.14 (0.21), relatively shorter front limbs HLL about 20 % of SVL (HLL = 25.7 % of SVL). The last 
<taxonomicName id="4C282BBA8436FF8DFCF9F8873EB1F8ED" baseAuthorityName="AmphibiaWeb" baseAuthorityYear="2022" box="[788,928,1836,1862]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B8436FF8DFCF9F8873EB1F8ED" box="[788,928,1836,1862]" italics="true" pageId="5" pageNumber="548">Bolitoglossa</emphasis>
</taxonomicName>
described in the Talamanca mountain range 
<taxonomicName id="4C282BBA8436FF8DFF7AF8FB3F59F8C1" authority="Kubicki and Arias" authorityName="Kubicki and Arias" box="[151,584,1872,1898]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="aurae">
<emphasis id="B95C8C2B8436FF8DFF7AF8FB3C7FF8C1" box="[151,366,1872,1898]" italics="true" pageId="5" pageNumber="548">Bolitoglossa aurae</emphasis>
Kubicki and Arias
</taxonomicName>
has a significantly different coloration, dorsum uniform light yellow with a thin dark brown middorsal stripe on the trunk, flanks light yellow to light brown, ventrum uniform translucent yellow and lacking any evident or contrasting light or dark dermal pigmentation or midventral stripe (pinkish flesh with flame scarlet speckles and ventral salmon color), relatively longer trunk in female AX = 58 % of SVL (51.3% in male 
<typeStatus id="5493EE9B8436FF8DFF1CF84B3C46F850" box="[241,343,2016,2043]" pageId="5" pageNumber="548" type="holotype">holotype</typeStatus>
), with 4.5 costal folds between adpressed limbs in female (
<quantity id="4CD0FDDC8436FF8DFC00F84B3909F851" box="[1005,1048,2016,2042]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="548" unit="in" value="3.0">3 in</quantity>
male 
<typeStatus id="5493EE9B8436FF8DFBB1F84B39D2F850" box="[1116,1219,2016,2043]" pageId="5" pageNumber="548" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C33203B28435FF80FF2AFF3D3F8BFA76" lastPageId="8" lastPageNumber="551" pageId="6" pageNumber="549" type="description">
<paragraph id="8B9750398435FF8EFF2AFF3D3F1AFE5E" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="549">
<emphasis id="B95C8C2B8435FF8EFF2AFF3D3F04FF1A" bold="true" box="[199,533,150,177]" pageId="6" pageNumber="549">
Description of the 
<typeStatus id="5493EE9B8435FF8EFE4AFF3C3F01FF1A" box="[423,528,151,177]" pageId="6" pageNumber="549" type="holotype">holotype</typeStatus>
.
</emphasis>
A moderately small species (SVL = 46.0 mm), head relatively broad compared to other members of genus; head long (HL about 29 % of SVL) and broad (HW = 21 % of SVL); tail long and prehensile, TL/SVL is 1.5; hands and feet broad (HAW = 10 %, HFW = 14% of SVL), and limbs moderate (HLL about 19 % of SVL, limb interval 3 costal folds between adpressed limbs); digital webbing well developed, only portions of distal-most phalanges of longest digits free (
<figureCitation id="13134CBC8435FF8EFCEEFE8C3E57FEEA" box="[771,838,295,321]" captionStart="FIGURE 3" captionStartId="6.[151,250,1936,1960]" captionTargetBox="[172,1414,669,1913]" captionTargetId="figure-274@6.[172,1414,669,1913]" captionTargetPageId="6" captionText="FIGURE 3. Details of hand and foot of preserved holotype of Bolitoglossa cathyledecae sp. nov. (MHCH 3240); A) Right hand, left: dorsal view, right: ventral view; B) Right foot, left: dorsal view, right: ventral view. Scale bars: 5 mm. Photographs taken by MP." figureDoi="http://doi.org/10.5281/zenodo.6504112" httpUri="https://zenodo.org/record/6504112/files/figure.png" pageId="6" pageNumber="549">Fig. 3</figureCitation>
). Fingers in order of decreasing length: 3, 4, 2, 1; toes in order of decreasing length: 3, 4, 5, 2, 1; subterminal pads present on longest digits; eyes relatively small; do not protrude beyond lateral margins of head and are not visible in ventral view; labial protuberances weakly developed; longer snout, ES = 43.75 % of HW. Teeth moderate in size and numerous (PMT 7, MT 78, VT 33). Digits of hands and feet easily distinguishable but extensively webbed; digital tips broadly rounded, hands and feet flattened; digits flattened, not cylindrical (
<figureCitation id="13134CBC8435FF8EFE54FE703CECFE5E" box="[441,509,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1936,1960]" captionTargetBox="[172,1414,669,1913]" captionTargetId="figure-274@6.[172,1414,669,1913]" captionTargetPageId="6" captionText="FIGURE 3. Details of hand and foot of preserved holotype of Bolitoglossa cathyledecae sp. nov. (MHCH 3240); A) Right hand, left: dorsal view, right: ventral view; B) Right foot, left: dorsal view, right: ventral view. Scale bars: 5 mm. Photographs taken by MP." figureDoi="http://doi.org/10.5281/zenodo.6504112" httpUri="https://zenodo.org/record/6504112/files/figure.png" pageId="6" pageNumber="549">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B9750398435FF8EFF2AFE543CA3FD2E" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="549">
<emphasis id="B95C8C2B8435FF8EFF2AFE5439BFFDB3" bold="true" box="[199,1198,510,537]" pageId="6" pageNumber="549">
Measurements (in mm), limb interval and tooth count of the male 
<typeStatus id="5493EE9B8435FF8EFC3CFE543929FDB2" box="[977,1080,511,537]" pageId="6" pageNumber="549" type="holotype">holotype</typeStatus>
(
<tableCitation id="C6AA65828435FF8EFBABFE55398EFDB3" box="[1094,1183,510,536]" captionStart="TABLE 2" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 2. Morphometric characters of Bolitoglossa cathyledecae sp. nov. holotype and paratypes. see methods for abbreviation explanations of abbreviations." httpUri="http://table.plazi.org/id/DF5700B18434FF8FFF7AFF3C3F74FF78" pageId="6" pageNumber="549" tableUuid="DF5700B18434FF8FFF7AFF3C3F74FF78">Table 2</tableCitation>
).
</emphasis>
SVL: 46.0, TL: 68.8, HW: 9.6, SG: 13.2, HD: 5, EW: 1.9, EL: 3, ES: 4.2, ED: 2.9, IC: 4.9, IO: 3.5, SF: 17.6, IN: 2.7, SP: 0.8, SW: 8.2, SAV: 18.5, AX: 26.9, LI: 3.5, FLL: 10.8, HLL: 8.6, HAW: 4.8, FW: 6.4, T3: 1.4, T5: 0.9, Numbers of teeth: PMT: 7, MT: 40/38, VT: 17/16.
</paragraph>
<caption id="DF5700B18435FF8EFF7AF83B3C0CF844" ID-DOI="http://doi.org/10.5281/zenodo.6504112" ID-Zenodo-Dep="6504112" httpUri="https://zenodo.org/record/6504112/files/figure.png" pageId="6" pageNumber="549" startId="6.[151,250,1936,1960]" targetBox="[172,1414,669,1913]" targetPageId="6">
<paragraph id="8B9750398435FF8EFF7AF83B3C0CF844" blockId="6.[151,1436,1936,2032]" pageId="6" pageNumber="549">
<emphasis id="B95C8C2B8435FF8EFF7AF83B3C04F803" bold="true" box="[151,277,1936,1960]" pageId="6" pageNumber="549">FIGURE 3.</emphasis>
Details of hand and foot of preserved holotype of 
<taxonomicName id="4C282BBA8435FF8EFCC6F83B3927F803" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[811,1078,1936,1960]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="549" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B8435FF8EFCC6F83B3927F803" box="[811,1078,1936,1960]" italics="true" pageId="6" pageNumber="549">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
<emphasis id="B95C8C2B8435FF8EFBD2F83B3983F803" bold="true" box="[1087,1170,1936,1960]" pageId="6" pageNumber="549">
<taxonomicNameLabel id="A26F31508435FF8EFBD2F83B3983F803" box="[1087,1170,1936,1960]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MHCH 3240); A) Right hand, left: dorsal view, right: ventral view; B) Right foot, left: dorsal view, right: ventral view. Scale bars: 5 mm. Photographs taken by MP.
