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<document id="3E760B38F715D4CFCD4A41FE936203B3" ID-DOI="10.5281/zenodo.5355008" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355008" ID-ZooBank="DF60AAD1-1B35-4328-9131-E3ADFC06B6AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630214904780" checkinUser="tatiana" docAuthor="Komai, Tomoyuki &amp; Rahayu, Dwi Listyo" docDate="2014" docId="03810513FF92015503C983FAFAC1FA54" docLanguage="en" docName="62rbz620-646.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Pagurus pergranulatus" docType="treatment" docVersion="3" lastPageNumber="629" masterDocId="FFB87D6BFF91015C005C872FFFADFFC1" masterDocTitle="New records and new species of the hermit crab genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Philippines" masterLastPageNumber="646" masterPageNumber="620" pageNumber="623" updateTime="1699189036804" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4C895080F3F51635E875560E369DA2D4">New records and new species of the hermit crab genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Philippines</mods:title>
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<mods:name id="A469B834B949C1802F22ACDBD1581A95" type="personal">
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<mods:namePart id="A39822106FCFF2A8CAB9F4870C057810">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="3A8C34F0B8151389BAB76CC22019F894">Rahayu, Dwi Listyo</mods:namePart>
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<mods:title id="DE062C03A8EF3A7D29CBC2418AC7758D">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="E81F5E569D4C234BB8C89AD9CD65C6A2">2014</mods:date>
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<mods:number id="EEC9AA9DA8464EB1FE29175956399DAC">2014-09-10</mods:number>
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<mods:number id="63F1804C84E481A83334580A3C293C7F">62</mods:number>
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<mods:classification id="BCACE2FDD5F98A42EF3F201336D6323B">journal article</mods:classification>
<mods:identifier id="DC218C418673AA5C13AB4A0605711239" type="DOI">10.5281/zenodo.5355008</mods:identifier>
<mods:identifier id="22AB8DA23F09ECB7EC6D722F9F2C1FDE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E0E1331032670A3D5AA88033779DD485" type="Zenodo-Dep">5355008</mods:identifier>
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<treatment id="03810513FF92015503C983FAFAC1FA54" LSID="urn:lsid:plazi:treatment:03810513FF92015503C983FAFAC1FA54" httpUri="http://treatment.plazi.org/id/03810513FF92015503C983FAFAC1FA54" lastPageId="9" lastPageNumber="629" pageId="3" pageNumber="623">
<subSubSection id="C332E78EFF92015F03C983FAFADDFB2C" box="[917,1392,1237,1261]" pageId="3" pageNumber="623" type="nomenclature">
<paragraph id="8B97B405FF92015F03C983FAFADDFB2C" blockId="3.[917,1392,1237,1293]" box="[917,1392,1237,1261]" pageId="3" pageNumber="623">
<heading id="D0DF0369FF92015F03C983FAFADDFB2C" bold="true" box="[917,1392,1237,1261]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="623" reason="2">
<taxonomicName id="4C28CF86FF92015F03C983FAFADDFB2C" ID-CoL="6ea5435d-03c4-41f0-b678-4b7940b9fbcc" authority="(Henderson, 1896)" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[917,1392,1237,1261]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF92015F03C983FAFADDFB2C" bold="true" box="[917,1392,1237,1261]" pageId="3" pageNumber="623">
<emphasis id="B95C6817FF92015F03C983FAFB3BFB2C" bold="true" box="[917,1174,1237,1261]" italics="true" pageId="3" pageNumber="623">Pagurus pergranulatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF92015F04F983FAFAC4FB2C" author="Henderson JR" box="[1189,1385,1237,1261]" pageId="3" pageNumber="623" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF92015F046C83DAFB7BFACC" box="[1072,1238,1269,1293]" pageId="3" pageNumber="623" type="description">
<paragraph id="8B97B405FF92015F046C83DAFB7BFACC" blockId="3.[917,1392,1237,1293]" box="[1072,1238,1269,1293]" pageId="3" pageNumber="623">
(
<figureCitation id="1313A880FF92015F046483DAFB35FACC" box="[1080,1176,1269,1293]" captionStart="Fig" captionStartId="2.[113,148,1976,1998]" captionTargetBox="[199,1368,622,1952]" captionTargetId="figure-372@2.[199,1368,622,1952]" captionTargetPageId="2" captionText="Fig. 1. A, Pagurus fungiformis Komai &amp; Rahayu, 2004, male (sl 2.7 mm), PANGLAO 2004, stn M51, Mayacabac, Panglao Island, ZRC 2013.0645; B, Pagurus kulkarnii Sankolli, 1962, male (sl mm), PANGLAO 2004, stn R66, Sungcolan inlet, Panglao Island, ZRC 2014.0300; C, Pagurus moluccensis Haig &amp; Ball, 1988, male (sl 3.2 mm), PANGLAO 2004, stn M2, west end of Alona Beach, Panglao Island, ZRC 2013.0644; D, Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301." figureDoi="http://doi.org/10.5281/zenodo.5355010" httpUri="https://zenodo.org/record/5355010/files/figure.png" pageId="3" pageNumber="623">Figs. 1D</figureCitation>
,
<figureCitation id="1313A880FF92015F04F983DAFB60FACC" box="[1189,1229,1269,1293]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="5.[136,171,2004,2026]" captionStartId-1="6.[113,148,2005,2027]" captionStartId-2="7.[136,171,2034,2056]" captionStartId-3="8.[113,148,2034,2056]" captionTargetBox-0="[150,1460,350,1966]" captionTargetBox-1="[138,1441,241,1964]" captionTargetBox-2="[147,1464,979,1994]" captionTargetBox-3="[124,1439,633,1994]" captionTargetId-0="figure-8@5.[127,1482,328,1987]" captionTargetId-1="figure-10@6.[105,1459,212,1985]" captionTargetId-2="figure-8@7.[113,1510,957,2013]" captionTargetId-3="figure-144@8.[98,1467,614,2005]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." captionText-1="Fig. 3. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right chela, dorsal view; B, right cheliped, mesial view (setae omitted); C, same, lateral view; D, carpus of right cheliped, dorsal view; E, close up of capsulate tubercle on right palm, dorsal view (setae omitted); F, same, obliquely lateral view. Scale bars = 2 mm for A–D; 0.5 mm for E, F." captionText-2="Fig. 4. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, chela and carpus of left cheliped, dorsal view; B, left cheliped, mesial view (setae omitted); C, same, lateral view. Scale bar = 2 mm." captionText-3="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5355012" figureDoi-1="http://doi.org/10.5281/zenodo.5355014" figureDoi-2="http://doi.org/10.5281/zenodo.5355016" figureDoi-3="http://doi.org/10.5281/zenodo.5355018" httpUri-0="https://zenodo.org/record/5355012/files/figure.png" httpUri-1="https://zenodo.org/record/5355014/files/figure.png" httpUri-2="https://zenodo.org/record/5355016/files/figure.png" httpUri-3="https://zenodo.org/record/5355018/files/figure.png" pageId="3" pageNumber="623">2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF92015F031F821BFA6EFA1B" pageId="3" pageNumber="623" type="reference_group">
<paragraph id="8B97B405FF92015F031F821BFA6FFA43" blockId="3.[835,1475,1332,1498]" pageId="3" pageNumber="623">
<treatmentCitationGroup id="AB38932BFF92015F031F821BFA6FFA43" pageId="3" pageNumber="623">
<treatmentCitation id="0A899214FF92015F031F821BFA81FA8B" author="Henderson JR" box="[835,1324,1332,1354]" page="520" pageId="3" pageNumber="623" year="1896">
<taxonomicName id="4C28CF86FF92015F031F821BFA81FA8B" ID-CoL="fa1499c4-b547-4307-a965-19734a08eace" authority="Henderson, 1896: 520" authorityName="Henderson" authorityPageNumber="520" authorityYear="1896" box="[835,1324,1332,1354]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF92015F031F821BFBEFFA88" box="[835,1090,1332,1353]" italics="true" pageId="3" pageNumber="623">Eupagurus pergranulatus</emphasis>
<bibRefCitation id="EFB9C9F4FF92015F0410821BFA81FA8B" author="Henderson JR" box="[1100,1324,1332,1354]" pageId="3" pageNumber="623" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896: 520</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="54930AA7FF92015F0567821BFAC5FA88" box="[1339,1384,1332,1353]" pageId="3" pageNumber="623">type</typeStatus>
locality: off east coast of
<collectingCountry id="F33FF495FF92015F044D827FFBFBFAA7" box="[1041,1110,1360,1382]" name="Sri Lanka" pageId="3" pageNumber="623">Ceylon</collectingCountry>
(
<collectingCountry id="F33FF495FF92015F043A827FFB61FAA7" box="[1126,1228,1360,1382]" name="Sri Lanka" pageId="3" pageNumber="623">Sri Lanka</collectingCountry>
