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<mods:title id="BAFE130F93C724A25A1CE8D7B38F5320">Unexpected diversity of the genus Collarina Jullien, 1886 (Bryozoa, Cheilostomatida) in the NE Atlantic-Mediterranean region: new species and reappraisal of C. balzaci (Audouin, 1826) and C. fayalensis Harmelin, 1978</mods:title>
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<taxonomicName id="4C2883DFFFEFFF85FC133ABCFAB32E7D" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[914,1285,503,530]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFEFFF85FC133ABCFB272E7D" bold="true" box="[914,1169,503,530]" italics="true" pageId="19" pageNumber="402">Collarina macaronensis</emphasis>
Harmelin
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,
<taxonomicNameLabel id="A26F9935FFEFFF85FA913AB2FAFC2E7C" box="[1296,1354,505,531]" pageId="19" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="1313E4D9FFEFFF85FCF63953FC7A2E5D" box="[887,972,536,562]" captionStart="FIG" captionStartId="4.[167,178,836,854]" captionTargetBox="[379,1214,221,790]" captionTargetId="figure@4.[338,1245,216,826]" captionTargetPageId="4" captionText="FIG. 1. — Savignys drawings (1817: pl. 9). Flustra balzaci Audouin, 1826, in Savignys figure 8, and Flustra jauberti Audouin, 1826, in Savignys figure 9." figureDoi="http://doi.org/10.5281/zenodo.3725958" httpUri="https://zenodo.org/record/3725958/files/figure.png" pageId="19" pageNumber="402">Figs 8B</figureCitation>
; 12; 15C; E; 17A, C; 18E;
<tableCitation id="C6AACDE7FFEFFF85FB723953FAD42E5D" box="[1267,1378,536,562]" captionStart-0="TABLE" captionStart-1="TABLE" captionStart-2="TABLE" captionStartId-0="9.[132,143,221,239]" captionStartId-1="29.[132,143,221,239]" captionStartId-2="29.[813,824,221,239]" captionText-0="TABLE 1. — Measurements of Collarina Jullien,1886 species:C. balzaci (Audouin,1826);C. speluncola Harmelin,n. sp.; C. denticulata Harmelin,n. sp.; C. gautieri Harmelin, n. sp.; C. macaronensis Harmelin, n. sp.; C. fayalensis Harmelin, 1978. Abbreviations: X, mean; SD, standard deviation; N, number of measurements and range; LZo, WZo, length and width of non ovicelled autozooid; LOr, WOr, of orifice of non ovicelled zooid; Lov, WOv, orifice of ovicell. Other abbreviations: Atl., Atlantic ocean; Kerk., Kerkennah Islands; Med., Mediterranean." captionText-1="TABLE 2. — Characterization of Collarina Jullien, 1886 species by contrasting character states. Abbreviations: C.b.: C. balzaci (Audouin, 1826); C.s.: C. speluncola Harmelin, n. sp.; C.d.: C. denticulata Harmelin, n. sp.; C.g.: C. gautieri Harmelin, n. sp.; C.m.: C. macaronensis Harmelin, n. sp.; C.f.: C. fayalensis Harmelin, 1978; Ov., ovicell; Kz, distal kenozooid; Az, distal autozooid." captionText-2="TABLE 3. — Features of ovicells in the six Collarina Jullien,1886 species (Mediterranean and Atlantic material for C. gautieri Harmelin, n. sp.). Frequency data in %: A, occurrence frequency of ovicelled autozooids within colonies; B, ooecium types: frequency of ooecia produced by a kenozooid (Kz) or a distal autozooid (Az), and frequency of ooecia bearing a distal avicularium (Av); C, type of ovicell closure.Origin of data C. balzaci (Audouin,1826):Turkey,Lebanon, Adriatic,France Planier,7 colonies,88 zooids,53 ooecia; C. speluncola Harmelin, n. sp.: Morgiou, Conger Cave, 8 colonies including types, 473 zooids, 89 ooecia; C. denticulata Harmelin, n. sp.: Port-Miou &amp; Renaire, 6 colonies, 342 zooids, 104 ooecia (63 Az, 41 Kz); C. gautieri Harmelin, n. sp., Mediterranean: Medes,Naples,Bonifacio,Port-Cros,Veyron (holotype),5 colonies,173 zooids, 92 ooecia; C. gautieri Harmelin, n. sp., Atlantic: Plymouth, Brittany (2 sites), Algarve, 9 colonies,191 zooids,100 ooecia; C. macaronensis Harmelin,n. sp.: Madeira (NHMUK 4 colonies + Baixo Is., holotype &amp; paratype), 298 zooids, 67 ooecia; C. fayalensis Harmelin, 1978: Sao Miguel, I. Villa Franca, 3 colonies, 433 zooids, 72 ooecia. Frequency numbers in %. n.d.: no data." httpUri-0="http://table.plazi.org/id/DF57A8D4FFF5FF9FFF053B96FD742D5C" httpUri-1="http://table.plazi.org/id/DF57A8D4FFE1FF8BFF053B96FD642D25" httpUri-2="http://table.plazi.org/id/DF57A8D4FFE1FF8BFCAC3B96FA8A2E2C" pageId="19" pageNumber="402" tableUuid-0="DF57A8D4FFF5FF9FFF053B96FD742D5C" tableUuid-1="DF57A8D4FFE1FF8BFF053B96FD642D25" tableUuid-2="DF57A8D4FFE1FF8BFCAC3B96FA8A2E2C">Tables 1-3</tableCitation>
)
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<emphasis id="B95C244EFFEFFF85FCAC39CCFC632EF0" box="[813,981,647,671]" italics="true" pageId="19" pageNumber="402">Cribrilina balzaci</emphasis>
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<treatmentCitation id="0A89DE4DFFEFFF85FC6F39C3FB322EF0" author="WATERS A. W." box="[1006,1156,647,671]" page="9" pageId="19" pageNumber="402" year="1899">
<bibRefCitation id="EFB985ADFFEFFF85FC6F39C3FB322EF0" author="WATERS A. W." box="[1006,1156,647,671]" pageId="19" pageNumber="402" pagination="6 - 16" refId="ref30475" refString="WATERS A. W. 1899. - Bryozoa from Madeira. Journal of the Royal Microscopical Society 1899: 6 - 16. https: // doi. org / 10.1111 / j. 1365 - 2818.1899. tb 00139. x" type="journal article" year="1899">Waters 1899: 9</bibRefCitation>
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, figs 31-32. —
<treatmentCitation id="0A89DE4DFFEFFF85FA9C39C3FCFE2ED5" author="NORMAN A. M." page="98" pageId="19" pageNumber="402" year="1903">
<bibRefCitation id="EFB985ADFFEFFF85FA9C39C3FCFE2ED5" author="NORMAN A. M." pageId="19" pageNumber="402" pagination="87 - 128" refId="ref29204" refString="NORMAN A. M. 1903. - Notes on the natural history of East Finmark. Annals and Magazine of Natural History Series 7 (12): 87 - 128. https: // doi. org / 10.1080 / 00222930308678831" type="journal article" year="1903">Norman 1903: 98</bibRefCitation>
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, pl. 9, fig. 6; 1909: 292.
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<emphasis id="B95C244EFFEFFF85FCAC399AFC6F2E86" box="[813,985,721,745]" italics="true" pageId="19" pageNumber="402">Collarina balzaci</emphasis>
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<bibRefCitation id="EFB985ADFFEFFF85FC77399AFB172E86" author="HARMELIN J. G." box="[1014,1185,721,745]" pageId="19" pageNumber="402" pagination="173 - 192" refId="ref28028" refString="HARMELIN J. G. 1978 a. - Sur quelques cribrimorphes (Bryozoa, Cheilostomata) de l'Atlantique oriental. Tethys 8: 173 - 192." type="journal article" year="1978">Harmelin 1978a</bibRefCitation>
(in part): 189
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, fig. 8 (not pl. I, fig. 7).