</paragraph>
</caption>
<caption id="DF5700B18434FF8FFF7AFF3C3F74FF78" ID-Table-UUID="DF5700B18434FF8FFF7AFF3C3F74FF78" httpUri="http://table.plazi.org/id/DF5700B18434FF8FFF7AFF3C3F74FF78" pageId="7" pageNumber="550" startId="7.[151,239,151,175]" targetBox="[166,1380,228,690]" targetIsTable="true" targetPageId="7">
<paragraph id="8B9750398434FF8FFF7AFF3C3F74FF78" blockId="7.[151,1436,151,211]" pageId="7" pageNumber="550">
<emphasis id="B95C8C2B8434FF8FFF7AFF3C3C1FFF04" bold="true" box="[151,270,151,175]" pageId="7" pageNumber="550">TABLE 2.</emphasis>
Morphometric characters of 
<taxonomicName id="4C282BBA8434FF8FFDB9FF3C3E7FFF04" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[596,878,151,175]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="550" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B8434FF8FFDB9FF3C3E7FFF04" box="[596,878,151,175]" italics="true" pageId="7" pageNumber="550">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
<emphasis id="B95C8C2B8434FF8FFC94FF333EC3FF1B" bold="true" box="[889,978,152,176]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="A26F31508434FF8FFC94FF333EC3FF1B" box="[889,978,152,176]" pageId="7" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype and paratypes. see methods for abbreviation explanations of abbreviations.
</paragraph>
</caption>
<paragraph id="8B9750398434FF8FFF4BFF4F39ECFD19" pageId="7" pageNumber="550">
<table id="F928A29984340077FF4BFF4F3875FD19" box="[166,1380,228,690]" gridcols="4" gridrows="11" pageId="7" pageNumber="550">
<tr id="3518527B84340077FF4BFF4F3875FF57" box="[166,1380,228,252]" gridrow="0" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFF4F3CB0FF57" box="[166,417,228,252]" gridcol="0" gridrow="0" pageId="7" pageNumber="550">Measurements (mm)</th>
<th id="76C93B0784340077FD5EFF4F3E58FF57" box="[691,841,228,252]" gridcol="1" gridrow="0" pageId="7" pageNumber="550">Holotype</th>
<th id="76C93B0784340077FC42FF4F3940FF57" box="[943,1105,228,252]" gridcol="2" gridrow="0" pageId="7" pageNumber="550">Paratype</th>
<th id="76C93B0784340077FB41FF4F3875FF57" box="[1196,1380,228,252]" gridcol="3" gridrow="0" pageId="7" pageNumber="550">Paratype</th>
</tr>
<tr id="3518527B84340077FF4BFEA73875FEE3" box="[166,1380,268,328]" gridrow="1" pageId="7" pageNumber="550" rowspan-0="1">
<td id="76C93B0784340077FD5EFEA73E58FEE3" box="[691,841,268,328]" gridcol="1" gridrow="1" pageId="7" pageNumber="550">MP-12 (MHCH 3240)</td>
<td id="76C93B0784340077FC42FEA73940FEE3" box="[943,1105,268,328]" gridcol="2" gridrow="1" pageId="7" pageNumber="550">MP-175 (MHCH 3241)</td>
<td id="76C93B0784340077FB41FEA73875FEE3" box="[1196,1380,268,328]" gridcol="3" gridrow="1" pageId="7" pageNumber="550">MP-185 (MHCH 3242)</td>
</tr>
<tr id="3518527B84340077FF4BFEF33875FEDB" box="[166,1380,344,368]" gridrow="2" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFEF33CB0FEDB" box="[166,417,344,368]" gridcol="0" gridrow="2" pageId="7" pageNumber="550">Sex</th>
<td id="76C93B0784340077FD5EFEF33E58FEDB" box="[691,841,344,368]" gridcol="1" gridrow="2" pageId="7" pageNumber="550">Male</td>
<td id="76C93B0784340077FC42FEF33940FEDB" box="[943,1105,344,368]" gridcol="2" gridrow="2" pageId="7" pageNumber="550">Male (subadult)</td>
<td id="76C93B0784340077FB41FEF33875FEDB" box="[1196,1380,344,368]" gridcol="3" gridrow="2" pageId="7" pageNumber="550">Female (subadult)</td>
</tr>
<tr id="3518527B84340077FF4BFE2B3875FE32" box="[166,1380,384,409]" gridrow="3" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFE2B3CB0FE32" box="[166,417,384,409]" gridcol="0" gridrow="3" pageId="7" pageNumber="550">Length total (LT)</th>
<td id="76C93B0784340077FD5EFE2B3E58FE32" box="[691,841,384,409]" gridcol="1" gridrow="3" pageId="7" pageNumber="550">115.9</td>
<td id="76C93B0784340077FC42FE2B3940FE32" box="[943,1105,384,409]" gridcol="2" gridrow="3" pageId="7" pageNumber="550">60.7</td>
<td id="76C93B0784340077FB41FE2B3875FE32" box="[1196,1380,384,409]" gridcol="3" gridrow="3" pageId="7" pageNumber="550">57.7</td>
</tr>
<tr id="3518527B84340077FF4BFE023875FE6A" box="[166,1380,425,449]" gridrow="4" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFE023CB0FE6A" box="[166,417,425,449]" gridcol="0" gridrow="4" pageId="7" pageNumber="550">Snout-vent length (SVL)</th>
<td id="76C93B0784340077FD5EFE023E58FE6A" box="[691,841,425,449]" gridcol="1" gridrow="4" pageId="7" pageNumber="550">46</td>
<td id="76C93B0784340077FC42FE023940FE6A" box="[943,1105,425,449]" gridcol="2" gridrow="4" pageId="7" pageNumber="550">30.9</td>
<td id="76C93B0784340077FB41FE023875FE6A" box="[1196,1380,425,449]" gridcol="3" gridrow="4" pageId="7" pageNumber="550">25</td>
</tr>
<tr id="3518527B84340077FF4BFE7A3875FE42" box="[166,1380,465,489]" gridrow="5" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFE7A3CB0FE42" box="[166,417,465,489]" gridcol="0" gridrow="5" pageId="7" pageNumber="550">Tail length (TL)</th>
<td id="76C93B0784340077FD5EFE7A3E58FE42" box="[691,841,465,489]" gridcol="1" gridrow="5" pageId="7" pageNumber="550">68.8</td>
<td id="76C93B0784340077FC42FE7A3940FE42" box="[943,1105,465,489]" gridcol="2" gridrow="5" pageId="7" pageNumber="550">27.8</td>
<td id="76C93B0784340077FB41FE7A3875FE42" box="[1196,1380,465,489]" gridcol="3" gridrow="5" pageId="7" pageNumber="550">32.6</td>
</tr>
<tr id="3518527B84340077FF4BFE523875FDBA" box="[166,1380,505,529]" gridrow="6" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFE523CB0FDBA" box="[166,417,505,529]" gridcol="0" gridrow="6" pageId="7" pageNumber="550">Axilla-groin (AG)</th>
<td id="76C93B0784340077FD5EFE523E58FDBA" box="[691,841,505,529]" gridcol="1" gridrow="6" pageId="7" pageNumber="550">26.9</td>
<td id="76C93B0784340077FC42FE523940FDBA" box="[943,1105,505,529]" gridcol="2" gridrow="6" pageId="7" pageNumber="550">18.9</td>
<td id="76C93B0784340077FB41FE523875FDBA" box="[1196,1380,505,529]" gridcol="3" gridrow="6" pageId="7" pageNumber="550">14.7</td>
</tr>
<tr id="3518527B84340077FF4BFD8A3875FD91" box="[166,1380,545,570]" gridrow="7" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFD8A3CB0FD91" box="[166,417,545,570]" gridcol="0" gridrow="7" pageId="7" pageNumber="550">Head width (HW)</th>
<td id="76C93B0784340077FD5EFD8A3E58FD91" box="[691,841,545,570]" gridcol="1" gridrow="7" pageId="7" pageNumber="550">9.6</td>