), 28 fathoms];
<bibRefCitation id="EFB9C9F4FF92015F053E827FFBA9FA43" author="Alcock A &amp; Anderson AR" pageId="3" pageNumber="623" pagination="28 - 32" refId="ref16412" refString="Alcock A &amp; Anderson AR (1897) Crustacea, Part V. In: Illustrations of the Zoology of the Royal Indian Marine Surveying Steamer Investigator under the Command of Commander C. F. Oldham, R. N. Trustees of the Indian Museum, Calcutta. Pls. 28 - 32." type="book chapter" year="1897">Alcock &amp; Anderson, 1897</bibRefCitation>
: pl. 31, fig. 1;
<treatmentCitation id="0A899214FF92015F04DA8242FAE9FA42" author="Alcock A" box="[1158,1348,1389,1411]" page="125" pageId="3" pageNumber="623" year="1905">
<bibRefCitation id="EFB9C9F4FF92015F04DA8242FAE9FA42" author="Alcock A" box="[1158,1348,1389,1411]" pageId="3" pageNumber="623" pagination="1 - 197" refId="ref16372" refString="Alcock A (1905 b) Anomura, Fascicules 1, Pagurides. Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, 2: i - xi, 1 - 197, 16 pls." type="book chapter" year="1905">Alcock, 1905b: 125</bibRefCitation>
</treatmentCitation>
, pl. 11, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B97B405FF92015F031F82A4FA6EFA1B" blockId="3.[835,1475,1332,1498]" pageId="3" pageNumber="623">
<treatmentCitationGroup id="AB38932BFF92015F031F82A4FA6EFA1B" pageId="3" pageNumber="623">
<taxonomicName id="4C28CF86FF92015F031F82A4FB8EFA5E" ID-CoL="6ea5435d-03c4-41f0-b678-4b7940b9fbcc" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[835,1059,1418,1440]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF92015F031F82A4FB8EFA5E" box="[835,1059,1418,1440]" italics="true" pageId="3" pageNumber="623">Pagurus pergranulatus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A899214FF92015F046182A5FB59FA5E" author="Gordan J" box="[1085,1268,1418,1440]" page="333" pageId="3" pageNumber="623" year="1956">
<bibRefCitation id="EFB9C9F4FF92015F046182A5FB59FA5E" author="Gordan J" box="[1085,1268,1418,1440]" pageId="3" pageNumber="623" pagination="253 - 352" refId="ref17065" refString="Gordan J (1956) A bibliography of pagurid crabs, exclusive of Alcock, 1905. Bulletin of the American Museum of Natural History, 108: 253 - 352." type="journal article" year="1956">Gordan, 1956: 333</bibRefCitation>
</treatmentCitation>
(bibliography);
<treatmentCitation id="0A899214FF92015F05CF82A5FB88FA7D" author="Haig J &amp; Ball E" page="190" pageId="3" pageNumber="623" year="1988">
<bibRefCitation id="EFB9C9F4FF92015F05CF82A5FB88FA7D" author="Haig J &amp; Ball E" pageId="3" pageNumber="623" pagination="151 - 196" refId="ref17097" refString="Haig J &amp; Ball E (1988) Hermit crabs from North Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40: 151 - 196." type="journal article" year="1988">Haig &amp; Ball, 1988: 190</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A899214FF92015F046F8288FB59FA7C" author="Rahayu DL" box="[1075,1268,1447,1469]" page="350" pageId="3" pageNumber="623" year="1996">
<bibRefCitation id="EFB9C9F4FF92015F046F8288FB59FA7C" author="Rahayu DL" box="[1075,1268,1447,1469]" pageId="3" pageNumber="623" pagination="335 - 355" refId="ref19633" refString="Rahayu DL (1996) Notes on littoral hermit crabs (excluding Coenobitidae) (Crustacea: Decapoda: Anomura) mainly from Singapore and Peninsular Malaysia. Raffles Bulletin of Zoology, 44: 335 - 355." type="journal article" year="1996">Rahayu, 1996: 350</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A899214FF92015F055E8288FC20FA1B" author="McLaughlin PA" page="526" pageId="3" pageNumber="623" year="1997">
<bibRefCitation id="EFB9C9F4FF92015F055E8288FC20FA1B" author="McLaughlin PA" pageId="3" pageNumber="623" pagination="433 - 572" refId="ref18396" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Vol. 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997: 526</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="0A899214FF92015F038F82EBFBDCFA18" author="Davie PJF" box="[979,1137,1476,1498]" page="84" pageId="3" pageNumber="623" year="2002">
<bibRefCitation id="EFB9C9F4FF92015F038F82EBFBDCFA18" author="Davie PJF" box="[979,1137,1476,1498]" pageId="3" pageNumber="623" refId="ref16627" refString="Davie PJF (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda -- Anomura, Brachyura. In: Wells A &amp; Houston WWK (eds.) Zoological Catalogue of Australia, Volume 19.3 B. CSIRO Publishing, Australia. xiv + 641 pp." type="book" year="2002">Davie, 2002: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A899214FF92015F042782EBFA25FA18" author="McLaughlin PA &amp; Komai T &amp; Lemaitre R &amp; Rahayu DL" box="[1147,1416,1476,1498]" page="34" pageId="3" pageNumber="623" year="2010">
<bibRefCitation id="EFB9C9F4FF92015F042782EBFA25FA18" author="McLaughlin PA &amp; Komai T &amp; Lemaitre R &amp; Rahayu DL" box="[1147,1416,1476,1498]" pageId="3" pageNumber="623" pagination="5 - 107" refId="ref18745" refString="McLaughlin PA, Komai T, Lemaitre R &amp; Rahayu DL (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part I. Lithodoidea, Lomisoidea and Paguroidea. Raffles Bulletin of Zoology, Supplement 23: 5 - 107." type="journal article" year="2010">McLaughlin et al., 2010: 34</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF92015F031F82D2FBACF9A8" pageId="3" pageNumber="623" type="materials_examined">
<paragraph id="8B97B405FF92015F031F82D2FBACF9A8" blockId="3.[835,1474,1533,1642]" pageId="3" pageNumber="623">
<emphasis id="B95C6817FF92015F031F82D2FBB0F9D2" bold="true" box="[835,1053,1533,1555]" pageId="3" pageNumber="623">Material examined.</emphasis>
<materialsCitation id="3B40BE58FF92015F047482D2FC50F9A8" collectingDate="2004-06-04" collectionCode="ZRC" elevation="14" latitude="9.518333" location="Black Forest" longLatPrecision="129" longitude="123.68833" municipality="Balicasag Island" pageId="3" pageNumber="623" specimenCode="ZRC 2014.0301" specimenCount="1">
1 ovigerous female (sl 6.0 mm) (
<specimenCode id="DB8E1C7EFF92015F05CC82D2FC00F9F1" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="623">ZRC 2014.0301</specimenCode>
), PANGLAO 2004, stn R16,
<location id="8EF7E2DEFF92015F048B8135FAF5F9F1" LSID="urn:lsid:plazi:treatment:03810513FF92015503C983FAFAC1FA54:8EF7E2DEFF92015F048B8135FAF5F9F1" box="[1239,1368,1562,1584]" latitude="9.518333" longLatPrecision="129" longitude="123.68833" municipality="Balicasag Island" name="Black Forest" pageId="3" pageNumber="623">Black Forest</location>
,
<collectingMunicipality id="6BF32E7FFF92015F05388135FC2DF98C" pageId="3" pageNumber="623">Balicasag Island</collectingMunicipality>
,
<geoCoordinate id="EE1CD2C2FF92015F03D78118FC43F98D" box="[907,1006,1591,1612]" degrees="09" direction="north" minutes="31.1" orientation="latitude" pageId="3" pageNumber="623" precision="92" value="9.518333">09°31.1'N</geoCoordinate>
,
<geoCoordinate id="EE1CD2C2FF92015F03A68118FBCBF98D" box="[1018,1126,1591,1612]" degrees="123" direction="east" minutes="41.3" orientation="longitude" pageId="3" pageNumber="623" precision="92" value="123.68833">123°41.3'E</geoCoordinate>
,
<quantity id="4CD019E0FF92015F042E8118FB13F98D" box="[1138,1214,1591,1613]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="3" pageNumber="623" unit="m" value="14.0" valueMax="22.0" valueMin="6.0">
<elevation id="00055336FF92015F042E8118FB13F98D" box="[1138,1214,1591,1613]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="3" pageNumber="623" unit="m" value="14.0" valueMax="22.0" valueMin="6.0">6–22 m</elevation>
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, edge of reef platform and slope,
<date id="FF9692C5FF92015F03DF817BFC50F9A8" box="[899,1021,1620,1641]" pageId="3" pageNumber="623" value="2004-06-04">
<collectingDate id="EFD26B2DFF92015F03DF817BFC50F9A8" box="[899,1021,1620,1641]" pageId="3" pageNumber="623" value="2004-06-04">4 June 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF920155031F81BDFDE7F9F7" lastPageId="9" lastPageNumber="629" pageId="3" pageNumber="623" type="description">
<paragraph id="8B97B405FF92015F031F81BDFAA3F96B" blockId="3.[835,1294,1682,1706]" box="[835,1294,1682,1706]" pageId="3" pageNumber="623">
<emphasis id="B95C6817FF92015F031F81BDFC63F96B" bold="true" box="[835,974,1682,1706]" pageId="3" pageNumber="623">Description.</emphasis>
Eleven pairs of biserial gills.
