treatments-xml/data/03/BA/DC/03BADC104901E114FC3914DBFB3B35E5.xml

<document id="BEB90E7F298C27D5C79281EDAC040987" ID-DOI="10.11606/1807-0205/2020.60.special-issue.31" ID-ISSN="1807-0205" ID-Zenodo-Dep="4615268" ID-ZooBank="127DDA45-D851-422F-947C-A35E18D7C631" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616009207957" checkinUser="carolina" docAuthor="Caron, Edilson, Navarro, Bruna V. &amp; Cajaiba, Reinaldo L." docDate="2020" docId="03BADC104901E114FC3914DBFB3B35E5" docLanguage="en" docName="PapAvZool.60.special-issue.e202060.31.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60 (31)" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.31" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Diochus parvulus Kraatz 1860" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="FF83A4684903E110FFD41501FFA23256" masterDocTitle="Diochus cleidecostae, a new species from the Brazilian Amazon and a discussion of the sexual dimorphism on sternum VIII (Coleoptera: Staphylinidae: Staphylininae: Diochini)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="3" updateTime="1698889457682" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="3F97CDADE5CE29446A2EBE5E035D5F5C">Diochus cleidecostae, a new species from the Brazilian Amazon and a discussion of the sexual dimorphism on sternum VIII (Coleoptera: Staphylinidae: Staphylininae: Diochini)</mods:title>
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<mods:namePart id="27C0A6F6A3BD0570263A489F8470A6D3">Caron, Edilson</mods:namePart>
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<mods:namePart id="675FD74ADAD8FBEB88F0A198E1297A13">Navarro, Bruna V.</mods:namePart>
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<mods:namePart id="5F11E701EFA2B67538D4EDB1340F4338">Cajaiba, Reinaldo L.</mods:namePart>
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<mods:title id="B994F9D3ABB7E273CE7AC7590CC0C65A">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="AFF7F0577744C6DABAD3E50BF687EC2E">2020</mods:date>
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<mods:title id="59D9251B43E42600593277707DA715BB">Pap. Avulsos Zool., S. Paulo</mods:title>
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<subSubSection id="C3093E8D4901E112FC3914DBFAEF33A5" box="[1005,1357,474,499]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BAC6D064901E112FC3914DBFAEF33A5" blockId="2.[1005,1357,474,531]" box="[1005,1357,474,499]" pageId="2" pageNumber="3">
<heading id="D0E4DA6A4901E112FC3914DBFAEF33A5" box="[1005,1357,474,499]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="5">
<taxonomicName id="4C1316854901E112FC3914DBFAEF33A5" ID-CoL="6CW7X" authority="Kraatz, 1860" authorityName="Kraatz" authorityYear="1860" box="[1005,1357,474,499]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FC3914DBFAEF33A5" bold="true" box="[1005,1357,474,499]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FC3914DBFB1633A5" bold="true" box="[1005,1204,474,499]" italics="true" pageId="2" pageNumber="3">Diochus parvulus</emphasis>
<bibRefCitation id="EF8210F74901E112FB6E14DBFAEF33A5" author="Kraatz, G." box="[1210,1357,474,499]" pageId="2" pageNumber="3" pagination="25 - 28" refId="ref4717" refString="Kraatz, G. 1860. Ueber die Gattung Diochus Er. Wiener Entomologische Monatschrift, 4: 25 - 28." type="journal article" year="1860">Kraatz, 1860</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3093E8D4901E112FB9314FBFB513045" box="[1095,1267,506,531]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BAC6D064901E112FB9314FBFB513045" blockId="2.[1005,1357,474,531]" box="[1095,1267,506,531]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FB9314FBFB513045" bold="true" box="[1095,1267,506,531]" pageId="2" pageNumber="3">
(
<figureCitation id="132871834901E112FB9A14FBFB3F3045" box="[1102,1181,506,531]" captionStart="Figures 4-6" captionStartId="2.[170,238,2042,2067]" captionTargetBox="[172,792,1370,2013]" captionTargetId="figure-668@2.[172,792,1370,2013]" captionTargetPageId="2" captionText="Figures 4-6. Habitus (dorsal). (4) Diochus santacatarinae Irmler; (5) D. parvulus Kraatz; (6) D. cleidecostae Caron &amp; Navarro sp. nov., paratype (female). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615272" httpUri="https://zenodo.org/record/4615272/files/figure.png" pageId="2" pageNumber="3">Figs. 5</figureCitation>
, 
<figureCitation id="132871834901E112FB7314FBFB493045" box="[1191,1259,506,531]" captionStart="Figures 7-17" captionStartId="3.[94,162,2010,2035]" captionTargetBox="[96,1417,657,1980]" captionTargetId="figure-238@3.[94,1417,657,1980]" captionTargetPageId="3" captionText="Figures 7-17. Dissected parts,microscope photographs.(7-9) Diochus santacatarinae Irmler:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) D.parvulus Kraatz: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) D. cleidecostae Caron &amp; Navarro sp. nov.: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="2" pageNumber="3">10-14</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4901E112FCB1173BFB503165" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="8BAC6D064901E112FCB1173BFB503165" blockId="2.[869,1494,570,819]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB034A284901E112FCB1173BFB503165" pageId="2" pageNumber="3">
<treatmentCitation id="0AB24B174901E112FCB1173BFB433005" author="Kraatz, G." box="[869,1249,570,595]" page="27" pageId="2" pageNumber="3" year="1860">
<taxonomicName id="4C1316854901E112FCB1173BFB433005" ID-CoL="6CW7X" authority="Kraatz, 1860: 27" authorityName="Kraatz" authorityPageNumber="27" authorityYear="1860" box="[869,1249,570,595]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FCB1173BFB803005" box="[869,1058,570,595]" italics="true" pageId="2" pageNumber="3">Diochus parvulus</emphasis>
<bibRefCitation id="EF8210F74901E112FBF8173BFB433005" author="Kraatz, G." box="[1068,1249,570,595]" pageId="2" pageNumber="3" pagination="25 - 28" refId="ref4717" refString="Kraatz, G. 1860. Ueber die Gattung Diochus Er. Wiener Entomologische Monatschrift, 4: 25 - 28." type="journal article" year="1860">Kraatz, 1860: 27</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description, 
<typeStatus id="54A8D3A44901E112FC5F175BFC1F3025" box="[907,957,602,627]" pageId="2" pageNumber="3">type</typeStatus>
locality: “ 
<collectingRegion id="49D7A3E44901E112FBF9175BFBD23025" box="[1069,1136,602,627]" country="Brazil" name="Bahia" pageId="2" pageNumber="3">Bahia</collectingRegion>
”); 
<treatmentCitation id="0AB24B174901E112FB58175BFC1630C5" author="Bernhauer, M. &amp; Schubert, K." page="319" pageId="2" pageNumber="3" year="1914">
<bibRefCitation id="EF8210F74901E112FB58175BFC1630C5" author="Bernhauer, M. &amp; Schubert, K." pageId="2" pageNumber="3" pagination="289 - 408" refId="ref4304" refString="Bernhauer, M. &amp; Schubert, K. 1914. Staphylinidae IV. In: Schenkling, S. (Ed.). Coleopterorum Catalogus. Berlin, Junk. p. 289 - 408." type="book chapter" year="1914">Bernhauer &amp; Schubert, 1914: 319</bibRefCitation>