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<treatmentCitationGroup id="AB38DF72FFEFFF85FCAC3850FB572F21" pageId="19" pageNumber="402">
<taxonomicName id="4C2883DFFFEFFF85FCAC3850FC552F5C" ID-CoL="ZFNQ" box="[813,995,795,819]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="punctata">
<emphasis id="B95C244EFFEFFF85FCAC3850FC552F5C" box="[813,995,795,819]" italics="true" pageId="19" pageNumber="402">Cribrilina punctata</emphasis>
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<treatmentCitation id="0A89DE4DFFEFFF85FC793850FCE32F21" author="FERNANDEZ PULPEIRO E. &amp; RODRIGUEZ BABIO C." page="136" pageId="19" pageNumber="402" year="1980">
<bibRefCitation id="EFB985ADFFEFFF85FC793850FCE32F21" author="FERNANDEZ PULPEIRO E. &amp; RODRIGUEZ BABIO C." pageId="19" pageNumber="402" pagination="119 - 168" refId="ref27585" refString="FERNANDEZ PULPEIRO E. &amp; RODRIGUEZ BABIO C. 1980. - Aportaciones al conocimiento de la fauna briozoologica del litoral de la ria de Vigo. Investigacion Pesquera 44 (1): 119 - 168." type="journal article" year="1980">Fernández Pulpeiro &amp; Rodriguez Babio 1980: 136</bibRefCitation>
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, fig. 2. —
<treatmentCitation id="0A89DE4DFFEFFF85FC3C387EFBC42F21" author="ALVAREZ J. A." box="[957,1138,821,847]" page="348" pageId="19" pageNumber="402" year="1988">
<bibRefCitation id="EFB985ADFFEFFF85FC3C387EFBC42F21" author="ALVAREZ J. A." box="[957,1138,821,847]" pageId="19" pageNumber="402" pagination="347 - 352" refId="ref25800" refString="ALVAREZ J. A. 1988. - La familia Cribrilinidae Hincks, 1880 (Bryozoa, Cheilostomida) en la Costa Vasca. Miscellanea Zoologica 12: 347 - 352." type="journal article" year="1988">Álvarez 1988: 348</bibRefCitation>
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, fig. 3B-D.
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<treatmentCitationGroup id="AB38DF72FFEFFF85FCAC382EFC632FF7" pageId="19" pageNumber="402">
<taxonomicName id="4C2883DFFFEFFF85FCAC382EFC542F12" ID-CoL="X2VF" authorityName="Harmelin" authorityYear="1978" box="[813,994,869,893]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="fayalensis">
<emphasis id="B95C244EFFEFFF85FCAC382EFC542F12" box="[813,994,869,893]" italics="true" pageId="19" pageNumber="402">Collarina fayalensis</emphasis>
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<bibRefCitation id="EFB985ADFFEFFF85FC78382EFB3F2F11" author="ARISTEGUI RUIZ J." box="[1017,1161,869,894]" pageId="19" pageNumber="402" pagination="1 - 524" refId="ref25832" refString="ARISTEGUI RUIZ J. 1984. - Briozoos quilostomados (Ectoprocta, Cheilostomata) de Canarias: Estudio sistematico, faunistico y biogeografico. Tesis Doctoral, Facultad de Biologia, Universidad de La Laguna, Las Palma, i-iii, 1 - 524." type="book chapter" year="1984">Arístegui 1984</bibRefCitation>
(in part): 230
</treatmentCitation>
, fig. 49b, pl. 11, fig. 6 (not fig. 7).
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TYPE LOCALITY. —
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,
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,
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Santo, Baixo Is
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.
</paragraph>
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<subSubSection id="C332ABD7FFEFFF85FCAC3894FB752A72" pageId="19" pageNumber="402" type="materials_examined">
<paragraph id="8B97F85CFFEFFF85FCAC3894FBDF2829" blockId="19.[813,1455,991,1200]" pageId="19" pageNumber="402">
<collectionCode id="ED396099FFEFFF85FCAC3894FCEB2F9B" box="[813,861,991,1015]" pageId="19" pageNumber="402">TYPE</collectionCode>
MATERIAL. —
<materialsCitation id="3B40F201FFEFFF85FC653894FBDF2829" ID-GBIF-Occurrence="2580041123" collectionCode="TYPE" collectorName="Baixo Is., R &amp; V Jean-Charcot" country="Portugal" elevation="7" latitude="33.00889" location="Madeira" longLatPrecision="20" longitude="-16.386944" pageId="19" pageNumber="402" specimenCount="1" stateProvince="Madeira" typeStatus="holotype">
<emphasis id="B95C244EFFEFFF85FC653894FBF72F98" bold="true" box="[996,1089,991,1015]" pageId="19" pageNumber="402">
<typeStatus id="549346FEFFEFFF85FC653894FBF72F98" box="[996,1089,991,1015]" pageId="19" pageNumber="402" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCountry id="F33FB8CCFFEFFF85FBCA3894FB282F98" box="[1099,1182,991,1015]" name="Portugal" pageId="19" pageNumber="402">Portugal</collectingCountry>
,
<collectingRegion id="49EC36BEFFEFFF85FB243894FB412F98" box="[1189,1271,991,1015]" country="Portugal" name="Madeira" pageId="19" pageNumber="402">Madeira</collectingRegion>
,
<collectingRegion id="49EC36BEFFEFFF85FB7E38ABFA852F98" box="[1279,1331,992,1015]" country="Portugal" name="Porto" pageId="19" pageNumber="402">Porto</collectingRegion>
Santo,
<collectorName id="26DD9D8AFFEFFF85FAF83894FCEA287D" pageId="19" pageNumber="402">Baixo Is., R</collectorName>
/
<collectorName id="26DD9D8AFFEFFF85FCE238B1FC58287D" box="[867,1006,1018,1042]" pageId="19" pageNumber="402">
V
<emphasis id="B95C244EFFEFFF85FCF838B1FC58287D" box="[889,1006,1018,1042]" italics="true" pageId="19" pageNumber="402">Jean-Charcot</emphasis>
</collectorName>
, ZARCO 1966 St. 23,
<geoCoordinate id="EE1C9E9BFFEFFF85FB4538B1FA9A287D" box="[1220,1324,1018,1042]" degrees="33" direction="north" minutes="0" orientation="latitude" pageId="19" pageNumber="402" precision="15" seconds="32" value="33.00889">33°032”N</geoCoordinate>
,
<geoCoordinate id="EE1C9E9BFFEFFF85FAB438B1FA1A287D" box="[1333,1452,1018,1042]" degrees="16" direction="west" minutes="23" orientation="longitude" pageId="19" pageNumber="402" precision="15" seconds="13" value="-16.386944">16°2313”W</geoCoordinate>
,
<quantity id="4CD055B9FFEFFF85FCAC3F5FFCDD2843" box="[813,875,1044,1068]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="19" pageNumber="402" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">
<elevation id="00051F6FFFEFFF85FCAC3F5FFCDD2843" box="[813,875,1044,1068]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="19" pageNumber="402" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">5-8 m</elevation>
</quantity>
, dark tunnel, 1 large ovicelled colony (&gt; 1000 zooids) on a pebble, MNHN-IB-2014-1924.
</materialsCitation>
</paragraph>
<paragraph id="8B97F85CFFEFFF85FCAC3F02FBF828DF" blockId="19.[813,1455,991,1200]" pageId="19" pageNumber="402">
<materialsCitation id="3B40F201FFEFFF85FCAC3F02FB2B2814" ID-GBIF-Occurrence="2580041195" collectionCode="MNHN-IB-" country="Afghanistan" pageId="19" pageNumber="402" specimenCount="1" typeStatus="paratype">
<emphasis id="B95C244EFFEFFF85FCAC3F02FC3B280E" bold="true" box="[813,909,1097,1121]" pageId="19" pageNumber="402">
<typeStatus id="549346FEFFEFFF85FCAC3F02FC3B280E" box="[813,909,1097,1121]" pageId="19" pageNumber="402" type="paratype">Paratypes</typeStatus>
</emphasis>
. Same origin as the holotype:
<collectionCode id="ED396099FFEFFF85FB3E3F02FA8A280E" box="[1215,1340,1097,1121]" pageId="19" pageNumber="402">MNHN-IB-</collectionCode>
2014-1925: 2 colonies together with the holotype.