</paragraph>
<paragraph id="8B97B405FF920158031F81FDFDBAFE57" blockId="3.[835,1474,1746,2090]" lastBlockId="4.[113,752,190,406]" lastPageId="4" lastPageNumber="624" pageId="3" pageNumber="623">
Shield (
<figureCitation id="1313A880FF92015F03C681FDFC5CF92B" box="[922,1009,1746,1770]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="3" pageNumber="623">Fig. 2A</figureCitation>
) approximately as long as broad; anterior margins between rostrum and lateral projections gently concave; anterolateral margins sloping; posterior margin roundly truncate; dorsal surface slightly convex transversely, with longitudinal row of tufts of moderately long setae on either side of midline; paragastric grooves faint. Rostrum acutely triangular, extending as far as level of lateral projections. Lateral projections roundly triangular, with small submarginal spine. Posterior carapace (not figured), measured along midline, distinctly shorter than shield; carapace lateral lobe moderately narrow, sclerotised but not strongly calcified; cardiac sulci parallel, reaching to midlength of posterior carapace; sulci cardiobranchiales also parallel, slightly falling short of midlength of posterior carapace; median area consisting of posteromedian and posterolateral plates not calcified but weakly sclerotised, with several paired tufts of stiff setae; branchiostegite membranous, with scattered tufts of short to moderately long, stiff setae.
</paragraph>
<paragraph id="8B97B405FF950158002D8691FE3AFD37" blockId="4.[113,753,446,758]" pageId="4" pageNumber="624">
Ocular peduncles (including corneas) (
<figureCitation id="1313A880FF95015802488691FDC4FE17" box="[532,617,446,470]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="4" pageNumber="624">Fig. 2A</figureCitation>
) moderately stout, about 0.8 times as long as shield; slightly inflated basally; cornea slightly dilated, its width about 0.4 of peduncular length; dorsal surface with longitudinal row of tufts of moderately short setae mesially. Ocular acicles narrowly subtriangular with rounded tips, separated basally by width of more than one acicle, with minute submarginal spine distally (left) or spine missing (right); dorsal surface shallowly concave. Interocular lobe partially overhung by rostrum, medially concave.
</paragraph>
<paragraph id="8B97B405FF950158002D8431FECEFBD7" blockId="4.[113,752,798,1046]" pageId="4" pageNumber="624">
Antennular peduncles (
<figureCitation id="1313A880FF950158013D8431FE1EFCF7" box="[353,435,798,822]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="4" pageNumber="624">Fig. 2A</figureCitation>
), when fully extended, slightly falling short of distal corneal margins. Ultimate segment about 1.3 times as long as penultimate segment, slightly widened distally in lateral view, with 1 short seta at dorsolateral distal angle. Basal segment with distolateral margin produced in rounded lobe; statocyst lobe weakly inflated, with prominent spine distally; ventromesial distal angle not produced, bearing tuft of long stiff setae.
</paragraph>
<paragraph id="8B97B405FF950158002D8311FD16F997" blockId="4.[113,753,1086,1622]" pageId="4" pageNumber="624">
Antennal peduncles (
<figureCitation id="1313A880FF95015801008311FE19FB97" box="[348,436,1086,1110]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="4" pageNumber="624">Fig. 2A</figureCitation>
) overreaching corneal bases but not reaching distal corneal margins, with supernumerary segmentation. Fifth segment with few setae laterally. Fourth segment with 1 prominent subdistal tuft of stiff setae mesially. Third segment unarmed on ventromesial distal angle, with several tufts of long stiff setae mesially. Second segment with dorsolateral distal angle slightly overreaching midlength of fourth segment, terminating in bifid spine; dorsomesial distal angle with small spine; mesial and lateral faces with short to long stiff setae. First segment with subterminal spinule on lateral face; ventrodistal margin with cluster of spinules just lateral to excretory pore. Antennal acicle overreaching base of cornea but not reaching distal corneal margin, gently arcuate, terminating in small spine obscured by long stiff setae; dorsomesial margin with several tufts of long stiff setae. Antennal flagellum about 3 times as long as shield; each article with 1 or 2 minute setae on distal margin.
</paragraph>
<paragraph id="8B97B405FF950158002D8151FEC0F857" blockId="4.[113,752,1662,1942]" pageId="4" pageNumber="624">
Mouthparts not dissected. Third maxilliped (
<figureCitation id="1313A880FF95015802D78151FD4BF957" box="[651,742,1662,1686]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="4" pageNumber="624">Fig. 2B</figureCitation>
) moderately slender; dactylus shorter than propodus; carpus unarmed on dorsodistal margin; merus unarmed on dorsodistal and ventral margins; ischium with crista dentata consisting of narrowly spaced, moderately small, blunt to subacute corneous teeth, and with 3 moderately strong accessory teeth (
<figureCitation id="1313A880FF95015801788011FED4F897" box="[292,377,1854,1878]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="4" pageNumber="624">Fig. 2C</figureCitation>
); basis-ischium fusion incomplete; basis with 2 denticles on mesial margin; exopod reaching distal margin of merus.
</paragraph>
<paragraph id="8B97B405FF950158002D8091FC57F8D7" blockId="4.[113,752,1982,2102]" lastBlockId="4.[813,1452,190,1814]" pageId="4" pageNumber="624">
Chelipeds grossly unequal. Right cheliped (
<figureCitation id="1313A880FF950158023E8091FD45F817" box="[610,744,1982,2006]" captionStart="Fig" captionStartId="6.[113,148,2005,2027]" captionTargetBox="[138,1441,241,1964]" captionTargetId="figure-10@6.[105,1459,212,1985]" captionTargetPageId="6" captionText="Fig. 3. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right chela, dorsal view; B, right cheliped, mesial view (setae omitted); C, same, lateral view; D, carpus of right cheliped, dorsal view; E, close up of capsulate tubercle on right palm, dorsal view (setae omitted); F, same, obliquely lateral view. Scale bars = 2 mm for A–D; 0.5 mm for E, F." figureDoi="http://doi.org/10.5281/zenodo.5355014" httpUri="https://zenodo.org/record/5355014/files/figure.png" pageId="4" pageNumber="624">Figs. 3A–D</figureCitation>
) with chela 1.5 times longer than greatest width at base of dactylus, generally suboval in outline in dorsal view. Dactylus subequal in length to palm and slightly overlapped by fixed finger, slightly curved ventrally; dorsal surface with sparse, short feathered setae and prominent median row of broadbased, closely set spines or spine-like tubercles; dorsomesial margin with closely set, moderately large tubercles decreasing in size distally; space between dorsal midline and dorsomesial margin with small, low tubercles each bearing central or subdistal, slender, proximally directed and curved capsule; ventral surface with row of low, moderately large protuberances adjacent to mesial margin and tufts of moderately short stiff setae arranged in longitudinal rows; cutting edge with row of broad calcareous teeth and adjacent row of tufts of stiff setae, terminating in large calcareous claw. Palm distinctly longer than carpus; dorsomesial margin delimited by single row of moderately large, stout spines, dorsomesial distal angle somewhat produced with few small spines; dorsolateral margin delimited by small, sharp, forwardly directed spines extending onto fixed finger and decreasing in size and sharpness distally; dorsal surface convex, with numerous, small, scattered capsulate spines and row of moderately small, acute spines on proximal half of midline; vast majority of capsulate spines bifid terminally, flanking base of each capsule, sometimes bearing tuft of short to moderately long feathered setae (
<figureCitation id="1313A880FF950158049D8451FA95FC57" box="[1217,1336,894,918]" captionStart="Fig" captionStartId="6.[113,148,2005,2027]" captionTargetBox="[138,1441,241,1964]" captionTargetId="figure-10@6.[105,1459,212,1985]" captionTargetPageId="6" captionText="Fig. 3. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right chela, dorsal view; B, right cheliped, mesial view (setae omitted); C, same, lateral view; D, carpus of right cheliped, dorsal view; E, close up of capsulate tubercle on right palm, dorsal view (setae omitted); F, same, obliquely lateral view. Scale bars = 2 mm for A–D; 0.5 mm for E, F." figureDoi="http://doi.org/10.5281/zenodo.5355014" httpUri="https://zenodo.org/record/5355014/files/figure.png" pageId="4" pageNumber="624">Fig. 3E, F</figureCitation>
); capsules similar to those on dactylus; mesial face with scattered, very low, small protuberances and tufts of short stiff setae; ventrolateral surface with scattered, very low, minute to small protuberances and sparse very short setae; ventral surface gently convex, with some low protuberances accompanied by tufts of long setae. Cutting edge of fixed finger with row of low calcareous teeth, terminating in large calcareous claw. Carpus slightly shorter than merus; dorsomesial margin delimited by row of large sharp spines accompanied by tufts of long stiff setae; dorsal surface with scattered, small to moderately large spines and tufts of short to long stiff setae, dorsodistal margin with row of small spines; dorsolateral margin delimited by row of small spines; lateral surface with some very low, small protuberances, all accompanied by tufts of short stiff setae, ventrolateral margin with 1 small distal spine; mesial surface shallowly concave medially, with some low protuberances, all accompanied by tufts of long stiff setae; ventral surface with distal margin strongly raised, fringed with long stiff setae. Merus unarmed on dorsodistal margin; dorsal surface glabrous; lateral surface nearly glabrous except for few tufts of short setae, ventrolateral margin with row of small sharp spines and some small, low tubercles proximally; mesial surface with sparse tufts of short setae, ventromesial margin slightly elevated, with row of closely spaced small spines or tubercles; ventral surface with some low, blister-like protuberances and tufts of moderately long setae. Ischium with tufts of short setae on all surfaces; ventromesial margin with few denticles.