<td id="76C93B0784340077FC42FD8A3940FD91" box="[943,1105,545,570]" gridcol="2" gridrow="7" pageId="7" pageNumber="550">6.7</td>
<td id="76C93B0784340077FB41FD8A3875FD91" box="[1196,1380,545,570]" gridcol="3" gridrow="7" pageId="7" pageNumber="550">5.4</td>
</tr>
<tr id="3518527B84340077FF4BFDE13875FDC9" box="[166,1380,586,610]" gridrow="8" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFDE13CB0FDC9" box="[166,417,586,610]" gridcol="0" gridrow="8" pageId="7" pageNumber="550">Hind limb length (HLL)</th>
<td id="76C93B0784340077FD5EFDE13E58FDC9" box="[691,841,586,610]" gridcol="1" gridrow="8" pageId="7" pageNumber="550">11.8</td>
<td id="76C93B0784340077FC42FDE13940FDC9" box="[943,1105,586,610]" gridcol="2" gridrow="8" pageId="7" pageNumber="550">6.1</td>
<td id="76C93B0784340077FB41FDE13875FDC9" box="[1196,1380,586,610]" gridcol="3" gridrow="8" pageId="7" pageNumber="550">4.6</td>
</tr>
<tr id="3518527B84340077FF4BFDD93875FD21" box="[166,1380,626,650]" gridrow="9" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFDD93CB0FD21" box="[166,417,626,650]" gridcol="0" gridrow="9" pageId="7" pageNumber="550">Maxillary teeth (MT)</th>
<td id="76C93B0784340077FD5EFDD93E58FD21" box="[691,841,626,650]" gridcol="1" gridrow="9" pageId="7" pageNumber="550">78 (40/38)</td>
<td id="76C93B0784340077FC42FDD93940FD21" box="[943,1105,626,650]" gridcol="2" gridrow="9" pageId="7" pageNumber="550">36 (17/19)</td>
<td id="76C93B0784340077FB41FDD93875FD21" box="[1196,1380,626,650]" gridcol="3" gridrow="9" pageId="7" pageNumber="550">25 (13/12)</td>
</tr>
<tr id="3518527B84340077FF4BFD313875FD19" box="[166,1380,666,690]" gridrow="10" pageId="7" pageNumber="550">
<th id="76C93B0784340077FF4BFD313CB0FD19" box="[166,417,666,690]" gridcol="0" gridrow="10" pageId="7" pageNumber="550">Vomerine teeth (VT)</th>
<td id="76C93B0784340077FD5EFD313E58FD19" box="[691,841,666,690]" gridcol="1" gridrow="10" pageId="7" pageNumber="550">33 (17/16)</td>
<td id="76C93B0784340077FC42FD313940FD19" box="[943,1105,666,690]" gridcol="2" gridrow="10" pageId="7" pageNumber="550">19 (10/11))</td>
<td id="76C93B0784340077FB41FD313875FD19" box="[1196,1380,666,690]" gridcol="3" gridrow="10" pageId="7" pageNumber="550">17 (8/9)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9750398434FF8FFF2AFD4A39D5FC27" blockId="7.[151,1437,737,1017]" pageId="7" pageNumber="550">
<emphasis id="B95C8C2B8434FF8FFF2AFD4A3F43FD50" bold="true" box="[199,594,737,764]" pageId="7" pageNumber="550">
Coloration of the 
<typeStatus id="5493EE9B8434FF8FFE72FD493F12FD57" box="[415,515,738,764]" pageId="7" pageNumber="550" type="holotype">holotype</typeStatus>
in life
</emphasis>
(
<figureCitation id="13134CBC8434FF8FFD88FD493FBAFD57" box="[613,683,738,764]" captionStart="FIGURE 4" captionStartId="7.[151,250,1962,1986]" captionTargetBox="[156,1431,1040,1939]" captionTargetId="figure-277@7.[156,1431,1040,1939]" captionTargetPageId="7" captionText="FIGURE 4. Holotype of Bolitoglossa cathyledecae sp. nov. (MHCH 3240) in life; A–B) Details of head; C–D) Body coloration in life. Photograph by MP." figureDoi="http://doi.org/10.5281/zenodo.6504114" httpUri="https://zenodo.org/record/6504114/files/figure.png" pageId="7" pageNumber="550">Fig. 4</figureCitation>
). Dorsal ground of the head, back color Pinkish Flesh (253), with Flame Scarlet (73) speckles, upper part of the extremities similar in color to dorsal surface. Lateral and ventral ground coloration Salmon Color (251), with Pinkish Flesh (253) speckles, in the gular and ventral region, lower surface of the hands, and feet Light Pinkish Rose (217). Most of the tail similar in color to the background dorsal color with the tip of the tail Spectrum Yellow (78). Iris Pale Sulphur Yellow (92) with reticulum Flame Scarlet (73).
</paragraph>
<paragraph id="8B9750398434FF8FFF2AFC3E39F6FC52" blockId="7.[151,1437,737,1017]" pageId="7" pageNumber="550">
<emphasis id="B95C8C2B8434FF8FFF2AFC3E3CDFFC1B" bold="true" box="[199,462,917,944]" pageId="7" pageNumber="550">Coloration in ethanol.</emphasis>
After three years in ethanol (70%), the dorsal coloration of the 
<typeStatus id="5493EE9B8434FF8FFB74FC3D39EAFC1B" box="[1177,1275,918,944]" pageId="7" pageNumber="550" type="holotype">holotype</typeStatus>
has darkened, the head and the anterior part of the trunk are Burnt Umber (48) while the rest of the trunk and tail are Amber (51). The ventral coloration of the specimen is mainly Clay Color (18) speckled with Drab (19) (
<figureCitation id="13134CBC8434FF8FFB78FC7539C8FC52" box="[1173,1241,990,1017]" captionStart="FIGURE 5" captionStartId="8.[151,250,1296,1320]" captionTargetBox="[151,1436,181,1272]" captionTargetId="figure-21@8.[151,1436,181,1272]" captionTargetPageId="8" captionText="FIGURE 5. Holotype after three years in 70 % ethanol; A) Dorsal view; B) ventral view. Scale bars= 2 cm. Photographs taken by MP." figureDoi="http://doi.org/10.5281/zenodo.6504116" httpUri="https://zenodo.org/record/6504116/files/figure.png" pageId="7" pageNumber="550">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF5700B18434FF8FFF7AF8013CB5F84D" ID-DOI="http://doi.org/10.5281/zenodo.6504114" ID-Zenodo-Dep="6504114" httpUri="https://zenodo.org/record/6504114/files/figure.png" pageId="7" pageNumber="550" startId="7.[151,250,1962,1986]" targetBox="[156,1431,1040,1939]" targetPageId="7">
<paragraph id="8B9750398434FF8FFF7AF8013CB5F84D" blockId="7.[151,1436,1962,2023]" pageId="7" pageNumber="550">
<emphasis id="B95C8C2B8434FF8FFF7AF8013C03F868" bold="true" box="[151,274,1962,1987]" pageId="7" pageNumber="550">FIGURE 4.</emphasis>
Holotype of 
<taxonomicName id="4C282BBA8434FF8FFE7BF8003F8FF868" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[406,670,1963,1987]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="550" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B8434FF8FFE7BF8003F8FF868" box="[406,670,1963,1987]" italics="true" pageId="7" pageNumber="550">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
<emphasis id="B95C8C2B8434FF8FFD49F8003FE2F868" bold="true" box="[676,755,1963,1987]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="A26F31508434FF8FFD49F8003FE2F868" box="[676,755,1963,1987]" pageId="7" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MHCH 3240) in life; A–B) Details of head; C–D) Body coloration in life. Photograph by MP.