</materialsCitation>
<materialsCitation id="3B40F201FFEFFF85FB413F28FBBD28F9" ID-GBIF-Occurrence="2580041102" country="Afghanistan" pageId="19" pageNumber="402" specimenCount="1">MNHN-IB-2014-1926: 4 colonies on 2 pebbles.</materialsCitation>
<materialsCitation id="3B40F201FFEFFF85FBAD3F35FBFE28DF" ID-GBIF-Occurrence="2580041127" country="Afghanistan" pageId="19" pageNumber="402" specimenCount="1">MNHN-IB-2014-1927: coated ovicelled colony for SEM examination</materialsCitation>
.
</paragraph>
<paragraph id="8B97F85CFFEFFF85FCAC3F83FB752A72" blockId="19.[813,1456,1224,1566]" pageId="19" pageNumber="402">
OTHER MATERIAL EXAMINED. —
<materialsCitation id="3B40F201FFEFFF85FBE43F83FCC72926" ID-GBIF-Occurrence="2580041148" collectingDate="1971-10-12" country="Italy" latitude="38.52" longitude="-28.701944" pageId="19" pageNumber="402" stateProvince="Azores" typeStatus="holotype">
<emphasis id="B95C244EFFEFFF85FBE43F83FB10288F" bold="true" box="[1125,1190,1224,1248]" pageId="19" pageNumber="402">
<collectingRegion id="49EC36BEFFEFFF85FBE43F83FB10288F" box="[1125,1190,1224,1248]" country="Portugal" name="Acores" pageId="19" pageNumber="402">Azores</collectingRegion>
</emphasis>
, Faial,
<collectingRegion id="49EC36BEFFEFFF85FB683F83FAC828B0" box="[1257,1406,1224,1248]" country="Portugal" name="Castelo Branco" pageId="19" pageNumber="402">Castelo Branco</collectingRegion>
, R/V
<emphasis id="B95C244EFFEFFF85FCAC3FA9FC112895" box="[813,935,1250,1274]" italics="true" pageId="19" pageNumber="402">Jean-Charcot</emphasis>
, BIAÇORES 1971 St. P.11,
<geoCoordinate id="EE1C9E9BFFEFFF85FB3A3FA8FA822894" box="[1211,1332,1251,1275]" degrees="38" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="402" precision="15" seconds="12" value="38.52">38°3112”N</geoCoordinate>
,
<geoCoordinate id="EE1C9E9BFFEFFF85FABC3FA8FA1A2894" box="[1341,1452,1251,1275]" degrees="28" direction="west" minutes="42" orientation="longitude" pageId="19" pageNumber="402" precision="15" seconds="7" value="-28.701944">28°427”W</geoCoordinate>
,
<quantity id="4CD055B9FFEFFF85FCAC3FB6FCDF297B" box="[813,873,1277,1301]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="19" pageNumber="402" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">5-7 m</quantity>
,
<date id="FF96DE9CFFEFFF85FCF33FB6FC68297A" box="[882,990,1277,1301]" pageId="19" pageNumber="402" value="1971-10-12">12.X.1971</date>
, coll. by H. Zibrowius, MNHN-IB-2008-7909: 2 colonies on pebble together with
<typeStatus id="549346FEFFEFFF85FBF93E5CFB7B2940" box="[1144,1229,1303,1327]" pageId="19" pageNumber="402" type="holotype">holotype</typeStatus>
and
<typeStatus id="549346FEFFEFFF85FB7B3E53FAE3295F" box="[1274,1365,1304,1328]" pageId="19" pageNumber="402" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C2883DFFFEFFF85FAF03E53FCDC2925" authorityName="Harmelin" authorityYear="1978" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="fayalensis">
<emphasis id="B95C244EFFEFFF85FAF03E53FCDC2925" italics="true" pageId="19" pageNumber="402">C. fayalensis</emphasis>
</taxonomicName>
.
</materialsCitation>
<materialsCitation id="3B40F201FFEFFF85FC1A3E79FB202910" ID-GBIF-Occurrence="2580041162" collectionCode="SEM" country="Italy" latitude="43.35611" longitude="-8.838056" pageId="19" pageNumber="402" stateProvince="Galicia">
<collectingRegion id="49EC36BEFFEFFF85FC1A3E79FC532925" box="[923,997,1330,1354]" country="Spain" name="Galicia" pageId="19" pageNumber="402">Galicia</collectingRegion>
, Sisargas Is.,
<geoCoordinate id="EE1C9E9BFFEFFF85FBEC3E79FB5D2925" box="[1133,1259,1330,1354]" degrees="43" direction="north" minutes="21" orientation="latitude" pageId="19" pageNumber="402" precision="15" seconds="22" value="43.35611">43°2122”N</geoCoordinate>
,
<geoCoordinate id="EE1C9E9BFFEFFF85FB763E79FADE2925" box="[1271,1384,1330,1354]" degrees="8" direction="west" minutes="50" orientation="longitude" pageId="19" pageNumber="402" precision="15" seconds="17" value="-8.838056">8°5017”W</geoCoordinate>
, depth unknown, coll. by Fernández-Pulpeiro &amp; Reverter leg., 1 colony (
<collectionCode id="ED396099FFEFFF85FCB23E2CFCD12910" box="[819,871,1383,1407]" pageId="19" pageNumber="402">SEM</collectionCode>
photos), MHNUSC-Bry-662.
