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<document ID-DOI="http://doi.org/10.5281/zenodo.4776261" ID-GBIF-Dataset="2ab49e31-d925-463d-891e-f7061492af8b" ID-GBIF-Taxon="182247911" ID-ISSN="1982-0224" ID-Zenodo-Dep="4776261" LSID="urn:lsid:zoobank.org:act:42F632D8-A127-4092-B09B-708882C0C3B9" checkinTime="1613739459940" checkinUser="carolina" docAuthor="Cramer, Christian Andreas &amp; Py-Daniel, Lúcia Helena Rapp" docDate="2015" docId="038187DCB901FFEA0FB47729FE6FC99B" docLanguage="en" docName="NeotropIchthyol.13.3.461-470.pdf" docOrigin="Neotropical Ichthyology (Neotrop. Ichthyol.) 13 (3)" docSource="http://dx.doi.org/10.1590/1982-0224-20140099" docStyle="DocumentStyle:7EA6CBAC9B92AC526D6A0354C904CC04.3:NeotropIchthyol.2014-2015.journal_article" docStyleId="7EA6CBAC9B92AC526D6A0354C904CC04" docStyleName="NeotropIchthyol.2014-2015.journal_article" docStyleVersion="3" docTitle="Panaqolus nix Cramer &amp; Py-Daniel 2015, new species" docType="treatment" docUuid="42F632D8-A127-4092-B09B-708882C0C3B9" docUuidSource="ZooBank" docVersion="7" lastPageId="7" lastPageNumber="468" masterDocId="FFB8FFA4B900FFED0C667566FF88CD5D" masterDocTitle="A new species of Panaqolus (Siluriformes: Loricariidae) from the rio Madeira basin with remarkable intraspecific color variation" masterLastPageNumber="470" masterPageNumber="461" pageId="1" pageNumber="462" updateTime="1636535390443" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Panaqolus (Siluriformes: Loricariidae) from the rio Madeira basin with remarkable intraspecific color variation</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cramer, Christian Andreas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Py-Daniel, Lúcia Helena Rapp</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="series">
<mods:title>Neotrop. Ichthyol.</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2015-08-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>13</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>461</mods:start>
<mods:end>470</mods:end>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1590/1982-0224-20140099</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/1982-0224-20140099</mods:identifier>
<mods:identifier type="GBIF-Dataset">2ab49e31-d925-463d-891e-f7061492af8b</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4551377</mods:identifier>
<mods:identifier type="ZooBank">12677D4B-E956-4303-AAB0-47C50106A099</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4776261" ID-GBIF-Taxon="182247911" ID-Zenodo-Dep="4776261" LSID="urn:lsid:zoobank.org:act:42F632D8-A127-4092-B09B-708882C0C3B9" httpUri="http://treatment.plazi.org/id/038187DCB901FFEA0FB47729FE6FC99B" lastPageId="7" lastPageNumber="468" pageId="1" pageNumber="462">
<subSubSection pageId="1" pageNumber="462" type="nomenclature">
<paragraph blockId="1.[978,1285,591,615]" box="[978,1285,591,615]" pageId="1" pageNumber="462">
<heading box="[978,1285,591,615]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="462" reason="2">
<emphasis bold="true" box="[978,1285,591,615]" pageId="1" pageNumber="462">
<taxonomicName authority="Cramer &amp; Py-Daniel, 2015" authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[978,1140,591,615]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="462" phylum="Chordata" rank="species" species="nix" status="new species">
<emphasis bold="true" box="[978,1140,591,615]" italics="true" pageId="1" pageNumber="462">Panaqolus nix</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1154,1285,591,615]" pageId="1" pageNumber="462" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[813,1451,655,741]" pageId="1" pageNumber="462">
u r n:l s i d:z o o b a n k.o r g:a c t:
<uuid pageId="1" pageNumber="462">42F632 D 8-A127- 4 0 9 2 -B0 9B - 708882C0C3B9</uuid>
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="463" pageId="1" pageNumber="462" type="description">
<paragraph blockId="1.[813,1451,655,741]" lastBlockId="2.[136,775,174,260]" lastPageId="2" lastPageNumber="463" pageId="1" pageNumber="462">
(
<figureCitation box="[1085,1177,717,741]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="1.[129,169,1871,1895]" captionStartId-1="4.[136,176,1422,1446]" captionStartId-2="5.[129,168,1016,1040]" captionStartId-3="5.[129,168,1542,1566]" captionStartId-4="6.[136,176,1046,1070]" captionTargetBox-0="[374,1206,786,1858]" captionTargetBox-1="[136,775,486,1408]" captionTargetBox-2="[166,1413,174,1006]" captionTargetBox-3="[208,1395,1113,1522]" captionTargetBox-4="[261,1333,174,1033]" captionTargetId-0="figure-277@1.[374,1206,786,1858]" captionTargetId-1="figure-85@4.[136,775,486,1408]" captionTargetId-2="figure-181@5.[166,1413,174,1006]" captionTargetId-3="figure-218@5.[166,1413,1104,1537]" captionTargetId-4="figure-10@6.[261,1333,174,1033]" captionTargetPageId-0="1" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Fig. 1. Dorsal, lateral, and ventral views of Panaqolus nix, holotype, INPA 39606, male, 110.1 mm SL, Brazil, Rondônia, rio Madeira, cofferdam at construction site of Santo Antônio hydroelectric power plant (former Santo Antônio rapids) (Photos by B. S. F. Barros)." captionText-1="Fig. 2. Details of the mouth of Panaqolus nix: the dentary tooth rows are in an acute angle (below) or even nearly parallel (above) (MZUSP 114009, 68.8 mm + 76.2 mm SL). Scale bar = 5 mm." captionText-2="Fig. 3. Variation in the color of Panaqolus nix: a) nearly white specimen (UFRO-I 10050, 90.4 mm SL) b), intermediately colored specimen (UFRO-I 7968, 81.4 mm SL) (Photos by B. S. F. Barros)." captionText-3="Fig. 4. Live photo of the holotype of Panaqolus nix (INPA 39606; 110.1 mm SL)." captionText-4="Fig. 5. Sexual dimorphic male of Panaqolus nix (INPA 41150 (29859); 105.1 mm SL) with elongated odontodes on pectoral spine, caudal peduncle and caudal fin." figureDoi-0="http://doi.org/10.5281/zenodo.4551379" figureDoi-1="http://doi.org/10.5281/zenodo.4551383" figureDoi-2="http://doi.org/10.5281/zenodo.4551385" figureDoi-3="http://doi.org/10.5281/zenodo.4551388" figureDoi-4="http://doi.org/10.5281/zenodo.4551390" httpUri-0="https://zenodo.org/record/4551379/files/figure.png" httpUri-1="https://zenodo.org/record/4551383/files/figure.png" httpUri-2="https://zenodo.org/record/4551385/files/figure.png" httpUri-3="https://zenodo.org/record/4551388/files/figure.png" httpUri-4="https://zenodo.org/record/4551390/files/figure.png" pageId="1" pageNumber="462">Figs. 1-5</figureCitation>
)
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[136,222,175,196]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="genus">
<emphasis box="[136,222,175,196]" italics="true" pageId="2" pageNumber="463">Panaque</emphasis>
</taxonomicName>
sp. 1: photo - Zawadzki &amp; Chamon, 2013: p. 312.
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[136,222,207,228]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="genus">
<emphasis box="[136,222,207,228]" italics="true" pageId="2" pageNumber="463">Panaque</emphasis>
</taxonomicName>
sp. 2: UFRO-I 6384, Zawadzki &amp; Chamon, 2013: p. 313
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551379" ID-Zenodo-Dep="4551379" httpUri="https://zenodo.org/record/4551379/files/figure.png" pageId="1" pageNumber="462" startId="1.[129,169,1871,1895]" targetBox="[374,1206,786,1858]" targetPageId="1">
<paragraph blockId="1.[129,1451,1871,1959]" pageId="1" pageNumber="462">
<emphasis bold="true" box="[129,198,1871,1895]" pageId="1" pageNumber="462">Fig. 1.</emphasis>
Dorsal, lateral, and ventral views of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[590,743,1871,1895]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="462" phylum="Chordata" rank="species" species="nix">
<emphasis box="[590,743,1871,1895]" italics="true" pageId="1" pageNumber="462">Panaqolus nix</emphasis>
</taxonomicName>
, holotype, INPA 39606, male, 110.1 mm SL, Brazil, Rondônia, rio Madeira, cofferdam at construction site of Santo Antônio hydroelectric power plant (former Santo Antônio rapids) (Photos by B. S. F. Barros).
</paragraph>
</caption>
<paragraph blockId="2.[136,775,174,260]" box="[174,540,238,260]" pageId="2" pageNumber="463">(see remarks for further explanation).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="463" type="materials_examined">
<paragraph blockId="2.[136,775,301,453]" pageId="2" pageNumber="463">
<materialsCitation ID-GBIF-Occurrence="3114482311" collectingDate="2012-02-14" collectionCode="INPA" collectorName="C. A. Cramer." country="Brazil" latitude="-8.801667" location="Santo Antonio" longLatPrecision="21" longitude="-63.95" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="INPA 39606" specimenCount="1" stateProvince="Rondonia" typeStatus="holotype">
<emphasis bold="true" box="[136,248,301,325]" pageId="2" pageNumber="463">
<typeStatus box="[136,243,301,325]" pageId="2" pageNumber="463">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[266,415,301,325]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">INPA 39606</specimenCode>
, male,
<quantity box="[513,627,301,325]" metricMagnitude="-1" metricUnit="m" metricValue="1.101" pageId="2" pageNumber="463" unit="mm" value="110.1">110.1 mm</quantity>
SL,
<collectingCountry box="[701,770,301,325]" name="Brazil" pageId="2" pageNumber="463">Brazil</collectingCountry>
,
<collectingRegion box="[136,243,333,357]" country="Brazil" name="Rondonia" pageId="2" pageNumber="463">Rondônia</collectingRegion>
,
<collectingMunicipality box="[258,393,333,357]" pageId="2" pageNumber="463">rio Madeira</collectingMunicipality>
, cofferdam at construction site of
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0CEE740BFEA4CCD8" box="[136,300,365,389]" country="Brazil" latitude="-8.801667" longLatPrecision="21" longitude="-63.95" municipality="rio Madeira" name="Santo Antonio" pageId="2" pageNumber="463" stateProvince="Rondonia">Santo Antônio</location>
hydroelectric power plant (former Santo Antônio rapids),
<geoCoordinate box="[321,446,397,421]" degrees="08" direction="south" minutes="48" orientation="latitude" pageId="2" pageNumber="463" precision="15" seconds="06" value="-8.801667">08°4806”S</geoCoordinate>
<geoCoordinate box="[454,589,397,421]" degrees="63" direction="west" minutes="57" orientation="longitude" pageId="2" pageNumber="463" precision="15" seconds="00" value="-63.95">63°5700”W</geoCoordinate>
,
<date box="[602,737,397,421]" pageId="2" pageNumber="463" value="2012-02-14">
<collectingDate box="[602,737,397,421]" pageId="2" pageNumber="463" value="2012-02-14">14 Feb 2012</collectingDate>
</date>
,
<collectorName pageId="2" pageNumber="463">C. A. Cramer.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="2.[136,776,493,1285]" pageId="2" pageNumber="463">
<materialsCitation ID-GBIF-Occurrence="3114482306" collectionCode="INPA" country="Brazil" location="Sao Lourenco" municipality="Mamore" pageId="2" pageNumber="463" specimenCode="INPA 39605, 3" specimenCount="48" stateProvince="Rondonia" typeStatus="paratype">
<emphasis bold="true" box="[136,259,493,517]" pageId="2" pageNumber="463">
<typeStatus box="[136,254,493,517]" pageId="2" pageNumber="463">Paratypes</typeStatus>
.