</paragraph>
</caption>
<caption id="DF5700B1843BFF80FF7AFABB3DF1FAE7" ID-DOI="http://doi.org/10.5281/zenodo.6504116" ID-Zenodo-Dep="6504116" httpUri="https://zenodo.org/record/6504116/files/figure.png" pageId="8" pageNumber="551" startId="8.[151,250,1296,1320]" targetBox="[151,1436,181,1272]" targetPageId="8">
<paragraph id="8B975039843BFF80FF7AFABB3DF1FAE7" blockId="8.[151,1436,1296,1356]" pageId="8" pageNumber="551">
<emphasis id="B95C8C2B843BFF80FF7AFABB3C02FA83" bold="true" box="[151,275,1296,1320]" pageId="8" pageNumber="551">FIGURE 5.</emphasis>
Holotype after three years in 70 % ethanol; 
<emphasis id="B95C8C2B843BFF80FD39FABB3FF7FA83" bold="true" box="[724,742,1296,1320]" pageId="8" pageNumber="551">A</emphasis>
) Dorsal view; 
<emphasis id="B95C8C2B843BFF80FC90FABB3E9FFA83" bold="true" box="[893,910,1296,1320]" pageId="8" pageNumber="551">B</emphasis>
) ventral view. Scale bars= 2 cm. Photographs taken by MP.
</paragraph>
</caption>
<paragraph id="8B975039843BFF80FF2AFAD13F8BFA76" blockId="8.[151,1437,1402,2041]" pageId="8" pageNumber="551">
<emphasis id="B95C8C2B843BFF80FF2AFAD13C29FA3F" bold="true" box="[199,312,1402,1428]" pageId="8" pageNumber="551">Variation</emphasis>
(
<figureCitation id="13134CBC843BFF80FEA4FAD13C9EFA3E" box="[329,399,1402,1429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1234,1258]" captionTargetBox="[198,1430,309,1211]" captionTargetId="figure-73@9.[157,1430,309,1211]" captionTargetPageId="9" captionText="FIGURE 6. Color variation in the paratypes.A) Dorsum in daytime coloration (MHCH 3242); B-C) Details of head and dorsum in daytime coloration (MHCH 3241); D) Paratype at the moment of encounter at night (MHCH 3242). Photographs taken by MP." figureDoi="http://doi.org/10.5281/zenodo.6504118" httpUri="https://zenodo.org/record/6504118/files/figure.png" pageId="8" pageNumber="551">Fig. 6</figureCitation>
). There are 
<specimenCount id="9D2E9BB0843BFF80FDF1FAD03FABFA3E" box="[540,698,1403,1429]" pageId="8" pageNumber="551" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(a male and a female) both juveniles, these specimens present a coloration like the 
<typeStatus id="5493EE9B843BFF80FE87FA353CDEFA13" box="[362,463,1438,1464]" pageId="8" pageNumber="551" type="holotype">holotype</typeStatus>
, but with the dorsal coloration Chrome Orange (74) and the smallest 
<typeStatus id="5493EE9B843BFF80FB34FA343828FA12" box="[1241,1337,1439,1465]" pageId="8" pageNumber="551" type="paratype">paratype</typeStatus>
presents the last third of the tail Spectrum Yellow (78).
</paragraph>
</subSubSection>
<subSubSection id="C33203B2843BFF80FF2AFA4C3C0BF9E3" pageId="8" pageNumber="551" type="etymology">
<paragraph id="8B975039843BFF80FF2AFA4C3C0BF9E3" blockId="8.[151,1437,1402,2041]" pageId="8" pageNumber="551">
<emphasis id="B95C8C2B843BFF80FF2AFA4C3C5FF9AA" bold="true" box="[199,334,1511,1537]" pageId="8" pageNumber="551">Etymology.</emphasis>
The specific epithet of this beautiful new species honors Cathy Ledec, a passionate conservationist and long-time supporter of conservational organizations working to preserve the habitat of salamanders in the Neotropics.
</paragraph>
</subSubSection>
<subSubSection id="C33203B2843BFF80FF2AF9F93ED6F8EF" pageId="8" pageNumber="551" type="biology_ecology">
<paragraph id="8B975039843BFF80FF2AF9F93ED6F8EF" blockId="8.[151,1437,1402,2041]" pageId="8" pageNumber="551">
<emphasis id="B95C8C2B843BFF80FF2AF9F93FD0F9C7" bold="true" box="[199,705,1618,1645]" pageId="8" pageNumber="551">Habitat and natural history observations.</emphasis>
<taxonomicName id="4C282BBA843BFF80FD27F9F93EE0F9C7" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[714,1009,1618,1644]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843BFF80FD27F9F93EE0F9C7" box="[714,1009,1618,1644]" italics="true" pageId="8" pageNumber="551">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
, was found in a montane rain forest, characterized by a short dry season (one to three months, 
<emphasis id="B95C8C2B843BFF80FCDDF9DC3E7FF93B" box="[816,878,1655,1680]" italics="true" pageId="8" pageNumber="551">sensu</emphasis>
<bibRefCitation id="EFB92DC8843BFF80FC95F9DD393FF93A" author="Holdridge, L. R." box="[888,1070,1654,1681]" pageId="8" pageNumber="551" refId="ref7051" refString="Holdridge, L. R. (1967) Life Zone Ecology. Tropical Science Center, San Jose, Costa Rica, 206 pp." type="book" year="1967">Holdridge 1967</bibRefCitation>
). Annual precipitation oscillates between 3600 and 
<quantity id="4CD0FDDC843BFF80FE84F9313CC5F91E" box="[361,468,1690,1717]" metricMagnitude="0" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="551" unit="mm" value="7500.0">7500 mm</quantity>
and the annual temperature between 12ºC and 17ºC. At this site trees are covered with epiphytes and bryophytes, with a dense undergrowth vegetation (
<figureCitation id="13134CBC843BFF80FC9FF9153EA7F972" box="[882,950,1726,1753]" captionStart="FIGURE 7" captionStartId="10.[151,250,1563,1587]" captionTargetBox="[151,1436,633,1539]" captionTargetId="figure-249@10.[151,1436,633,1539]" captionTargetPageId="10" captionText="FIGURE 7. Habitat, of Bolitoglossa cathyledecae sp. nov. from the cloud forest at the Serranía de Talamanca Panamá around 1900 m. A-C) Habitat of B. cathyledecae; D) field work made during the expedition where the new species was found. A-C Photographs taken by MP, D by Carlos Castillo." figureDoi="http://doi.org/10.5281/zenodo.6504120" httpUri="https://zenodo.org/record/6504120/files/figure.png" pageId="8" pageNumber="551">Fig. 7</figureCitation>
). The 
<typeStatus id="5493EE9B843BFF80FC13F9153971F973" box="[1022,1120,1726,1752]" pageId="8" pageNumber="551" type="holotype">holotype</typeStatus>
was found on 
<date id="FF9676F9843BFF80FAEBF914388DF972" box="[1286,1436,1727,1753]" pageId="8" pageNumber="551" value="2017-05-20">20 May 2017</date>
at 19:30 h. active on a feather bamboo of the genus 
<taxonomicName id="4C282BBA843BFF80FD3CF9493E76F956" box="[721,871,1762,1789]" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" order="Poales" pageId="8" pageNumber="551" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B95C8C2B843BFF80FD3CF9493E50F957" box="[721,833,1762,1788]" italics="true" pageId="8" pageNumber="551">Chusquea</emphasis>
sp.
</taxonomicName>
at 
<quantity id="4CD0FDDC843BFF80FC6AF9483EA3F956" box="[903,946,1763,1789]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="551" unit="m" value="1.0">1 m</quantity>
above the ground, there was a light rain that day. 
<typeStatus id="5493EE9B843BFF80FF21F8AC3C28F88A" box="[204,313,1799,1825]" pageId="8" pageNumber="551" type="paratype">Paratypes</typeStatus>
were found on 
<date id="FF9676F9843BFF80FE0AF8AD3FD6F88A" box="[487,711,1798,1825]" pageId="8" pageNumber="551" value="2017-09-31">September 31, 2017</date>
between 20:00‒23:30 h active on ferns approximately 
<quantity id="4CD0FDDC843BFF80FAC2F8AC3842F88A" box="[1327,1363,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="551" unit="m" value="2.0">2m</quantity>
above the ground. The night was humid due to rain between 16:00 and 18:00 h.