</materialsCitation>
<materialsCitation id="3B40F201FFEFFF85FB3D3E2CFB092A72" ID-GBIF-Occurrence="2580041139" collectionCode="NHMUK" country="Italy" pageId="19" pageNumber="402" stateProvince="Madeira">
Material from
<collectionCode id="ED396099FFEFFF85FACD3E2CFA192911" box="[1356,1455,1383,1406]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="402" type="Museum">NHMUK</collectionCode>
collections, SEM photos (
<collectionCode id="ED396099FFEFFF85FB9D3ECAFBFA29F6" box="[1052,1100,1409,1433]" pageId="19" pageNumber="402">MSJ</collectionCode>
):
<collectionCode id="ED396099FFEFFF85FBDB3EC9FB0D29F6" box="[1114,1211,1410,1433]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="402" type="Museum">NHMUK</collectionCode>
1899.7.1.1344 (
<taxonomicName id="4C2883DFFFEFFF85FAD93ECAFCD929DB" authorityName=": Harmelin" authorityYear="1978" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFEFFF85FAD93ECAFCD929DB" italics="true" pageId="19" pageNumber="402">Collarina balzaci</emphasis>
</taxonomicName>
),
<collectingRegion id="49EC36BEFFEFFF85FC043ED7FC6F29DB" box="[901,985,1436,1460]" country="Portugal" name="Madeira" pageId="19" pageNumber="402">Madeira</collectingRegion>
, J. Y. Johnson leg., ovicelled colony;
<collectionCode id="ED396099FFEFFF85FACD3ED7FA1929DC" box="[1356,1455,1436,1459]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="402" type="Museum">NHMUK</collectionCode>
1899.7.1.2419 (
<taxonomicName id="4C2883DFFFEFFF85FC483EFDFB9429A1" authorityName="Jullien" authorityYear="1886" box="[969,1058,1462,1486]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="genus">
<emphasis id="B95C244EFFEFFF85FC483EFDFB9429A1" box="[969,1058,1462,1486]" italics="true" pageId="19" pageNumber="402">Collarina</emphasis>
</taxonomicName>
sp.),
<collectingRegion id="49EC36BEFFEFFF85FBD43EFDFB1E29A1" box="[1109,1192,1462,1486]" country="Portugal" name="Madeira" pageId="19" pageNumber="402">Madeira</collectingRegion>
, J. Y. Johnson leg., ovicelled colony;
<collectionCode id="ED396099FFEFFF85FCFF3E9AFC542987" box="[894,994,1489,1512]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="402" type="Museum">NHMUK</collectionCode>
1899.7.1.2105 (
<taxonomicName id="4C2883DFFFEFFF85FB0F3E9BFB5F2987" authorityName="Jullien" authorityYear="1886" box="[1166,1257,1488,1512]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="genus">
<emphasis id="B95C244EFFEFFF85FB0F3E9BFB5F2987" box="[1166,1257,1488,1512]" italics="true" pageId="19" pageNumber="402">Collarina</emphasis>
</taxonomicName>
sp.),
<collectingRegion id="49EC36BEFFEFFF85FAA53E9AFACF2986" box="[1316,1401,1489,1513]" country="Portugal" name="Madeira" pageId="19" pageNumber="402">Madeira</collectingRegion>
, J. Y. Johnson leg., ovicelled colony. 3d;
<collectionCode id="ED396099FFEFFF85FBFC3EA7FB562A6C" box="[1149,1248,1516,1539]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="402" type="Museum">NHMUK</collectionCode>
1911.10.1.705 (
<taxonomicName id="4C2883DFFFEFFF85FA073EA0FC182A72" authorityName=": Harmelin" authorityYear="1978" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="402" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFEFFF85FA073EA0FC182A72" italics="true" pageId="19" pageNumber="402">Collarina balzaci</emphasis>
</taxonomicName>
),
<collectingRegion id="49EC36BEFFEFFF85FC423D4EFBA12A72" box="[963,1047,1541,1565]" country="Portugal" name="Madeira" pageId="19" pageNumber="402">Madeira</collectingRegion>
, ovicelled colony
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C332ABD7FFEFFF85FCAC3D7EFC042AED" pageId="19" pageNumber="402" type="etymology">
<paragraph id="8B97F85CFFEFFF85FCAC3D7EFC042AED" blockId="19.[813,1455,1589,1666]" pageId="19" pageNumber="402">
ETYMOLOGY. — From Macaronesia, biogeographical area including the
<collectingRegion id="49EC36BEFFEFFF85FCF63D1BFBD02A08" box="[887,1126,1615,1640]" isEnumeration="true" pageId="19" pageNumber="402">Canary Islands, Madeira</collectingRegion>
and the
<collectingRegion id="49EC36BEFFEFFF85FB3C3D1BFB4B2A07" box="[1213,1277,1616,1640]" country="Portugal" name="Acores" pageId="19" pageNumber="402">Azores</collectingRegion>
where this species was recorded.
</paragraph>
</subSubSection>
<subSubSection id="C332ABD7FFEFFF85FCAC3DD2FBAB2B04" pageId="19" pageNumber="402" type="diagnosis">
<paragraph id="8B97F85CFFEFFF85FCAC3DD2FBAB2B04" blockId="19.[813,1456,1689,1899]" pageId="19" pageNumber="402">DIAGNOSIS. — Colonies medium- to large-sized; autozooids with broad marginal gymnocyst, costate shield with a lower central area, predominantly composed of 6-7 costae with smooth surface and long basal part; apertural bar moderately raised, forming an arch; proximal edge of orifice smooth and slightly concave; 3-4 oral spines; paired adventitious avicularia with marked disto-lateral orientation; ooecium formed by distal kenozooid or autozooid; ancestrula with 6 spines.</paragraph>
</subSubSection>
<subSubSection id="C332ABD7FFEFFF83FCAC3CD8FD33285F" lastPageId="21" lastPageNumber="404" pageId="19" pageNumber="402" type="description">
<paragraph id="8B97F85CFFEFFF85FCAC3CD8FC0B2BC4" blockId="19.[813,1455,1939,2030]" box="[813,957,1939,1965]" pageId="19" pageNumber="402">DESCRIPTION</paragraph>
<paragraph id="8B97F85CFFEFFF83FCAC3CF8FD33285F" blockId="19.[813,1455,1939,2030]" lastBlockId="21.[132,775,217,1072]" lastPageId="21" lastPageNumber="404" pageId="19" pageNumber="402">
Colony encrusting, pluriserial, unilaminar, reaching large size (&gt; 1000 zooids) (
<figureCitation id="1313E4D9FFEFFF85FC5F3C98FB8B2B82" box="[990,1085,2003,2029]" captionStart="FIG" captionStartId="30.[132,143,1749,1767]" captionTargetBox="[136,1453,215,1699]" captionTargetId="figure@30.[135,1453,213,1699]" captionTargetPageId="30" captionText="FIG. 18. — Habitats of Collarina species:A, C. balzaci (Audouin,1826), bundles of Posidonia leaves; B, C. speluncola Harmelin,n. sp., wall of dark submarine cave with freshwater seepage; C, C. denticulata Harmelin, n. sp., underside of pottery fragment, Cassis, Port-Miou, 18 m; D, C. gautieri Harmelin, n. sp., underside of Pinna shell, paratype MNHN-IB-2014-1921, Veyron, 24 m; E, C. macaronensis Harmelin, n. sp., underside of pebble, paratype MNHN-IB-2014-1926, Madeira, Porto Santo, Baixo Is.; F, C. fayalensis Harmelin, 1978, underside of pebble, Azores, Saô Miguel, Vila Franca, 15 m. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3725994" httpUri="https://zenodo.org/record/3725994/files/figure.png" pageId="19" pageNumber="402">Fig. 18E</figureCitation>
). Zooids subpentagonal with distal edge rounded. Gymnocyst visible in frontal view, of variable width laterally, clearly much wider in the proximal part, with some gymnocystal pseudopores (
<figureCitation id="1313E4D9FFE9FF83FE693A52FD8D2D5C" box="[488,571,281,307]" captionStart="FIG" captionStartId="4.[167,178,836,854]" captionTargetBox="[379,1214,221,790]" captionTargetId="figure@4.[338,1245,216,826]" captionTargetPageId="4" captionText="FIG. 1. — Savignys drawings (1817: pl. 9). Flustra balzaci Audouin, 1826, in Savignys figure 8, and Flustra jauberti Audouin, 1826, in Savignys figure 9." figureDoi="http://doi.org/10.5281/zenodo.3725958" httpUri="https://zenodo.org/record/3725958/files/figure.png" pageId="21" pageNumber="404">Fig. 8B</figureCitation>
; 12C, H). Costate shield with central part lower than the periphery, composed of 4-9 (6-7:&gt; 60%) short and thick costae, with each ascending basal part forming a clearly distinct lobe, prominent on the gymnocyst and bearing a large pelma, a second, smaller pelma in inner position, before the central, lower part of the shield. Intercostal lacunae 2-3 between adjacent costae, small, irregularly slot-shaped (
<figureCitation id="1313E4D9FFE9FF83FE073AB3FE632E7D" box="[390,469,504,530]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Fig. 12</figureCitation>
D-H). Apertural bar arched below the orifice, with a moderately raised tip, a pair of medium-sized pseudopores on both sides of the tip and two larger lateral pelmata, near the base of the avicularia (
<figureCitation id="1313E4D9FFE9FF83FDF83913FD782E1D" box="[633,718,600,626]" captionStart="FIG" captionStartId="4.[167,178,836,854]" captionTargetBox="[379,1214,221,790]" captionTargetId="figure@4.[338,1245,216,826]" captionTargetPageId="4" captionText="FIG. 1. — Savignys drawings (1817: pl. 9). Flustra balzaci Audouin, 1826, in Savignys figure 8, and Flustra jauberti Audouin, 1826, in Savignys figure 9." figureDoi="http://doi.org/10.5281/zenodo.3725958" httpUri="https://zenodo.org/record/3725958/files/figure.png" pageId="21" pageNumber="404">Figs 8B</figureCitation>