</emphasis>
<specimenCount box="[271,426,493,517]" pageId="2" pageNumber="463" type="generic">48 specimens</specimenCount>
.
<collectingCountry box="[440,510,493,517]" name="Brazil" pageId="2" pageNumber="463">Brazil</collectingCountry>
:
<collectingRegion box="[525,699,493,517]" country="Brazil" name="Rondonia" pageId="2" pageNumber="463">Rondônia State</collectingRegion>
:
<specimenCode collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">INPA 39605</specimenCode>
,
<specimenCode box="[213,227,525,549]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">3</specimenCode>
,
<quantity box="[238,383,525,549]" metricMagnitude="-2" metricUnit="m" metricValue="6.415" metricValueMax="7.37" metricValueMin="5.46" pageId="2" pageNumber="463" unit="mm" value="64.15" valueMax="73.7" valueMin="54.6">54.6-73.7 mm</quantity>
SL,
<collectingMunicipality box="[432,526,525,549]" pageId="2" pageNumber="463">Mamoré</collectingMunicipality>
, near the
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0E17776BFC8FCF78" box="[625,775,525,549]" country="Brazil" municipality="Mamore" name="Sao Lourenco" pageId="2" pageNumber="463" stateProvince="Rondonia">São Lourenço</location>
community,
<collectingMethod box="[271,365,557,581]" pageId="2" pageNumber="463">trawl net</collectingMethod>
fishing
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482313" collectingDate="2010-05-30" collectionCode="INPA" collectorName="L. H. Rapp Py-Daniel." country="Brazil" latitude="-11.723361" location="rio Karipunas" longLatPrecision="1" longitude="-65.19214" pageId="2" pageNumber="463" specimenCode="INPA 41148, 2, 95.3" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<geoCoordinate box="[457,606,557,581]" degrees="11" direction="south" minutes="43" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="24.10" value="-11.723361">11°4324.10”S</geoCoordinate>
<geoCoordinate box="[612,770,557,581]" degrees="65" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="31.70" value="-65.19214">65°1131.70”W</geoCoordinate>
,
<date box="[136,282,589,614]" pageId="2" pageNumber="463" value="2010-05-30">
<collectingDate box="[136,282,589,614]" pageId="2" pageNumber="463" value="2010-05-30">30 May 2010</collectingDate>
</date>
,
<collectorName box="[295,544,589,613]" pageId="2" pageNumber="463">L. H. Rapp Py-Daniel.</collectorName>
<specimenCode box="[552,687,589,614]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">INPA 41148</specimenCode>
,
<specimenCode box="[700,716,589,613]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">2</specimenCode>
,
<specimenCode box="[729,775,589,613]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">95.3</specimenCode>
and
<quantity box="[184,281,621,645]" metricMagnitude="-2" metricUnit="m" metricValue="9.68" pageId="2" pageNumber="463" unit="mm" value="96.8">96.8 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0D36770BFE60CFD8" box="[336,488,621,645]" country="Brazil" latitude="-11.723361" longLatPrecision="1" longitude="-65.19214" name="rio Karipunas" pageId="2" pageNumber="463" stateProvince="Rondonia">rio Karipunas</location>
near mouth
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482304" collectingDate="2010-10-04" collectionCode="INPA" country="Brazil" latitude="-9.196278" location="rio Karipunas" longLatPrecision="1" longitude="-64.62242" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="INPA 41149, 1" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<geoCoordinate box="[636,774,621,645]" degrees="09" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="46.6" value="-9.196278">09°1146.6”S</geoCoordinate>
<geoCoordinate box="[136,290,653,677]" degrees="64" direction="west" minutes="37" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="20.7" value="-64.62242">64°3720.7”W</geoCoordinate>
,
<date box="[297,409,653,677]" pageId="2" pageNumber="463" value="2010-10-04">
<collectingDate box="[297,409,653,677]" pageId="2" pageNumber="463" value="2010-10-04">4 Oct 2010</collectingDate>
</date>
, NaturaeEnvironmentalCompany technicians.
<specimenCode box="[272,403,685,710]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">INPA 41149</specimenCode>
,
<specimenCode box="[414,427,685,709]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">1</specimenCode>
, 85.0 mm SL,
<collectingMunicipality box="[586,717,685,709]" pageId="2" pageNumber="463">rio Madeira</collectingMunicipality>
, near mouth of
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0C9C77ABFE10CFB8" box="[250,408,717,741]" country="Brazil" latitude="-9.196278" longLatPrecision="1" longitude="-64.62242" municipality="rio Madeira" name="rio Karipunas" pageId="2" pageNumber="463" stateProvince="Rondonia">rio Karipunas</location>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482318" collectingDate="2010-10-30" collectionCode="INPA" collectorName="Naturae Environmental Company" country="Brazil" county="rio Madeira" latitude="-9.20525" location="Jirau" longLatPrecision="1" longitude="-64.619" municipality="Jirau" pageId="2" pageNumber="463" specimenCode="INPA 41150, 28" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<geoCoordinate box="[424,565,717,741]" degrees="09" direction="south" minutes="12" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="18.9" value="-9.20525">09°1218.9”S</geoCoordinate>
<geoCoordinate box="[577,732,717,741]" degrees="64" direction="west" minutes="37" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="08.4" value="-64.619">64°3708.4”W</geoCoordinate>
,
<date pageId="2" pageNumber="463" value="2010-10-30">
<collectingDate pageId="2" pageNumber="463" value="2010-10-30">30 Oct 2010</collectingDate>
</date>
,
<collectorName box="[257,633,749,773]" pageId="2" pageNumber="463">Naturae Environmental Company</collectorName>
technicians.
<specimenCode box="[136,268,781,806]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">INPA 41150</specimenCode>
,
<specimenCode box="[277,306,781,805]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="463">28</specimenCode>
,
<quantity box="[316,475,781,805]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="11.22" metricValueMin="3.28" pageId="2" pageNumber="463" unit="mm" value="72.5" valueMax="112.2" valueMin="32.8">32.8-112.2 mm</quantity>
SL,
<collectingCounty box="[525,655,781,805]" pageId="2" pageNumber="463">rio Madeira</collectingCounty>
, cofferdam at the construction site of
<collectingMunicipality box="[431,484,813,837]" pageId="2" pageNumber="463">Jirau</collectingMunicipality>
hydroelectric power plant (former
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0C81762BFE94CE38" box="[231,284,845,869]" country="Brazil" county="rio Madeira" latitude="-9.20525" longLatPrecision="1" longitude="-64.619" municipality="Jirau" name="Jirau" pageId="2" pageNumber="463" stateProvince="Rondonia">Jirau</location>
rapids)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482308" collectingDate="2011-11-11" collectingDateMax="2011-11-13" collectingDateMin="2011-11-11" collectionCode="MZUSP" collectorName="Naturae Environmental Company" country="Brazil" county="rio Madeira" latitude="-9.254361" location="Jirau" longLatPrecision="1" longitude="-64.64733" municipality="Jirau" pageId="2" pageNumber="463" specimenCode="MZUSP 114009, 3, UFRO-I 6384, 5" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<geoCoordinate box="[392,532,845,869]" degrees="09" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="15.7" value="-9.254361">09°1515.7”S</geoCoordinate>
<geoCoordinate box="[546,699,845,869]" degrees="64" direction="west" minutes="38" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="50.4" value="-64.64733">64°3850.4”W</geoCoordinate>
,
<date pageId="2" pageNumber="463" value="2011-11-11" valueMax="2011-11-13" valueMin="2011-11-11">
<collectingDate pageId="2" pageNumber="463" value="2011-11-11" valueMax="2011-11-13" valueMin="2011-11-11">11-13 Nov 2011</collectingDate>
</date>
,
<collectorName box="[259,634,877,901]" pageId="2" pageNumber="463">Naturae Environmental Company</collectorName>
technicians.
<specimenCode box="[136,310,909,933]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="463">MZUSP 114009</specimenCode>
,
<specimenCode box="[319,333,909,933]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="463">3</specimenCode>
,
<quantity box="[345,492,909,933]" metricMagnitude="-2" metricUnit="m" metricValue="7.234999999999999" metricValueMax="7.62" metricValueMin="6.85" pageId="2" pageNumber="463" unit="mm" value="72.35" valueMax="76.2" valueMin="68.5">68.5-76.2 mm</quantity>
SL;
<specimenCode box="[532,685,909,933]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 6384</specimenCode>
,
<specimenCode box="[695,709,909,933]" collectionCode="UFRO-I" pageId="2" pageNumber="463">5</specimenCode>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="6.575" metricValueMax="8.07" metricValueMin="5.08" pageId="2" pageNumber="463" unit="mm" value="65.75" valueMax="80.7" valueMin="50.8">50.8- 80.7 mm</quantity>
SL, same data as INPA
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3114482301" country="Iceland" location="Bufalo" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 7968, 1" specimenCount="1" typeStatus="paratype">
39605.
<specimenCode box="[589,743,941,965]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 7968</specimenCode>
,
<specimenCode box="[756,769,941,965]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
,
<quantity box="[136,232,973,997]" metricMagnitude="-2" metricUnit="m" metricValue="8.14" pageId="2" pageNumber="463" unit="mm" value="81.4">81.4 mm</quantity>
SL,
<collectingMunicipality box="[291,425,973,997]" pageId="2" pageNumber="463">rio Madeira</collectingMunicipality>
, near
<collectingCountry box="[496,539,973,997]" name="Iceland" pageId="2" pageNumber="463">Ilha</collectingCountry>
do
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0E2D76ABFD1ECEB8" box="[587,662,973,997]" country="Iceland" municipality="rio Madeira" name="Bufalo" pageId="2" pageNumber="463">Búfalo</location>
,
<collectingMethod box="[676,775,973,997]" pageId="2" pageNumber="463">trawl net</collectingMethod>
fishing
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482307" collectingDate="2010-10-23" collectorName="A. Ribeiro." country="Iceland" latitude="-9.147528" location="Santo Antonio" longLatPrecision="1" longitude="-64.542694" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 9974, 1" specimenCount="1" typeStatus="paratype">
<geoCoordinate box="[231,384,1005,1029]" degrees="09" direction="south" minutes="08" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="51.10" value="-9.147528">09°0851.10”S</geoCoordinate>
<geoCoordinate box="[399,564,1005,1029]" degrees="64" direction="west" minutes="32" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="33.70" value="-64.542694">64°3233.70”W</geoCoordinate>
,
<date box="[582,730,1005,1029]" pageId="2" pageNumber="463" value="2010-10-23">
<collectingDate box="[582,730,1005,1029]" pageId="2" pageNumber="463" value="2010-10-23">23 Oct 2010</collectingDate>
</date>
,
<collectorName pageId="2" pageNumber="463">A. Ribeiro.</collectorName>
<specimenCode box="[231,383,1037,1061]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 9974</specimenCode>
,
<specimenCode box="[394,407,1037,1061]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
,
<quantity box="[421,515,1037,1061]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="2" pageNumber="463" unit="mm" value="53.6">53.6 mm</quantity>
SL,
<collectingMunicipality box="[568,699,1037,1061]" pageId="2" pageNumber="463">rio Madeira</collectingMunicipality>
, below
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0CEE714BFEA6C918" box="[136,302,1069,1093]" country="Iceland" latitude="-9.147528" longLatPrecision="1" longitude="-64.542694" municipality="rio Madeira" name="Santo Antonio" pageId="2" pageNumber="463">Santo Antônio</location>
rapids,
<collectingMethod box="[407,511,1069,1093]" pageId="2" pageNumber="463">trawl net</collectingMethod>
fishing
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482305" collectingDate="2011-07-23" collectorName="L. Nogueira." country="Iceland" latitude="-8.776697" location="Santo Antonio" longLatPrecision="1" longitude="-63.923916" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 10050, 1" specimenCount="1" typeStatus="paratype">
<geoCoordinate box="[623,775,1069,1093]" degrees="08" direction="south" minutes="46" orientation="latitude" pageId="2" pageNumber="463" precision="1" seconds="36.11" value="-8.776697">08°4636.11”S</geoCoordinate>
<geoCoordinate box="[136,302,1101,1125]" degrees="63" direction="west" minutes="55" orientation="longitude" pageId="2" pageNumber="463" precision="1" seconds="26.09" value="-63.923916">63°5526.09”W</geoCoordinate>
,
<date box="[315,443,1101,1125]" pageId="2" pageNumber="463" value="2011-07-23">
<collectingDate box="[315,443,1101,1125]" pageId="2" pageNumber="463" value="2011-07-23">23 Jul 2011</collectingDate>
</date>
,
<collectorName box="[456,596,1101,1125]" pageId="2" pageNumber="463">L. Nogueira.</collectorName>
<specimenCode box="[604,771,1101,1125]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 10050</specimenCode>
,
<specimenCode box="[136,149,1133,1157]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
,
<quantity box="[163,258,1133,1157]" metricMagnitude="-2" metricUnit="m" metricValue="9.04" pageId="2" pageNumber="463" unit="mm" value="90.4">90.4 mm</quantity>
SL, same data as INPA
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3114482314" collectingDate="2012-02-14" collectionCode="INPA" collectorName="C. A. Cramer." country="Brazil" latitude="-8.801667" location="Santo Antonio" longLatPrecision="21" longitude="-63.95" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 13039, 1" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
39605.