</paragraph>
</subSubSection>
<subSubSection id="C33203B2843BFF80FF2AF8E53C1BF81A" pageId="8" pageNumber="551" type="discussion">
<paragraph id="8B975039843BFF80FF2AF8E53C1BF81A" blockId="8.[151,1437,1402,2041]" pageId="8" pageNumber="551">
Other amphibian species collected in the vicinity of the 
<typeStatus id="5493EE9B843BFF80FCDFF8E43E73F8C2" box="[818,866,1871,1897]" pageId="8" pageNumber="551">type</typeStatus>
series include 
<taxonomicName id="4C282BBA843BFF80FBEAF8E53817F8C3" box="[1031,1286,1870,1896]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="compacta">
<emphasis id="B95C8C2B843BFF80FBEAF8E53817F8C3" box="[1031,1286,1870,1896]" italics="true" pageId="8" pageNumber="551">Bolitoglossa compacta</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843BFF80FAFEF8E43887F8C3" box="[1299,1430,1870,1896]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="minutula">
<emphasis id="B95C8C2B843BFF80FAFEF8E43887F8C3" box="[1299,1430,1870,1896]" italics="true" pageId="8" pageNumber="551">B. minutula</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843BFF80FF7AF8D83C71F827" baseAuthorityName="Barbour" baseAuthorityYear="1928" box="[151,352,1906,1933]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="blairi">
<emphasis id="B95C8C2B843BFF80FF7AF8D83C71F827" box="[151,352,1906,1933]" italics="true" pageId="8" pageNumber="551">Craugastor blairi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843BFF80FE9CF8D83FB3F827" baseAuthorityName="Cope" baseAuthorityYear="1875" box="[369,674,1906,1932]" class="Amphibia" family="Eleutherodactylidae" genus="Diasporus" isUncertain="true" kingdom="Animalia" order="Anura" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="hylaeformis">
<emphasis id="B95C8C2B843BFF80FE9CF8D83CF7F827" box="[369,486,1907,1932]" italics="true" pageId="8" pageNumber="551">Diasporus</emphasis>
aff. 
<emphasis id="B95C8C2B843BFF80FDF3F8D93FB3F827" box="[542,674,1906,1932]" italics="true" pageId="8" pageNumber="551">hylaeformis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843BFF80FD5EF8D93E67F827" baseAuthorityName="Cope" baseAuthorityYear="1875" box="[691,886,1906,1932]" class="Amphibia" family="Bufonidae" genus="Incilius" kingdom="Animalia" order="Anura" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="epioticus">
<emphasis id="B95C8C2B843BFF80FD5EF8D93E67F827" box="[691,886,1906,1932]" italics="true" pageId="8" pageNumber="551">Incilius epioticus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843BFF80FC6AF8D839C1F827" baseAuthorityName="Barbour" baseAuthorityYear="1928" box="[903,1232,1906,1932]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="caryophyllaceus">
<emphasis id="B95C8C2B843BFF80FC6AF8D839C1F827" box="[903,1232,1906,1932]" italics="true" pageId="8" pageNumber="551">Pristimantis caryophyllaceus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C282BBA843BFF80FAFEF8D83C13F81B" baseAuthorityName="Ibanez, Jaramillo &amp; Arosemena" baseAuthorityYear="1994" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="museosus">
<emphasis id="B95C8C2B843BFF80FAFEF8D83C13F81B" italics="true" pageId="8" pageNumber="551">Pristimantis museosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33203B2843BFF81FF2AF8113C9AFEB6" lastPageId="9" lastPageNumber="552" pageId="8" pageNumber="551" type="conservation">
<paragraph id="8B975039843BFF81FF2AF8113C9AFEB6" blockId="8.[151,1437,1402,2041]" lastBlockId="9.[151,1437,151,285]" lastPageId="9" lastPageNumber="552" pageId="8" pageNumber="551">
<emphasis id="B95C8C2B843BFF80FF2AF8113CAFF87E" bold="true" box="[199,446,1978,2005]" pageId="8" pageNumber="551">Conservation status.</emphasis>
This species is restricted to a small area of a high elevation habitat in the Talamanca Mountain range, suggesting it be classified as Critically Endangered (A3c, B2a [ii, iii, iv], according to the IUCN Red List categories and criteria (IUCN Standards and Petitions Committee. 2019). Anthropogenic pressure around the area will probably lead to further declines of populations through habitat deterioration and reduction of the area of occupancy. This species will also be allocated to “High” level of environmental vulnerability using the EVS of 
<bibRefCitation id="EFB92DC8843AFF81FF7AFEA83C96FEB6" author="Johnson, J. D. &amp; Mata-Silva, V. &amp; Wilson, L. D." box="[151,391,258,285]" pageId="9" pageNumber="552" pagination="1 - 94" refId="ref7293" refString="Johnson, J. D., Mata-Silva, V. &amp; Wilson, L. D. (2015) A conservation reassessment of the Central American herpetofauna based on the EVS measure. Amphibian &amp; Reptile Conservation, 9, 1 - 94." type="journal article" year="2015">
Johnson 
<emphasis id="B95C8C2B843AFF81FF17FEA83C23FEB6" box="[250,306,259,285]" italics="true" pageId="9" pageNumber="552">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5700B1843AFF81FF7AFB793DD1FA99" ID-DOI="http://doi.org/10.5281/zenodo.6504118" ID-Zenodo-Dep="6504118" httpUri="https://zenodo.org/record/6504118/files/figure.png" pageId="9" pageNumber="552" startId="9.[151,250,1234,1258]" targetBox="[198,1430,309,1211]" targetPageId="9">
<paragraph id="8B975039843AFF81FF7AFB793DD1FA99" blockId="9.[151,1436,1234,1330]" pageId="9" pageNumber="552">
<emphasis id="B95C8C2B843AFF81FF7AFB793C03FB41" bold="true" box="[151,274,1234,1258]" pageId="9" pageNumber="552">FIGURE 6.</emphasis>
Color variation in the paratypes. A) Dorsum in daytime coloration (MHCH 3242); B-C) Details of head and dorsum in daytime coloration (MHCH 3241); D) Paratype at the moment of encounter at night (MHCH 3242). Photographs taken by MP.