</treatmentCitation>
(catalogue, as valid species); 
<treatmentCitation id="0AB24B174901E112FAD1177BFC1630E5" author="Blackwelder, R. E." page="456" pageId="2" pageNumber="3" year="1943">
<bibRefCitation id="EF8210F74901E112FAD1177BFC1630E5" author="Blackwelder, R. E." pageId="2" pageNumber="3" pagination="1 - 658" refId="ref4342" refString="Blackwelder, R. E. 1943. Monograph of the West Indian beetles of the family Staphylinidae. United States National Museum Bulletin, 1 - 658." type="book chapter" year="1943">Blackwelder, 1943: 456</bibRefCitation>
</treatmentCitation>
(mention as synonym of 
<taxonomicName id="4C1316854901E112FB22179BFA6E30E5" ID-CoL="369Z6" authority="Erichson" authorityName="Erichson" box="[1270,1484,666,691]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B967B1144901E112FB22179BFAF830E5" box="[1270,1370,666,691]" italics="true" pageId="2" pageNumber="3">D. nanus</emphasis>
Erichson
</taxonomicName>
); 
<treatmentCitation id="0AB24B174901E112FC5F17BBFB363085" author="Blackwelder, R. E." box="[907,1172,698,723]" page="143" pageId="2" pageNumber="3" year="1944">
<bibRefCitation id="EF8210F74901E112FC5F17BBFB363085" author="Blackwelder, R. E." box="[907,1172,698,723]" pageId="2" pageNumber="3" pagination="1 - 188" refId="ref4371" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin United States National Museum, 185: xii + 1 - 188." type="journal article" year="1944">Blackwelder, 1944: 143</bibRefCitation>
</treatmentCitation>
(catalogue, as synonym of 
<taxonomicName id="4C1316854901E112FC5F17DBFBFA30A5" ID-CoL="369Z6" authority="Erichson" authorityName="Erichson" box="[907,1112,730,755]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B967B1144901E112FC5F17DBFC4E30A5" box="[907,1004,730,755]" italics="true" pageId="2" pageNumber="3">D. nanus</emphasis>
Erichson
</taxonomicName>
); 
<treatmentCitation id="0AB24B174901E112FBBF17DBFAF330A5" author="Herman, L. H." box="[1131,1361,730,755]" page="2447" pageId="2" pageNumber="3" year="2001">
<bibRefCitation id="EF8210F74901E112FBBF17DBFAF330A5" author="Herman, L. H." box="[1131,1361,730,755]" pageId="2" pageNumber="3" pagination="1 - 4218" refId="ref4596" refString="Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millenium, parts I-VII. Bulletin of the American Museum of Natural History, 167: 1 - 4218." type="journal article" year="2001">Herman, 2001: 2447</bibRefCitation>
</treatmentCitation>
(catalogue, as synonym of 
<taxonomicName id="4C1316854901E112FBEB17FBFAAC3145" ID-CoL="369Z6" authority="Erichson" authorityName="Erichson" box="[1087,1294,762,787]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B967B1144901E112FBEB17FBFB023145" box="[1087,1184,762,787]" italics="true" pageId="2" pageNumber="3">D. nanus</emphasis>
Erichson
</taxonomicName>
); 
<treatmentCitation id="0AB24B174901E112FAF617FBFA763145" author="Irmler, U." box="[1314,1492,762,787]" httpUri="http://treatment.plazi.org/id/3A060C6249056C34FF71779DFE66FDDD" page="39" pageId="2" pageNumber="3" year="2017">
<bibRefCitation id="EF8210F74901E112FAF617FBFA763145" author="Irmler, U." box="[1314,1492,762,787]" pageId="2" pageNumber="3" pagination="1 - 62" refId="ref4675" refString="Irmler, U. 2017. A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Contributions to Entomology: Beitrage zur Entomologie, 67 (1): 1 - 62. DOI" type="journal article" year="2017">Irmler, 2017: 39</bibRefCitation>
</treatmentCitation>
(re-description, as valid species).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4901E112FC5F165BFBE336A5" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BAC6D064901E112FC5F165BFB1231E5" blockId="2.[869,1494,858,947]" pageId="2" pageNumber="3">
<materialsCitation id="3B7B675B4901E112FC5F165BFB1231E5" collectionCode="SDEI" country="Germany" location="Senckenberg Deutsches Entomologisches Institut" municipality="Muncheberg" pageId="2" pageNumber="3" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A8D3A44901E112FC5F165BFC503125" box="[907,1010,858,883]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
(not seen) deposited in 
<location id="8ECC3BDD4901E112FA97165BFA8031C5" LSID="urn:lsid:plazi:treatment:03BADC104901E114FC3914DBFB3B35E5:8ECC3BDD4901E112FA97165BFA8031C5" country="Germany" municipality="Muncheberg" name="Senckenberg Deutsches Entomologisches Institut" pageId="2" pageNumber="3">Senckenberg Deutsches Entomologisches Institut</location>
, 
<collectingMunicipality id="6BC8F77C4901E112FA95167BFA7031C5" box="[1345,1490,890,915]" pageId="2" pageNumber="3">Müncheberg</collectingMunicipality>
, 
<collectingCountry id="F3042D964901E112FCB1169BFC6931E5" box="[869,971,922,947]" name="Germany" pageId="2" pageNumber="3">Germany</collectingCountry>
(
<collectionCode id="ED02F5C34901E112FC02169BFBAE31E5" box="[982,1036,922,947]" pageId="2" pageNumber="3">SDEI</collectionCode>
) (
<bibRefCitation id="EF8210F74901E112FBCA169BFB0431E5" author="Irmler, U." box="[1054,1190,922,947]" pageId="2" pageNumber="3" pagination="1 - 62" refId="ref4675" refString="Irmler, U. 2017. A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Contributions to Entomology: Beitrage zur Entomologie, 67 (1): 1 - 62. DOI" type="journal article" year="2017">Irmler, 2017</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAC6D064901E112FCB116DBFBE336A5" blockId="2.[869,1494,986,1267]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FCB116DBFBEF31A5" bold="true" box="[869,1101,986,1011]" pageId="2" pageNumber="3">Material examined:</emphasis>
“ 
<collectingCountry id="F3042D964901E112FB8E16DBFB3B31A5" box="[1114,1177,986,1011]" name="Brazil" pageId="2" pageNumber="3">Brasil</collectingCountry>
, Pará, Uruará / Mata [Native vegetation], 
<date id="FFAD4BC64901E112FC2316FBFBEC3645" box="[1015,1102,1018,1043]" pageId="2" pageNumber="3" value="2015-06">VI-2015</date>
, / Intermediário [collection information], / Pitfall banana / Reinaldo, Wully, Lucas, / Sidi, col.” [white label, printed in black] (
<specimenCount id="9D15A68F4901E112FB32113BFAAB3607" box="[1254,1289,1082,1107]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
, dissected, 
<collectionCode id="ED02F5C34901E112FA5C113BFA6E3605" box="[1416,1484,1082,1107]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="3" type="University or college">DZUP</collectionCode>
). “ 
<collectingCountry id="F3042D964901E112FCB9115BFC0E3625" box="[877,940,1114,1139]" name="Brazil" pageId="2" pageNumber="3">Brasil</collectingCountry>
, Pará, Uruará / Pastagem [Pasture for extensive livestock], 
<date id="FFAD4BC64901E112FC30117BFB9936C5" box="[996,1083,1146,1171]" pageId="2" pageNumber="3" value="2015-06">VI-2015</date>
, / Intermediário [collection information], / Serrapilheira [Forest litter] / Reinaldo, Wully, Idielson, / col.” [white label, printed in black] (