<specimenCode box="[608,771,1133,1157]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 13039</specimenCode>
,
<specimenCode box="[136,149,1165,1189]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
,
<quantity box="[164,259,1165,1189]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" pageId="2" pageNumber="463" unit="mm" value="49.5">49.5 mm</quantity>
SL, same locality as holotype
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3114482303" collectingDate="2011-12-01" collectionCode="INPA" collectorName="C. A. Cramer." country="Brazil" latitude="-8.801667" location="Santo Antonio" longLatPrecision="21" longitude="-63.95" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 13040, 1" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<date box="[612,738,1165,1189]" pageId="2" pageNumber="463" value="2011-12-01">
<collectingDate box="[612,738,1165,1189]" pageId="2" pageNumber="463" value="2011-12-01">1 Dec 2011</collectingDate>
</date>
,
<collectorName pageId="2" pageNumber="463">C. A. Cramer.</collectorName>
<specimenCode box="[269,437,1197,1221]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 13040</specimenCode>
,
<specimenCode box="[451,464,1197,1221]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
,
<quantity box="[479,573,1197,1221]" metricMagnitude="-2" metricUnit="m" metricValue="5.77" pageId="2" pageNumber="463" unit="mm" value="57.7">57.7 mm</quantity>
SL, same data as UFRO-I
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3114482319" collectingDate="2012-02-14" collectionCode="INPA" collectorName="C. A. Cramer." country="Brazil" latitude="-8.801667" location="Santo Antonio" longLatPrecision="21" longitude="-63.95" municipality="rio Madeira" pageId="2" pageNumber="463" specimenCode="UFRO-I 19646, 1" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
13039.
<specimenCode box="[319,489,1229,1253]" collectionCode="UFRO-I" pageId="2" pageNumber="463">UFRO-I 19646</specimenCode>
,
<specimenCode box="[505,518,1229,1253]" collectionCode="UFRO-I" pageId="2" pageNumber="463">1</specimenCode>
, female,
<quantity box="[626,725,1229,1253]" metricMagnitude="-2" metricUnit="m" metricValue="9.73" pageId="2" pageNumber="463" unit="mm" value="97.3">97.3 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[136,775,1326,1414]" pageId="2" pageNumber="463">
<emphasis bold="true" box="[136,261,1326,1350]" pageId="2" pageNumber="463">Non-types.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3114482317" accessionNumber="ROM 92440" collectingDate="2010-08-16" collectionCode="ROM" collectorName="K. Roach &amp; A. Jackson" country="Peru" latitude="-12.8" location="Rio Tambopata" longLatPrecision="1292" longitude="-69.28333" pageId="2" pageNumber="463" specimenCode="1" specimenCount="1" stateProvince="Madre de Dios">
<specimenCount box="[266,391,1326,1350]" pageId="2" pageNumber="463" type="generic">1 specimen</specimenCount>
.
<collectingCountry box="[400,454,1326,1349]" name="Peru" pageId="2" pageNumber="463">Peru</collectingCountry>
:
<accessionNumber box="[462,597,1326,1350]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM92440" pageId="2" pageNumber="463">
<collectionCode box="[462,521,1326,1350]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="2" pageNumber="463" type="Museum">ROM</collectionCode>
92440
</accessionNumber>
,
<specimenCode box="[606,619,1326,1350]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="2" pageNumber="463" type="Museum">1</specimenCode>
, 117.0 mm SL,
<location LSID="urn:lsid:plazi:treatment:038187DCB901FFEA0FB47729FE6FC99B:8EF76011B902FFEF0CEE7028FEBEC83B" box="[136,310,1358,1382]" country="Peru" latitude="-12.8" longLatPrecision="1292" longitude="-69.28333" name="Rio Tambopata" pageId="2" pageNumber="463" stateProvince="Madre de Dios">Río Tambopata</location>
,
<collectingRegion box="[325,495,1358,1382]" country="Peru" name="Madre de Dios" pageId="2" pageNumber="463">Madre de Dios</collectingRegion>
drainage, approximately
<geoCoordinate box="[136,220,1390,1414]" degrees="12" direction="south" minutes="48" orientation="latitude" pageId="2" pageNumber="463" precision="925" value="-12.8">12°48S</geoCoordinate>
<geoCoordinate box="[227,321,1390,1414]" degrees="69" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="463" precision="925" value="-69.28333">69°17W</geoCoordinate>
,
<date box="[332,469,1390,1414]" pageId="2" pageNumber="463" value="2010-08-16">
<collectingDate box="[332,469,1390,1414]" pageId="2" pageNumber="463" value="2010-08-16">16 Aug 2010</collectingDate>
</date>
,
<collectorName box="[480,581,1390,1414]" pageId="2" pageNumber="463">K. Roach</collectorName>
and
<collectorName box="[633,755,1390,1414]" pageId="2" pageNumber="463">A. Jackson</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="463" type="diagnosis">
<paragraph blockId="2.[136,775,1454,1959]" lastBlockId="2.[820,1460,174,1702]" pageId="2" pageNumber="463">
<emphasis bold="true" box="[136,255,1454,1478]" pageId="2" pageNumber="463">Diagnosis.</emphasis>
The presence of small white dots on the whole body and fins distinguish
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[413,567,1486,1510]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[413,567,1486,1510]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
from all congeners except
<taxonomicName baseAuthorityName="Kanazawa" baseAuthorityYear="1958" box="[215,406,1518,1542]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="albomaculatus">
<emphasis box="[215,406,1518,1542]" italics="true" pageId="2" pageNumber="463">P. albomaculatus</emphasis>
</taxonomicName>
. These dots are frequently faded in preserved specimens (typically, at least the dorsal and caudal fins show some traces of dots).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[570,727,1582,1606]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[570,727,1582,1606]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Kanazawa" baseAuthorityYear="1958" box="[393,587,1614,1638]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="albomaculatus">
<emphasis box="[393,587,1614,1638]" italics="true" pageId="2" pageNumber="463">P. albomaculatus</emphasis>
</taxonomicName>
by having more dots on the trunk (on nearly all plates having one dot each
<emphasis box="[136,165,1679,1702]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
one dot each on less than half of the plates), a longer head-eye length (35.1-41.5% HL [mean 37.8]
<emphasis box="[622,651,1711,1734]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
31.9-37.2% HL [mean 34.4]), a smaller orbital diameter (12.5-16.8% HL [mean 14.3]
<emphasis box="[311,341,1775,1798]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
16.0-19.9% HL [mean 17.7]), and longer rictal barbels (9.7-19.6% HL [mean 13.8]
<emphasis box="[593,623,1807,1830]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
1.6-8.9% HL [mean 4.9]). Only two other
<taxonomicName authorityName="Isbrucker &amp; Schraml" authorityYear="2001" box="[454,569,1838,1862]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="genus">
<emphasis box="[454,569,1838,1862]" italics="true" pageId="2" pageNumber="463">Panaqolus</emphasis>
</taxonomicName>
species (
<taxonomicName baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[673,775,1838,1862]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="dentex">
<emphasis box="[673,775,1838,1862]" italics="true" pageId="2" pageNumber="463">P. dentex</emphasis>
</taxonomicName>
and
<taxonomicName box="[183,264,1870,1894]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="koko">
<emphasis box="[183,264,1870,1894]" italics="true" pageId="2" pageNumber="463">P. koko</emphasis>
</taxonomicName>
) lack alternating dark and light bands on fins.
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[136,298,1902,1926]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[136,298,1902,1926]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[628,735,1902,1926]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="dentex">
<emphasis box="[628,735,1902,1926]" italics="true" pageId="2" pageNumber="463">P. dentex</emphasis>
</taxonomicName>
by lacking lighter colored saddles (
<emphasis box="[503,532,1935,1958]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
three lighter colored saddles between dorsal-fin origin and caudal fin), by having a shorter head-pectoral length (22.4-28.2% SL [mean 25.5]
<emphasis box="[820,849,239,262]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
27.2-38.1% SL [mean 29.2]), a lower caudal peduncle depth (9.3-11.4% SL [mean 10.5]
<emphasis box="[1188,1218,271,294]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
11.0-13.3% SL [mean 12.2]), a smaller orbital diameter (12.5-16.8% HL [mean 14.3]
<emphasis box="[884,913,335,358]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
16.4-22.8% HL [mean 17.8]), and a larger adult body size (more than
<quantity box="[1056,1145,366,390]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="463" unit="mm" value="100.0">100 mm</quantity>
SL
<emphasis box="[1191,1221,367,390]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
less than
<quantity box="[1331,1407,366,390]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="463" unit="mm" value="80.0">80 mm</quantity>
SL).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[820,974,398,422]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[820,974,398,422]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[1117,1197,398,422]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="koko">
<emphasis box="[1117,1197,398,422]" italics="true" pageId="2" pageNumber="463">P. koko</emphasis>
</taxonomicName>
in the shape of the main tooth cusp (round and without or with very small lateral cusp
<emphasis box="[820,849,463,486]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
quadrangular and with strong lateral cusp), a smaller orbital diameter (12.5-16.8% HL
<emphasis box="[1179,1208,495,518]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
18.9-20.8% HL [n=7]), and a larger interorbital width (34.3-39.3% HL
<emphasis box="[1364,1393,527,550]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
29.4- 33.2% HL).
<taxonomicName authorityName="Lujan, Steele and Velasquez" authorityYear="2013" box="[952,1188,558,582]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="albivermis">
<emphasis box="[952,1188,558,582]" italics="true" pageId="2" pageNumber="463">Panaqolus albivermis</emphasis>
</taxonomicName>
typically has alternating light bands and rows of light dots or short lines on its body.
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[820,974,622,646]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[820,974,622,646]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
can be separated from this species by having a shorter dorsal spine (26.2-32.5% SL
<emphasis box="[1257,1286,655,678]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
33.0-36.0% SL [n=4]), a shallower caudal peduncle (9.3-11.4% SL
<emphasis box="[1368,1397,687,710]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
12.2- 14.9% SL [n=4]), and a higher adipose-anal depth (17.9- 22.2% SL
<emphasis box="[942,971,751,774]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
15.8-17.9% SL [n=4]).