</paragraph>
</caption>
<subSubSection id="C33203B2843AFF81FF2AFACA3F71FA4C" pageId="9" pageNumber="552" type="distribution">
<paragraph id="8B975039843AFF81FF2AFACA3F71FA4C" blockId="9.[151,1437,1377,2015]" pageId="9" pageNumber="552">
<emphasis id="B95C8C2B843AFF81FF2AFACA3C70FAD0" bold="true" box="[199,353,1377,1403]" pageId="9" pageNumber="552">Distribution.</emphasis>
<taxonomicName id="4C282BBA843AFF81FE85FACA3F9FFAD0" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[360,654,1377,1403]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843AFF81FE85FACA3F9FFAD0" box="[360,654,1377,1403]" italics="true" pageId="9" pageNumber="552">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
is known only from one site within the montane rain forest life zone (
<emphasis id="B95C8C2B843AFF81FF4CFA2D3DCEFA34" box="[161,223,1414,1439]" italics="true" pageId="9" pageNumber="552">sensu</emphasis>
<bibRefCitation id="EFB92DC8843AFF81FF09FA2E3C85FA0B" author="Holdridge, L. R." box="[228,404,1413,1440]" pageId="9" pageNumber="552" refId="ref7051" refString="Holdridge, L. R. (1967) Life Zone Ecology. Tropical Science Center, San Jose, Costa Rica, 206 pp." type="book" year="1967">Holdridge 1967</bibRefCitation>
) along the northeastern mid-elevation slopes of the Cordillera de Talamanca, in the vicinity of the continental divide between 
<collectingRegion id="49EC9EDB843AFF81FE19FA023F40FA68" box="[500,593,1449,1475]" country="Panama" name="Chiriqui" pageId="9" pageNumber="552">Chiriquí</collectingRegion>
and 
<collectingRegion id="49EC9EDB843AFF81FD6AFA023E20FA68" box="[647,817,1449,1475]" country="Panama" name="Bocas del Toro" pageId="9" pageNumber="552">Bocas del Toro</collectingRegion>
, within the La Amistad National Park, district Boquete, at about 
<quantity id="4CD0FDDC843AFF81FF15FA663C5CFA4C" box="[248,333,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="552" unit="m" value="1900.0">1900 m</quantity>
above sea level (
<figureCitation id="13134CBC843AFF81FDE3FA663F5EFA4C" box="[526,591,1485,1511]" captionStart="FIGURE 1" captionStartId="1.[151,250,1981,2005]" captionTargetBox="[155,1427,981,1955]" captionTargetId="figure-302@1.[151,1436,973,1958]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the type locality for Bolitoglossa cathyledecae sp. nov. (red hexagon) and paratypes record (blue hexagon), in the Cordillera de Talamanca." figureDoi="http://doi.org/10.5281/zenodo.6504106" httpUri="https://zenodo.org/record/6504106/files/figure.png" pageId="9" pageNumber="552">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33203B2843AFF82FF2AFA5A3833FDCA" lastPageId="10" lastPageNumber="553" pageId="9" pageNumber="552" type="discussion">
<paragraph id="8B975039843AFF82FF2AFA5A3CCAFECE" blockId="9.[151,1437,1377,2015]" lastBlockId="10.[151,1437,151,609]" lastPageId="10" lastPageNumber="553" pageId="9" pageNumber="552">
<emphasis id="B95C8C2B843AFF81FF2AFA5A3C2AF9A0" bold="true" box="[199,315,1521,1547]" pageId="9" pageNumber="552">Remarks.</emphasis>
The striking color pattern and the differences in molecular makeup, support the recognition of 
<taxonomicName id="4C282BBA843AFF81FF7AF9BE3CD0F984" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[151,449,1557,1583]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843AFF81FF7AF9BE3CD0F984" box="[151,449,1557,1583]" italics="true" pageId="9" pageNumber="552">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
as a species new to science in lower Central America, placed within the subgenus 
<taxonomicName id="4C282BBA843AFF81FF7AF9923CDFF9F8" authority="(Parra et al. 2004)" baseAuthorityName="Parra" baseAuthorityYear="2004" box="[151,462,1593,1619]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B843AFF81FF7AF9923DEDF9F8" box="[151,252,1593,1619]" italics="true" pageId="9" pageNumber="552">Eladinea</emphasis>
(Parra 
<emphasis id="B95C8C2B843AFF81FEA0F9913C95F9F8" box="[333,388,1593,1619]" italics="true" pageId="9" pageNumber="552">et al.</emphasis>
2004)
</taxonomicName>
. 
<taxonomicName id="4C282BBA843AFF81FE35F9923FEDF9F8" authority="Ponce &amp; Navarro &amp; Morales &amp; Batista, 2022" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[472,764,1593,1619]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae" status="sp. nov.">
<emphasis id="B95C8C2B843AFF81FE35F9923FEDF9F8" box="[472,764,1593,1619]" italics="true" pageId="9" pageNumber="552">Bolitoglossa cathyledecae</emphasis>
</taxonomicName>
<emphasis id="B95C8C2B843AFF81FCEFF9923E4BF9F8" bold="true" box="[770,858,1593,1619]" pageId="9" pageNumber="552">
<taxonomicNameLabel id="A26F3150843AFF81FCEFF9923E4BF9F8" box="[770,858,1593,1619]" pageId="9" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is the 23rd 
<taxonomicName id="4C282BBA843AFF81FC0CF99239D5F9F8" box="[993,1220,1593,1619]" pageId="9" pageNumber="552">
<emphasis id="B95C8C2B843AFF81FC0CF992397CF9F8" box="[993,1133,1593,1619]" italics="true" pageId="9" pageNumber="552">Bolitoglossa</emphasis>
species
</taxonomicName>
present in 
<collectingCountry id="F33F10A9843AFF81FAD0F9923888F9F8" box="[1341,1433,1593,1619]" name="Panama" pageId="9" pageNumber="552">Panama</collectingCountry>
. The new species is phylogenetically related to the 
<taxonomicName id="4C282BBA843AFF81FD2AF9F63E31F9DC" box="[711,800,1629,1655]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B843AFF81FD2AF9F63E31F9DC" box="[711,800,1629,1655]" italics="true" pageId="9" pageNumber="552">epimela</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C282BBA843AFF81FCB1F9F63ECEF9DC" box="[860,991,1629,1655]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B843AFF81FCB1F9F63ECEF9DC" box="[860,991,1629,1655]" italics="true" pageId="9" pageNumber="552">subpalmata</emphasis>
</taxonomicName>
species group (
<emphasis id="B95C8C2B843AFF81FB63F9F539DDF9DC" box="[1166,1228,1630,1655]" italics="true" pageId="9" pageNumber="552">sensu</emphasis>
<bibRefCitation id="EFB92DC8843AFF81FB3FF9F63DC0F930" author="Garcia-Paris, M. &amp; Parra-Olea, G. &amp; Wake, D. B." pageId="9" pageNumber="552" pagination="23 - 31" refId="ref6883" refString="Garcia-Paris, M., Parra-Olea, G., &amp; Wake, D. B. (2008) Description of a new species of the Bolitoglossa subpalmata group (Caudata: Plethodontidae) from Costa Rica. The Herpetological Journal, 18, 23 - 31." type="journal article" year="2008">
Garcia-Paris 
<emphasis id="B95C8C2B843AFF81FA88F9F5388CF9DC" box="[1381,1437,1629,1655]" italics="true" pageId="9" pageNumber="552">et al.</emphasis>
2008
</bibRefCitation>
) and shows the lowest genetic distance to 
<taxonomicName id="4C282BBA843AFF81FD59F9293E32F930" box="[692,803,1665,1691]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="bramei">
<emphasis id="B95C8C2B843AFF81FD59F9293E32F930" box="[692,803,1665,1691]" italics="true" pageId="9" pageNumber="552">B. bramei</emphasis>
</taxonomicName>
, a species of the 
<taxonomicName id="4C282BBA843AFF81FC04F92A397DF930" box="[1001,1132,1665,1691]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B843AFF81FC04F92A397DF930" box="[1001,1132,1665,1691]" italics="true" pageId="9" pageNumber="552">subpalmata</emphasis>
</taxonomicName>
group, with distribution in southwestern 
<collectingCountry id="F33F10A9843AFF81FEDCF90E3CBBF914" box="[305,426,1701,1727]" name="Costa Rica" pageId="9" pageNumber="552">Costa Rica</collectingCountry>
and western 
<collectingCountry id="F33F10A9843AFF81FDD1F90E3F89F914" box="[572,664,1701,1727]" name="Panama" pageId="9" pageNumber="552">Panama</collectingCountry>
. Considering that the estimated average evolutionary divergence over sequence pairs within the groups mentioned above is as high as 3.5% (3.5% for 
<taxonomicName id="4C282BBA843AFF81FBE0F9623977F948" box="[1037,1126,1737,1763]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B843AFF81FBE0F9623977F948" box="[1037,1126,1737,1763]" italics="true" pageId="9" pageNumber="552">epimela</emphasis>