</paragraph>
<paragraph id="8B97B405FF95015B03718011FAFFFD54" blockId="4.[813,1452,1854,2102]" lastBlockId="7.[836,1474,189,661]" lastPageId="7" lastPageNumber="627" pageId="4" pageNumber="624">
Left cheliped (
<figureCitation id="1313A880FF95015803898011FBE2F897" box="[981,1103,1854,1878]" captionStart="Fig" captionStartId="7.[136,171,2034,2056]" captionTargetBox="[147,1464,979,1994]" captionTargetId="figure-8@7.[113,1510,957,2013]" captionTargetPageId="7" captionText="Fig. 4. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, chela and carpus of left cheliped, dorsal view; B, left cheliped, mesial view (setae omitted); C, same, lateral view. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355016" httpUri="https://zenodo.org/record/5355016/files/figure.png" pageId="4" pageNumber="624">Fig. 4A–C</figureCitation>
) slightly overreaching base of dactylus of right cheliped, moderately compressed laterally. Chela elongate subovate in dorsal view, 2.7 times longer than greatest width at base of dactylus. Dactylus distinctly longer than palm, nearly straight with ventrally curved tip; dorsal surface sloping mesially, unarmed but with sparse tufts of short feathered setae; all surfaces with tufts of moderately short to long stiff setae; cutting edge with row of minute, subacute calcareous denticles in proximal 0.4 and row of minute, closely set corneous teeth in distal 0.6, terminating in small corneous claw. Palm about half length of carpus, subtriangular in cross section; dorsal surface elevated in midline but not forming distinct ridge or crest, with double row consisting of capsulate tubercles or small spines extending onto base of fixed finger; lateral margin with row of small acute spines; dorsolateral surface with capsulate tubercles arranged in irregular longitudinal rows and extending onto proximal half of fixed finger; dorsomesial surface sloping to mesial surface, with few low protuberances accompanied by tufts of long stiff setae; ventral surface with tufts of long setae. Fixed finger with cutting edge bearing row of minute calcareous denticles in proximal 0.7, and bearing corneous plate, fused with terminal claw, in distal 0.3. Carpus subequal in length to merus; dorsolateral margin with row of small sharp spines and low protuberances and with tufts of short feathered setae, no spine located at distal angle; dorsomesial margin with row of moderately small to large spines, 3 clustered spines located at distal angle; lateral surface with some very low protuberances accompanied by tufts of setae, ventrodistal angle produced, bearing 4 small spines; mesial surface also with very low protuberances accompanied by tufts of long stiff setae, ventrodistal angle with 1 small spine; ventral surface with prominent row of tufts of long feathered setae laterally and scattered tufts of long stiff setae on remaining part. Merus unarmed on dorsodistal margin; dorsal surface with sparse short stiff setae; lateral surface nearly glabrous except for few proximoventral tufts of setae, ventrolateral margin with row of slender, sharp spines and long stiff setae; mesial surface also nearly glabrous except for row of tufts of stiff setae adjacent to dorsal margin and few similar tufts near ventral margin, ventromesial margin with row of long setae and 3 proximal spines; ventral surface with 1 small spine medially and 1 prominent protuberance laterally. Ischium with 4 minute to tiny denticles on ventromesial margin; surfaces unarmed but with tufts of short to moderately long stiff setae.
</paragraph>
<caption id="DF57E48DFF94015900D480FBFC76F781" ID-DOI="http://doi.org/10.5281/zenodo.5355012" ID-Zenodo-Dep="5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="5" pageNumber="625" startId="5.[136,171,2004,2026]" targetBox="[150,1460,350,1966]" targetPageId="5">
<paragraph id="8B97B405FF94015900D480FBFC76F781" blockId="5.[136,1474,2004,2113]" pageId="5" pageNumber="625">
Fig. 2.
<taxonomicName id="4C28CF86FF940159008E80FAFDD8F82B" authority="(Henderson, 1896)" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[210,629,2004,2026]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="625" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF940159008E80FAFE1EF828" box="[210,435,2004,2026]" italics="true" pageId="5" pageNumber="625">Pagurus pergranulatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF940159019F80FBFDC3F82B" author="Henderson JR" box="[451,622,2004,2026]" pageId="5" pageNumber="625" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896</bibRefCitation>
)
</taxonomicName>
, PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G.
</paragraph>
</caption>
<caption id="DF57E48DFF97015A002D80FAFD95F780" ID-DOI="http://doi.org/10.5281/zenodo.5355014" ID-Zenodo-Dep="5355014" httpUri="https://zenodo.org/record/5355014/files/figure.png" pageId="6" pageNumber="626" startId="6.[113,148,2005,2027]" targetBox="[138,1441,241,1964]" targetPageId="6">
<paragraph id="8B97B405FF97015A002D80FAFD95F780" blockId="6.[113,1452,2005,2113]" pageId="6" pageNumber="626">
Fig. 3.
<taxonomicName id="4C28CF86FF97015A00E780F9FDF3F82A" authority="(Henderson, 1896)" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[187,606,2005,2027]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="626" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF97015A00E780F9FE31F82B" box="[187,412,2005,2027]" italics="true" pageId="6" pageNumber="626">Pagurus pergranulatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF97015A01F080FAFDFAF82A" author="Henderson JR" box="[428,599,2005,2027]" pageId="6" pageNumber="626" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896</bibRefCitation>
)
</taxonomicName>
, PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right chela, dorsal view; B, right cheliped, mesial view (setae omitted); C, same, lateral view; D, carpus of right cheliped, dorsal view; E, close up of capsulate tubercle on right palm, dorsal view (setae omitted); F, same, obliquely lateral view. Scale bars = 2 mm for A–D; 0.5 mm for E, F.