<taxonomicName baseAuthorityName="Chockley &amp; Armbruster" baseAuthorityYear="2002" box="[1235,1452,750,774]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="changae">
<emphasis box="[1235,1452,750,774]" italics="true" pageId="2" pageNumber="463">Panaqolus changae</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[820,938,783,806]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="gnomus">
<emphasis box="[820,938,783,806]" italics="true" pageId="2" pageNumber="463">P. gnomus</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[956,1071,783,806]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="maccus">
<emphasis box="[956,1071,783,806]" italics="true" pageId="2" pageNumber="463">P. maccus</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[1089,1232,783,806]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nocturnus">
<emphasis box="[1089,1232,783,806]" italics="true" pageId="2" pageNumber="463">P. nocturnus</emphasis>
</taxonomicName>
, and
<taxonomicName baseAuthorityName="La Monte" baseAuthorityYear="1935" box="[1301,1458,783,806]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="purusiensis">
<emphasis box="[1301,1458,783,806]" italics="true" pageId="2" pageNumber="463">P. purusiensis</emphasis>
</taxonomicName>
have alternating dark and light bands on the body (except
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[820,961,847,870]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nocturnus">
<emphasis box="[820,961,847,870]" italics="true" pageId="2" pageNumber="463">P. nocturnus</emphasis>
</taxonomicName>
and adult
<taxonomicName baseAuthorityName="La Monte" baseAuthorityYear="1935" box="[1083,1238,847,870]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="purusiensis">
<emphasis box="[1083,1238,847,870]" italics="true" pageId="2" pageNumber="463">P. purusiensis</emphasis>
</taxonomicName>
) and fins (
<emphasis box="[1361,1391,847,870]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
never showing bands on body or fins in
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1208,1272,879,902]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1208,1272,879,902]" italics="true" pageId="2" pageNumber="463">P. nix</emphasis>
</taxonomicName>
). Further,
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1394,1459,879,902]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1394,1459,879,902]" italics="true" pageId="2" pageNumber="463">P. nix</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Chockley &amp; Armbruster" baseAuthorityYear="2002" box="[1105,1224,910,934]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="changae">
<emphasis box="[1105,1224,910,934]" italics="true" pageId="2" pageNumber="463">P. changae</emphasis>
</taxonomicName>
by a narrower ventral cleithral width (29.0-33.8% SL [mean 31.0]
<emphasis box="[1321,1350,943,966]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
33.4-37.4 [mean 34.8), a shorter head-pectoral length (22.4-28.2% SL [mean 25.5]
<emphasis box="[1003,1033,1007,1030]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
28.0-31.1% SL [mean 29.9), a smaller orbital diameter (12.5-16.8% HL [mean 14.3]
<emphasis box="[1303,1332,1039,1062]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
16.2-20.9% HL [mean 18.2]), and a larger adult body size (more than
<quantity box="[820,909,1102,1126]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="463" unit="mm" value="100.0">100 mm</quantity>
SL
<emphasis box="[954,983,1103,1126]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
less than
<quantity box="[1091,1167,1102,1126]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="463" unit="mm" value="90.0">90 mm</quantity>
SL).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1226,1381,1102,1126]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1226,1381,1102,1126]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
differs from
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[885,1004,1135,1158]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="gnomus">
<emphasis box="[885,1004,1135,1158]" italics="true" pageId="2" pageNumber="463">P. gnomus</emphasis>
</taxonomicName>
by a lower dorsal-pectoral depth (26.9- 30.3% SL [mean 28.6]
<emphasis box="[1080,1110,1167,1190]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
30.3-36.8% SL [mean 32.9]), a smaller interorbital width (34.4-39.3% HL
<emphasis box="[1300,1329,1199,1222]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
39.7-44.7% HL), and a larger adult body size (more than
<quantity box="[1299,1386,1230,1254]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="463" unit="mm" value="100.0">100 mm</quantity>
SL
<emphasis box="[1430,1459,1231,1254]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
less than
<quantity box="[922,998,1262,1286]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="463" unit="mm" value="80.0">80 mm</quantity>
SL).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1056,1212,1262,1286]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1056,1212,1262,1286]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
can be separated from
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[820,936,1295,1318]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="maccus">
<emphasis box="[820,936,1295,1318]" italics="true" pageId="2" pageNumber="463">P. maccus</emphasis>
</taxonomicName>
by a shorter predorsal length (40.1-44.5% SL [mean 42.3]
<emphasis box="[954,983,1327,1350]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
44.3-49.2% SL [mean 46.0), a shorter headpectoral length (22.4-28.2% SL [mean 25.5]
<emphasis box="[1302,1332,1359,1382]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
27.2-44.1% SL [mean 30.8]), and a larger adult body size (more than
<quantity box="[820,912,1422,1446]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="463" unit="mm" value="100.0">100 mm</quantity>
SL
<emphasis box="[963,992,1423,1446]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
less than
<quantity box="[1109,1188,1422,1446]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="463" unit="mm" value="90.0">90 mm</quantity>
SL).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1253,1411,1422,1446]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1253,1411,1422,1446]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Schaefer &amp; Stewart" baseAuthorityYear="1993" box="[1065,1202,1455,1478]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nocturnus">
<emphasis box="[1065,1202,1455,1478]" italics="true" pageId="2" pageNumber="463">P. nocturnus</emphasis>
</taxonomicName>
by the angle of dentary tooth rows (less than 50° to nearly parallel
<emphasis box="[1270,1299,1487,1510]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
approximately 70°).
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[878,1032,1518,1542]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="nix">
<emphasis box="[878,1032,1518,1542]" italics="true" pageId="2" pageNumber="463">Panaqolus nix</emphasis>
</taxonomicName>
differs from
<taxonomicName baseAuthorityName="La Monte" baseAuthorityYear="1935" box="[1175,1328,1519,1542]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="463" phylum="Chordata" rank="species" species="purusiensis">
<emphasis box="[1175,1328,1519,1542]" italics="true" pageId="2" pageNumber="463">P. purusiensis</emphasis>
</taxonomicName>
by having a lower dorsal-pectoral depth (26.9-30.3% SL [mean 28.6]
<emphasis box="[1430,1459,1551,1574]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
29.8-32.9% SL [mean 30.9;]), a lower caudal peduncle depth (9.3-11.4% SL [mean 10.6]
<emphasis box="[1115,1144,1615,1638]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
10.9-14.0% SL [mean 12.1]), and a smaller mouth width (34.4-46.4% HL [mean 39.9]
<emphasis box="[1430,1459,1647,1670]" italics="true" pageId="2" pageNumber="463">vs.</emphasis>
41.6-50.3% HL [mean 44.1]).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="465" pageId="2" pageNumber="463" type="description">
<paragraph blockId="2.[820,1459,1742,1959]" lastBlockId="3.[813,1451,174,486]" lastPageId="3" lastPageNumber="464" pageId="2" pageNumber="463">
<emphasis bold="true" box="[820,961,1742,1766]" pageId="2" pageNumber="463">Description.</emphasis>
Counts and proportional measurements in
<tableCitation box="[820,903,1775,1799]" captionStart="Table 1" captionStartId="3.[129,192,533,557]" captionTargetPageId="3" captionText="Table 1. Selected morphometrics of Panaqolus nix. Values are given as percents of standard length or head length. SD" httpUri="http://table.plazi.org/id/DF576642B903FFEE0CE77773FA23CF70" pageId="2" pageNumber="463" tableUuid="DF576642B903FFEE0CE77773FA23CF70">Table 1</tableCitation>
. Medium-sized loricariid with standard length of measured specimens up to
<quantity box="[1118,1225,1807,1831]" metricMagnitude="-1" metricUnit="m" metricValue="1.122" pageId="2" pageNumber="463" unit="mm" value="112.2">112.2 mm</quantity>
SL. Dorsal profile of head and snout strongly convex from snout tip to posterior tip of supraoccipital, straight and posteroventrally slanted between dorsal-fin origin and adipose-fin origin, gently concave through caudal peduncle to posterior tip of procurrent caudal-fin ray. Dorsal orbit margin only slightly raised, forming gentle ridge, narrowing anteriorly, from anterior orbit margin to area lateral to nares. Dorsal surface of trunk transversely flattened from dorsal-fin origin to adipose-fin base. Ventral profile of head and body flat from oral disk to anal-fin origin. Caudal peduncle oval in crosssection. Greatest body depth at dorsal-fin origin. Pectoralfin origin just posterior to orbit; pelvic-fin origin at vertical through origin of second dorsal-fin ray; anal-fin origin slightly posterior to vertical through origin of last dorsal- fin ray. Adipose fin with well-ossified leading spine bearing odontodes.
</paragraph>
<paragraph blockId="3.[813,1451,174,486]" pageId="3" pageNumber="464">Head and body covered by odontodes of uniform size and distribution. Enlarged odontodes on anterodorsal border of pectoral-fin spine. Cheek odontodes hypertrophied with anteriorly curved hooks on the tips; longest odontode extending to posterior cleithrum margin. Interorbital space flat or slightly convex. Eye dorsolaterally placed; orbit diameter 12.5-16.8% HL. Iris operculum present. Nares small and ovoid, slightly longer than wide.</paragraph>
<caption ID-Table-UUID="DF576642B903FFEE0CE77773FA23CF70" box="[129,1451,532,557]" httpUri="http://table.plazi.org/id/DF576642B903FFEE0CE77773FA23CF70" pageId="3" pageNumber="464" startId="3.[129,192,533,557]" targetBox="[144,1393,645,1947]" targetIsTable="true" targetPageId="3">
<paragraph blockId="3.[128,1451,532,621]" box="[129,1451,532,557]" pageId="3" pageNumber="464">
<emphasis bold="true" box="[129,221,532,557]" pageId="3" pageNumber="464">Table 1.</emphasis>
Selected morphometrics of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[534,692,533,557]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="464" phylum="Chordata" rank="species" species="nix">
<emphasis box="[534,692,533,557]" italics="true" pageId="3" pageNumber="464">Panaqolus nix</emphasis>
</taxonomicName>
. Values are given as percents of standard length or head length. SD
</paragraph>
</caption>
<paragraph blockId="3.[128,1451,532,621]" box="[128,1451,565,589]" pageId="3" pageNumber="464">
= standard deviation, n = number of specimens, H =
<typeStatus box="[718,816,565,589]" pageId="3" pageNumber="464">holotype</typeStatus>
. Interlandmarks (ILM) are the two points between which
</paragraph>
<paragraph blockId="3.[128,1451,532,621]" box="[128,683,597,621]" pageId="3" pageNumber="464">
measurements were taken (from Luj
<emphasis box="[518,598,597,621]" italics="true" pageId="3" pageNumber="464">an et al</emphasis>
., 2010).