<specimenCount id="9D15A68F4901E112FA8211BBFAD73687" box="[1366,1397,1210,1235]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
, dissect- ed, no head, 
<collectionCode id="ED02F5C34901E112FC2711DBFB9536A5" box="[1011,1079,1242,1267]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="3" type="University or college">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4901E112FCB1101BFA103765" box="[869,1458,1306,1331]" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BAC6D064901E112FCB1101BFA103765" blockId="2.[869,1458,1306,1331]" box="[869,1458,1306,1331]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FCB1101BFB033765" bold="true" box="[869,1185,1306,1331]" pageId="2" pageNumber="3">Diagnosis and description:</emphasis>
See 
<bibRefCitation id="EF8210F74901E112FB00101BFA0C3765" author="Irmler, U." box="[1236,1454,1306,1331]" pageId="2" pageNumber="3" pagination="1 - 62" refId="ref4675" refString="Irmler, U. 2017. A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Contributions to Entomology: Beitrage zur Entomologie, 67 (1): 1 - 62. DOI" type="journal article" year="2017">Irmler (2017: 39-41)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4901E112FCB1105BFAB935A5" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BAC6D064901E112FCB1105BFABE3405" blockId="2.[869,1493,1370,1907]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FCB1105BFC713725" bold="true" box="[869,979,1370,1395]" pageId="2" pageNumber="3">Remarks:</emphasis>
The specimens from 
<collectingRegion id="49D7A3E44901E112FB6D105BFB4E3725" box="[1209,1260,1370,1395]" country="Suriname" name="Para" pageId="2" pageNumber="3">Pará</collectingRegion>
, identified as 
<emphasis id="B967B1144901E112FAAA105BFC6A37C5" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C1316854901E112FAAA105BFC6637C5" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">Diochus parvulus</taxonomicName>
,
</emphasis>
have similar aedeagus (mainly parameres) and spermatheca as those illustrated by 
<bibRefCitation id="EF8210F74901E112FA96109BFA7737E5" author="Irmler, U." box="[1346,1493,1434,1459]" pageId="2" pageNumber="3" pagination="1 - 62" refId="ref4675" refString="Irmler, U. 2017. A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Contributions to Entomology: Beitrage zur Entomologie, 67 (1): 1 - 62. DOI" type="journal article" year="2017">Irmler (2017)</bibRefCitation>
(
<figureCitation id="132871834901E112FCBF10BBFC4F3785" box="[875,1005,1466,1491]" captionStart="Figures 7-17" captionStartId="3.[94,162,2010,2035]" captionTargetBox="[96,1417,657,1980]" captionTargetId="figure-238@3.[94,1417,657,1980]" captionTargetPageId="3" captionText="Figures 7-17. Dissected parts,microscope photographs.(7-9) Diochus santacatarinae Irmler:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) D.parvulus Kraatz: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) D. cleidecostae Caron &amp; Navarro sp. nov.: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="2" pageNumber="3">Figs. 10-14</figureCitation>
). However, similarly to 
<emphasis id="B967B1144901E112FAD410BBFC0337A5" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C1316854901E112FAD410BBFC3F37A5" authorityName="Irmler" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="santacatarinae">Diochus santacatarinae</taxonomicName>
,
</emphasis>
the male of 
<taxonomicName id="4C1316854901E112FBE110DBFB1237A5" box="[1077,1200,1498,1523]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FBE110DBFB1237A5" box="[1077,1200,1498,1523]" italics="true" pageId="2" pageNumber="3">D. parvulus</emphasis>
</taxonomicName>
has the sternum VIII with posterior margin emarginate at the middle, whereas the female has the sternum VIII with posterior margin broad curved and slightly acute at the middle.
</paragraph>
<paragraph id="8BAC6D064901E112FC5F135BFBC33485" blockId="2.[869,1493,1370,1907]" pageId="2" pageNumber="3">
The specimens of 
<taxonomicName id="4C1316854901E112FBB4135BFB7E3425" box="[1120,1244,1626,1651]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FBB4135BFB7E3425" box="[1120,1244,1626,1651]" italics="true" pageId="2" pageNumber="3">D. parvulus</emphasis>
</taxonomicName>
were collected using the winkler method in forest litter of pasture for extensive livestock and using pitfalls baited with banana in native vegetation (
<figureCitation id="132871834901E112FBCC13BBFBF43485" box="[1048,1110,1722,1747]" captionStart="Figures1-3" captionStartId="1.[794,862,1978,2003]" captionTargetBox="[794,1418,1081,1948]" captionTargetId="figure-521@1.[794,1418,1079,1948]" captionTargetPageId="1" captionText="Figures1-3. (1) Municipalityof Uruará,state of Pará,northern Brazil(PF,primary forest;SF-15,secondary forest with15 years of regeneration;SF-5,secondary forest with five years of regeneration;Ag = agriculture;Pa = pasture for extensive livestock). (2) Detail of primary forest. (3) Detail of agriculture (cocoa plantations,Theobroma cacao L.)." figureDoi="http://doi.org/10.5281/zenodo.4615270" httpUri="https://zenodo.org/record/4615270/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BAC6D064901E112FC5F13DBFBFB3525" blockId="2.[869,1493,1370,1907]" pageId="2" pageNumber="3">
<taxonomicName id="4C1316854901E112FC5F13DBFBF634A5" box="[907,1108,1754,1779]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FC5F13DBFBF634A5" box="[907,1108,1754,1779]" italics="true" pageId="2" pageNumber="3">Diochus parvulus</emphasis>
</taxonomicName>
was previously known from 
<collectingCountry id="F3042D964901E112FCB113FBFC783545" box="[869,986,1786,1811]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
, 
<collectingCountry id="F3042D964901E112FC3613FBFB9C3545" box="[994,1086,1786,1811]" name="Ecuador" pageId="2" pageNumber="3">Ecuador</collectingCountry>
, 
<collectingCountry id="F3042D964901E112FB9113FBFB0F3545" box="[1093,1197,1786,1811]" name="Paraguay" pageId="2" pageNumber="3">Paraguay</collectingCountry>
, 
<collectingCountry id="F3042D964901E112FB6113FBFA813545" box="[1205,1315,1786,1811]" name="Argentina" pageId="2" pageNumber="3">Argentina</collectingCountry>
and 
<collectingCountry id="F3042D964901E112FA8C13FBFA363545" box="[1368,1428,1786,1811]" name="Brazil" pageId="2" pageNumber="3">Brazil</collectingCountry>
(from the states of Amazonas, 
<collectingRegion id="49D7A3E44901E112FBA6121BFB5D3565" box="[1138,1279,1818,1843]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="3">Mato Grosso</collectingRegion>
, 
<collectingRegion id="49D7A3E44901E112FADC121BFA063565" box="[1288,1444,1818,1843]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="3">Rio de Janeiro</collectingRegion>
and 
<collectingRegion id="49D7A3E44901E112FCB1123BFBA93505" box="[869,1035,1850,1875]" country="Brazil" name="Santa Catarina" pageId="2" pageNumber="3">Santa Catarina</collectingRegion>
). The species is recorded for the first time from the state of Pará.