</taxonomicName>
, and 3.3% for 
<taxonomicName id="4C282BBA843AFF81FAE4F962389DF948" box="[1289,1420,1737,1763]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B843AFF81FAE4F962389DF948" box="[1289,1420,1737,1763]" italics="true" pageId="9" pageNumber="552">subpalmata</emphasis>
</taxonomicName>
), contrary to the 
<emphasis id="B95C8C2B843AFF81FEAEF9463CB9F8AC" box="[323,424,1773,1799]" italics="true" pageId="9" pageNumber="552">adspersa</emphasis>
species group where the genetic distances between some species are higher (up to 11.9%, 
<bibRefCitation id="EFB92DC8843AFF81FF7AF8BA3C63F880" author="Batista, A. &amp; Kohler, G. &amp; Mebert, K. &amp; Vesely, M." box="[151,370,1809,1835]" pageId="9" pageNumber="552" pagination="97 - 121" refId="ref6299" refString="Batista, A., Kohler, G., Mebert, K. &amp; Vesely, M. (2014) A new species of Bolitoglossa (Amphibia: Plethodontidae) from eastern Panama, with comments on other species of the adspersa species group from eastern Panama. Mesoamerican Herpetology, 1, 97 - 121." type="journal article" year="2014">
Batista 
<emphasis id="B95C8C2B843AFF81FF1DF8B93C34F880" box="[240,293,1809,1835]" italics="true" pageId="9" pageNumber="552">et al</emphasis>
. 2014
</bibRefCitation>
; 
<bibRefCitation id="EFB92DC8843AFF81FE6CF8BA3F66F880" author="Jaramillo, A. F. &amp; De La Riva, I. &amp; Guayasamin, J. M. &amp; Chaparro, J. C. &amp; Gagliardi-Urrutia, G. &amp; Gutierrez, R. C. &amp; Castroviejo-Fisher, S." box="[385,631,1809,1835]" pageId="9" pageNumber="552" pagination="106841" refId="ref7205" refString="Jaramillo, A. F., De La Riva, I., Guayasamin, J. M., Chaparro, J. C., Gagliardi-Urrutia, G., Gutierrez, R. C. &amp; Castroviejo-Fisher, S. (2020) Vastly underestimated species richness of Amazonian salamanders (Plethodontidae: Bolitoglossa) and implications about plethodontid diversification. Molecular Phylogenetics and Evolution, 149, 106841. https: // doi. org / 10.1016 / j. ympev. 2020.106841" type="journal article" year="2020">
Jaramillo 
<emphasis id="B95C8C2B843AFF81FE19F8B93F21F880" box="[500,560,1809,1835]" italics="true" pageId="9" pageNumber="552">et al.</emphasis>
2020
</bibRefCitation>
), along with the morphological distinctions found in 
<taxonomicName id="4C282BBA843AFF81FB0AF8B9388DF880" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[1255,1436,1809,1835]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843AFF81FB0AF8B9388DF880" box="[1255,1436,1809,1835]" italics="true" pageId="9" pageNumber="552">B. cathyledecae</emphasis>
</taxonomicName>
compared to other species groups in the subgenus 
<taxonomicName id="4C282BBA843AFF81FD24F89E3E3FF8E4" box="[713,814,1845,1871]" class="Amphibia" family="Plethodontidae" genus="Eladinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="genus">
<emphasis id="B95C8C2B843AFF81FD24F89E3E3FF8E4" box="[713,814,1845,1871]" italics="true" pageId="9" pageNumber="552">Eladinea</emphasis>
</taxonomicName>
; we refrain from assigning 
<taxonomicName id="4C282BBA843AFF81FB89F89D3805F8E4" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[1124,1300,1845,1871]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843AFF81FB89F89D3805F8E4" box="[1124,1300,1845,1871]" italics="true" pageId="9" pageNumber="552">B. cathyledecae</emphasis>
</taxonomicName>
to a species group at this time. Although the Talamancan clade in the phylogenetic analysis, with the 
<taxonomicName id="4C282BBA843AFF81FB42F8F23819F8D8" box="[1199,1288,1881,1907]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B843AFF81FB42F8F23819F8D8" box="[1199,1288,1881,1907]" italics="true" pageId="9" pageNumber="552">epimela</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA843AFF81FAF7F8F2388CF8D8" box="[1306,1437,1881,1907]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B843AFF81FAF7F8F2388CF8D8" box="[1306,1437,1881,1907]" italics="true" pageId="9" pageNumber="552">subpalmata</emphasis>
</taxonomicName>
species group and 
<taxonomicName id="4C282BBA843AFF81FE9DF8D63F13F83C" authorityName="Ponce &amp; Navarro &amp; Morales &amp; Batista" authorityYear="2022" box="[368,514,1917,1943]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="cathyledecae">
<emphasis id="B95C8C2B843AFF81FE9DF8D63F13F83C" box="[368,514,1917,1943]" italics="true" pageId="9" pageNumber="552">cathyledecae</emphasis>
</taxonomicName>
was not supported by the ML analysis, it got strong support with the Bayesian analysis (in part). In both phylogenetic hypothesis, there is a paraphyly, having species of the 
<taxonomicName id="4C282BBA843AFF81FB48F80A39EFF810" box="[1189,1278,1953,1979]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B843AFF81FB48F80A39EFF810" box="[1189,1278,1953,1979]" italics="true" pageId="9" pageNumber="552">epimela</emphasis>
</taxonomicName>
species group nested within species of the 
<taxonomicName id="4C282BBA843AFF81FE20F86E3F41F874" box="[461,592,1989,2015]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B843AFF81FE20F86E3F41F874" box="[461,592,1989,2015]" italics="true" pageId="9" pageNumber="552">subpalmata</emphasis>
</taxonomicName>
species group (
<figureCitation id="13134CBC843AFF81FD16F86E3E2FF874" box="[763,830,1989,2015]" captionStart="FIGURE 2" captionStartId="5.[151,250,901,925]" captionTargetBox="[170,1419,185,875]" captionTargetId="figure-19@5.[168,1419,181,878]" captionTargetPageId="5" captionText="FIGURE 2. Maximum likelihood trees based on 16S mtDNA sequence of Bolitoglossa spp. of the Eladinea subgenus. SHaLRT (Shimodaira and Hasegawa approximate likelihood ratio test) support (%) / ultrafast bootstrap support (%) are shown on the internodes. Outgroups not shown." figureDoi="http://doi.org/10.5281/zenodo.6504108" httpUri="https://zenodo.org/record/6504108/files/figure.png" pageId="9" pageNumber="552">Fig. 2</figureCitation>
), although there is one evident clade, with the northern 
<taxonomicName id="4C282BBA8439FF82FF7AFF3C3C0BFF1A" box="[151,282,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8439FF82FF7AFF3C3C0BFF1A" box="[151,282,151,177]" italics="true" pageId="10" pageNumber="553">subpalmata</emphasis>
</taxonomicName>
species from 
<collectingCountry id="F33F10A98439FF82FE50FF3C3F28FF1A" box="[445,569,151,177]" name="Costa Rica" pageId="10" pageNumber="553">Costa Rica</collectingCountry>
(
<taxonomicName id="4C282BBA8439FF82FDA1FF3C3FE1FF1A" box="[588,752,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8439FF82FDA1FF3C3FE1FF1A" box="[588,752,151,177]" italics="true" pageId="10" pageNumber="553">B. subpalmata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8439FF82FCEDFF3C3E5AFF1B" box="[768,843,151,176]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="tica">
<emphasis id="B95C8C2B8439FF82FCEDFF3C3E5AFF1B" box="[768,843,151,176]" italics="true" pageId="10" pageNumber="553">B. tica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8439FF82FCB1FF3C3EC2FF1A" box="[860,979,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B95C8C2B8439FF82FCB1FF3C3EC2FF1A" box="[860,979,151,177]" italics="true" pageId="10" pageNumber="553">B. gracilis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8439FF82FC0EFF3C3960FF1A" box="[995,1137,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="splendida">
<emphasis id="B95C8C2B8439FF82FC0EFF3C3960FF1A" box="[995,1137,151,177]" italics="true" pageId="10" pageNumber="553">B. splendida</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8439FF82FB6CFF3C3819FF1A" box="[1153,1288,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B95C8C2B8439FF82FB6CFF3C3819FF1A" box="[1153,1288,151,177]" italics="true" pageId="10" pageNumber="553">B. pesrubra</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C282BBA8439FF82FAF5FF3C3893FF1A" box="[1304,1410,151,177]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B95C8C2B8439FF82FAF5FF3C3893FF1A" box="[1304,1410,151,177]" italics="true" pageId="10" pageNumber="553">B. kamuk</emphasis>