</paragraph>
<paragraph pageId="3" pageNumber="464">
<table box="[144,1393,645,1947]" gridcols="6" gridrows="36" pageId="3" pageNumber="464">
<tr box="[144,1393,645,664]" gridrow="0" pageId="3" pageNumber="464">
<th box="[144,199,645,664]" gridcol="0" gridrow="0" pageId="3" pageNumber="464">ILM</th>
<th box="[268,590,645,664]" gridcol="1" gridrow="0" pageId="3" pageNumber="464">Measurement</th>
<th box="[669,911,645,664]" gridcol="2" gridrow="0" pageId="3" pageNumber="464">H</th>
<th box="[961,982,645,664]" gridcol="3" gridrow="0" pageId="3" pageNumber="464">n</th>
<th box="[1094,1197,645,664]" gridcol="4" gridrow="0" pageId="3" pageNumber="464">range</th>
<th box="[1306,1393,645,664]" gridcol="5" gridrow="0" pageId="3" pageNumber="464">mean±SD</th>
</tr>
<tr box="[144,1393,682,701]" gridrow="1" pageId="3" pageNumber="464">
<th box="[144,199,682,701]" gridcol="0" gridrow="1" pageId="3" pageNumber="464">1-20</th>
<td box="[268,590,682,701]" gridcol="1" gridrow="1" pageId="3" pageNumber="464">Standard length (mm)</td>
<td box="[669,911,682,701]" gridcol="2" gridrow="1" pageId="3" pageNumber="464">110.1</td>
<td box="[961,982,682,701]" gridcol="3" gridrow="1" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,682,701]" gridcol="4" gridrow="1" pageId="3" pageNumber="464">32.84-112.2</td>
<td box="[1306,1393,682,701]" gridcol="5" gridrow="1" pageId="3" pageNumber="464">82.44</td>
</tr>
<tr box="[144,1393,718,738]" gridrow="2" pageId="3" pageNumber="464" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[669,911,718,738]" gridcol="2" gridrow="2" pageId="3" pageNumber="464">Percents of Standard Length</td>
</tr>
<tr box="[144,1393,755,774]" gridrow="3" pageId="3" pageNumber="464">
<th box="[144,199,755,774]" gridcol="0" gridrow="3" pageId="3" pageNumber="464">1-10</th>
<td box="[268,590,755,774]" gridcol="1" gridrow="3" pageId="3" pageNumber="464">Predorsal length</td>
<td box="[669,911,755,774]" gridcol="2" gridrow="3" pageId="3" pageNumber="464">43.1</td>
<td box="[961,982,755,774]" gridcol="3" gridrow="3" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,755,774]" gridcol="4" gridrow="3" pageId="3" pageNumber="464">40.1-44.5</td>
<td box="[1306,1393,755,774]" gridcol="5" gridrow="3" pageId="3" pageNumber="464">42.3±1.0</td>
</tr>
<tr box="[144,1393,791,811]" gridrow="4" pageId="3" pageNumber="464">
<th box="[144,199,791,811]" gridcol="0" gridrow="4" pageId="3" pageNumber="464">1-7</th>
<td box="[268,590,791,811]" gridcol="1" gridrow="4" pageId="3" pageNumber="464">Head length</td>
<td box="[669,911,791,811]" gridcol="2" gridrow="4" pageId="3" pageNumber="464">35.7</td>
<td box="[961,982,791,811]" gridcol="3" gridrow="4" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,791,811]" gridcol="4" gridrow="4" pageId="3" pageNumber="464">33.2-39.3</td>
<td box="[1306,1393,791,811]" gridcol="5" gridrow="4" pageId="3" pageNumber="464">35.2±1.3</td>
</tr>
<tr box="[144,1393,828,847]" gridrow="5" pageId="3" pageNumber="464">
<th box="[144,199,828,847]" gridcol="0" gridrow="5" pageId="3" pageNumber="464">8-9</th>
<td box="[268,590,828,847]" gridcol="1" gridrow="5" pageId="3" pageNumber="464">Cleithral width</td>
<td box="[669,911,828,847]" gridcol="2" gridrow="5" pageId="3" pageNumber="464">31.5</td>
<td box="[961,982,828,847]" gridcol="3" gridrow="5" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,828,847]" gridcol="4" gridrow="5" pageId="3" pageNumber="464">30.8-34.3</td>
<td box="[1306,1393,828,847]" gridcol="5" gridrow="5" pageId="3" pageNumber="464">32.3±0.9</td>
</tr>
<tr box="[144,1393,865,884]" gridrow="6" pageId="3" pageNumber="464">
<th box="[144,199,865,884]" gridcol="0" gridrow="6" pageId="3" pageNumber="464">8'-9'</th>
<td box="[268,590,865,884]" gridcol="1" gridrow="6" pageId="3" pageNumber="464">Cleithral width ventral</td>
<td box="[669,911,865,884]" gridcol="2" gridrow="6" pageId="3" pageNumber="464">29.5</td>
<td box="[961,982,865,884]" gridcol="3" gridrow="6" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,865,884]" gridcol="4" gridrow="6" pageId="3" pageNumber="464">29.0-33.8</td>
<td box="[1306,1393,865,884]" gridcol="5" gridrow="6" pageId="3" pageNumber="464">31.0±1.2</td>
</tr>
<tr box="[144,1393,901,921]" gridrow="7" pageId="3" pageNumber="464">
<th box="[144,199,901,921]" gridcol="0" gridrow="7" pageId="3" pageNumber="464">1-12</th>
<td box="[268,590,901,921]" gridcol="1" gridrow="7" pageId="3" pageNumber="464">Head-pectoral length</td>
<td box="[669,911,901,921]" gridcol="2" gridrow="7" pageId="3" pageNumber="464">27.0</td>
<td box="[961,982,901,921]" gridcol="3" gridrow="7" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,901,921]" gridcol="4" gridrow="7" pageId="3" pageNumber="464">22.4-28.2</td>
<td box="[1306,1393,901,921]" gridcol="5" gridrow="7" pageId="3" pageNumber="464">25.5±1.3</td>
</tr>
<tr box="[144,1393,938,957]" gridrow="8" pageId="3" pageNumber="464">
<th box="[144,199,938,957]" gridcol="0" gridrow="8" pageId="3" pageNumber="464">12-13</th>
<td box="[268,590,938,957]" gridcol="1" gridrow="8" pageId="3" pageNumber="464">Thorax length</td>
<td box="[669,911,938,957]" gridcol="2" gridrow="8" pageId="3" pageNumber="464">25.3</td>
<td box="[961,982,938,957]" gridcol="3" gridrow="8" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,938,957]" gridcol="4" gridrow="8" pageId="3" pageNumber="464">22.7-30.7</td>
<td box="[1306,1393,938,957]" gridcol="5" gridrow="8" pageId="3" pageNumber="464">26.4±2.0</td>
</tr>
<tr box="[144,1393,975,994]" gridrow="9" pageId="3" pageNumber="464">
<th box="[144,199,975,994]" gridcol="0" gridrow="9" pageId="3" pageNumber="464">12-29</th>
<td box="[268,590,975,994]" gridcol="1" gridrow="9" pageId="3" pageNumber="464">Pectoral-spine length</td>
<td box="[669,911,975,994]" gridcol="2" gridrow="9" pageId="3" pageNumber="464">31.1</td>
<td box="[961,982,975,994]" gridcol="3" gridrow="9" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,975,994]" gridcol="4" gridrow="9" pageId="3" pageNumber="464">29.5-34.9</td>
<td box="[1306,1393,975,994]" gridcol="5" gridrow="9" pageId="3" pageNumber="464">32.0±1.4</td>
</tr>
<tr box="[144,1393,1011,1031]" gridrow="10" pageId="3" pageNumber="464">
<th box="[144,199,1011,1031]" gridcol="0" gridrow="10" pageId="3" pageNumber="464">13-14</th>
<td box="[268,590,1011,1031]" gridcol="1" gridrow="10" pageId="3" pageNumber="464">Abdominal length</td>
<td box="[669,911,1011,1031]" gridcol="2" gridrow="10" pageId="3" pageNumber="464">23.8</td>
<td box="[961,982,1011,1031]" gridcol="3" gridrow="10" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1011,1031]" gridcol="4" gridrow="10" pageId="3" pageNumber="464">20.5-24.3</td>
<td box="[1306,1393,1011,1031]" gridcol="5" gridrow="10" pageId="3" pageNumber="464">22.4±0.9</td>
</tr>
<tr box="[144,1393,1048,1067]" gridrow="11" pageId="3" pageNumber="464">
<th box="[144,199,1048,1067]" gridcol="0" gridrow="11" pageId="3" pageNumber="464">13-30</th>
<td box="[268,590,1048,1067]" gridcol="1" gridrow="11" pageId="3" pageNumber="464">Pelvic-spine length</td>
<td box="[669,911,1048,1067]" gridcol="2" gridrow="11" pageId="3" pageNumber="464">27.8</td>
<td box="[961,982,1048,1067]" gridcol="3" gridrow="11" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1048,1067]" gridcol="4" gridrow="11" pageId="3" pageNumber="464">25.7-31.2</td>
<td box="[1306,1393,1048,1067]" gridcol="5" gridrow="11" pageId="3" pageNumber="464">28.1±1.1</td>
</tr>
<tr box="[144,1393,1085,1104]" gridrow="12" pageId="3" pageNumber="464">
<th box="[144,199,1085,1104]" gridcol="0" gridrow="12" pageId="3" pageNumber="464">13-13'</th>
<td box="[268,590,1085,1104]" gridcol="1" gridrow="12" pageId="3" pageNumber="464">Pelvic girdle width</td>
<td box="[669,911,1085,1104]" gridcol="2" gridrow="12" pageId="3" pageNumber="464">20.2</td>
<td box="[961,982,1085,1104]" gridcol="3" gridrow="12" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1085,1104]" gridcol="4" gridrow="12" pageId="3" pageNumber="464">19.4-22.7</td>
<td box="[1306,1393,1085,1104]" gridcol="5" gridrow="12" pageId="3" pageNumber="464">20.9±0.7</td>
</tr>
<tr box="[144,1393,1121,1141]" gridrow="13" pageId="3" pageNumber="464">
<th box="[144,199,1121,1141]" gridcol="0" gridrow="13" pageId="3" pageNumber="464">14-15</th>
<td box="[268,590,1121,1141]" gridcol="1" gridrow="13" pageId="3" pageNumber="464">Postanal length</td>
<td box="[669,911,1121,1141]" gridcol="2" gridrow="13" pageId="3" pageNumber="464">34.8</td>
<td box="[961,982,1121,1141]" gridcol="3" gridrow="13" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1121,1141]" gridcol="4" gridrow="13" pageId="3" pageNumber="464">31.7-37.6</td>
<td box="[1306,1393,1121,1141]" gridcol="5" gridrow="13" pageId="3" pageNumber="464">34.9±1.3</td>
</tr>
<tr box="[144,1393,1158,1177]" gridrow="14" pageId="3" pageNumber="464">
<th box="[144,199,1158,1177]" gridcol="0" gridrow="14" pageId="3" pageNumber="464">14-31</th>
<td box="[268,590,1158,1177]" gridcol="1" gridrow="14" pageId="3" pageNumber="464">Anal-fin spine length</td>
<td box="[669,911,1158,1177]" gridcol="2" gridrow="14" pageId="3" pageNumber="464">18.3</td>
<td box="[961,982,1158,1177]" gridcol="3" gridrow="14" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1158,1177]" gridcol="4" gridrow="14" pageId="3" pageNumber="464">12.8-18.3</td>
<td box="[1306,1393,1158,1177]" gridcol="5" gridrow="14" pageId="3" pageNumber="464">15.3±1.2</td>
</tr>
<tr box="[144,1393,1195,1214]" gridrow="15" pageId="3" pageNumber="464">
<th box="[144,199,1195,1214]" gridcol="0" gridrow="15" pageId="3" pageNumber="464">10-12</th>
<td box="[268,590,1195,1214]" gridcol="1" gridrow="15" pageId="3" pageNumber="464">Dorsal-pectoral depth</td>
<td box="[669,911,1195,1214]" gridcol="2" gridrow="15" pageId="3" pageNumber="464">29.7</td>
<td box="[961,982,1195,1214]" gridcol="3" gridrow="15" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1195,1214]" gridcol="4" gridrow="15" pageId="3" pageNumber="464">26.9-30.3</td>
<td box="[1306,1393,1195,1214]" gridcol="5" gridrow="15" pageId="3" pageNumber="464">28.6±1.0</td>
</tr>
<tr box="[144,1393,1231,1250]" gridrow="16" pageId="3" pageNumber="464">
<th box="[144,199,1231,1250]" gridcol="0" gridrow="16" pageId="3" pageNumber="464">10-11</th>