; 12F, G; 15C). Adventitious avicularia typically directed laterodistally, almost always paired, rostrum with slightly hooked tip (
<figureCitation id="1313E4D9FFE9FF83FF0A39FCFF692EBD" box="[139,223,695,722]" captionStart="FIG" captionStartId="4.[167,178,836,854]" captionTargetBox="[379,1214,221,790]" captionTargetId="figure@4.[338,1245,216,826]" captionTargetPageId="4" captionText="FIG. 1. — Savignys drawings (1817: pl. 9). Flustra balzaci Audouin, 1826, in Savignys figure 8, and Flustra jauberti Audouin, 1826, in Savignys figure 9." figureDoi="http://doi.org/10.5281/zenodo.3725958" httpUri="https://zenodo.org/record/3725958/files/figure.png" pageId="21" pageNumber="404">Figs 8B</figureCitation>
; 12B, D, H; 15C), nested cystids relatively frequent (
<figureCitation id="1313E4D9FFE9FF83FF0D399CFEBC2E9D" box="[140,266,727,754]" captionStart="FIG" captionStartId="28.[132,143,1119,1137]" captionTargetBox="[133,1455,216,1071]" captionTargetId="figure@28.[133,1456,214,1071]" captionTargetPageId="28" captionText="FIG. 17. — Adventitious avicularia with nested cystids in Collarina Jullien, 1886 species: A, C. macaronensis Harmelin, n. sp., ovicelled zooid; B, C. gautieri Harmelin, n. sp., four piles of nested avicularian cystids adventitious on two autozooids and an ooecium; C, C. macaronensis Harmelin, n. sp.; D, C. fayalensis Harmelin, 1978; E, C. speluncola Harmelin, n. sp. Origin: A, Madeira, NHMUK 1899.7.1.2105; B, Galicia, Ria de Ferrol, MHNUSC-Bry-113; C, Galicia, Sisargas Is., MHNUSC-Bry-662; D, Azores, Saô Miguel, Vila Franca; E, Marseille, Conger Cave. Scale bar: A, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3725990" httpUri="https://zenodo.org/record/3725990/files/figure.png" pageId="21" pageNumber="404">Fig. 17A, C</figureCitation>
). Orifice wider than long, dimorphic, broader in ovicelled zooids; proximal edge slightly or noticeably concave; condyles triangular, moderately prominent (
<figureCitation id="1313E4D9FFE9FF83FDE8385CFD762F5E" box="[617,704,791,817]" captionStart="FIG" captionStartId="4.[167,178,836,854]" captionTargetBox="[379,1214,221,790]" captionTargetId="figure@4.[338,1245,216,826]" captionTargetPageId="4" captionText="FIG. 1. — Savignys drawings (1817: pl. 9). Flustra balzaci Audouin, 1826, in Savignys figure 8, and Flustra jauberti Audouin, 1826, in Savignys figure 9." figureDoi="http://doi.org/10.5281/zenodo.3725958" httpUri="https://zenodo.org/record/3725958/files/figure.png" pageId="21" pageNumber="404">Figs 8B</figureCitation>
; 12D, F, G). Spines thin, with small conical base,
<quantity id="4CD055B9FFE9FF83FDE0387CFD3B2F3E" box="[609,653,823,849]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="21" pageNumber="404" unit="in" value="3.0">3 in</quantity>
most cases or 4, 2 arched in ovicelled zooids (
<figureCitation id="1313E4D9FFE9FF83FE7E381CFD122F1D" box="[511,676,855,882]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Fig. 12B, D, F</figureCitation>
). Ovicell prominent, seemingly acleithral (
<figureCitation id="1313E4D9FFE9FF83FE62383CFDF22FFD" box="[483,580,887,914]" captionStart="FIG" captionStartId="26.[132,143,1397,1415]" captionTargetBox="[132,1452,215,1351]" captionTargetId="figure@26.[130,1453,214,1351]" captionTargetPageId="26" captionText="FIG. 15. — Types of ooecia in Collarina Jullien, 1886 species: A-C, kenozooidal ooecia with and without distal avicularium (A, C. speluncola Harmelin, n. sp; B, C. fayalensis Harmelin, 1978; C, C. macaronensis Harmelin, n. sp.); D, C. denticulata Harmelin, n. sp., colony portion with two kenozooidal ooecia (left) and three ooecia incorporated in distal autozooid (right); E, C. macaronensis Harmelin, n. sp., 2 ovicelled zooids with kenozooidal ooecium (left) vs ooecium incorporated in distal autozooid (right). Origin: A, Marseille, Conger Cave; B, Azores, Saô Miguel, Vila Franca Is.; C, E, Madeira, NHMUK 1911.10.1.705; D, Catalonia, Medes Is. Scale bars: A-C, E 100 µm, D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3725986" httpUri="https://zenodo.org/record/3725986/files/figure.png" pageId="21" pageNumber="404">Fig. 15C</figureCitation>
); ooecium formed by a distal kenozooid at the colony growing edge or by a distal daughter autozooid (
<figureCitation id="1313E4D9FFE9FF83FEE638FCFE002FBE" box="[359,438,951,977]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Figs 12</figureCitation>
A-C; 15C, E), slightly broader than long, ectooecium with relatively smooth surface, punctured with a dozen medium-sized pseudopores. Ancestrula cribrimorph, with small spinocyst and 6 spines.
</paragraph>
</subSubSection>
<caption id="DF57A8D4FFE8FF82FF073C23FC1A2B84" ID-DOI="http://doi.org/10.5281/zenodo.3725980" ID-Zenodo-Dep="3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="20" pageNumber="403" startId="20.[134,145,1896,1914]" targetBox="[134,1457,215,1743]" targetPageId="20">
<paragraph id="8B97F85CFFE8FF82FF073C23FC1A2B84" blockId="20.[134,1459,1896,2027]" pageId="20" pageNumber="403">
FIG. 12. —
<taxonomicName id="4C2883DFFFE8FF82FF5E3C23FE472B15" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[223,497,1896,1914]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="403" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE8FF82FF5E3C23FE162B15" box="[223,416,1896,1914]" italics="true" pageId="20" pageNumber="403">Collarina macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE8FF82FE793C23FD9C2B15" box="[504,554,1896,1914]" pageId="20" pageNumber="403" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B95C244EFFE8FF82FDB13C23FD8B2B15" bold="true" box="[560,573,1896,1914]" pageId="20" pageNumber="403">A</emphasis>
,
<emphasis id="B95C244EFFE8FF82FDC73C23FDE52B15" bold="true" box="[582,595,1896,1914]" pageId="20" pageNumber="403">B</emphasis>
, colony edge,general view and detail with ovicelled and non-ovicelled zooids;
<emphasis id="B95C244EFFE8FF82FB5F3C23FB5A2B15" bold="true" box="[1246,1260,1896,1914]" pageId="20" pageNumber="403">C</emphasis>
, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;
<emphasis id="B95C244EFFE8FF82FDF03C34FDC92BFE" bold="true" box="[625,639,1919,1937]" pageId="20" pageNumber="403">D</emphasis>
,
<emphasis id="B95C244EFFE8FF82FD093C34FD202BFE" bold="true" box="[648,662,1919,1937]" pageId="20" pageNumber="403">G</emphasis>
, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst;
<emphasis id="B95C244EFFE8FF82FC703CDDFC4B2BC7" bold="true" box="[1009,1021,1942,1960]" pageId="20" pageNumber="403">E</emphasis>
,
<taxonomicName id="4C2883DFFFE8FF82FB893CDEFAAA2BC8" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[1032,1308,1941,1959]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="403" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE8FF82FB893CDEFB262BC8" box="[1032,1168,1941,1959]" italics="true" pageId="20" pageNumber="403">Collarina balzaci</emphasis>
(
<bibRefCitation id="EFB985ADFFE8FF82FB1B3CDEFAA12BC8" author="AUDOUIN V." box="[1178,1303,1941,1959]" pageId="20" pageNumber="403" pagination="225 - 244" refId="ref25877" refString="AUDOUIN V. 1826. - Explication sommaire des planches de polypes de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, membre de l'Institut; offrant un expose des caracteres naturels des genres avec la distinction des especes, in JOMARD E. F. (ed.), Description de l'Egypte. Histoire naturelle. Imprimerie nationale, Paris: 225 - 244. http: // bibliotheques. mnhn. fr / EXPLOITATION / infodoc / digitalCollections / viewerpopup. aspx? seid = MNHN _ FOLRES 43" type="book chapter" year="1826">Audouin, 1826</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFB985ADFFE8FF82FAA23CDEFA192BC8" author="HARMELIN J. G." box="[1315,1455,1941,1959]" pageId="20" pageNumber="403" pagination="173 - 192" refId="ref28028" refString="HARMELIN J. G. 1978 a. - Sur quelques cribrimorphes (Bryozoa, Cheilostomata) de l'Atlantique oriental. Tethys 8: 173 - 192." type="journal article" year="1978">Harmelin (1978a</bibRefCitation>
, fig. 8), Azores, Faial;
<emphasis id="B95C244EFFE8FF82FEB53CE7FE892BD1" bold="true" box="[308,319,1964,1982]" pageId="20" pageNumber="403">F</emphasis>
, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines;
<emphasis id="B95C244EFFE8FF82FA8C3CE7FAAD2BD1" bold="true" box="[1293,1307,1964,1982]" pageId="20" pageNumber="403">H</emphasis>
, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm.