</paragraph>
<paragraph id="8BAC6D064901E112FC4512BBFAB935A5" blockId="2.[913,1449,1978,2035]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FC4512BBFAB935A5" bold="true" pageId="2" pageNumber="3">
<taxonomicName id="4C1316854901E112FC4512BBFAE83585" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[913,1354,1978,2003]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144901E112FC4512BBFB233585" bold="true" box="[913,1153,1978,2003]" italics="true" pageId="2" pageNumber="3">Diochus cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4901E112FA9B12BBFA0B3585" box="[1359,1449,1978,2003]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="132871834901E112FBF212DBFBD735A5" box="[1062,1141,2010,2035]" captionStart="Figures 4-6" captionStartId="2.[170,238,2042,2067]" captionTargetBox="[172,792,1370,2013]" captionTargetId="figure-668@2.[172,792,1370,2013]" captionTargetPageId="2" captionText="Figures 4-6. Habitus (dorsal). (4) Diochus santacatarinae Irmler; (5) D. parvulus Kraatz; (6) D. cleidecostae Caron &amp; Navarro sp. nov., paratype (female). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615272" httpUri="https://zenodo.org/record/4615272/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
, 
<figureCitation id="132871834901E112FBAB12DBFB6735A5" box="[1151,1221,2010,2035]" captionStart="Figures 7-17" captionStartId="3.[94,162,2010,2035]" captionTargetBox="[96,1417,657,1980]" captionTargetId="figure-238@3.[94,1417,657,1980]" captionTargetPageId="3" captionText="Figures 7-17. Dissected parts,microscope photographs.(7-9) Diochus santacatarinae Irmler:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) D.parvulus Kraatz: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) D. cleidecostae Caron &amp; Navarro sp. nov.: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="2" pageNumber="3">15-17</figureCitation>
, 
<figureCitation id="132871834901E112FB1B12DBFAB135A5" box="[1231,1299,2010,2035]" captionStart="Figures 18-22" captionStartId="4.[170,238,1103,1128]" captionTargetBox="[170,794,204,1069]" captionTargetId="figure-662@4.[170,794,204,1073]" captionTargetPageId="4" captionText="Figures 18-22. Dissected parts, drawings.(18-19) D. santacatarinae Irmler: (18) Sternite 8 of male, ventral; (19) Sternite 8 of female, ventral; (20- 22) D. cleidecostae Caron &amp; Navarro sp. nov.: (20) antenna; (21) Aedeagus, dorsal;(22) Aedeagus,ventral.Scale bars:0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4615276" httpUri="https://zenodo.org/record/4615276/files/figure.png" pageId="2" pageNumber="3">20-22</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF6C3D8E4901E112FF7E12FBFE983A05" ID-DOI="http://doi.org/10.5281/zenodo.4615272" ID-Zenodo-Dep="4615272" httpUri="https://zenodo.org/record/4615272/files/figure.png" pageId="2" pageNumber="3" startId="2.[170,238,2042,2067]" targetBox="[172,792,1370,2013]" targetPageId="2">
<paragraph id="8BAC6D064901E112FF7E12FBFE983A05" blockId="2.[170,795,2042,2131]" pageId="2" pageNumber="3">
Figures 4-6. Habitus (dorsal). (4) 
<taxonomicName id="4C1316854901E112FE1C12FBFD1F3A45" authority="Irmler" authorityName="Irmler" authorityYear="2017" box="[456,701,2042,2067]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="santacatarinae">
<emphasis id="B967B1144901E112FE1C12FBFD263A45" bold="true" box="[456,644,2042,2067]" italics="true" pageId="2" pageNumber="3">Diochus santacatarinae</emphasis>
Irmler
</taxonomicName>
; (5) 
<taxonomicName id="4C1316854901E112FD3712FBFEAD3A65" authority="Kraatz" authorityName="Kraatz" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144901E112FD3712FBFF763A65" bold="true" italics="true" pageId="2" pageNumber="3">D. parvulus</emphasis>
Kraatz
</taxonomicName>
; (6) 
<taxonomicName id="4C1316854901E112FEE01D1BFD913A65" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[308,563,2074,2099]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144901E112FEE01D1BFE0B3A65" bold="true" box="[308,425,2074,2099]" italics="true" pageId="2" pageNumber="3">D. cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4901E112FDEC1D1BFDDB3A65" box="[568,633,2074,2099]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (female). Scale bars:1 mm.