<td box="[268,590,1231,1250]" gridcol="1" gridrow="16" pageId="3" pageNumber="464">Dorsal-spine length</td>
<td box="[669,911,1231,1250]" gridcol="2" gridrow="16" pageId="3" pageNumber="464">27.5</td>
<td box="[961,982,1231,1250]" gridcol="3" gridrow="16" pageId="3" pageNumber="464">39</td>
<td box="[1094,1197,1231,1250]" gridcol="4" gridrow="16" pageId="3" pageNumber="464">26.2-32.5</td>
<td box="[1306,1393,1231,1250]" gridcol="5" gridrow="16" pageId="3" pageNumber="464">29.5±1.5</td>
</tr>
<tr box="[144,1393,1268,1287]" gridrow="17" pageId="3" pageNumber="464">
<th box="[144,199,1268,1287]" gridcol="0" gridrow="17" pageId="3" pageNumber="464">10-16</th>
<td box="[268,590,1268,1287]" gridcol="1" gridrow="17" pageId="3" pageNumber="464">Dorsal-fin base length</td>
<td box="[669,911,1268,1287]" gridcol="2" gridrow="17" pageId="3" pageNumber="464">27.5</td>
<td box="[961,982,1268,1287]" gridcol="3" gridrow="17" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1268,1287]" gridcol="4" gridrow="17" pageId="3" pageNumber="464">20.5-28.8</td>
<td box="[1306,1393,1268,1287]" gridcol="5" gridrow="17" pageId="3" pageNumber="464">25.3±1.3</td>
</tr>
<tr box="[144,1393,1305,1324]" gridrow="18" pageId="3" pageNumber="464">
<th box="[144,199,1305,1324]" gridcol="0" gridrow="18" pageId="3" pageNumber="464">16-17</th>
<td box="[268,590,1305,1324]" gridcol="1" gridrow="18" pageId="3" pageNumber="464">Dorsal-adipose distance</td>
<td box="[669,911,1305,1324]" gridcol="2" gridrow="18" pageId="3" pageNumber="464">17.7</td>
<td box="[961,982,1305,1324]" gridcol="3" gridrow="18" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1305,1324]" gridcol="4" gridrow="18" pageId="3" pageNumber="464">13.7-19.4</td>
<td box="[1306,1393,1305,1324]" gridcol="5" gridrow="18" pageId="3" pageNumber="464">17.2±1.2</td>
</tr>
<tr box="[144,1393,1341,1360]" gridrow="19" pageId="3" pageNumber="464">
<th box="[144,199,1341,1360]" gridcol="0" gridrow="19" pageId="3" pageNumber="464">17-18</th>
<td box="[268,590,1341,1360]" gridcol="1" gridrow="19" pageId="3" pageNumber="464">Adipose-spine length</td>
<td box="[669,911,1341,1360]" gridcol="2" gridrow="19" pageId="3" pageNumber="464">5.7</td>
<td box="[961,982,1341,1360]" gridcol="3" gridrow="19" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1341,1360]" gridcol="4" gridrow="19" pageId="3" pageNumber="464">5.7-10.0</td>
<td box="[1306,1393,1341,1360]" gridcol="5" gridrow="19" pageId="3" pageNumber="464">8.2±0.7</td>
</tr>
<tr box="[144,1393,1378,1397]" gridrow="20" pageId="3" pageNumber="464">
<th box="[144,199,1378,1397]" gridcol="0" gridrow="20" pageId="3" pageNumber="464">17-19</th>
<td box="[268,590,1378,1397]" gridcol="1" gridrow="20" pageId="3" pageNumber="464">Adipose-upper caudal distance</td>
<td box="[669,911,1378,1397]" gridcol="2" gridrow="20" pageId="3" pageNumber="464">12.0</td>
<td box="[961,982,1378,1397]" gridcol="3" gridrow="20" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1378,1397]" gridcol="4" gridrow="20" pageId="3" pageNumber="464">12.0-20.7</td>
<td box="[1306,1393,1378,1397]" gridcol="5" gridrow="20" pageId="3" pageNumber="464">15.8±1.4</td>
</tr>
<tr box="[144,1393,1415,1434]" gridrow="21" pageId="3" pageNumber="464">
<th box="[144,199,1415,1434]" gridcol="0" gridrow="21" pageId="3" pageNumber="464">15-19</th>
<td box="[268,590,1415,1434]" gridcol="1" gridrow="21" pageId="3" pageNumber="464">Caudal peduncle depth</td>
<td box="[669,911,1415,1434]" gridcol="2" gridrow="21" pageId="3" pageNumber="464">10.8</td>
<td box="[961,982,1415,1434]" gridcol="3" gridrow="21" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1415,1434]" gridcol="4" gridrow="21" pageId="3" pageNumber="464">9.3-11.4</td>
<td box="[1306,1393,1415,1434]" gridcol="5" gridrow="21" pageId="3" pageNumber="464">10.6±0.4</td>
</tr>
<tr box="[144,1393,1451,1471]" gridrow="22" pageId="3" pageNumber="464">
<th box="[144,199,1451,1471]" gridcol="0" gridrow="22" pageId="3" pageNumber="464">20-32</th>
<td box="[268,590,1451,1471]" gridcol="1" gridrow="22" pageId="3" pageNumber="464">Caudal peduncle-middle caudal ray</td>
<td box="[669,911,1451,1471]" gridcol="2" gridrow="22" pageId="3" pageNumber="464">23.5</td>
<td box="[961,982,1451,1471]" gridcol="3" gridrow="22" pageId="3" pageNumber="464">42</td>
<td box="[1094,1197,1451,1471]" gridcol="4" gridrow="22" pageId="3" pageNumber="464">9.5-26.0</td>
<td box="[1306,1393,1451,1471]" gridcol="5" gridrow="22" pageId="3" pageNumber="464">17.7±4.1</td>
</tr>
<tr box="[144,1393,1488,1507]" gridrow="23" pageId="3" pageNumber="464">
<th box="[144,199,1488,1507]" gridcol="0" gridrow="23" pageId="3" pageNumber="464">20-33</th>
<td box="[268,590,1488,1507]" gridcol="1" gridrow="23" pageId="3" pageNumber="464">Caudal peduncle-dorsal caudal spine</td>
<td box="[669,911,1488,1507]" gridcol="2" gridrow="23" pageId="3" pageNumber="464">47.0</td>
<td box="[961,982,1488,1507]" gridcol="3" gridrow="23" pageId="3" pageNumber="464">17</td>
<td box="[1094,1197,1488,1507]" gridcol="4" gridrow="23" pageId="3" pageNumber="464">33.7-81.7</td>
<td box="[1306,1393,1488,1507]" gridcol="5" gridrow="23" pageId="3" pageNumber="464">51.8±13.0</td>
</tr>
<tr box="[144,1393,1525,1544]" gridrow="24" pageId="3" pageNumber="464">
<th box="[144,199,1525,1544]" gridcol="0" gridrow="24" pageId="3" pageNumber="464">15-17</th>
<td box="[268,590,1525,1544]" gridcol="1" gridrow="24" pageId="3" pageNumber="464">Adipose-lower caudal depth</td>
<td box="[669,911,1525,1544]" gridcol="2" gridrow="24" pageId="3" pageNumber="464">20.5</td>
<td box="[961,982,1525,1544]" gridcol="3" gridrow="24" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1525,1544]" gridcol="4" gridrow="24" pageId="3" pageNumber="464">20.5-25.2</td>
<td box="[1306,1393,1525,1544]" gridcol="5" gridrow="24" pageId="3" pageNumber="464">22.6±1.1</td>
</tr>
<tr box="[144,1393,1561,1581]" gridrow="25" pageId="3" pageNumber="464" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[669,911,1561,1581]" gridcol="2" gridrow="25" pageId="3" pageNumber="464">Percents of Head Length</td>
</tr>
<tr box="[144,1393,1598,1617]" gridrow="26" pageId="3" pageNumber="464">
<th box="[144,199,1598,1617]" gridcol="0" gridrow="26" pageId="3" pageNumber="464">5-7</th>
<td box="[268,590,1598,1617]" gridcol="1" gridrow="26" pageId="3" pageNumber="464">Head-eye length</td>
<td box="[669,911,1598,1617]" gridcol="2" gridrow="26" pageId="3" pageNumber="464">35.6</td>
<td box="[961,982,1598,1617]" gridcol="3" gridrow="26" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1598,1617]" gridcol="4" gridrow="26" pageId="3" pageNumber="464">35.2-41.5</td>
<td box="[1306,1393,1598,1617]" gridcol="5" gridrow="26" pageId="3" pageNumber="464">37.8±1.3</td>
</tr>
<tr box="[144,1393,1634,1654]" gridrow="27" pageId="3" pageNumber="464">
<th box="[144,199,1634,1654]" gridcol="0" gridrow="27" pageId="3" pageNumber="464">4-5</th>
<td box="[268,590,1634,1654]" gridcol="1" gridrow="27" pageId="3" pageNumber="464">Orbit diameter</td>
<td box="[669,911,1634,1654]" gridcol="2" gridrow="27" pageId="3" pageNumber="464">13.5</td>
<td box="[961,982,1634,1654]" gridcol="3" gridrow="27" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1634,1654]" gridcol="4" gridrow="27" pageId="3" pageNumber="464">12.5-16.8</td>
<td box="[1306,1393,1634,1654]" gridcol="5" gridrow="27" pageId="3" pageNumber="464">14.3±1.0</td>
</tr>
<tr box="[144,1393,1671,1690]" gridrow="28" pageId="3" pageNumber="464">
<th box="[144,199,1671,1690]" gridcol="0" gridrow="28" pageId="3" pageNumber="464">1-4</th>
<td box="[268,590,1671,1690]" gridcol="1" gridrow="28" pageId="3" pageNumber="464">Snout length</td>
<td box="[669,911,1671,1690]" gridcol="2" gridrow="28" pageId="3" pageNumber="464">61.0</td>
<td box="[961,982,1671,1690]" gridcol="3" gridrow="28" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1671,1690]" gridcol="4" gridrow="28" pageId="3" pageNumber="464">49.7-63.7</td>
<td box="[1306,1393,1671,1690]" gridcol="5" gridrow="28" pageId="3" pageNumber="464">59.5±2.2</td>
</tr>
<tr box="[144,1393,1708,1727]" gridrow="29" pageId="3" pageNumber="464">
<th box="[144,199,1708,1727]" gridcol="0" gridrow="29" pageId="3" pageNumber="464">2-3</th>
<td box="[268,590,1708,1727]" gridcol="1" gridrow="29" pageId="3" pageNumber="464">Internares width</td>
<td box="[669,911,1708,1727]" gridcol="2" gridrow="29" pageId="3" pageNumber="464">11.8</td>
<td box="[961,982,1708,1727]" gridcol="3" gridrow="29" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1708,1727]" gridcol="4" gridrow="29" pageId="3" pageNumber="464">10.5-14.7</td>
<td box="[1306,1393,1708,1727]" gridcol="5" gridrow="29" pageId="3" pageNumber="464">12.2±0.9</td>
</tr>
<tr box="[144,1393,1744,1764]" gridrow="30" pageId="3" pageNumber="464">
<th box="[144,199,1744,1764]" gridcol="0" gridrow="30" pageId="3" pageNumber="464">5-6</th>
<td box="[268,590,1744,1764]" gridcol="1" gridrow="30" pageId="3" pageNumber="464">Interorbital width</td>
<td box="[669,911,1744,1764]" gridcol="2" gridrow="30" pageId="3" pageNumber="464">51.8</td>
<td box="[961,982,1744,1764]" gridcol="3" gridrow="30" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1744,1764]" gridcol="4" gridrow="30" pageId="3" pageNumber="464">46.5-54.6</td>
<td box="[1306,1393,1744,1764]" gridcol="5" gridrow="30" pageId="3" pageNumber="464">49.6±1.6</td>
</tr>
<tr box="[144,1393,1781,1800]" gridrow="31" pageId="3" pageNumber="464">
<th box="[144,199,1781,1800]" gridcol="0" gridrow="31" pageId="3" pageNumber="464">5'-6'</th>
<td box="[268,590,1781,1800]" gridcol="1" gridrow="31" pageId="3" pageNumber="464">Dorsal interorbital width</td>
<td box="[669,911,1781,1800]" gridcol="2" gridrow="31" pageId="3" pageNumber="464">35.0</td>
<td box="[961,982,1781,1800]" gridcol="3" gridrow="31" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1781,1800]" gridcol="4" gridrow="31" pageId="3" pageNumber="464">34.4-39.3</td>
<td box="[1306,1393,1781,1800]" gridcol="5" gridrow="31" pageId="3" pageNumber="464">36.7±1.2</td>
</tr>
<tr box="[144,1393,1818,1837]" gridrow="32" pageId="3" pageNumber="464">
<th box="[144,199,1818,1837]" gridcol="0" gridrow="32" pageId="3" pageNumber="464">7-12</th>