</paragraph>
</caption>
<paragraph id="8B97B405FF96015403188592FBFFFE54" blockId="7.[836,1474,701,917]" lastBlockId="8.[812,1452,189,405]" lastPageId="8" lastPageNumber="628" pageId="7" pageNumber="627">
Ambulatory legs (
<figureCitation id="1313A880FF96015B04518592FB25FD14" box="[1037,1160,701,725]" captionStart="Fig" captionStartId="8.[113,148,2034,2056]" captionTargetBox="[124,1439,633,1994]" captionTargetId="figure-144@8.[98,1467,614,2005]" captionTargetPageId="8" captionText="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="7" pageNumber="627">Fig. 5A, D</figureCitation>
) moderately long and stout, right second pereopod overreaching tip of extended right cheliped by half length of dactylus. Dactyli 1.0–1.3 times as long as propodi, 6.3–6.5 times longer than broad, in dorsal view nearly straight, in lateral view slightly curving ventrally; dorsal margins each with row of tufts of short to moderately long setae; lateral faces each without median sulcus; mesial faces also devoid of median sulcus, armed with 1 row (second;
<figureCitation id="1313A880FF990154010A87F2FE05FF35" box="[342,424,220,245]" captionStart="Fig" captionStartId="8.[113,148,2034,2056]" captionTargetBox="[124,1439,633,1994]" captionTargetId="figure-144@8.[98,1467,614,2005]" captionTargetPageId="8" captionText="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="8" pageNumber="628">Fig. 5B</figureCitation>
) or 2 rows (third;
<figureCitation id="1313A880FF990154022B87F2FD64FF35" box="[631,713,220,245]" captionStart="Fig" captionStartId="8.[113,148,2034,2056]" captionTargetBox="[124,1439,633,1994]" captionTargetId="figure-144@8.[98,1467,614,2005]" captionTargetPageId="8" captionText="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="8" pageNumber="628">Fig. 5E</figureCitation>
) of corneous spinules adjacent to dorsal margin; ventral margins each with row of 7 small corneous spines increasing in size distally. Propodi not narrowing distally; dorsal surfaces unarmed but with row of long stiff setae; lateral and mesial surfaces with few tufts of long stiff setae; ventral surfaces with few tufts of long stiff setae at least distally and with 1 or 2 corneous spinules on distal half, ventrodistal margins with 4 (second) or 2 (third) spinules mesially. Carpi each with small dorsodistal spine; dorsal margin with 1 additional spine at midlength (second;
<figureCitation id="1313A880FF99015401F88532FDB7FDF4" box="[420,538,540,565]" captionStart="Fig" captionStartId="8.[113,148,2034,2056]" captionTargetBox="[124,1439,633,1994]" captionTargetId="figure-144@8.[98,1467,614,2005]" captionTargetPageId="8" captionText="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="8" pageNumber="628">Fig. 5A, C</figureCitation>
) or unarmed (third;
<figureCitation id="1313A880FF990154002D8512FF65FD95" box="[113,200,572,597]" captionStart="Fig" captionStartId="8.[113,148,2034,2056]" captionTargetBox="[124,1439,633,1994]" captionTargetId="figure-144@8.[98,1467,614,2005]" captionTargetPageId="8" captionText="Fig. 5. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="8" pageNumber="628">Fig. 5D</figureCitation>
) and with row of tufts of long stiff setae; lateral surface convex, with row of tufts of long stiff setae along midline. Meri each with row of tufts of long stiff setae on dorsal surface; lateral surface almost smooth, with few tufts of long stiff setae distally; ventral surface unarmed but with row of tufts of short to long stiff setae, ventrodistal lateral margin unarmed. Ischia unarmed but with tufts of setae on dorsal and ventral margins.
</paragraph>
<caption id="DF57E48DFF96015B00D480DDFED6F780" ID-DOI="http://doi.org/10.5281/zenodo.5355016" ID-Zenodo-Dep="5355016" httpUri="https://zenodo.org/record/5355016/files/figure.png" pageId="7" pageNumber="627" startId="7.[136,171,2034,2056]" targetBox="[147,1464,979,1994]" targetPageId="7">
<paragraph id="8B97B405FF96015B00D480DDFED6F780" blockId="7.[136,1474,2034,2113]" pageId="7" pageNumber="627">
Fig. 4.
<taxonomicName id="4C28CF86FF96015B008E80DDFDD8F7C9" authority="(Henderson, 1896)" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[210,629,2034,2056]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="627" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF96015B008E80DDFE1EF7C6" box="[210,435,2034,2055]" italics="true" pageId="7" pageNumber="627">Pagurus pergranulatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF96015B019F80DDFDC3F7C9" author="Henderson JR" box="[451,622,2034,2056]" pageId="7" pageNumber="627" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896</bibRefCitation>
)
</taxonomicName>
, PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, chela and carpus of left cheliped, dorsal view; B, left cheliped, mesial view (setae omitted); C, same, lateral view. Scale bar = 2 mm.
</paragraph>
</caption>
<paragraph id="8B97B405FF99015503708692FF5AFE94" blockId="8.[812,1452,445,597]" lastBlockId="9.[136,775,189,341]" lastPageId="9" lastPageNumber="629" pageId="8" pageNumber="628">
Fourth pereopods (
<figureCitation id="1313A880FF99015403A58692FBE3FE14" box="[1017,1102,445,469]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="8" pageNumber="628">Fig. 2D</figureCitation>
) semichelate, similar from right to left, but left slightly shorter than right. Dactylus slightly curved ventrally, terminating in elongate corneous claw (about half-length of corps of dactylus); distal 0.6 of opposable surface with lateral row of long corneous teeth and dense mat of setae increasing in length distally, proximal 0.4 glabrous; preungual process rod-like, reaching midlength of terminal claw. Propodal rasp consisting of about 8 rows of corneous scales. All segments with dorsal and/or ventral tufts of long stiff setae.
</paragraph>
<caption id="DF57E48DFF990154002D80DDFAEFF783" ID-DOI="http://doi.org/10.5281/zenodo.5355018" ID-Zenodo-Dep="5355018" httpUri="https://zenodo.org/record/5355018/files/figure.png" pageId="8" pageNumber="628" startId="8.[113,148,2034,2056]" targetBox="[124,1439,633,1994]" targetPageId="8">
<paragraph id="8B97B405FF990154002D80DDFAEFF783" blockId="8.[113,1452,2034,2114]" pageId="8" pageNumber="628">
Fig. 5.
<taxonomicName id="4C28CF86FF99015400E780DCFDF3F7C9" authority="(Henderson, 1896)" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[187,606,2034,2056]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="628" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF99015400E780DCFE31F7C6" box="[187,412,2034,2056]" italics="true" pageId="8" pageNumber="628">Pagurus pergranulatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF99015401F080DDFDFAF7C9" author="Henderson JR" box="[428,599,2034,2056]" pageId="8" pageNumber="628" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson, 1896</bibRefCitation>
)
</taxonomicName>
, PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, right second pereopod, lateral view; B, same, dactylus, mesial view (setae omitted); C, same, carpus, mesial view (setae omitted); D, left third pereopod, lateral view; E, same, dactylus, mesial view (setae omitted). Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8B97B405FF98015500D48652FD4BFDB4" blockId="9.[136,775,381,629]" pageId="9" pageNumber="629">
Thoracic sternite 3 with anterior margin nearly straight, with microscopic denticle on either side of midpoint; ventral surface with prominent tuft of setae medially. Anterior lobe of thoracic sternite 6 (
<figureCitation id="1313A880FF980155012A86F2FE6AFE34" box="[374,455,477,501]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="9" pageNumber="629">Fig. 2E</figureCitation>
) subsemicircular, bisected by blunt transverse ridge, sloping anterior surface masked by numerous long setae. Thoracic sternite 8 divided in 2 lobes by shallow median groove, each anterolateral angle slightly produced, with prominent tuft of moderately long setae.
</paragraph>
<paragraph id="8B97B405FF98015500D485B2FDF7FD34" blockId="9.[136,775,669,757]" pageId="9" pageNumber="629">
Pleon dextrally twisted (
<figureCitation id="1313A880FF98015501C685B2FE59FD74" box="[410,500,669,693]" captionStart="Fig" captionStartId="2.[113,148,1976,1998]" captionTargetBox="[199,1368,622,1952]" captionTargetId="figure-372@2.[199,1368,622,1952]" captionTargetPageId="2" captionText="Fig. 1. A, Pagurus fungiformis Komai &amp; Rahayu, 2004, male (sl 2.7 mm), PANGLAO 2004, stn M51, Mayacabac, Panglao Island, ZRC 2013.0645; B, Pagurus kulkarnii Sankolli, 1962, male (sl mm), PANGLAO 2004, stn R66, Sungcolan inlet, Panglao Island, ZRC 2014.0300; C, Pagurus moluccensis Haig &amp; Ball, 1988, male (sl 3.2 mm), PANGLAO 2004, stn M2, west end of Alona Beach, Panglao Island, ZRC 2013.0644; D, Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301." figureDoi="http://doi.org/10.5281/zenodo.5355010" httpUri="https://zenodo.org/record/5355010/files/figure.png" pageId="9" pageNumber="629">Fig. 1D</figureCitation>
). Female second to fifth pleopods greatly unequally biramous with much elongate exopods; length fourth&gt;fifth&gt;third&gt;second.