</paragraph>
</caption>
<subSubSection id="C3093E8D4901E113FCB11D1BFDA93025" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BAC6D064901E112FCB11D1BFA763A05" blockId="2.[869,1494,2074,2131]" pageId="2" pageNumber="3">
<emphasis id="B967B1144901E112FCB11D1BFBB83A65" bold="true" box="[869,1050,2074,2099]" pageId="2" pageNumber="3">Type material:</emphasis>
<materialsCitation id="3B7B675B4901E112FBF81D1BFA763A05" collectionCode="DZUP" country="Brazil" county="Cap" location="Para" municipality="Uruara" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A8D3A44901E112FBF81D1BFB073A65" box="[1068,1189,2074,2099]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="9D15A68F4901E112FB6A1D1BFB733A67" box="[1214,1233,2074,2097]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
, 
<collectionCode id="ED02F5C34901E112FB331D1BFA893A65" box="[1255,1323,2074,2099]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="3" type="University or college">DZUP</collectionCode>
): “ 
<collectingCountry id="F3042D964901E112FA981D1BFA293A65" box="[1356,1419,2074,2099]" name="Brazil" pageId="2" pageNumber="3">Brasil</collectingCountry>
, 
<location id="8ECC3BDD4901E112FA741D1BFA713A65" LSID="urn:lsid:plazi:treatment:03BADC104901E114FC3914DBFB3B35E5:8ECC3BDD4901E112FA741D1BFA713A65" box="[1440,1491,2074,2099]" country="Brazil" county="Cap" municipality="Uruara" name="Para" pageId="2" pageNumber="3">Pará</location>
, 
<collectingMunicipality id="6BC8F77C4901E112FCB11D3BFC123A05" box="[869,944,2106,2131]" pageId="2" pageNumber="3">Uruará</collectingMunicipality>
/ 
<collectingCounty id="62CD158A4901E112FC1F1D3BFC5A3A05" box="[971,1016,2106,2131]" pageId="2" pageNumber="3">Cap</collectingCounty>
5 [secondary vegetation with five years of
</materialsCitation>
</paragraph>
<paragraph id="8BAC6D064900E113FF8A15DBFDA93025" blockId="3.[94,719,218,627]" pageId="3" pageNumber="4">
<materialsCitation id="3B7B675B4900E113FF8A15DBFE753365" collectingDate="2015-06" collectorName="Sidi" county="Wully" location="Intermediario" municipality="Reinaldo" pageId="3" pageNumber="4" specimenCount="1">
regeneration], 
<date id="FFAD4BC64900E113FED715DBFEF832A5" box="[259,346,218,243]" pageId="3" pageNumber="4" value="2015-06">
<collectingDate id="EFE9B22E4900E113FED715DBFEF832A5" box="[259,346,218,243]" pageId="3" pageNumber="4" value="2015-06">VI-2015</collectingDate>
</date>
, / 
<location id="8ECC3BDD4900E113FEA215DBFDAC32A5" LSID="urn:lsid:plazi:treatment:03BADC104901E114FC3914DBFB3B35E5:8ECC3BDD4900E113FEA215DBFDAC32A5" box="[374,526,218,243]" county="Wully" municipality="Reinaldo" name="Intermediario" pageId="3" pageNumber="4">Intermediário</location>
[collection information], 
<location id="8ECC3BDD4900E113FF1415FBFEEE3345" LSID="urn:lsid:plazi:treatment:03BADC104901E114FC3914DBFB3B35E5:8ECC3BDD4900E113FF1415FBFEEE3345" box="[192,332,250,275]" county="Wully" municipality="Reinaldo" name="Serrapilheira" pageId="3" pageNumber="4">Serrapilheira</location>
[Forest litter] / 
<collectingMunicipality id="6BC8F77C4900E113FE3B15FBFDF13345" box="[495,595,250,275]" pageId="3" pageNumber="4">Reinaldo</collectingMunicipality>
, 
<collectingCounty id="62CD158A4900E113FD8D15FBFD3B3345" box="[601,665,250,275]" pageId="3" pageNumber="4">Wully</collectingCounty>
, 
<collectorName id="26E608D04900E113FD7415FBFD693345" box="[672,715,250,275]" pageId="3" pageNumber="4">Sidi</collectorName>
, col.” [white label, printed in black]
</materialsCitation>
; “ 
<typeStatus id="54A8D3A44900E113FE32141BFDC33365" box="[486,609,282,307]" pageId="3" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C1316854900E113FDA3141BFE383305" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae">
<emphasis id="B967B1144900E113FDA3141BFF433305" italics="true" pageId="3" pageNumber="4">Diochus cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
/ det. 
<determinerName id="7E7081C24900E113FE08143BFD9A3305" box="[476,568,314,339]" pageId="3" pageNumber="4">E. Caron</determinerName>
,2019” [red label, printed in black]. 
<typeStatus id="54A8D3A44900E113FE9C145BFE6B3325" box="[328,457,346,371]" pageId="3" pageNumber="4" type="paratype">PARATYPES</typeStatus>
:the first label as the 
<typeStatus id="54A8D3A44900E113FD7C145BFF0533C5" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
; 
<materialsCitation id="3B7B675B4900E113FF79147BFE7E3065" collectionCode="DZUP, MZSP" collectorName="Sidi" country="Brazil" county="Pitfall-carne" determinerName="E. Caron" location="Mata" municipality="Verao" pageId="3" pageNumber="4" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
“ 
<typeStatus id="54A8D3A44900E113FF60147BFE8533C5" box="[180,295,378,403]" pageId="3" pageNumber="4" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C1316854900E113FEED147BFD6C33C5" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[313,718,378,403]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae">
<emphasis id="B967B1144900E113FEED147BFDB533C5" box="[313,535,378,403]" italics="true" pageId="3" pageNumber="4">Diochus cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
/ det. 
<determinerName id="7E7081C24900E113FF4A149BFF5E33E5" box="[158,252,410,435]" pageId="3" pageNumber="4">E. Caron</determinerName>
, 2019” [yellow label, printed in black] (
<specimenCount id="9D15A68F4900E113FD75149BFD6B33E7" box="[673,713,410,435]" pageId="3" pageNumber="4" type="male">2 ♂</specimenCount>
, 
<specimenCount id="9D15A68F4900E113FF8A14BBFF233387" box="[94,129,442,467]" pageId="3" pageNumber="4" type="female">2 ♀</specimenCount>
, 
<collectionCode id="ED02F5C34900E113FF5F14BBFF693385" box="[139,203,442,467]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="3" pageNumber="4" type="University or college">DZUP</collectionCode>
and 
<specimenCount id="9D15A68F4900E113FED514BBFE853387" box="[257,295,442,467]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D15A68F4900E113FEE514BBFEF13387" box="[305,339,442,467]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED02F5C34900E113FE8A14BBFE003385" box="[350,418,442,467]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
).“ 
<collectingCountry id="F3042D964900E113FE6114BBFE563385" box="[437,500,442,467]" name="Brazil" pageId="3" pageNumber="4">Brasil</collectingCountry>
, Pará, Uruará / 
<location id="8ECC3BDD4900E113FD4214BBFD6C3385" LSID="urn:lsid:plazi:treatment:03BADC104901E114FC3914DBFB3B35E5:8ECC3BDD4900E113FD4214BBFD6C3385" box="[662,718,442,467]" country="Brazil" county="Pitfall-carne" municipality="Verao" name="Mata" pageId="3" pageNumber="4">Mata</location>
[native vegetation], IX-X-2015, / 
<collectingMunicipality id="6BC8F77C4900E113FE1014DBFDA133A5" box="[452,515,474,499]" pageId="3" pageNumber="4">Verão</collectingMunicipality>
[summer season], 
<collectingCounty id="62CD158A4900E113FF8A14FBFF413045" box="[94,227,506,531]" pageId="3" pageNumber="4">Pitfall-carne</collectingCounty>
[fresh meat] / Reinaldo, Wully, Lucas, / 
<collectorName id="26E608D04900E113FD7414FBFD693045" box="[672,715,506,531]" pageId="3" pageNumber="4">Sidi</collectorName>
, col.” [white label, printed in black]
</materialsCitation>
; 
<materialsCitation id="3B7B675B4900E113FE31171BFDA53025" collectionCode="DZUP" collectorName="Sidi" country="Brazil" county="Pitfall-carne" determinerName="E. Caron" isTailEnumeration="true" location="Mata" municipality="Verao" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
“ 
<typeStatus id="54A8D3A44900E113FE3F171BFDFC3065" box="[491,606,538,563]" pageId="3" pageNumber="4" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C1316854900E113FDA3171BFE013005" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae">
<emphasis id="B967B1144900E113FDA3171BFF433005" italics="true" pageId="3" pageNumber="4">Diochus cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
/ det. 