</paragraph>
</caption>
<subSubSection id="C332ABD7FFE9FF83FF053F1EFC562E5C" pageId="21" pageNumber="404" type="discussion">
<paragraph id="8B97F85CFFE9FF83FF053F1EFF5E2801" blockId="21.[132,775,1109,2030]" box="[132,232,1109,1135]" pageId="21" pageNumber="404">REMARKS</paragraph>
<paragraph id="8B97F85CFFE9FF83FF053F3DFE8A2B01" blockId="21.[132,775,1109,2030]" pageId="21" pageNumber="404">
<taxonomicName id="4C2883DFFFE9FF83FF053F3DFE5D28FF" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[132,491,1142,1168]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FF053F3DFECE28FF" box="[132,376,1142,1168]" italics="true" pageId="21" pageNumber="404">Collarina macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FE763F3CFD8428FE" box="[503,562,1143,1169]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
is closely related to
<taxonomicName id="4C2883DFFFE9FF83FF053FDDFEB428DF" authorityName="Harmelin" authorityYear="1978" box="[132,258,1174,1200]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="fayalensis">
<emphasis id="B95C244EFFE9FF83FF053FDDFEB428DF" box="[132,258,1174,1200]" italics="true" pageId="21" pageNumber="404">C. fayalensis</emphasis>
</taxonomicName>
in having orifices with a concave proximal edge, costae with similarly shaped ascending base bearing a large pelma and well-developed gymnocyst. However, in
<taxonomicName id="4C2883DFFFE9FF83FD213F9DFE8D2960" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FD213F9DFF7D2960" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FEC53FBDFEC8297F" box="[324,382,1270,1296]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
, the paired adventitious avicularia are more distinctly directed disto-laterally and the costate shield involves more costae. Specimens from Faial,
<collectingRegion id="49EC36BEFFE9FF83FDDC3E7DFD12293F" box="[605,676,1334,1360]" country="Portugal" name="Acores" pageId="21" pageNumber="404">Azores</collectingRegion>
recorded and figured as
<taxonomicName id="4C2883DFFFE9FF83FEA33E1EFE632900" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[290,469,1365,1391]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FEA33E1EFE632900" box="[290,469,1365,1391]" italics="true" pageId="21" pageNumber="404">Collarina balzaci</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB985ADFFE9FF83FE7F3E1EFD0C2900" author="HARMELIN J. G." box="[510,698,1365,1391]" pageId="21" pageNumber="404" pagination="173 - 192" refId="ref28028" refString="HARMELIN J. G. 1978 a. - Sur quelques cribrimorphes (Bryozoa, Cheilostomata) de l'Atlantique oriental. Tethys 8: 173 - 192." type="journal article" year="1978">Harmelin (1978a</bibRefCitation>
: fig. 8; reproduced here: fig 12E) present these typical characters and belong undoubtedly to
<taxonomicName id="4C2883DFFFE9FF83FEFD3EDEFD2429C0" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[380,658,1429,1455]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FEFD3EDEFD9429C0" box="[380,546,1429,1455]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FD1D3EDDFD6329DF" box="[668,725,1430,1456]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
This Azorean sample consisted in two colonies on a pebble bearing also the
<typeStatus id="549346FEFFE9FF83FE873E9EFE822980" box="[262,308,1493,1519]" pageId="21" pageNumber="404">type</typeStatus>
series of
<emphasis id="B95C244EFFE9FF83FE163E9EFDA92980" box="[407,543,1493,1519]" italics="true" pageId="21" pageNumber="404">
<taxonomicName id="4C2883DFFFE9FF83FE163E9EFDAD2980" authorityName="Harmelin" authorityYear="1978" box="[407,539,1493,1519]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="fayalensis">C. fayalensis</taxonomicName>
.
</emphasis>
The co-occurrence of these two species on the same small substrate confirms they are not ecomorphs of the same species. The discriminant features of
<taxonomicName id="4C2883DFFFE9FF83FF5A3D7EFE432A21" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[219,501,1588,1615]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FF5A3D7EFE352A21" box="[219,387,1589,1615]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FE7E3D7EFD8F2A20" box="[511,569,1589,1615]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
are recognizable in part of the figured material from
<collectingRegion id="49EC36BEFFE9FF83FE6A3D1FFD3D2A01" box="[491,651,1620,1646]" country="Spain" name="Canarias" pageId="21" pageNumber="404">Canary Islands</collectingRegion>
ascribed to
<taxonomicName id="4C2883DFFFE9FF83FF053D3EFEB22AE1" authorityName="Harmelin" authorityYear="1978" box="[132,260,1652,1679]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="fayalensis">
<emphasis id="B95C244EFFE9FF83FF053D3EFEB22AE1" box="[132,260,1652,1679]" italics="true" pageId="21" pageNumber="404">C. fayalensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB985ADFFE9FF83FEAF3D3FFE612AE1" author="ARISTEGUI RUIZ J." box="[302,471,1652,1678]" pageId="21" pageNumber="404" pagination="1 - 524" refId="ref25832" refString="ARISTEGUI RUIZ J. 1984. - Briozoos quilostomados (Ectoprocta, Cheilostomata) de Canarias: Estudio sistematico, faunistico y biogeografico. Tesis Doctoral, Facultad de Biologia, Universidad de La Laguna, Las Palma, i-iii, 1 - 524." type="book chapter" year="1984">Arístegui (1984</bibRefCitation>
: fig. 49b; pl. 11, fig. 6) and also in figures of colonies ascribed to
<taxonomicName id="4C2883DFFFE9FF83FD983DDFFD502AC1" box="[537,742,1684,1710]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="punctata">
<emphasis id="B95C244EFFE9FF83FD983DDFFD502AC1" box="[537,742,1684,1710]" italics="true" pageId="21" pageNumber="404">Cribrilina punctata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB985ADFFE9FF83FF053DFFFDCF2AA1" author="FERNANDEZ PULPEIRO E. &amp; RODRIGUEZ BABIO C." box="[132,633,1716,1742]" pageId="21" pageNumber="404" pagination="119 - 168" refId="ref27585" refString="FERNANDEZ PULPEIRO E. &amp; RODRIGUEZ BABIO C. 1980. - Aportaciones al conocimiento de la fauna briozoologica del litoral de la ria de Vigo. Investigacion Pesquera 44 (1): 119 - 168." type="journal article" year="1980">Fernández Pulpeiro &amp; Rodriguez Babio (1980</bibRefCitation>
:
<collectingRegion id="49EC36BEFFE9FF83FD093DFFFD612AA1" box="[648,727,1716,1742]" country="Spain" name="Galicia" pageId="21" pageNumber="404">Galicia</collectingRegion>
, Ria de Vigo) and
<bibRefCitation id="EFB985ADFFE9FF83FE9F3D98FE0F2A81" author="ALVAREZ J. A." box="[286,441,1747,1775]" pageId="21" pageNumber="404" pagination="347 - 352" refId="ref25800" refString="ALVAREZ J. A. 1988. - La familia Cribrilinidae Hincks, 1880 (Bryozoa, Cheilostomida) en la Costa Vasca. Miscellanea Zoologica 12: 347 - 352." type="journal article" year="1988">Álvarez (1988</bibRefCitation>
: Basque region). SEM examination of specimens from Sisargas Islands,
<collectingRegion id="49EC36BEFFE9FF83FDD83DBFFD132B61" box="[601,677,1780,1806]" country="Spain" name="Galicia" pageId="21" pageNumber="404">Galicia</collectingRegion>
(coll. by Fernández Pulpeiro, courtesy of O. Reverter) confirmed the occurrence of
<taxonomicName id="4C2883DFFFE9FF83FE993C7FFD9A2B22" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[280,556,1843,1870]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FE993C7FFE0A2B22" box="[280,444,1844,1870]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FDB43C7FFDD82B21" box="[565,622,1844,1870]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
on the Iberian coast (
<figureCitation id="1313E4D9FFE9FF83FF483C18FE862B01" box="[201,304,1875,1902]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Fig. 12H</figureCitation>
).