</paragraph>
<paragraph id="8B97B405FF98015500D48432FD50FC54" blockId="9.[136,775,797,917]" pageId="9" pageNumber="629">
Telson (
<figureCitation id="1313A880FF98015500828432FE82FCF4" box="[222,303,797,821]" captionStart="Fig" captionStartId="5.[136,171,2004,2026]" captionTargetBox="[150,1460,350,1966]" captionTargetId="figure-8@5.[127,1482,328,1987]" captionTargetPageId="5" captionText="Fig. 2. Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view (setae omitted); D, left fourth pereopod, lateral view (setae omitted); E, thoracic sternite 6, ventral view; F, telson, dorsal view; G, same, terminal margins, posterodorsal view. Scale bars = 2 mm for A; 1 mm for B–G." figureDoi="http://doi.org/10.5281/zenodo.5355012" httpUri="https://zenodo.org/record/5355012/files/figure.png" pageId="9" pageNumber="629">Fig. 2F</figureCitation>
) with distinct lateral indentations; posterior lobes strongly produced, rounded, unequal, median cleft very deep; each terminal margin with partially double row of numerous slender spines extending onto lateral margin.
</paragraph>
<paragraph id="8B97B405FF98015500D48492FDE7F9F7" blockId="9.[136,775,957,1590]" pageId="9" pageNumber="629">
<emphasis id="B95C6817FF98015500D48492FEFEFC14" bold="true" box="[136,339,957,981]" pageId="9" pageNumber="629">Coloration in life.</emphasis>
<figureCitation id="1313A880FF98015501068492FE1DFC14" box="[346,432,957,981]" captionStart="Fig" captionStartId="2.[113,148,1976,1998]" captionTargetBox="[199,1368,622,1952]" captionTargetId="figure-372@2.[199,1368,622,1952]" captionTargetPageId="2" captionText="Fig. 1. A, Pagurus fungiformis Komai &amp; Rahayu, 2004, male (sl 2.7 mm), PANGLAO 2004, stn M51, Mayacabac, Panglao Island, ZRC 2013.0645; B, Pagurus kulkarnii Sankolli, 1962, male (sl mm), PANGLAO 2004, stn R66, Sungcolan inlet, Panglao Island, ZRC 2014.0300; C, Pagurus moluccensis Haig &amp; Ball, 1988, male (sl 3.2 mm), PANGLAO 2004, stn M2, west end of Alona Beach, Panglao Island, ZRC 2013.0644; D, Pagurus pergranulatus (Henderson, 1896), PANGLAO 2004, stn R16, Black Forest, Balicasag Island, ovigerous female (sl 6.0 mm), ZRC 2014.0301." figureDoi="http://doi.org/10.5281/zenodo.5355010" httpUri="https://zenodo.org/record/5355010/files/figure.png" pageId="9" pageNumber="629">Fig. 1D</figureCitation>
. Shield mottled pale purple and white, with complex pattern of thin lines of dark purple. Ocular peduncles basically whitish; narrow tan ring at base of cornea and broad orange band medially; cornea mottled silver. Antennular peduncles orange, ultimate segment with white distal and proximal rings, penultimate segment with white distal ring. Antennal peduncle generally pale purple; second segment with 3 dark purple longitudinal stripes on dorsal surface; flagellum with alternating bands of purple (3 or 4 articles length) and white (1 or 2 articles length). Right cheliped mottled purple and white in general; mesial surfaces of carpus and merus with irregular patterns of purple on white background. Left cheliped with thin purple lines on white background on chela, carpus and merus. Ambulatory legs with thin purple lines on propodi, carpi, meri and ischia; dactyli each with 1 purple median line and 2 broad purple bands (1 subdistal and 1 proximal). Narrow purple lines on chelipeds and ambulatory legs usually not reaching ends of segments, somewhat irregular in form, occasionally broadening and becoming thinner again on each segment.
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF98015500D48172FEC5F914" pageId="9" pageNumber="629" type="distribution">
<paragraph id="8B97B405FF98015500D48172FEC5F914" blockId="9.[136,775,1629,1749]" pageId="9" pageNumber="629">
<emphasis id="B95C6817FF98015500D48172FE89F9B4" bold="true" box="[136,292,1629,1653]" pageId="9" pageNumber="629">Distribution.</emphasis>
<collectingCountry id="F33FF495FF98015501738172FE08F9B4" box="[303,421,1629,1653]" name="Sri Lanka" pageId="9" pageNumber="629">Sri Lanka</collectingCountry>
, Andaman Islands, Peninsular
<collectingCountry id="F33FF495FF98015500D48152FF5DF954" box="[136,240,1661,1685]" name="Malaysia" pageId="9" pageNumber="629">Malaysia</collectingCountry>
,
<collectingCountry id="F33FF495FF98015500A28152FEC4F954" box="[254,361,1661,1685]" name="Indonesia" pageId="9" pageNumber="629">Indonesia</collectingCountry>
(Seram),
<collectingCountry id="F33FF495FF98015501818152FDEFF954" box="[477,578,1661,1685]" name="Australia" pageId="9" pageNumber="629">Australia</collectingCountry>
(
<collectingRegion id="49EC7AE7FF98015502088152FD71F954" box="[596,732,1661,1685]" country="Australia" name="Queensland" pageId="9" pageNumber="629">Queensland</collectingRegion>
), at depths of
<quantity id="4CD019E0FF98015500A881B2FEC9F974" box="[244,356,1693,1717]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="10.0" metricValueMin="1.5" pageId="9" pageNumber="629" unit="m" value="57.5" valueMax="100.0" valueMin="15.0">
15–
<specimenCount id="9D2E7F8CFF980155014781B2FEC9F974" box="[283,356,1693,1717]" pageId="9" pageNumber="629" type="generic">100 m</specimenCount>
</quantity>
. Newly recorded from the
<collectingCountry id="F33FF495FF98015502D481B2FCAEF974" box="[648,771,1693,1717]" name="Philippines" pageId="9" pageNumber="629">Philippines</collectingCountry>
, at depths of
<quantity id="4CD019E0FF98015501538192FEC9F914" box="[271,356,1725,1749]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="9" pageNumber="629" unit="m" value="14.0" valueMax="22.0" valueMin="6.0">
6–
<specimenCount id="9D2E7F8CFF98015501768192FEC9F914" box="[298,356,1725,1749]" pageId="9" pageNumber="629" type="generic">22 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C332E78EFF98015500D481D2FAC1FA54" pageId="9" pageNumber="629" type="discussion">
<paragraph id="8B97B405FF98015500D481D2FB9FFEF4" blockId="9.[136,778,1789,2101]" lastBlockId="9.[835,1474,189,309]" pageId="9" pageNumber="629">
<emphasis id="B95C6817FF98015500D481D2FF5AF8D4" bold="true" box="[136,247,1789,1813]" pageId="9" pageNumber="629">Remarks.</emphasis>
The presence of capsules on the chelae is known in eight species in
<taxonomicName id="4C28CF86FF98015501138031FE04F8F4" authorityName="Fabricius" authorityYear="1775" box="[335,425,1822,1845]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="genus">
<emphasis id="B95C6817FF98015501138031FE04F8F4" box="[335,425,1822,1845]" italics="true" pageId="9" pageNumber="629">Pagurus</emphasis>
</taxonomicName>
, viz.,
<taxonomicName id="4C28CF86FF98015501B08031FF6EF894" authority="McLaughlin, 1997" authorityName="McLaughlin" authorityYear="1997" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="capsularis">
<emphasis id="B95C6817FF98015501B08031FDD8F8F4" box="[492,629,1821,1845]" italics="true" pageId="9" pageNumber="629">P. capsularis</emphasis>
<bibRefCitation id="EFB9C9F4FF98015502208032FF6EF894" author="McLaughlin PA" pageId="9" pageNumber="629" pagination="433 - 572" refId="ref18396" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Vol. 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C28CF86FF980155008E8011FEC1F894" authorityName="Miers" authorityYear="1880" box="[210,364,1853,1877]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="hirtimanus">