<td box="[268,590,1818,1837]" gridcol="1" gridrow="32" pageId="3" pageNumber="464">Head depth</td>
<td box="[669,911,1818,1837]" gridcol="2" gridrow="32" pageId="3" pageNumber="464">70.3</td>
<td box="[961,982,1818,1837]" gridcol="3" gridrow="32" pageId="3" pageNumber="464">47</td>
<td box="[1094,1197,1818,1837]" gridcol="4" gridrow="32" pageId="3" pageNumber="464">61.7-75.4</td>
<td box="[1306,1393,1818,1837]" gridcol="5" gridrow="32" pageId="3" pageNumber="464">69.3±2.2</td>
</tr>
<tr box="[144,1393,1854,1874]" gridrow="33" pageId="3" pageNumber="464">
<th box="[144,199,1854,1874]" gridcol="0" gridrow="33" pageId="3" pageNumber="464">1-24</th>
<td box="[268,590,1854,1874]" gridcol="1" gridrow="33" pageId="3" pageNumber="464">Mouth length</td>
<td box="[669,911,1854,1874]" gridcol="2" gridrow="33" pageId="3" pageNumber="464">46.2</td>
<td box="[961,982,1854,1874]" gridcol="3" gridrow="33" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1854,1874]" gridcol="4" gridrow="33" pageId="3" pageNumber="464">36.2-50.5</td>
<td box="[1306,1393,1854,1874]" gridcol="5" gridrow="33" pageId="3" pageNumber="464">42.7±3.7</td>
</tr>
<tr box="[144,1393,1891,1910]" gridrow="34" pageId="3" pageNumber="464">
<th box="[144,199,1891,1910]" gridcol="0" gridrow="34" pageId="3" pageNumber="464">21-22</th>
<td box="[268,590,1891,1910]" gridcol="1" gridrow="34" pageId="3" pageNumber="464">Mouth width</td>
<td box="[669,911,1891,1910]" gridcol="2" gridrow="34" pageId="3" pageNumber="464">42.0</td>
<td box="[961,982,1891,1910]" gridcol="3" gridrow="34" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1891,1910]" gridcol="4" gridrow="34" pageId="3" pageNumber="464">34.4-46.4</td>
<td box="[1306,1393,1891,1910]" gridcol="5" gridrow="34" pageId="3" pageNumber="464">39.9±3.2</td>
</tr>
<tr box="[144,1393,1927,1947]" gridrow="35" pageId="3" pageNumber="464">
<th box="[144,199,1927,1947]" gridcol="0" gridrow="35" pageId="3" pageNumber="464">22-23</th>
<td box="[268,590,1927,1947]" gridcol="1" gridrow="35" pageId="3" pageNumber="464">Barbel length</td>
<td box="[669,911,1927,1947]" gridcol="2" gridrow="35" pageId="3" pageNumber="464">13.6</td>
<td box="[961,982,1927,1947]" gridcol="3" gridrow="35" pageId="3" pageNumber="464">46</td>
<td box="[1094,1197,1927,1947]" gridcol="4" gridrow="35" pageId="3" pageNumber="464">9.7-19.6</td>
<td box="[1306,1393,1927,1947]" gridcol="5" gridrow="35" pageId="3" pageNumber="464">13.8±2.5</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[136,775,174,454]" lastBlockId="4.[820,1459,174,1031]" pageId="4" pageNumber="465">
Oral disk round, maxillary barbels of moderate length (10-20% HL). Lips papillate; small patch of elongate fleshy papillae behind each dentary tooth row (
<figureCitation box="[606,675,238,262]" captionStart="Fig" captionStartId="4.[136,176,1422,1446]" captionTargetBox="[136,775,486,1408]" captionTargetId="figure-85@4.[136,775,486,1408]" captionTargetPageId="4" captionText="Fig. 2. Details of the mouth of Panaqolus nix: the dentary tooth rows are in an acute angle (below) or even nearly parallel (above) (MZUSP 114009, 68.8 mm + 76.2 mm SL). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4551383" httpUri="https://zenodo.org/record/4551383/files/figure.png" pageId="4" pageNumber="465">Fig. 2</figureCitation>
). Border of lips smooth, without papillae. Teeth spoon-shaped and unicuspidate or with very small lateral cusp. Premaxillary teeth 3-7 per ramus (mode 4), mandibular teeth 4-7 per ramus (mode 5). Premaxillary tooth rows angled at approximately 90°, dentary tooth rows acutely angled at approximately 50° or nearly parallel (
<figureCitation box="[545,610,430,454]" captionStart="Fig" captionStartId="4.[136,176,1422,1446]" captionTargetBox="[136,775,486,1408]" captionTargetId="figure-85@4.[136,775,486,1408]" captionTargetPageId="4" captionText="Fig. 2. Details of the mouth of Panaqolus nix: the dentary tooth rows are in an acute angle (below) or even nearly parallel (above) (MZUSP 114009, 68.8 mm + 76.2 mm SL). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4551383" httpUri="https://zenodo.org/record/4551383/files/figure.png" pageId="4" pageNumber="465">Fig. 2</figureCitation>
). small plates. Body with pronounced lateral ridge extending from cleithrum to posterior margin of fifth or sixth plate of the inframedian plate row, decreasing in prominence posteriorly. Trunk without elevated ridges. 7-8 plates on dorsal-fin base (mode 7), 5-7 plates between dorsal and adipose fin (mode 6), usually 1 azygous preadipose plate, 6-8 plates between adipose and caudal fin (mode 7), 2 plates on anal-fin base, 11-12 scutes between anal and caudal fin (mode 11), and 24-25 lateral plates (mode 25).
</paragraph>
<paragraph blockId="4.[820,1459,174,1031]" pageId="4" pageNumber="465">
Dorsal fin II,7-8 (
<typeStatus box="[1038,1135,462,486]" pageId="4" pageNumber="465">holotype</typeStatus>
II,7), pectoral fin I,6, pelvic fin i,5, anal fin i,4, caudal fin i,13-14,i (
<typeStatus box="[1196,1293,494,518]" pageId="4" pageNumber="465">holotype</typeStatus>
i,14,i). Spinelet triangular, dorsal-fin spine lock functional, posterior fin margin straight, margin of last two rays rounded. Dorsalfin origin closer to vertical line passing through pelvicfin origin than to vertical line passing through pectoralfin origin; not reaching adipose fin when adpressed. Last dorsal-fin ray without prolonged membrane. Adipose fin triangular; adipose-fin spine slanted posteroventrally, tip straight to curved ventrally, pointed; posterior margin of adipose-fin membrane concave to nearly vertical. Pectoralfin spine robust, membrane between spine and first ray without fleshy extension, distal fin margin straight. Pectoral fin, when depressed reaching 1/3 of pelvic fin. Pelvic-fin spine robust, distal margin slightly rounded, when adpressed reaching mid-length of anal fin. Caudal fin strongly forked; caudal-fin spines usually tipped with filaments of the length of the longest branched caudal-fin rays.
</paragraph>
<paragraph blockId="4.[820,1459,1070,1351]" pageId="4" pageNumber="465">
<emphasis bold="true" box="[820,1009,1070,1095]" pageId="4" pageNumber="465">Color in alcohol.</emphasis>
Coloration of head, body and fins without bands or saddles. Base coloration varying from uniformly yellowish-white over light brown to dark brown or nearly black in different specimens. Small white dots (about 1/3 of the eye diameter) present on whole body and all fins; approximately one dot per plate; dots smaller on head. Dots faded to absent in preserved specimens, especially on trunk, but usually visible on fins. Caudal fin showing distinct dots or diffuse lighter markings.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551383" ID-Zenodo-Dep="4551383" httpUri="https://zenodo.org/record/4551383/files/figure.png" pageId="4" pageNumber="465" startId="4.[136,176,1422,1446]" targetBox="[136,775,486,1408]" targetPageId="4">
<paragraph blockId="4.[136,775,1422,1542]" pageId="4" pageNumber="465">
<emphasis bold="true" box="[136,209,1422,1446]" pageId="4" pageNumber="465">Fig. 2.</emphasis>
Details of the mouth of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[480,635,1422,1446]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="465" phylum="Chordata" rank="species" species="nix">
<emphasis box="[480,635,1422,1446]" italics="true" pageId="4" pageNumber="465">Panaqolus nix</emphasis>
</taxonomicName>
: the dentary tooth rows are in an acute angle (below) or even nearly parallel (above) (MZUSP 114009, 68.8 mm + 76.2 mm SL). Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph blockId="4.[136,776,1582,1958]" lastBlockId="4.[820,1459,1391,1671]" pageId="4" pageNumber="465">
Head and body plated dorsally, except for small naked area around dorsal-fin base; some specimens with naked area at tip of snout. Supraoccipital bordered posteriorly by 2-3 (mode 3) plates on each side. Abdomen of adults ranging from incompletely plated (plates only in pectoral girdle, along sides, and posterior to pelvic fin) to nearly completely covered by small irregularly arranged platelets, with a small naked area around pelvic-fin origin and the urogenital orifice. Abdomen of juveniles of
<quantity box="[417,494,1838,1862]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="465" unit="mm" value="70.0">70 mm</quantity>
SL or less naked or with only very few plates on the border with inframedian plates. Large naked area dorsally to pelvic-fin base, below ventral margin of inframedian plate row; sometimes showing 1-3
<emphasis bold="true" box="[820,964,1391,1415]" pageId="4" pageNumber="465">Color in life.</emphasis>
Color of head, body and fins without bands or saddles. Base coloration varying from uniformly yellowishwhite over pale brown (
<figureCitation box="[1071,1132,1455,1479]" captionStart="Fig" captionStartId="5.[129,168,1016,1040]" captionTargetBox="[166,1413,174,1006]" captionTargetId="figure-181@5.[166,1413,174,1006]" captionTargetPageId="5" captionText="Fig. 3. Variation in the color of Panaqolus nix: a) nearly white specimen (UFRO-I 10050, 90.4 mm SL) b), intermediately colored specimen (UFRO-I 7968, 81.4 mm SL) (Photos by B. S. F. Barros)." figureDoi="http://doi.org/10.5281/zenodo.4551385" httpUri="https://zenodo.org/record/4551385/files/figure.png" pageId="4" pageNumber="465">Fig. 3</figureCitation>
) to dark brown or nearly black in different specimens. In an aquarium, with clear water, light colored fish become dark in few days (
<figureCitation box="[1320,1388,1519,1543]" captionStart="Fig" captionStartId="5.[129,168,1542,1566]" captionTargetBox="[208,1395,1113,1522]" captionTargetId="figure-218@5.[166,1413,1104,1537]" captionTargetPageId="5" captionText="Fig. 4. Live photo of the holotype of Panaqolus nix (INPA 39606; 110.1 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4551388" httpUri="https://zenodo.org/record/4551388/files/figure.png" pageId="4" pageNumber="465">Fig. 4</figureCitation>
). Live specimens with small white dots (more or less 1/3 of the eye diameter) on whole body and all fins; approximately one dot per plate. Dots smaller on head. Caudal fin showing distinct dots or diffuse lighter markings.