</paragraph>
<paragraph id="8B97F85CFFE9FF83FF053C38FC562E5C" blockId="21.[132,775,1109,2030]" lastBlockId="21.[813,1456,217,563]" pageId="21" pageNumber="404">
SEM examination of three specimens kept at the NHMUK, collected by J. Y. Johnson at
<collectingRegion id="49EC36BEFFE9FF83FE383CD8FDA22BC2" box="[441,532,1939,1965]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
and labelled
<taxonomicName id="4C2883DFFFE9FF83FD233CD8FF642BA2" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FD233CD8FF642BA2" italics="true" pageId="21" pageNumber="404">Collarina balzaci</emphasis>
</taxonomicName>
or
<taxonomicName id="4C2883DFFFE9FF83FF7F3CF8FEDE2BA2" authorityName="Jullien" authorityYear="1886" box="[254,360,1971,1997]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="genus">
<emphasis id="B95C244EFFE9FF83FF7F3CF8FEDE2BA2" box="[254,360,1971,1997]" italics="true" pageId="21" pageNumber="404">Collarina</emphasis>
</taxonomicName>
sp., has proved that they belong to
<taxonomicName id="4C2883DFFFE9FF83FF053C98FE292B82" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[132,415,2003,2029]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FF053C98FE9A2B82" box="[132,300,2003,2029]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FE283C9FFE552B81" box="[425,483,2004,2030]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="1313E4D9FFE9FF83FE713C98FDD92B81" box="[496,623,2003,2030]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Fig. 12A, B</figureCitation>
). As indicated by
<bibRefCitation id="EFB985ADFFE9FF83FCCE3B91FC482C9B" author="NORMAN A. M." box="[847,1022,217,244]" pageId="21" pageNumber="404" pagination="275 - 314" refId="ref29247" refString="NORMAN A. M. 1909. - The Polyzoa of Madeira and neighbouring islands. Journal of the Linnean Society Zoology 30: 275 - 314, pl. 33 - 42. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 02407. x" type="journal article" year="1909">Norman (1909)</bibRefCitation>
in his introduction the Johnson collection was examined by G. Busk,
<bibRefCitation id="EFB985ADFFE9FF83FB073BB1FA942D7B" author="WATERS A. W." box="[1158,1314,249,276]" pageId="21" pageNumber="404" pagination="6 - 16" refId="ref30475" refString="WATERS A. W. 1899. - Bryozoa from Madeira. Journal of the Royal Microscopical Society 1899: 6 - 16. https: // doi. org / 10.1111 / j. 1365 - 2818.1899. tb 00139. x" type="journal article" year="1899">Waters (1899)</bibRefCitation>
and himself.
<bibRefCitation id="EFB985ADFFE9FF83FCAC3A51FC7B2D5B" author="WATERS A. W." box="[813,973,281,308]" pageId="21" pageNumber="404" pagination="6 - 16" refId="ref30475" refString="WATERS A. W. 1899. - Bryozoa from Madeira. Journal of the Royal Microscopical Society 1899: 6 - 16. https: // doi. org / 10.1111 / j. 1365 - 2818.1899. tb 00139. x" type="journal article" year="1899">Waters (1899)</bibRefCitation>
ascribed this material to
<taxonomicName id="4C2883DFFFE9FF83FB6C3A52FA062D5C" box="[1261,1456,281,307]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FB6C3A52FA062D5C" box="[1261,1456,281,307]" italics="true" pageId="21" pageNumber="404">Cribrilina balzaci</emphasis>
</taxonomicName>
and noted thereafter (
<bibRefCitation id="EFB985ADFFE9FF83FBA33A72FB182D3C" author="WATERS A. W." box="[1058,1198,313,339]" pageId="21" pageNumber="404" pagination="545 - 573" refId="ref30518" refString="WATERS A. W. 1923. - Mediterranean and other Cribrilinidae, together with their relationship to Cretaceous forms. Annals and Magazine of Natural History, Ninth Series 12: 545 - 573. https: // doi. org / 10.1080 / 00222932308632977" type="book chapter" year="1923">Waters 1923</bibRefCitation>
) that “There are specimens from
<collectingRegion id="49EC36BEFFE9FF83FC2D3A12FBBC2D1C" box="[940,1034,345,371]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
, which I took for
<taxonomicName id="4C2883DFFFE9FF83FB553A12FAA82D1C" box="[1236,1310,345,371]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FB553A12FAA82D1C" box="[1236,1310,345,371]" italics="true" pageId="21" pageNumber="404">balzaci</emphasis>
</taxonomicName>
, which must perhaps be considered a variety”. The figure of the spinocyst edge of a Madeiran specimen named
<taxonomicName id="4C2883DFFFE9FF83FB4C3AD3FA3A2DDD" box="[1229,1420,408,434]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FB4C3AD3FA3A2DDD" box="[1229,1420,408,434]" italics="true" pageId="21" pageNumber="404">Cribrilina balzaci</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB985ADFFE9FF83FCAC3AF2FC622DBC" author="NORMAN A. M." box="[813,980,441,467]" pageId="21" pageNumber="404" pagination="87 - 128" refId="ref29204" refString="NORMAN A. M. 1903. - Notes on the natural history of East Finmark. Annals and Magazine of Natural History Series 7 (12): 87 - 128. https: // doi. org / 10.1080 / 00222930308678831" type="journal article" year="1903">Norman (1903</bibRefCitation>
: pl. 9, fig. 6; same specimen shown here as
<figureCitation id="1313E4D9FFE9FF83FCAC3A93FC382D9D" box="[813,910,472,498]" captionStart="FIG" captionStartId="20.[134,145,1896,1914]" captionTargetBox="[134,1457,215,1743]" captionTargetId="figure@20.[134,1457,214,1743]" captionTargetPageId="20" captionText="FIG. 12. — Collarina macaronensis Harmelin, n. sp.: A, B, colony edge,general view and detail with ovicelled and non-ovicelled zooids; C, oblique view: structure of costate shield,apertural bar,paired and apical avicularia;D, G, non-ovicelled zooids with typical traits: concave orifice poster,avicularia directed disto-laterally, costate shield with lower central part and digitate margin, large pelmata also present on the gymnocyst; E, Collarina balzaci (Audouin, 1826): Harmelin (1978a, fig. 8), Azores, Faial; F, distal part of non-ovicelled zooid with typical orifice, apertural bar with large and small pseudopores, and 3 spines; H, specimen from a continental area. Origin: A, B, NHMUK 1899.7.1.2105, Madeira, J.Y. Johnson leg.; C, NHMUK 1911.10.1.705, Madeira; D, F, G, MNHN-IB-2014-1927, Madeira, Baixo Is.; H, Galicia, Sisargas Is. Scale bars: A, 400 µm; B, C, 200 µm; D, H 100 µm; F, G, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3725980" httpUri="https://zenodo.org/record/3725980/files/figure.png" pageId="21" pageNumber="404">Fig. 12C</figureCitation>