</taxonomicName>
)). The evidence of the paraphyly found among the species in the 
<taxonomicName id="4C282BBA8439FF82FCB4FF103ECDFF7E" box="[857,988,187,213]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8439FF82FCB4FF103ECDFF7E" box="[857,988,187,213]" italics="true" pageId="10" pageNumber="553">subpalmata</emphasis>
</taxonomicName>
species group found in this study could be an artifact of the use of a single genetic marker, given that, the 
<taxonomicName id="4C282BBA8439FF82FC65FF743EF0FF52" box="[904,993,223,249]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="epimela">
<emphasis id="B95C8C2B8439FF82FC65FF743EF0FF52" box="[904,993,223,249]" italics="true" pageId="10" pageNumber="553">epimela</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C282BBA8439FF82FBF7FF74398CFF52" box="[1050,1181,223,249]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B95C8C2B8439FF82FBF7FF74398CFF52" box="[1050,1181,223,249]" italics="true" pageId="10" pageNumber="553">subpalmata</emphasis>
</taxonomicName>
clades have been well supported in other studies when using more than one molecular marker (
<bibRefCitation id="EFB92DC88439FF82FC3FFEA839F7FEB6" author="Kubicki, B. &amp; Arias, E." box="[978,1254,259,285]" pageId="10" pageNumber="553" pagination="329 - 346" refId="ref7471" refString="Kubicki, B. &amp; Arias, E. (2016) A beautiful new yellow salamander, genus Bolitoglossa (Caudata: Plethodontidae), from the northeastern slopes of the Cordillera de Talamanca, Costa Rica. Zootaxa, 4184 (2), 329 - 346. https: // doi. org / 10.11646 / zootaxa. 4184.2.5" type="journal article" year="2016">Kubicki and Arias 2016</bibRefCitation>
, 
<bibRefCitation id="EFB92DC88439FF82FB1FFEA83DC2FEEA" author="Jaramillo, A. F. &amp; De La Riva, I. &amp; Guayasamin, J. M. &amp; Chaparro, J. C. &amp; Gagliardi-Urrutia, G. &amp; Gutierrez, R. C. &amp; Castroviejo-Fisher, S." pageId="10" pageNumber="553" pagination="106841" refId="ref7205" refString="Jaramillo, A. F., De La Riva, I., Guayasamin, J. M., Chaparro, J. C., Gagliardi-Urrutia, G., Gutierrez, R. C. &amp; Castroviejo-Fisher, S. (2020) Vastly underestimated species richness of Amazonian salamanders (Plethodontidae: Bolitoglossa) and implications about plethodontid diversification. Molecular Phylogenetics and Evolution, 149, 106841. https: // doi. org / 10.1016 / j. ympev. 2020.106841" type="journal article" year="2020">
Jaramillo 
<emphasis id="B95C8C2B8439FF82FA8EFEA8388CFEB6" box="[1379,1437,259,285]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2020
</bibRefCitation>
), although in those studies 
<taxonomicName id="4C282BBA8439FF82FDECFE8C3F8AFEEA" box="[513,667,295,321]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="anthracina">
<emphasis id="B95C8C2B8439FF82FDECFE8C3F8AFEEA" box="[513,667,295,321]" italics="true" pageId="10" pageNumber="553">B. anthracina</emphasis>
</taxonomicName>
has not been included which could give a different framework of the relationship between groups.
</paragraph>
<paragraph id="8B9750398439FF82FF2AFEC43833FDCA" blockId="10.[151,1437,151,609]" pageId="10" pageNumber="553">
Taxonomic work is still needed in the highlands between 
<collectingCountry id="F33F10A98439FF82FC8CFEC43ECCFE22" box="[865,989,367,393]" name="Costa Rica" pageId="10" pageNumber="553">Costa Rica</collectingCountry>
and 
<collectingCountry id="F33F10A98439FF82FBF5FEC43960FE22" box="[1048,1137,367,393]" name="Panama" pageId="10" pageNumber="553">Panamá</collectingCountry>
(
<bibRefCitation id="EFB92DC88439FF82FB6FFEC43886FE22" author="Garcia-Paris, M. &amp; Parra-Olea, G. &amp; Wake, D. B." box="[1154,1431,367,393]" pageId="10" pageNumber="553" pagination="23 - 31" refId="ref6883" refString="Garcia-Paris, M., Parra-Olea, G., &amp; Wake, D. B. (2008) Description of a new species of the Bolitoglossa subpalmata group (Caudata: Plethodontidae) from Costa Rica. The Herpetological Journal, 18, 23 - 31." type="journal article" year="2008">
Garcia-Paris 
<emphasis id="B95C8C2B8439FF82FAF4FEC43842FE22" box="[1305,1363,367,393]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2008
</bibRefCitation>
; 
<bibRefCitation id="EFB92DC88439FF82FF7AFE383CB7FE06" author="Bolanos, F. &amp; Wake, D. B." box="[151,422,403,429]" pageId="10" pageNumber="553" pagination="57 - 68" refId="ref6466" refString="Bolanos, F. &amp; Wake, D. B. (2009) Two new species of mountain web-footed salamanders (Plethodontidae: Bolitoglossa) from the Costa Rica-Panama border region. Zootaxa, 1981 (1), 57 - 68. https: // doi. org / 10.11646 / zootaxa. 1981.1.5" type="journal article" year="2009">
Bolaños and 
<collectingCountry id="F33F10A98439FF82FECBFE383C74FE06" box="[294,357,403,429]" name="Wake Island" pageId="10" pageNumber="553">Wake</collectingCountry>
2009
</bibRefCitation>
; Boza-Oviedo 
<emphasis id="B95C8C2B8439FF82FDA1FE383F92FE06" box="[588,643,403,429]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2012; 
<bibRefCitation id="EFB92DC88439FF82FD23FE383EC9FE05" author="Kubicki, B. &amp; Arias, E." box="[718,984,403,430]" pageId="10" pageNumber="553" pagination="329 - 346" refId="ref7471" refString="Kubicki, B. &amp; Arias, E. (2016) A beautiful new yellow salamander, genus Bolitoglossa (Caudata: Plethodontidae), from the northeastern slopes of the Cordillera de Talamanca, Costa Rica. Zootaxa, 4184 (2), 329 - 346. https: // doi. org / 10.11646 / zootaxa. 4184.2.5" type="journal article" year="2016">Kubicki and Arias 2016</bibRefCitation>
), with vast remote areas still unexplored, mainly along the Serranía de Talamanca. Cloud forests are vulnerable to climate change due to the low range of mobility of the upper highland endemic species (
<bibRefCitation id="EFB92DC88439FF82FD5DFE703E6CFE5E" author="Davies, K. F. &amp; Margules, C. R. &amp; Lawrence, J. F." box="[688,893,475,501]" pageId="10" pageNumber="553" pagination="265 - 271" refId="ref6698" refString="Davies, K. F., Margules, C. R. &amp; Lawrence, J. F. (2004) A synergistic effect puts rare, specialized species at greater risk of extinction. Ecology, 85, 265 - 271. https: // doi. org / 10.1890 / 03 - 0110" type="journal article" year="2004">
Davies 
<emphasis id="B95C8C2B8439FF82FCE8FE703E2DFE5E" box="[773,828,475,501]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2004
</bibRefCitation>
; 
<bibRefCitation id="EFB92DC88439FF82FC6AFE703968FE5E" author="Paaijmans, K. P. &amp; Heinig, R. L. &amp; Seliga, R. A. &amp; Blanford, J. I. &amp; Blanford, S. &amp; Murdock, C. C. &amp; Thomas, M. B." box="[903,1145,475,501]" pageId="10" pageNumber="553" pagination="2373 - 2380" refId="ref7782" refString="Paaijmans, K. P., Heinig, R. L., Seliga, R. A., Blanford, J. I., Blanford, S., Murdock, C. C. &amp; Thomas, M. B. (2013) Temperature variation makes ectotherms more sensitive to climate change. Global Change Biology, 19, 2373 - 2380. https: // doi. org / 10.1111 / gcb. 12240" type="journal article" year="2013">
Paaijmans 
<emphasis id="B95C8C2B8439FF82FC12FE703927FE5E" box="[1023,1078,475,501]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2013
</bibRefCitation>
). Consequently, species or populations restricted to such habitats are enormously susceptible to slight changes in the environment and thus face an elevated risk of extinction (Batista 
<emphasis id="B95C8C2B8439FF82FDD7FD883F60FD96" box="[570,625,547,573]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2020). An urgent conservation plan is required to protect the surrounding cloudforest and the entire La Amistad International Park to preserve its unique pool of endemic species.
</paragraph>
</subSubSection>
</treatment>
</document>