<determinerName id="7E7081C24900E113FE3B173BFDEC3005" box="[495,590,570,595]" pageId="3" pageNumber="4">E. Caron</determinerName>
, 2019” [yellow label, printed in black] (
<specimenCount id="9D15A68F4900E113FE46175BFE103027" box="[402,434,602,627]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED02F5C34900E113FE69175BFDA33025" box="[445,513,602,627]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="3" pageNumber="4" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4900E113FCCE15DBFC533045" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BAC6D064900E113FCCE15DBFC533045" blockId="3.[794,1418,218,531]" pageId="3" pageNumber="4">
<emphasis id="B967B1144900E113FCCE15DBFC3432A5" bold="true" box="[794,918,218,243]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
Within 
<taxonomicName id="4C1316854900E113FC2D15DBFBF232A5" authorityName="Erichson" authorityYear="1839" box="[1017,1104,218,243]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B967B1144900E113FC2D15DBFBF232A5" box="[1017,1104,218,243]" italics="true" pageId="3" pageNumber="4">Diochus</emphasis>
</taxonomicName>
nanus-group, 
<taxonomicName id="4C1316854900E113FB2B15DBFBA23345" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144900E113FB2B15DBFC9E3345" italics="true" pageId="3" pageNumber="4">D. cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4900E113FBDC15FBFBFD3345" box="[1032,1119,250,275]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C1316854900E113FB2615FBFC203365" authority="Cameron" authorityName="Cameron" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apicipennis">
<emphasis id="B967B1144900E113FB2615FBFA2B3345" box="[1266,1417,250,275]" italics="true" pageId="3" pageNumber="4">D. apicipennis</emphasis>
Cameron
</taxonomicName>
, 
<taxonomicName id="4C1316854900E113FC58141BFC483365" box="[908,1002,282,307]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B967B1144900E113FC58141BFC483365" box="[908,1002,282,307]" italics="true" pageId="3" pageNumber="4">D. nanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1316854900E113FBF0141BFAB63365" authority="Cameron" authorityName="Cameron" box="[1060,1300,282,307]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="perplexus">
<emphasis id="B967B1144900E113FBF0141BFB053365" box="[1060,1191,282,307]" italics="true" pageId="3" pageNumber="4">D. perplexus</emphasis>
Cameron
</taxonomicName>
by having an aedeagus with clearing trifurcate apex of parameres (parameres bifurcate at the apex in 
<emphasis id="B967B1144900E113FB38145BFCDB33C5" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C1316854900E113FB38145BFA273325" box="[1260,1413,346,371]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apicipennis">D. apicipennis</taxonomicName>
, 
<taxonomicName id="4C1316854900E113FCCE147BFCDB33C5" box="[794,889,378,403]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nanus">D. nanus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C1316854900E113FC62147BFB9833C5" box="[950,1082,378,403]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="perplexus">
<emphasis id="B967B1144900E113FC62147BFB9833C5" box="[950,1082,378,403]" italics="true" pageId="3" pageNumber="4">D. perplexus</emphasis>
</taxonomicName>
), and differs from 
<taxonomicName id="4C1316854900E113FADB147BFA2B33C5" box="[1295,1417,378,403]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144900E113FADB147BFA2B33C5" box="[1295,1417,378,403]" italics="true" pageId="3" pageNumber="4">D. parvulus</emphasis>
</taxonomicName>
by having three apical long setae of parameres distributed on ventral and dorsal lobes, and without long setae on medial lobe (one of three apical long setae on medial lobe in 
<taxonomicName id="4C1316854900E113FCB814FBFC463045" box="[876,996,506,531]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144900E113FCB814FBFC463045" box="[876,996,506,531]" italics="true" pageId="3" pageNumber="4">D. parvulus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4900E114FCCE173BFC7A3425" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="8BAC6D064900E113FCCE173BFA2B3025" blockId="3.[794,1418,570,627]" pageId="3" pageNumber="4">
<emphasis id="B967B1144900E113FCCE173BFC553005" bold="true" box="[794,1015,570,595]" pageId="3" pageNumber="4">Description: Male:</emphasis>
Maximum body length: 3.00 mm; maximum elytral width: 
<quantity id="4CEBC0E34900E113FBE4175BFB353025" box="[1072,1175,602,627]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="4" unit="mm" value="0.56">0.56 mm</quantity>
. Body light brown to
</paragraph>
<caption id="DF6C3D8E4900E113FF8A12DBFE063A05" ID-DOI="http://doi.org/10.5281/zenodo.4615274" ID-Zenodo-Dep="4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="3" pageNumber="4" startId="3.[94,162,2010,2035]" targetBox="[96,1417,657,1980]" targetPageId="3">
<paragraph id="8BAC6D064900E113FF8A12DBFE063A05" blockId="3.[94,1419,2010,2131]" pageId="3" pageNumber="4">
Figures 7-17. Dissected parts,microscope photographs.(7-9) 
<taxonomicName id="4C1316854900E113FD8A12DBFCF235A5" authority="Irmler" authorityName="Irmler" authorityYear="2017" box="[606,848,2010,2035]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="santacatarinae">
<emphasis id="B967B1144900E113FD8A12DBFCBA35A5" bold="true" box="[606,792,2010,2035]" italics="true" pageId="3" pageNumber="4">Diochus santacatarinae</emphasis>
Irmler
</taxonomicName>
:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) 
<taxonomicName id="4C1316854900E113FA8012DBFF673A45" authority="Kraatz" authorityName="Kraatz" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B967B1144900E113FA8012DBFF2A3A45" bold="true" italics="true" pageId="3" pageNumber="4">D.parvulus</emphasis>
Kraatz
</taxonomicName>
: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) 
<taxonomicName id="4C1316854900E113FE351D1BFD443A65" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[481,742,2074,2099]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144900E113FE351D1BFDF53A65" bold="true" box="[481,599,2074,2099]" italics="true" pageId="3" pageNumber="4">D. cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4900E113FD381D1BFC8C3A65" box="[748,814,2074,2099]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm.
</paragraph>
</caption>
<caption id="DF6C3D8E4907E114FF7E114EFDE5369E" ID-DOI="http://doi.org/10.5281/zenodo.4615276" ID-Zenodo-Dep="4615276" httpUri="https://zenodo.org/record/4615276/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,238,1103,1128]" targetBox="[170,794,204,1069]" targetPageId="4">
<paragraph id="8BAC6D064907E114FF7E114EFDE5369E" blockId="4.[170,795,1103,1224]" pageId="4" pageNumber="5">
Figures 18-22. Dissected parts, drawings.(18-19) 
<taxonomicName id="4C1316854907E114FD85114EFCB5363E" authority="Irmler" authorityName="Irmler" authorityYear="2017" box="[593,791,1103,1128]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="santacatarinae">
<emphasis id="B967B1144907E114FD85114EFD7C363E" bold="true" box="[593,734,1103,1128]" italics="true" pageId="4" pageNumber="5">D. santacatarinae</emphasis>
Irmler
</taxonomicName>
: (18) Sternite 8 of male, ventral; (19) Sternite 8 of female, ventral; (20- 22) 
<taxonomicName id="4C1316854907E114FF18118EFE7036FE" authority="Caron &amp; Navarro" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[204,466,1167,1192]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144907E114FF18118EFEE036FE" bold="true" box="[204,322,1167,1192]" italics="true" pageId="4" pageNumber="5">D. cleidecostae</emphasis>
Caron &amp; Navarro
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4907E114FE0C118EFDB936FE" box="[472,539,1167,1192]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
: (20) antenna; (21) Aedeagus, dorsal;(22) Aedeagus,ventral.Scale bars:0.5mm.