) shows costae with basal parts protruding greatly from the gymnocyst, a typical character of
<taxonomicName id="4C2883DFFFE9FF83FA843AB2FC2C2E5D" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FA843AB2FA192E7D" box="[1285,1455,505,531]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FC243952FC562E5C" box="[933,992,537,563]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C332ABD7FFE9FF83FCAC3913FBC4283F" pageId="21" pageNumber="404" type="distribution">
<paragraph id="8B97F85CFFE9FF83FCAC3913FB022E1D" blockId="21.[813,1456,600,1105]" box="[813,1204,600,626]" pageId="21" pageNumber="404">
<emphasis id="B95C244EFFE9FF83FCAC3913FB022E1D" box="[813,1204,600,626]" italics="true" pageId="21" pageNumber="404">Habitat and geographical distribution</emphasis>
</paragraph>
<paragraph id="8B97F85CFFE9FF83FCAC3933FBC4283F" blockId="21.[813,1456,600,1105]" pageId="21" pageNumber="404">
In both the
<collectingRegion id="49EC36BEFFE9FF83FC313933FC412EFD" box="[944,1015,632,658]" country="Portugal" name="Acores" pageId="21" pageNumber="404">Azores</collectingRegion>
and
<collectingRegion id="49EC36BEFFE9FF83FBAF3933FB3E2EFD" box="[1070,1160,632,658]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
<taxonomicName id="4C2883DFFFE9FF83FB0E3933FA1D2EFD" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[1167,1451,632,658]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FB0E3933FA8E2EFE" box="[1167,1336,632,658]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FCAC39D3FCD32EDD" box="[813,869,664,690]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
was found on pebbles and cobbles in shaded conditions (
<figureCitation id="1313E4D9FFE9FF83FCB539FCFC232EBD" box="[820,917,695,722]" captionStart="FIG" captionStartId="30.[132,143,1749,1767]" captionTargetBox="[136,1453,215,1699]" captionTargetId="figure@30.[135,1453,213,1699]" captionTargetPageId="30" captionText="FIG. 18. — Habitats of Collarina species:A, C. balzaci (Audouin,1826), bundles of Posidonia leaves; B, C. speluncola Harmelin,n. sp., wall of dark submarine cave with freshwater seepage; C, C. denticulata Harmelin, n. sp., underside of pottery fragment, Cassis, Port-Miou, 18 m; D, C. gautieri Harmelin, n. sp., underside of Pinna shell, paratype MNHN-IB-2014-1921, Veyron, 24 m; E, C. macaronensis Harmelin, n. sp., underside of pebble, paratype MNHN-IB-2014-1926, Madeira, Porto Santo, Baixo Is.; F, C. fayalensis Harmelin, 1978, underside of pebble, Azores, Saô Miguel, Vila Franca, 15 m. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3725994" httpUri="https://zenodo.org/record/3725994/files/figure.png" pageId="21" pageNumber="404">Fig. 18E</figureCitation>
). On the other hand, its absence on rocky walls of dark caves at
<collectingRegion id="49EC36BEFFE9FF83FC41399CFBAC2E9E" box="[960,1050,727,753]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
despite an extensive sampling survey (JGH,
<date id="FF96DE9CFFE9FF83FCFB39BCFB9A2F7E" box="[890,1068,759,785]" pageId="21" pageNumber="404" value="2000-09">September 2000</date>
) suggests that this habitat is unsuitable to this
<taxonomicName id="4C2883DFFFE9FF83FC25385CFBB12F5E" authorityName="Jullien" authorityYear="1886" box="[932,1031,791,817]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="genus">
<emphasis id="B95C244EFFE9FF83FC25385CFBB12F5E" box="[932,1031,791,817]" italics="true" pageId="21" pageNumber="404">Collarina</emphasis>
</taxonomicName>
species, in contrast to several
<taxonomicName id="4C2883DFFFE9FF83FABC385CFA192F5E" box="[1341,1455,791,817]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilaria" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="genus">
<emphasis id="B95C244EFFE9FF83FABC385CFA192F5E" box="[1341,1455,791,817]" italics="true" pageId="21" pageNumber="404">Cribrilaria</emphasis>
</taxonomicName>
species which were very abundant (JGH, unpublished data). Small substrates in contact with a flat bottom at shallow depth may be an ecological requirement of this species. It is worth noting that this species was noticed by
<bibRefCitation id="EFB985ADFFE9FF83FA8638DCFA1F2FDE" author="NORMAN A. M." box="[1287,1449,919,945]" pageId="21" pageNumber="404" pagination="275 - 314" refId="ref29247" refString="NORMAN A. M. 1909. - The Polyzoa of Madeira and neighbouring islands. Journal of the Linnean Society Zoology 30: 275 - 314, pl. 33 - 42. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 02407. x" type="journal article" year="1909">Norman (1909</bibRefCitation>
, as
<taxonomicName id="4C2883DFFFE9FF83FCC838FCFC052FBE" box="[841,947,951,977]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="balzaci">
<emphasis id="B95C244EFFE9FF83FCC838FCFC052FBE" box="[841,947,951,977]" italics="true" pageId="21" pageNumber="404">C. balzaci</emphasis>
</taxonomicName>
) on stones between tide-marks at
<collectingRegion id="49EC36BEFFE9FF83FAAD38FCFA3C2FBE" box="[1324,1418,951,977]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
. At the present state of knowledge, the geographical range of
<taxonomicName id="4C2883DFFFE9FF83FCAC38BCFBFD287F" authority="Harmelin" authorityName="Harmelin" authorityYear="2019" box="[813,1099,1014,1041]" class="Gymnolaemata" family="Cribrilinidae" genus="Collarina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="404" phylum="Bryozoa" rank="species" species="macaronensis" status="sp. nov.">
<emphasis id="B95C244EFFE9FF83FCAC38BCFC61287F" box="[813,983,1015,1041]" italics="true" pageId="21" pageNumber="404">C. macaronensis</emphasis>
Harmelin
</taxonomicName>
,
<taxonomicNameLabel id="A26F9935FFE9FF83FBD938BCFB23287E" box="[1112,1173,1015,1041]" pageId="21" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
includes the NE Atlantic archipelagoes (
<collectingRegion id="49EC36BEFFE9FF83FC4B3F5DFB9A285F" box="[970,1068,1046,1072]" country="Spain" name="Canarias" pageId="21" pageNumber="404">Canaries</collectingRegion>
,
<collectingRegion id="49EC36BEFFE9FF83FBB63F5DFB22285F" box="[1079,1172,1046,1072]" country="Portugal" name="Madeira" pageId="21" pageNumber="404">Madeira</collectingRegion>
,
<collectingRegion id="49EC36BEFFE9FF83FB1E3F5CFB5E285E" box="[1183,1256,1047,1073]" country="Portugal" name="Acores" pageId="21" pageNumber="404">Azores</collectingRegion>
) and the northern coast of the Iberian Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>