</paragraph>
<paragraph blockId="4.[820,1466,1711,1960]" pageId="4" pageNumber="465">
<emphasis bold="true" box="[820,1052,1711,1735]" pageId="4" pageNumber="465">Sexual dimorphism.</emphasis>
Mature males have longer odontodes on the pectoral-fin spine and on the caudal peduncle (
<figureCitation box="[1383,1445,1744,1768]" captionStart="Fig" captionStartId="6.[136,176,1046,1070]" captionTargetBox="[261,1333,174,1033]" captionTargetId="figure-10@6.[261,1333,174,1033]" captionTargetPageId="6" captionText="Fig. 5. Sexual dimorphic male of Panaqolus nix (INPA 41150 (29859); 105.1 mm SL) with elongated odontodes on pectoral spine, caudal peduncle and caudal fin." figureDoi="http://doi.org/10.5281/zenodo.4551390" httpUri="https://zenodo.org/record/4551390/files/figure.png" pageId="4" pageNumber="465">Fig. 5</figureCitation>
). The illustrated specimen was dissected and its gender was confirmed based on gonads examination. A large female (UFRO-I 19646) that was kept in an aquarium did not develop any conspicuously elongated odontodes over time. Some eggs were released during the process of preservation shortly after she died in captivity.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551385" ID-Zenodo-Dep="4551385" httpUri="https://zenodo.org/record/4551385/files/figure.png" pageId="5" pageNumber="466" startId="5.[129,168,1016,1040]" targetBox="[166,1413,174,1006]" targetPageId="5">
<paragraph blockId="5.[129,1451,1016,1073]" pageId="5" pageNumber="466">
<emphasis bold="true" box="[129,200,1016,1040]" pageId="5" pageNumber="466">Fig. 3.</emphasis>
Variation in the color of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[474,629,1017,1041]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="466" phylum="Chordata" rank="species" species="nix">
<emphasis box="[474,629,1017,1041]" italics="true" pageId="5" pageNumber="466">Panaqolus nix</emphasis>
</taxonomicName>
: a) nearly white specimen (UFRO-I 10050, 90.4 mm SL) b), intermediately colored specimen (UFRO-I 7968, 81.4 mm SL) (Photos by B. S. F. Barros).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551388" ID-Zenodo-Dep="4551388" box="[129,1001,1542,1567]" httpUri="https://zenodo.org/record/4551388/files/figure.png" pageId="5" pageNumber="466" startId="5.[129,168,1542,1566]" targetBox="[208,1395,1113,1522]" targetPageId="5">
<paragraph blockId="5.[129,1001,1542,1567]" box="[129,1001,1542,1567]" pageId="5" pageNumber="466">
<emphasis bold="true" box="[129,198,1542,1566]" pageId="5" pageNumber="466">Fig. 4.</emphasis>
Live photo of the holotype of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[525,679,1543,1567]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="466" phylum="Chordata" rank="species" species="nix">
<emphasis box="[525,679,1543,1567]" italics="true" pageId="5" pageNumber="466">Panaqolus nix</emphasis>
</taxonomicName>
(INPA 39606; 110.1 mm SL).
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="466" type="distribution">
<paragraph blockId="5.[129,767,1614,1959]" lastBlockId="5.[813,1452,1615,1767]" pageId="5" pageNumber="466">
<emphasis bold="true" box="[129,442,1614,1639]" pageId="5" pageNumber="466">Distribution and Habitat.</emphasis>
The known distribution of this species is the Madeira basin, including the Madeira, Mamoré, and Tambopata (
<collectingRegion box="[419,585,1679,1703]" country="Peru" name="Madre de Dios" pageId="5" pageNumber="466">Madre de Dios</collectingRegion>
drainage) rivers (
<figureCitation box="[137,208,1711,1735]" captionStart="Fig" captionStartId="6.[136,176,1901,1925]" captionTargetBox="[274,1327,1146,1877]" captionTargetId="figure-38@6.[265,1330,1134,1887]" captionTargetPageId="6" captionText="Fig. 6. Distribution of Panaqolus nix. Open symbol: type locality. Symbols may represent more than one locality or lot. Ji = former Jirau rapids, SA = former Santo Antônio rapids." figureDoi="http://doi.org/10.5281/zenodo.4551392" httpUri="https://zenodo.org/record/4551392/files/figure.png" pageId="5" pageNumber="466">Fig. 6</figureCitation>
). Most of the specimens have been collected in cofferdams at the construction sites of the hydroelectric power plants of Santo Antônio and Jirau (former Santo Antônio and Jirau rapids), on the rio Madeira, by several technicians and ichthyologists hired by the two Consulting Environmental Companies responsible for the biological inventories in the area. Most of the remaining specimens have been caught in depths of
<quantity box="[468,591,1935,1959]" metricMagnitude="0" metricUnit="m" metricValue="7.05" metricValueMax="11.0" metricValueMin="3.1" pageId="5" pageNumber="466" unit="m" value="7.05" valueMax="11.0" valueMin="3.1">3.1 to 11 m</quantity>
(UFRO-I 6384, INPA 39605, MZUSP 114009) and
<quantity box="[1208,1342,1615,1639]" metricMagnitude="0" metricUnit="m" metricValue="6.3" metricValueMax="8.5" metricValueMin="4.1" pageId="5" pageNumber="466" unit="m" value="6.3" valueMax="8.5" valueMin="4.1">4.1 to 8.5 m</quantity>
(UFRO-I 6384) using trawl nets. This indicates that this species prefers deep-water habitats with strong current, which might help to explain its relative rarity in scientific collections.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="466" type="etymology">
<paragraph blockId="5.[813,1451,1807,1959]" pageId="5" pageNumber="466">
<emphasis bold="true" box="[813,943,1807,1831]" pageId="5" pageNumber="466">Etymology.</emphasis>
From the Latin
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[1139,1172,1808,1831]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="466" phylum="Chordata" rank="species" species="nix">
<emphasis box="[1139,1172,1808,1831]" italics="true" pageId="5" pageNumber="466">nix</emphasis>
</taxonomicName>
meaning snow, alluding to the color: in dark individuals the dots look like falling snowflakes, while pale individuals look like they have the whole body covered by snow; treated as a noun in apposition.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551390" ID-Zenodo-Dep="4551390" httpUri="https://zenodo.org/record/4551390/files/figure.png" pageId="6" pageNumber="467" startId="6.[136,176,1046,1070]" targetBox="[261,1333,174,1033]" targetPageId="6">
<paragraph blockId="6.[136,1459,1046,1102]" pageId="6" pageNumber="467">
<emphasis bold="true" box="[136,199,1046,1071]" pageId="6" pageNumber="467">Fig. 5</emphasis>
. Sexual dimorphic male of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[494,648,1046,1070]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="467" phylum="Chordata" rank="species" species="nix">
<emphasis box="[494,648,1046,1070]" italics="true" pageId="6" pageNumber="467">Panaqolus nix</emphasis>
</taxonomicName>
(INPA 41150 (#29859); 105.1 mm SL) with elongated odontodes on pectoral spine, caudal peduncle and caudal fin.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4551392" ID-Zenodo-Dep="4551392" httpUri="https://zenodo.org/record/4551392/files/figure.png" pageId="6" pageNumber="467" startId="6.[136,176,1901,1925]" targetBox="[274,1327,1146,1877]" targetPageId="6">
<paragraph blockId="6.[136,1459,1901,1957]" pageId="6" pageNumber="467">
<emphasis bold="true" box="[136,210,1901,1925]" pageId="6" pageNumber="467">Fig. 6.</emphasis>
Distribution of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[387,546,1901,1925]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="467" phylum="Chordata" rank="species" species="nix">
<emphasis box="[387,546,1901,1925]" italics="true" pageId="6" pageNumber="467">Panaqolus nix</emphasis>
</taxonomicName>
. Open symbol: type locality. Symbols may represent more than one locality or lot. Ji = former Jirau rapids, SA = former Santo Antônio rapids.
</paragraph>
</caption>
<subSubSection pageId="7" pageNumber="468" type="discussion">
<paragraph blockId="7.[128,768,174,582]" pageId="7" pageNumber="468">
<emphasis bold="true" box="[129,240,174,198]" pageId="7" pageNumber="468">Remarks.</emphasis>
<accessionNumber box="[247,382,174,198]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM92440" pageId="7" pageNumber="468">ROM 92440</accessionNumber>
is not being considered in the
<typeStatus box="[719,767,174,198]" pageId="7" pageNumber="468">type</typeStatus>
material as it is a dried specimen that disintegrated into several pieces. Zawadzki &amp; Chamon (2013), in their list of
<taxonomicName authorityName="Kner" authorityYear="1853" box="[129,288,270,294]" class="Actinopterygii" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="subFamily" subFamily="Hypostominae">Hypostominae</taxonomicName>
of the rio Madeira, illustrated
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[616,711,270,293]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="genus">
<emphasis box="[616,711,270,293]" italics="true" pageId="7" pageNumber="468">Panaque</emphasis>
</taxonomicName>
sp.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="468" unit="in" value="1.0">1 in</quantity>
page 312, and cited the lot UFRO-I 6384 as
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[634,729,302,325]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="genus">
<emphasis box="[634,729,302,325]" italics="true" pageId="7" pageNumber="468">Panaque</emphasis>
</taxonomicName>
sp. 2, in page 315. In fact, there was a mistake in assigning the lots to the images. The illustration of
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[539,634,366,389]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="genus">
<emphasis box="[539,634,366,389]" italics="true" pageId="7" pageNumber="468">Panaque</emphasis>
</taxonomicName>
sp. 1 shows in fact a specimen of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[357,512,398,422]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="species" species="nix">
<emphasis box="[357,512,398,422]" italics="true" pageId="7" pageNumber="468">Panaqolus nix</emphasis>
</taxonomicName>
and the correct number of this lot is UFRO-I 6384. The second illustration,
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[672,767,430,453]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="genus">
<emphasis box="[672,767,430,453]" italics="true" pageId="7" pageNumber="468">Panaque</emphasis>
</taxonomicName>
sp. 2, is a different species and the correct numbers of the lots for this species are UFRO-I 6383 and UFRO-I 13109. So,
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[172,329,526,550]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="species" species="nix">
<emphasis box="[172,329,526,550]" italics="true" pageId="7" pageNumber="468">Panaqolus nix</emphasis>
</taxonomicName>
was already known, but not named, as part of the ichthyofauna of the rio Madeira.
</paragraph>
<paragraph blockId="7.[128,768,622,1222]" pageId="7" pageNumber="468">
<emphasis bold="true" box="[128,369,622,646]" pageId="7" pageNumber="468">Conservation status.</emphasis>
The species seems to be widely distributed along rio Madeira, Mamoré and Tambopata as well. Despite the fact that part of its distribution includes two hydroelectric power dams, Jirau and Santo Antônio, the area of occurrence of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[311,464,750,774]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="species" species="nix">
<emphasis box="[311,464,750,774]" italics="true" pageId="7" pageNumber="468">Panaqolus nix</emphasis>
</taxonomicName>
goes well beyond the dams. Thus, considering that important threats to the species were not detected yet,
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" box="[307,368,815,838]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="species" species="nix">
<emphasis box="[307,368,815,838]" italics="true" pageId="7" pageNumber="468">P. nix</emphasis>
</taxonomicName>
should be classified as Least Concern (LC) according to the International Union for Conservation of Nature (IUCN) categories and criteria (IUCN Standards and Petitions Subcommittee, 2014). In the last years parts of the known habitats have been severely altered by the construction of these hydroelectric power plants, destroying the former Santo Antônio and Jirau rapids and transforming more than
<quantity box="[241,323,1038,1062]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="468" unit="km" value="200.0">200 km</quantity>
of the rio Madeira, enlarging the riverbed and reducing the current. Additional collecting efforts should be conducted in that region in order to better understand the impacts of these constructions on the population of
<taxonomicName authorityName="Cramer &amp; Py-Daniel" authorityYear="2015" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="468" phylum="Chordata" rank="species" species="nix">
<emphasis italics="true" pageId="7" pageNumber="468">Panaqolus nix</emphasis>
</taxonomicName>
and to be able to adjust (if necessary) the evaluation of the conservation status of the species.
</paragraph>
</subSubSection>
</treatment>
</document>