</paragraph>
</caption>
<paragraph id="8BAC6D064907E114FF7E11FBFC5336A5" blockId="4.[170,795,1274,1843]" lastBlockId="4.[869,1494,218,1267]" pageId="4" pageNumber="5">
brown, elytra darker (
<figureCitation id="132871834907E114FE7411FBFE433745" box="[416,481,1274,1299]" captionStart="Figures 4-6" captionStartId="2.[170,238,2042,2067]" captionTargetBox="[172,792,1370,2013]" captionTargetId="figure-668@2.[172,792,1370,2013]" captionTargetPageId="2" captionText="Figures 4-6. Habitus (dorsal). (4) Diochus santacatarinae Irmler; (5) D. parvulus Kraatz; (6) D. cleidecostae Caron &amp; Navarro sp. nov., paratype (female). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615272" httpUri="https://zenodo.org/record/4615272/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
); appendices yellowish, antennae slightly darker than other appendices; antennomere 11 with apical half lighter than basal half. 
<emphasis id="B967B1144907E114FD00103BFCB83705" bold="true" box="[724,794,1338,1363]" pageId="4" pageNumber="5">Head:</emphasis>
Head longer than wide, length 
<quantity id="4CEBC0E34907E114FDD5105BFDC73725" box="[513,613,1370,1395]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="5" unit="mm" value="0.4">0.40 mm</quantity>
, width 
<quantity id="4CEBC0E34907E114FD67105BFCB53725" box="[691,791,1370,1395]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="5" unit="mm" value="0.34">0.34 mm</quantity>
; eyes prominent, 
<quantity id="4CEBC0E34907E114FEBF107BFE7237C5" box="[363,464,1402,1427]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="5" unit="mm" value="0.11">0.11 mm</quantity>
long; temples twice as long as eyes; EL: HL = 0.28; temples parallel, PW: EW = 1.18; posterior angles rounded. Dorsal surface of head slightly microsculptured, vertex impunctate midline, convergent row of four punctures from anterior edge of eyes to middle of vertex, postocular area and at posterior marginal area more setiferous punctures. Antennae slightly longer than head; antennomere 1 nearly as long as 2-3 combined; 2 as long as 3; 3 one-fourth longer than 4; 4-10 wider than long, with same length, slightly increasing in width; 10 around twice wider than long; 11 nearly wid- er than 10 and length of 9-10 combined (
<figureCitation id="132871834907E114FDAB13DBFD6F34A5" box="[639,717,1754,1779]" captionStart="Figures 18-22" captionStartId="4.[170,238,1103,1128]" captionTargetBox="[170,794,204,1069]" captionTargetId="figure-662@4.[170,794,204,1073]" captionTargetPageId="4" captionText="Figures 18-22. Dissected parts, drawings.(18-19) D. santacatarinae Irmler: (18) Sternite 8 of male, ventral; (19) Sternite 8 of female, ventral; (20- 22) D. cleidecostae Caron &amp; Navarro sp. nov.: (20) antenna; (21) Aedeagus, dorsal;(22) Aedeagus,ventral.Scale bars:0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4615276" httpUri="https://zenodo.org/record/4615276/files/figure.png" pageId="4" pageNumber="5">Fig. 20</figureCitation>
). Gular plate with sides divergent towards anterior margin. 
<emphasis id="B967B1144907E114FF7E121BFEA63565" bold="true" box="[170,260,1818,1843]" pageId="4" pageNumber="5">Thorax:</emphasis>
Pronotum longer than wide, 
<quantity id="4CEBC0E34907E114FD99121BFD0C3565" box="[589,686,1818,1843]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="mm" value="0.5">0.50 mm</quantity>
long and 
<quantity id="4CEBC0E34907E114FCB115DBFC6B32A5" box="[869,969,218,243]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="5" unit="mm" value="0.44">0.44 mm</quantity>
wide; anterior and posterior angles rounded; anterior third strongly convergent. Dorsal surface of pronotum glossy; adjacent to wide impunctate midline with row of three punctures; interstice between anterior pair twice as wide as interstices between posterior pairs; four additional punctures laterad; several setiferous punctures along lateral margin. Elytra wider than long, 
<quantity id="4CEBC0E34907E114FCB114BBFC6B3385" box="[869,969,442,467]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="5" unit="mm" value="0.44">0.44 mm</quantity>
long, 
<quantity id="4CEBC0E34907E114FBC014BBFBD53385" box="[1044,1143,442,467]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="5" unit="mm" value="0.56">0.56 mm</quantity>
wide; sides divergent towards posterior margin; posterior margin slightly emarginate. Elytra glossy, setiferous punctures in irregular rows; first row of five punctures adjacent to suture line; two additional rows laterad; interstice between two rows on disc much closer than between row at suture and first row on disc.Hind wings developed.Tarsal formula 5-5-5; fore tarsomeres 1-4 strongly dilated; mid and hind tarsomeres 1-4 clearing decreasing in length; mid and hind tarsomeres 5 nearly the length of 3-4 combined. 
<emphasis id="B967B1144907E114FA8D17DBFA7730A5" bold="true" box="[1369,1493,730,755]" pageId="4" pageNumber="5">Abdomen:</emphasis>
Abdominal segments III-VI with sides subparallel, VII-VIII clearing convergent to posterior margin; densely pubescent. Posterior margin of tergum VIII truncate at middle two-fourths. Posterior margin of sternum VIII emarginate at the middle. Tergum X separating tergum IX in two equal parts. Tergum IX with elongate ventral struts. Posterior margin of sternum IX emarginate at the middle. Posterior margin of tergum X rounded. Aedeagus twice as long as wide (
<figureCitation id="132871834907E114FBA116FBFB563645" box="[1141,1268,1018,1043]" captionStart="Figures 18-22" captionStartId="4.[170,238,1103,1128]" captionTargetBox="[170,794,204,1069]" captionTargetId="figure-662@4.[170,794,204,1073]" captionTargetPageId="4" captionText="Figures 18-22. Dissected parts, drawings.(18-19) D. santacatarinae Irmler: (18) Sternite 8 of male, ventral; (19) Sternite 8 of female, ventral; (20- 22) D. cleidecostae Caron &amp; Navarro sp. nov.: (20) antenna; (21) Aedeagus, dorsal;(22) Aedeagus,ventral.Scale bars:0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4615276" httpUri="https://zenodo.org/record/4615276/files/figure.png" pageId="4" pageNumber="5">Figs. 21-22</figureCitation>
); seminal vesical of endophallus diagonal, 0.62 times as long as central lobe; basal inner duct shortly looped; complex of sclerotised apical forceps; central lobe with acute and crossed apical hooks. Parameres well-developed, each widened and forming large plate at the apex, trifurcate; three apical long setae, two on ventral lobe and one on dorsal lobe (
<figureCitation id="132871834907E114FCBF11DBFC4736A5" box="[875,997,1242,1267]" captionStart="Figures 7-17" captionStartId="3.[94,162,2010,2035]" captionTargetBox="[96,1417,657,1980]" captionTargetId="figure-238@3.[94,1417,657,1980]" captionTargetPageId="3" captionText="Figures 7-17. Dissected parts,microscope photographs.(7-9) Diochus santacatarinae Irmler:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) D.parvulus Kraatz: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) D. cleidecostae Caron &amp; Navarro sp. nov.: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="4" pageNumber="5">Figs. 15-16</figureCitation>
).