<emphasis id="B95C6817FF980155008E8011FEC1F894" box="[210,364,1853,1877]" italics="true" pageId="9" pageNumber="629">P. hirtimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C28CF86FF98015501238011FD4FF894" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[383,738,1853,1877]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B95C6817FF98015501238011FDA0F894" box="[383,525,1854,1877]" italics="true" pageId="9" pageNumber="629">P. japonicus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF980155027C8012FD77F894" author="Stimpson W" box="[544,730,1853,1877]" pageId="9" pageNumber="629" pagination="31 - 40" refId="ref19950" refString="Stimpson W (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione and Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars IV. Crustacea Cancroidea et Corystoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 31 - 40." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C28CF86FF98015502AE8011FEB0F8B4" baseAuthorityName="Henderson" baseAuthorityYear="1896" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF98015502AE8011FEB0F8B4" italics="true" pageId="9" pageNumber="629">P. pergranulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C28CF86FF98015501758071FD80F8B4" authority="Komai, 2003 b" authorityName="Komai" authorityYear="2003" box="[297,557,1885,1909]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="rubrior">
<emphasis id="B95C6817FF98015501758071FE22F8B4" box="[297,399,1885,1909]" italics="true" pageId="9" pageNumber="629">P. rubrior</emphasis>
<bibRefCitation id="EFB9C9F4FF98015501C88072FD80F8B4" author="Komai T" box="[404,557,1885,1909]" pageId="9" pageNumber="629" pagination="377 - 411" refId="ref17638" refString="Komai T (2003 b) Identities of Pagurus japonicus (Stimpson, 1858) and P. similis (Ortmann, 1892), with description of a new species of Pagurus. Zoosystema, 25: 377 - 411." type="journal article" year="2003">Komai, 2003b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C28CF86FF980155026A8071FF66F854" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B95C6817FF980155026A8071FD39F8B4" box="[566,660,1885,1909]" italics="true" pageId="9" pageNumber="629">P. similis</emphasis>
(
<bibRefCitation id="EFB9C9F4FF98015502FC8072FF6EF854" author="Ortmann AE" pageId="9" pageNumber="629" pagination="241 - 326" refId="ref19299" refString="Ortmann AE (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen, IV. Die Abtheilungen Galatheidea und Paguridea. Zoologischen Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6: 241 - 326, pls. 11, 12." type="book chapter" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C28CF86FF98015500868051FD8BF854" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[218,550,1917,1941]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="B95C6817FF98015500868051FEF8F854" box="[218,341,1918,1941]" italics="true" pageId="9" pageNumber="629">P. sinuatus</emphasis>
(
<bibRefCitation id="EFB9C9F4FF980155013B8052FDB3F854" author="Stimpson W" box="[359,542,1917,1941]" pageId="9" pageNumber="629" pagination="31 - 40" refId="ref19950" refString="Stimpson W (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione and Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars IV. Crustacea Cancroidea et Corystoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 31 - 40." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C28CF86FF980155023B8051FEB5F874" authority="Osawa, 2012" authorityName="Osawa" authorityYear="2012" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="tabataorum">
<emphasis id="B95C6817FF980155023B8051FCAAF854" box="[615,775,1917,1941]" italics="true" pageId="9" pageNumber="629">P. tabataorum</emphasis>
<bibRefCitation id="EFB9C9F4FF98015500D480B2FEB5F874" author="Osawa M" box="[136,280,1949,1973]" pageId="9" pageNumber="629" pagination="155 - 164" refId="ref19368" refString="Osawa M (2012) A new species of the genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan. In: Naruse T, Chan TY, Tan HH, Ahyong ST &amp; Reimer JD (eds.) Scientific Results of the Marine Biodiversity Expedition - KUMEJIMA 2009. Zootaxa, 3367: 155 - 164." type="journal article" year="2012">Osawa, 2012</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFB9C9F4FF980155017580B2FE57F874" author="McLaughlin PA" box="[297,506,1949,1973]" pageId="9" pageNumber="629" pagination="433 - 572" refId="ref18396" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Vol. 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
;
<bibRefCitation id="EFB9C9F4FF980155025480B2FD04F874" author="Komai T" box="[520,681,1949,1973]" pageId="9" pageNumber="629" pagination="377 - 411" refId="ref17638" refString="Komai T (2003 b) Identities of Pagurus japonicus (Stimpson, 1858) and P. similis (Ortmann, 1892), with description of a new species of Pagurus. Zoosystema, 25: 377 - 411." type="journal article" year="2003">Komai, 2003b</bibRefCitation>
;
<bibRefCitation id="EFB9C9F4FF98015502EB80B2FF6FF814" author="Osawa M" pageId="9" pageNumber="629" pagination="155 - 164" refId="ref19368" refString="Osawa M (2012) A new species of the genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan. In: Naruse T, Chan TY, Tan HH, Ahyong ST &amp; Reimer JD (eds.) Scientific Results of the Marine Biodiversity Expedition - KUMEJIMA 2009. Zootaxa, 3367: 155 - 164." type="journal article" year="2012">Osawa, 2012</bibRefCitation>
). All the eight species exclusively occur in the Indo- West Pacific region.
<bibRefCitation id="EFB9C9F4FF980155012480F2FDBAF834" author="Osawa M" box="[376,535,2013,2037]" pageId="9" pageNumber="629" pagination="155 - 164" refId="ref19368" refString="Osawa M (2012) A new species of the genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan. In: Naruse T, Chan TY, Tan HH, Ahyong ST &amp; Reimer JD (eds.) Scientific Results of the Marine Biodiversity Expedition - KUMEJIMA 2009. Zootaxa, 3367: 155 - 164." type="journal article" year="2012">Osawa (2012)</bibRefCitation>
suggested that these species are phylogenetically related based on this possibly synapomorphic character, although no informal species group has been proposed for them. These species also share a rather uncommon feature in the genus, the possession of a left unpaired second pleopod in males (
<bibRefCitation id="EFB9C9F4FF980155054687D2FA10FED4" author="Komai T" box="[1306,1469,253,277]" pageId="9" pageNumber="629" pagination="377 - 411" refId="ref17638" refString="Komai T (2003 b) Identities of Pagurus japonicus (Stimpson, 1858) and P. similis (Ortmann, 1892), with description of a new species of Pagurus. Zoosystema, 25: 377 - 411." type="journal article" year="2003">Komai, 2003b</bibRefCitation>
; personal observation).
</paragraph>
<paragraph id="8B97B405FF980155031F8672FBB3FC94" blockId="9.[835,1474,349,853]" pageId="9" pageNumber="629">
Although no detailed morphological description in modern standard has been published for
<taxonomicName id="4C28CF86FF98015504C98651FAEEFE54" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[1173,1347,381,405]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF98015504C98651FAEEFE54" box="[1173,1347,381,405]" italics="true" pageId="9" pageNumber="629">P. pergranulatus</emphasis>
</taxonomicName>
, the present specimen generally agrees with the previous descriptions by
<bibRefCitation id="EFB9C9F4FF98015503358692FB9FFE14" author="Henderson JR" box="[873,1074,445,469]" pageId="9" pageNumber="629" pagination="516 - 536" refId="ref17293" refString="Henderson JR (1896) Natural history notes from H. M. ' Investigator' Commander C. F. Oldham, R. N., commanding -- Series II, No. 24. Report on the Paguridae collected during the season 1893 - 94. Journal of the Asiatic Society of Bengal, 65: 516 - 536." type="journal article" year="1896">Henderson (1896)</bibRefCitation>
and
<bibRefCitation id="EFB9C9F4FF98015504308692FAB6FE14" author="Alcock A" box="[1132,1307,445,469]" pageId="9" pageNumber="629" pagination="1 - 197" refId="ref16372" refString="Alcock A (1905 b) Anomura, Fascicules 1, Pagurides. Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, 2: i - xi, 1 - 197, 16 pls." type="book chapter" year="1905">Alcock (1905b)</bibRefCitation>
except for the ventromesial margin of the merus of the right cheliped, which develops into a distinct, somewhat wing-like, ventrally produced lobe and bears a dense tuft of setae. However, no such development of a distinct ventromesial lobe of the merus of the right cheliped is seen in our female specimen.