</paragraph>
<paragraph id="8BAC6D064907E114FCB1101BFB013785" blockId="4.[869,1494,1306,1491]" pageId="4" pageNumber="5">
<emphasis id="B967B1144907E114FCB1101BFC603765" bold="true" box="[869,962,1306,1331]" pageId="4" pageNumber="5">Female:</emphasis>
Similar to male, except for posterior margin of sternum VIII broadly rounded; tergum IX without ventral struts; sternum IX consisting of pair of hemisternites; spermatheca small with capsule of subspherical shape; bursa copulatrix in lateral position to coiled approximately circular duct (
<figureCitation id="132871834907E114FB9910BBFB3A3785" box="[1101,1176,1466,1491]" captionStart="Figures 7-17" captionStartId="3.[94,162,2010,2035]" captionTargetBox="[96,1417,657,1980]" captionTargetId="figure-238@3.[94,1417,657,1980]" captionTargetPageId="3" captionText="Figures 7-17. Dissected parts,microscope photographs.(7-9) Diochus santacatarinae Irmler:(7) Paramere;(8) Apex of aedeagus;(9) Spermatheca;(10-14) D.parvulus Kraatz: (10) Paramere; (11) Spermatheca; (12) Apex of paramere, focusing on ventral lobe; (13) Apex of paramere, focusing on median lobe; (14) Apex of paramere, focusing on dorsal lobe. (15-17) D. cleidecostae Caron &amp; Navarro sp. nov.: (15) Paramere (dashed line indicates lost seta); (16) Apex of paramere; (17) Spermatheca.Scale bars:0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4615274" httpUri="https://zenodo.org/record/4615274/files/figure.png" pageId="4" pageNumber="5">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BAC6D064907E114FCB110FBFC7A3425" blockId="4.[869,1493,1530,1651]" pageId="4" pageNumber="5">
<emphasis id="B967B1144907E114FCB110FBFB973445" bold="true" box="[869,1077,1530,1555]" pageId="4" pageNumber="5">Biological notes:</emphasis>
The specimens of 
<taxonomicName id="4C1316854907E114FAF910FBFA773445" authority="Caron &amp; Navarro &amp; Cajaiba, 2020" authorityName="Caron &amp; Navarro &amp; Cajaiba" authorityYear="2020" box="[1325,1493,1530,1555]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
<emphasis id="B967B1144907E114FAF910FBFA773445" box="[1325,1493,1530,1555]" italics="true" pageId="4" pageNumber="5">D. cleidecostae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2540C6F4907E114FCB1131BFC1D3465" box="[869,959,1562,1587]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
were found in forest litter secondary vegetation (five years of regeneration) and native vegetation (
<figureCitation id="132871834907E114FCBF135BFC6F3425" box="[875,973,1626,1651]" captionStart="Figures1-3" captionStartId="1.[794,862,1978,2003]" captionTargetBox="[794,1418,1081,1948]" captionTargetId="figure-521@1.[794,1418,1079,1948]" captionTargetPageId="1" captionText="Figures1-3. (1) Municipalityof Uruará,state of Pará,northern Brazil(PF,primary forest;SF-15,secondary forest with15 years of regeneration;SF-5,secondary forest with five years of regeneration;Ag = agriculture;Pa = pasture for extensive livestock). (2) Detail of primary forest. (3) Detail of agriculture (cocoa plantations,Theobroma cacao L.)." figureDoi="http://doi.org/10.5281/zenodo.4615270" httpUri="https://zenodo.org/record/4615270/files/figure.png" pageId="4" pageNumber="5">Figs. 1, 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4907E114FCB1139BFA8A3565" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BAC6D064907E114FCB1139BFA8A3565" blockId="4.[869,1493,1690,1843]" pageId="4" pageNumber="5">
<emphasis id="B967B1144907E114FCB1139BFC4F34E5" bold="true" box="[869,1005,1690,1715]" pageId="4" pageNumber="5">Etymology:</emphasis>
The specific name “cleidecostae” is formed from a personal name and treated as noun in the genitive case (ICZN, 1999, art. 31.1). The specific name honors the Brazilian coleopterologist Dr Cleide Costa, from MZSP and specialist of immature insects.
</paragraph>
</subSubSection>
<subSubSection id="C3093E8D4907E114FE82127BFB3B35E5" pageId="4" pageNumber="5" type="key">
<paragraph id="8BAC6D064907E114FE82127BFB3B35E5" blockId="4.[342,1321,1914,1971]" pageId="4" pageNumber="5">
<emphasis id="B967B1144907E114FE82127BFB3B35E5" bold="true" pageId="4" pageNumber="5">
Key to males of the Neotropical species of 
<taxonomicName id="4C1316854907E114FC92127BFC0135C5" ID-CoL="8VRMP" authorityName="Erichson" authorityYear="1839" box="[838,931,1914,1939]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B967B1144907E114FC92127BFC0135C5" bold="true" box="[838,931,1914,1939]" italics="true" pageId="4" pageNumber="5">Diochus</emphasis>
</taxonomicName>
belonging to the 
<taxonomicName id="4C1316854907E114FBA0127BFB7A35C5" ID-CoL="369Z6" box="[1140,1240,1914,1939]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B967B1144907E114FBA0127BFB7A35C5" bold="true" box="[1140,1240,1914,1939]" italics="true" pageId="4" pageNumber="5">D. nanus</emphasis>
</taxonomicName>
-group (Adapted from 
<bibRefCitation id="EF8210F74907E114FD4D129BFC8835E5" author="Irmler, U." box="[665,810,1946,1971]" pageId="4" pageNumber="5" pagination="1 - 62" refId="ref4675" refString="Irmler, U. 2017. A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Contributions to Entomology: Beitrage zur Entomologie, 67 (1): 1 - 62. DOI" type="journal article" year="2017">Irmler, 2017</bibRefCitation>
, through the couplets 27 to 29)
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>