<bibRefCitation id="EFB9C9F4FF98015505258572FC35FD54" author="Komai T" pageId="9" pageNumber="629" pagination="377 - 411" refId="ref17638" refString="Komai T (2003 b) Identities of Pagurus japonicus (Stimpson, 1858) and P. similis (Ortmann, 1892), with description of a new species of Pagurus. Zoosystema, 25: 377 - 411." type="journal article" year="2003">Komai (2003b)</bibRefCitation>
showed that the structure of the ventral part of the merus of the right cheliped could be sexually dimorphic in
<taxonomicName id="4C28CF86FF980155031F8591FC68FD14" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[835,965,702,725]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B95C6817FF980155031F8591FC68FD14" box="[835,965,702,725]" italics="true" pageId="9" pageNumber="629">P. japonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C28CF86FF980155038E8591FB96FD14" authorityName="Komai" authorityYear="2003" box="[978,1083,701,725]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="rubrior">
<emphasis id="B95C6817FF980155038E8591FB96FD14" box="[978,1083,701,725]" italics="true" pageId="9" pageNumber="629">P. rubrior</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28CF86FF98015504328591FB63FD14" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[1134,1230,701,725]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B95C6817FF98015504328591FB63FD14" box="[1134,1230,701,725]" italics="true" pageId="9" pageNumber="629">P. similis</emphasis>
</taxonomicName>
, and it is reasonable to consider that the discrepancy could be attributable to sexual difference also in
<taxonomicName id="4C28CF86FF980155045885D1FB19FCD4" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[1028,1204,765,789]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF980155045885D1FB19FCD4" box="[1028,1204,765,789]" italics="true" pageId="9" pageNumber="629">P. pergranulatus</emphasis>
</taxonomicName>
. The coloration in life of our specimen also agrees well with the description given by
<bibRefCitation id="EFB9C9F4FF980155031F8412FBB7FC94" author="Haig J &amp; Ball E" box="[835,1050,829,853]" pageId="9" pageNumber="629" pagination="151 - 196" refId="ref17097" refString="Haig J &amp; Ball E (1988) Hermit crabs from North Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40: 151 - 196." type="journal article" year="1988">Haig &amp; Ball (1988)</bibRefCitation>
.
</paragraph>
<paragraph id="8B97B405FF980155031F8451FAC1FA54" blockId="9.[835,1478,893,1429]" pageId="9" pageNumber="629">
<taxonomicName id="4C28CF86FF980155031F8451FB98FC54" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[835,1077,893,917]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF980155031F8451FB98FC54" box="[835,1077,893,917]" italics="true" pageId="9" pageNumber="629">Pagurus pergranulatus</emphasis>
</taxonomicName>
is immediately distinguished from the other seven related species shown above by the characteristic armature of the right palm: the dorsomesial distal angle of the right palm is somewhat produced; vast majority of the capsulate spines on the dorsal surface are distally bifid. In the other seven species, the distomesial margin of the right palm is not produced; capsulate tubercles on the dorsal surface are variable in the shape, but never bifid in the six species other than
<taxonomicName id="4C28CF86FF98015503248351FBA3FB54" authorityName="Osawa" authorityYear="2012" box="[888,1038,1149,1173]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="tabataorum">
<emphasis id="B95C6817FF98015503248351FBA3FB54" box="[888,1038,1149,1173]" italics="true" pageId="9" pageNumber="629">P. tabataorum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C28CF86FF98015504418351FB1EFB54" authorityName="Osawa" authorityYear="2012" box="[1053,1203,1149,1173]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="tabataorum">
<emphasis id="B95C6817FF98015504418351FB1EFB54" box="[1053,1203,1149,1173]" italics="true" pageId="9" pageNumber="629">P. tabataorum</emphasis>
</taxonomicName>
has transverse scutes but no spines on chelae. The coloration in life of
<taxonomicName id="4C28CF86FF980155054983B1FA6FFB74" baseAuthorityName="Henderson" baseAuthorityYear="1896" box="[1301,1474,1181,1205]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="pergranulatus">
<emphasis id="B95C6817FF980155054983B1FA6FFB74" box="[1301,1474,1181,1205]" italics="true" pageId="9" pageNumber="629">P. pergranulatus</emphasis>
</taxonomicName>
is also quite characteristic, particularly in having thin lines or stripes of dark purple on the shield, chelipeds and ambulatory legs. No such lines or stripes are seen in the other seven species (cf.
<bibRefCitation id="EFB9C9F4FF98015503908232FBCBFAF4" author="Miyake S" box="[972,1126,1309,1333]" pageId="9" pageNumber="629" refId="ref18954" refString="Miyake S (1982) Japanese Crustacean Decapods and Stomatopods in Color, Volume I. First printing. Hoikusha Publishing Co., Ltd., Osaka, vii + 261 pp., 56 pls. [In Japanese]" type="book" year="1982">Miyake, 1982</bibRefCitation>
for
<taxonomicName id="4C28CF86FF98015504C78231FA8EFAF4" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1179,1315,1310,1333]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B95C6817FF98015504C78231FA8EFAF4" box="[1179,1315,1310,1333]" italics="true" pageId="9" pageNumber="629">P. japonicus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB9C9F4FF98015505698232FCD5FA94" author="McLaughlin PA" pageId="9" pageNumber="629" pagination="433 - 572" refId="ref18396" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Vol. 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
for
<taxonomicName id="4C28CF86FF98015503F48211FB9EFA94" authorityName="McLaughlin" authorityYear="1997" box="[936,1075,1341,1365]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="capsularis">
<emphasis id="B95C6817FF98015503F48211FB9EFA94" box="[936,1075,1341,1365]" italics="true" pageId="9" pageNumber="629">P. capsularis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB9C9F4FF980155041F8212FB70FA94" author="Komai T" box="[1091,1245,1341,1365]" pageId="9" pageNumber="629" pagination="377 - 411" refId="ref17638" refString="Komai T (2003 b) Identities of Pagurus japonicus (Stimpson, 1858) and P. similis (Ortmann, 1892), with description of a new species of Pagurus. Zoosystema, 25: 377 - 411." type="journal article" year="2003">Komai, 2003b</bibRefCitation>
for
<taxonomicName id="4C28CF86FF98015505518211FAD5FA94" authorityName="Komai" authorityYear="2003" box="[1293,1400,1341,1365]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="rubrior">
<emphasis id="B95C6817FF98015505518211FAD5FA94" box="[1293,1400,1341,1365]" italics="true" pageId="9" pageNumber="629">P. rubrior</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28CF86FF98015505F28211FC24FAB4" baseAuthorityName="Ortmann" baseAuthorityYear="1892" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B95C6817FF98015505F28211FC24FAB4" italics="true" pageId="9" pageNumber="629">P. similis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB9C9F4FF98015503CB8271FBBAFAB4" author="Poore GCB" box="[919,1047,1373,1397]" pageId="9" pageNumber="629" refId="ref19515" refString="Poore GCB (2004) Marine Decapod Crustacea of Southern Australia. A Guide to Identification. CSIRO Publishing, Australia, ix + 574 pp." type="book" year="2004">Poore, 2004</bibRefCitation>
for
<taxonomicName id="4C28CF86FF980155041E8271FB18FAB4" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1090,1205,1374,1397]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="B95C6817FF980155041E8271FB18FAB4" box="[1090,1205,1374,1397]" italics="true" pageId="9" pageNumber="629">P. sinuatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB9C9F4FF980155049F8272FA6FFAB4" author="McLaughlin PA &amp; Komai T &amp; Lemaitre R &amp; Rahayu DL" box="[1219,1474,1373,1397]" pageId="9" pageNumber="629" pagination="5 - 107" refId="ref18745" refString="McLaughlin PA, Komai T, Lemaitre R &amp; Rahayu DL (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part I. Lithodoidea, Lomisoidea and Paguroidea. Raffles Bulletin of Zoology, Supplement 23: 5 - 107." type="journal article" year="2010">McLaughlin et al., 2010</bibRefCitation>
for
<taxonomicName id="4C28CF86FF98015503368251FC57FA54" authorityName="Miers" authorityYear="1880" box="[874,1018,1405,1429]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="hirtimanus">
<emphasis id="B95C6817FF98015503368251FC57FA54" box="[874,1018,1405,1429]" italics="true" pageId="9" pageNumber="629">P. hirtimanus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB9C9F4FF98015504558252FB38FA54" author="Osawa M" box="[1033,1173,1405,1429]" pageId="9" pageNumber="629" pagination="155 - 164" refId="ref19368" refString="Osawa M (2012) A new species of the genus Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan. In: Naruse T, Chan TY, Tan HH, Ahyong ST &amp; Reimer JD (eds.) Scientific Results of the Marine Biodiversity Expedition - KUMEJIMA 2009. Zootaxa, 3367: 155 - 164." type="journal article" year="2012">Osawa, 2012</bibRefCitation>
for
<taxonomicName id="4C28CF86FF98015504998251FAF1FA54" authorityName="Osawa" authorityYear="2012" box="[1221,1372,1405,1429]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="629" phylum="Arthropoda" rank="species" species="tabataorum">
<emphasis id="B95C6817FF98015504998251FAF1FA54" box="[1221,1372,1405,1429]" italics="true" pageId="9" pageNumber="629">P. tabataorum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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